278 lines
26 KiB
XML
278 lines
26 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1044.62245" ID-GBIF-Dataset="71bbf3b9-a613-49cc-8971-e04a31a448b8" ID-PMC="PMC8222211" ID-Pensoft-Pub="1313-2970-1044-41" ID-Pensoft-UUID="535A79F3B89F5699A71E2B83B65C4C21" ID-PubMed="34183875" ID-ZooBank="C6AE8C5BC5D64A09A26E00FAADBF86E1" ModsDocID="1313-2970-1044-41" checkinTime="1623875478834" checkinUser="pensoft" docAuthor="Kavanaugh, David H., Maddison, David R., Simison, W. Brian, Schoville, Sean D., Schmidt, Joachim, Faille, Arnaud, Moore, Wendy, Pflug, James M., Archambeault, Sophie L., Hoang, Tinya & Chen, Jei-Ying" docDate="2021" docId="E1AC85327E405F19924E8D07745F9C82" docLanguage="en" docName="ZooKeys 1044: 41-152" docOrigin="ZooKeys 1044" docPubDate="2021-06-16" docSource="http://dx.doi.org/10.3897/zookeys.1044.62245" docTitle="Nebria (Nivalonebria) Kavanaugh 2021, subgen. nov." docType="treatment" docUuid="BDE4FBFA-633D-4D30-A57A-10D8A9B294AD" docUuidSource="ZooBank" docVersion="5" id="535A79F3B89F5699A71E2B83B65C4C21" lastPageNumber="41" masterDocId="535A79F3B89F5699A71E2B83B65C4C21" masterDocTitle="Phylogeny of the supertribe Nebriitae (Coleoptera, Carabidae) based on analyses of DNA sequence data" masterLastPageNumber="152" masterPageNumber="41" pageNumber="41" updateTime="1668150468173" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Phylogeny of the supertribe Nebriitae (Coleoptera, Carabidae) based on analyses of DNA sequence data</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kavanaugh, David H.</mods:namePart>
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<mods:affiliation>Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:nameIdentifier type="email">dkavanaugh@calacademy.org</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Maddison, David R.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7152-3824</mods:nameIdentifier>
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<mods:affiliation>Department of Integrative Biology, Oregon State University, Corvallis, OR 97331, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Simison, W. Brian</mods:namePart>
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<mods:affiliation>Center for Comparative Genomics, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Schoville, Sean D.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7364-434X</mods:nameIdentifier>
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<mods:affiliation>Department of Entomology, University of Wisconsin, Madison, WI 53706, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Schmidt, Joachim</mods:namePart>
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<mods:affiliation>Institute of Biosciences, University of Rostock, Universitaetsplatz 2, D- 18055 Rostock, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Faille, Arnaud</mods:namePart>
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<mods:affiliation>Department of Entomology, Coleoptera, Stuttgart State Museum of Natural History, Rosenstein 1, 70191 Stuttgart, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Moore, Wendy</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4441-6203</mods:nameIdentifier>
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<mods:affiliation>Department of Entomology, University of Arizona, Tucson, AZ 85721 - 0036, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pflug, James M.</mods:namePart>
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<mods:affiliation>Department of Integrative Biology, Oregon State University, Corvallis, OR 97331, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Archambeault, Sophie L.</mods:namePart>
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<mods:affiliation>Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA & University of California, Berkeley, 142 Weill Hall # 3200, Berkeley, CA 94720, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hoang, Tinya</mods:namePart>
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<mods:affiliation>Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Chen, Jei-Ying</mods:namePart>
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<mods:affiliation>Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA & University of California, Santa Cruz, Long Marine Lab, 117 McAllister Way, Santa Cruz, CA 95060, USA</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-06-16</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1044</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>41</mods:start>
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<mods:end>152</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1044.62245</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1044.62245</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1044-41</mods:identifier>
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<mods:identifier type="ZooBank">C6AE8C5BC5D64A09A26E00FAADBF86E1</mods:identifier>
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<mods:identifier type="Pensoft-UUID">535A79F3B89F5699A71E2B83B65C4C21</mods:identifier>
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<treatment ID-GBIF-Taxon="182754133" LSID="urn:lsid:zoobank.org:act:BDE4FBFA-633D-4D30-A57A-10D8A9B294AD" httpUri="http://treatment.plazi.org/id/E1AC85327E405F19924E8D07745F9C82" lastPageNumber="41" pageId="0" pageNumber="41">
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<subSubSection pageId="0" pageNumber="41" type="nomenclature">
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<paragraph pageId="0" pageNumber="41">
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Subgenus
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<taxonomicName LSID="http://zoobank.org/BDE4FBFA-633D-4D30-A57A-10D8A9B294AD" authority="Kavanaugh," authorityName="Kavanaugh" authorityYear="2021" class="Insecta" family="Carabidae" genus="Nebria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nebria (Nivalonebria)" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Nivalonebria">Nebria Nivalonebria Kavanaugh,</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="41">subgen. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="41" type="reference_group">
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<paragraph pageId="0" pageNumber="41">
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<taxonomicName authority="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen, 2021" authorityName="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen" authorityYear="2021" class="Insecta" family="Carabidae" genus="Nebria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nebria ovipennis" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="ovipennis" status="subgen. nov.">Nebria ovipennis</taxonomicName>
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= "
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<taxonomicName authority="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen, 2021" authorityName="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen" authorityYear="2021" class="Insecta" family="Carabidae" genus="Nebria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nebria ovipennis" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="ovipennis" status="subgen. nov.">Nebria ovipennis</taxonomicName>
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group" (in part) sensu
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<bibRefCitation author="Lindroth, CH" journalOrPublisher="Proceedings of a symposium (28 August, 1996, Florence, Italy). XX International Congress of Entomology. Museo Regionale di Scienze Naturali, Torino" pageId="0" pageNumber="41" refId="B127" refString="Lindroth, CH, 1961. The ground-beetles (Carabidae, excl. Cicindelinae) of Canada and Alaska. Part 2. Opuscula Entomologica Supplementum No.20: 1-200." title="The ground-beetles (Carabidae, excl. Cicindelinae) of Canada and Alaska. Part 2. Opuscula Entomologica Supplementum No. 20: 1 - 200." year="1961">Lindroth 1961</bibRefCitation>
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</paragraph>
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<paragraph pageId="0" pageNumber="41">
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<taxonomicName authority="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen, 2021" authorityName="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen" authorityYear="2021" class="Insecta" family="Carabidae" genus="Nakanebria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nakanebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus" status="subgen. nov.">Nakanebria</taxonomicName>
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Ledoux & Roux, 2005 (in part)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="41" type="type species">
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<paragraph pageId="0" pageNumber="41">Type species.</paragraph>
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<paragraph pageId="0" pageNumber="41">
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<taxonomicName authorityName="Darlington" authorityYear="1931" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" lsidName="Nebria paradisi" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="paradisi">
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<emphasis italics="true" pageId="0" pageNumber="41">Nebria paradisi</emphasis>
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</taxonomicName>
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Darlington, 1931:24, by present designation.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="41" type="diagnosis">
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<paragraph pageId="0" pageNumber="41">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="41">
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Body size small to medium, SBL = 8.7 to 11.3 mm. Head markedly broadened, not constricted behind eyes; vertex with a single medial pale area and a single pair of supraorbital setae. Eyes moderate in size, moderately convex. Antennal scape with one subapicodorsal seta; antennomeres 3 and 4 not laterally compressed, without extra setae. Labrum with three pairs of apical setae. Maxillary stipes typical for genus, with setae inserted flush on smooth surface. Penultimate labial palpomere with three setae. Pronotum with one midlateral and one basolateral seta present on each side. Elytral intervals smooth, without macrosculpture, interval 3 with one to seven setiferous pores, interval 5 with zero to two and interval 7 with zero to two or more setiferous punctures, intervals with setae only faintly catenate; elytral medial margins diverge slightly and arcuate from the midline apically. Hindwings reduced to short, slender strap-like vestiges. Metepisterna smooth, impunctate. Protarsomeres 1-3 expanded in males; mesotarsomeres 2-4 longer than their apical width; tarsi dorsally glabrous. Abdominal sternites IV-VI with one to four pairs of posterior paramedial setae, without paralateral setae. Median lobe of male aedeagus sclerotized dorsally at least to midlength on shaft, symmetrical in dorsal aspect; basal bulb expanded, quadrate, broadly open basally and closed dorsally, without a sagittal aileron present at base or with only a lightly sclerotized collar; mid-shaft parallel-sided in lateral aspect, slightly compressed in cross-section, with right lateral face unmodified; apical orifice markedly deflected right. Right paramere slender and very long. Female valvifers without vestiture; gonopods VIII fused to dorsomedial bases of gonocoxae; gonocoxae with ventral diagonal row of setiform setae and mediodorsal row of setae present. Bursa copulatrix with a pair of large dorsal paramedial sclerotized plates in vestibular chamber, with its longitudinal axis moderately sigmoid dorsally in lateral aspect; spermathecal chamber broadly cordate in dorsal aspect, without dorsal sclerites (in some
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<taxonomicName lsidName="N. paradisi" pageId="0" pageNumber="41" rank="species" species="paradisi">
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<emphasis italics="true" pageId="0" pageNumber="41">N. paradisi</emphasis>
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</taxonomicName>
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) or with a small midline plate (other
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<taxonomicName lsidName="N. paradisi" pageId="0" pageNumber="41" rank="species" species="paradisi">
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<emphasis italics="true" pageId="0" pageNumber="41">N. paradisi</emphasis>
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</taxonomicName>
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) or large and broad dorsal plate (in
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<taxonomicName lsidName="N. turmaduodecima" pageId="0" pageNumber="41" rank="species" species="turmaduodecima">
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<emphasis italics="true" pageId="0" pageNumber="41">N. turmaduodecima</emphasis>
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</taxonomicName>
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); spermathecal duct very long and of uniform diameter throughout or nearly so, inserted basodorsally on spermathecal chamber; spermathecal reservoir of medium length.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="41" type="etymology">
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<paragraph pageId="0" pageNumber="41">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="41">
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The subgeneric epithet is a noun of feminine gender and a combination of the Latin word,
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<taxonomicName baseAuthorityName="Paykull" baseAuthorityYear="1790" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" lsidName="Nebria nivalis" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="nivalis">
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<emphasis italics="true" pageId="0" pageNumber="41">Nebria nivalis</emphasis>
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</taxonomicName>
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, meaning snow, and the genus name,
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<taxonomicName class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" lsidName="Nebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="41">Nebria</emphasis>
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</taxonomicName>
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, in reference to the occurrence of all known members of this clade in the vicinity of montane snowfields and glaciers.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="41" type="remarks">
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<paragraph pageId="0" pageNumber="41">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="41">
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Members of this subgenus can be distinguished from those of subgenus
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<taxonomicName class="Insecta" family="Carabidae" genus="Nivalonebria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nivalonebria (Nakanebria)" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="subGenus" subGenus="Nakanebria">Nivalonebria Nakanebria</taxonomicName>
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, to which they were assigned by
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<bibRefCitation author="Ledoux, G" journalOrPublisher="Museum - Centre de Conservation et d'Etude des Collections et Societe Linneenne de Lyon, Lyon" pageId="0" pageNumber="41" refId="B121" refString="Ledoux, G, Roux, P, 2005. Nebria (Coleoptera, Nebriidae). Faune mondiale. Museum - Centre de Conservation et d'Etude des Collections et Societe Linneenne de Lyon, Lyon" title="Nebria (Coleoptera, Nebriidae). Faune mondiale." year="2005">Ledoux and Roux (2005)</bibRefCitation>
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, by several external and internal morphological features. Externally, the head is markedly broad, the medial margins of the elytra are slightly and arcuately divergent apically, and the ventroapical margin of metatarsomere 4 is markedly lobate laterally in members of
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<taxonomicName class="Insecta" family="Carabidae" genus="Nivalonebria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nivalonebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="41">Nivalonebria</emphasis>
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</taxonomicName>
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species, whereas the head is of moderate width, the medial margins of the elytra are sinuately divergent apically, and the ventroapical margin of metatarsomere 4 is extremely lobate laterally in
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<taxonomicName authorityName="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen" authorityYear="2021" class="Insecta" family="Carabidae" genus="Nakanebria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nakanebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="41">Nakanebria</emphasis>
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</taxonomicName>
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members. Internally, the apical orifice of the median lobe of male genitalia is markedly deflected right in
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<taxonomicName class="Insecta" family="Carabidae" genus="Nivalonebria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nivalonebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="41">Nivalonebria</emphasis>
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</taxonomicName>
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members, only slightly so in
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<taxonomicName authorityName="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen" authorityYear="2021" class="Insecta" family="Carabidae" genus="Nakanebria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nakanebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="41">Nakanebria</emphasis>
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</taxonomicName>
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members. Females of
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<taxonomicName class="Insecta" family="Carabidae" genus="Nivalonebria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nivalonebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="41">Nivalonebria</emphasis>
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</taxonomicName>
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species have valvifers without vestiture, a bursa copulatrix with a pair of large dorsal paramedial sclerotized plates in the vestibular chamber and with its longitudinal axis moderately sigmoid dorsally in lateral aspect and a spermathecal duct markedly long and inserted basodorsally on the chamber. In contrast,
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<taxonomicName authorityName="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen" authorityYear="2021" class="Insecta" family="Carabidae" genus="Nakanebria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nakanebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="41">Nakanebria</emphasis>
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</taxonomicName>
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females have valvifers with vestiture, a bursa copulatrix without sclerotized plates in the vestibular chamber and with its longitudinal axis only slightly deflected apicodorsally, and a spermathecal duct slightly to markedly short and inserted medioventrally on the chamber.
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<taxonomicName class="Insecta" family="Carabidae" genus="Nivalonebria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nivalonebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="41">Nivalonebria</emphasis>
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</taxonomicName>
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members are distinguished from those of the other subgenera in the
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<taxonomicName authorityName="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen" authorityYear="2021" class="Insecta" family="Carabidae" genus="Catonebria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Catonebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="41">Catonebria</emphasis>
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</taxonomicName>
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complex externally in having a single medial pale area on the vertex (as opposed to a distinct pair of paramedial pale spots as seen in the other subgenera) and the medial margins of the elytra slightly and arcuately divergent apically. In members of
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<taxonomicName class="Insecta" family="Carabidae" genus="Neaptenonebria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neaptenonebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="41">Neaptenonebria</emphasis>
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</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Carabidae" genus="Palaptenonebria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Palaptenonebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="41">Palaptenonebria</emphasis>
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</taxonomicName>
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|
and all
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|
<taxonomicName authorityName="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen" authorityYear="2021" class="Insecta" family="Carabidae" genus="Catonebria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Catonebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="41">Catonebria</emphasis>
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|
</taxonomicName>
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|
(except
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|
<taxonomicName authorityName="Kavanaugh" authorityYear="2015" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" lsidName="Nebria baumanni" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="baumanni">
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|
<emphasis italics="true" pageId="0" pageNumber="41">Nebria baumanni</emphasis>
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</taxonomicName>
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|
Kavanaugh, 2015) the medial margins of the elytra diverge apically and a distinct angle. Internally, they are distinguished in having males with the right paramere of the aedeagus very long and slender and females with a bursa copulatrix with a pair of large dorsal paramedial sclerotized plates in the vestibular chamber and a very long spermathecal duct. Males of
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|
<taxonomicName class="Insecta" family="Carabidae" genus="Neaptenonebria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neaptenonebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
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|
<emphasis italics="true" pageId="0" pageNumber="41">Neaptenonebria</emphasis>
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|
</taxonomicName>
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|
,
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|
<taxonomicName class="Insecta" family="Carabidae" genus="Palaptenonebria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Palaptenonebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
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|
<emphasis italics="true" pageId="0" pageNumber="41">Palaptenonebria</emphasis>
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|
</taxonomicName>
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|
and
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|
<taxonomicName authorityName="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen" authorityYear="2021" class="Insecta" family="Carabidae" genus="Catonebria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Catonebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
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|
<emphasis italics="true" pageId="0" pageNumber="41">Catonebria</emphasis>
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|
</taxonomicName>
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|
have the right paramere slightly or moderately shorter, and females have a bursa without sclerites in the vestibular chamber and with a shorter spermathecal duct. The right lateral face of the mid-shaft of the median lobe of
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|
<taxonomicName class="Insecta" family="Carabidae" genus="Nivalonebria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nivalonebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
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|
<emphasis italics="true" pageId="0" pageNumber="41">Nivalonebria</emphasis>
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|
</taxonomicName>
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|
males is smoothly convex and lacks any of the depressions or invaginations seen in this region in
|
|
<taxonomicName class="Insecta" family="Carabidae" genus="Neaptenonebria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neaptenonebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="41">Neaptenonebria</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName class="Insecta" family="Carabidae" genus="Palaptenonebria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Palaptenonebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="41">Palaptenonebria</emphasis>
|
|
</taxonomicName>
|
|
, and some
|
|
<taxonomicName authorityName="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen" authorityYear="2021" class="Insecta" family="Carabidae" genus="Catonebria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Catonebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
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|
<emphasis italics="true" pageId="0" pageNumber="41">Catonebria</emphasis>
|
|
</taxonomicName>
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|
males.
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|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="41" type="distribution">
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|
<paragraph pageId="0" pageNumber="41">Known distribution and diversity.</paragraph>
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|
<paragraph pageId="0" pageNumber="41">
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|
The geographical range of this clade includes two disjunct areas of western North America, each with a single endemic species: (1) the northern Cascade Range of Washington and northern Oregon, from North Cascades National Park in the north to Mount Hood in the south (with
|
|
<taxonomicName lsidName="N. paradisi" pageId="0" pageNumber="41" rank="species" species="paradisi">
|
|
<emphasis italics="true" pageId="0" pageNumber="41">N. paradisi</emphasis>
|
|
</taxonomicName>
|
|
endemic to that area); and (2) the Trinity Alps of northwestern California (with
|
|
<taxonomicName lsidName="N. turmaduodecima" pageId="0" pageNumber="41" rank="species" species="turmaduodecima">
|
|
<emphasis italics="true" pageId="0" pageNumber="41">N. turmaduodecima</emphasis>
|
|
</taxonomicName>
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|
endemic there).
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</paragraph>
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|
</subSubSection>
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</treatment>
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</document> |