602 lines
63 KiB
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602 lines
63 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1139.96639" ID-Pensoft-Pub="1313-2970-1139-33" ID-Pensoft-UUID="A2C101C84CB5568A92BA5F0DB93564D2" ID-ZooBank="0A828EAE0A7E4EF690869572E8F15E21" ModsDocID="1313-2970-1139-33" checkinTime="1673408730709" checkinUser="pensoft" docAuthor="Gao, Zhizhong, Hou, Yanmeng & Zhang, Feng" docDate="2023" docId="B8502145B46C58B48237EED883C95229" docLanguage="en" docName="ZooKeys 1139: 33-69" docOrigin="ZooKeys 1139" docPubDate="2023-01-09" docSource="http://dx.doi.org/10.3897/zookeys.1139.96639" docTitle="Spelaeochthonius wulibeiensis Gao & Hou & Zhang 2023, sp. nov." docType="treatment" docUuid="296D1EC6-9D37-43A4-B971-6330674C6711" docUuidSource="ZooBank" docVersion="1" id="A2C101C84CB5568A92BA5F0DB93564D2" lastPageNumber="33" masterDocId="A2C101C84CB5568A92BA5F0DB93564D2" masterDocTitle="Four new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Pseudotyrannochthoniidae) from Guizhou, China" masterLastPageNumber="69" masterPageNumber="33" pageNumber="33" updateTime="1673408730709" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Four new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Pseudotyrannochthoniidae) from Guizhou, China</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gao, Zhizhong</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6666-8746</mods:nameIdentifier>
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<mods:affiliation>Department of Biology, Xinzhou Teachers University, Xinzhou, Shanxi 034000, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hou, Yanmeng</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0059-3419</mods:nameIdentifier>
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<mods:affiliation>The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zhang, Feng</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3347-1031</mods:nameIdentifier>
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<mods:affiliation>The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<mods:nameIdentifier type="email">dudu06042001@163.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-01-09</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1139</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>33</mods:start>
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<mods:end>69</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1139.96639</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1139.96639</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1139-33</mods:identifier>
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<mods:identifier type="ZooBank">0A828EAE0A7E4EF690869572E8F15E21</mods:identifier>
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<mods:identifier type="Pensoft-UUID">A2C101C84CB5568A92BA5F0DB93564D2</mods:identifier>
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<subSection pageId="0" pageNumber="33" type="taxonomy">
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<treatment LSID="urn:lsid:zoobank.org:act:296D1EC6-9D37-43A4-B971-6330674C6711" httpUri="http://treatment.plazi.org/id/B8502145B46C58B48237EED883C95229" lastPageNumber="33" pageId="0" pageNumber="33">
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<subSubSection pageId="0" pageNumber="33" type="nomenclature">
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<paragraph pageId="0" pageNumber="33">
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<taxonomicName LSID="https://zoobank.org/296D1EC6-9D37-43A4-B971-6330674C6711" authority="Gao & Hou & Zhang, 2023" authorityName="Gao & Hou & Zhang" authorityYear="2023" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Spelaeochthonius wulibeiensis" order="Pseudoscorpiones" pageId="0" pageNumber="33" rank="species" species="wulibeiensis" status="sp. nov.">Spelaeochthonius wulibeiensis</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="33">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="33" type="description">
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<paragraph pageId="0" pageNumber="33">
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Study area with indication of cave locations representing the type localities. Each color represents an administrative region (red: Guizhou Province; purple: Weining County; green: Xishui County; blue: Fenggang County) A Wulibei Cave (Spelaeochthonius wulibeiensis sp. nov.) B Liangfeng Cave (Allochthonius bainiensis sp. nov.) C Daozuo Cave (A. pandus sp. nov.) D Sanjie Cave (A. xinqiaoensis sp. nov.)." figureDoi="10.3897/zookeys.1139.96639.figure1" httpUri="https://binary.pensoft.net/fig/793039" pageId="0" pageNumber="33">Figs 1A</figureCitation>
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<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Wulibei Cave, type locality of Spelaeochthonius wulibeiensis sp. nov. A cave location (red arrow) B entrance C inside the cave entrance D live male of S. wulibeiensis sp. nov. in its natural environment." figureDoi="10.3897/zookeys.1139.96639.figure18" httpUri="https://binary.pensoft.net/fig/793056" pageId="0" pageNumber="33">, 18</figureCitation>
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<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Spelaeochthonius wulibeiensis sp. nov. A holotype male, habitus (minus left chelicera, pedipalp, legs I and IV, dorsal view) B paratype female, habitus (dorsal view). Scale bars: 0.50 mm." figureDoi="10.3897/zookeys.1139.96639.figure19" httpUri="https://binary.pensoft.net/fig/793057" pageId="0" pageNumber="33">, 19</figureCitation>
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<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Spelaeochthonius wulibeiensis sp. nov., holotype male A left chela (lateral view) B carapace (dorsal view) C left chelicera (dorsal view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure20" httpUri="https://binary.pensoft.net/fig/793058" pageId="0" pageNumber="33">, 20</figureCitation>
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Spelaeochthonius wulibeiensis sp. nov., holotype male (A, B), paratype female (C) A coxae (ventral view) B male genital area (ventral view) C female genital area (ventral view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure21" httpUri="https://binary.pensoft.net/fig/793059" pageId="0" pageNumber="33">, 21</figureCitation>
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<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Spelaeochthonius wulibeiensis sp. nov., holotype male A carapace (dorsal view), with a detail of anterior margin B left chelicera (dorsal view), with details of teeth C coxal spines on coxae I (ventral view) D left pedipalp (minus chela, dorsal view) E rallum. Scale bars: 0.20 mm (A, B, D); 0.10 mm (C, E)." figureDoi="10.3897/zookeys.1139.96639.figure22" httpUri="https://binary.pensoft.net/fig/793060" pageId="0" pageNumber="33">, 22</figureCitation>
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<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Spelaeochthonius wulibeiensis sp. nov., holotype male A left chela (lateral view), with details of teeth and trichobothrial pattern B left chela (dorsal view) C leg I (lateral view) D leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1139.96639.figure23" httpUri="https://binary.pensoft.net/fig/793061" pageId="0" pageNumber="33">
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, 23 Chinese name
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<normalizedToken originalValue="五里碑穴伪蝎">五里碑穴伪蝎</normalizedToken>
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</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="33" type="type material">
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<paragraph pageId="0" pageNumber="33">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="33">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="33">Holotype</emphasis>
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: China • ♂; Guizhou Province, Weining County, Yancang Town, Yangguan Village, Wulibei Cave;
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<geoCoordinate degrees="26" direction="north" minutes="53.82" orientation="latitude" precision="9" value="26.897">26°53.82'N</geoCoordinate>
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,
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<geoCoordinate degrees="104" direction="east" minutes="19.36" orientation="longitude" precision="9" value="104.32267">104°19.36'E</geoCoordinate>
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; 2425 m a.s.l.; 07 Aug. 2019; Zegang Feng, Zhaoyi Li and Chen Zhang leg.; under a stone in the deep zone; Ps.-MHBU-GZC19080701 (Figs
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Study area with indication of cave locations representing the type localities. Each color represents an administrative region (red: Guizhou Province; purple: Weining County; green: Xishui County; blue: Fenggang County) A Wulibei Cave (Spelaeochthonius wulibeiensis sp. nov.) B Liangfeng Cave (Allochthonius bainiensis sp. nov.) C Daozuo Cave (A. pandus sp. nov.) D Sanjie Cave (A. xinqiaoensis sp. nov.)." figureDoi="10.3897/zookeys.1139.96639.figure1" httpUri="https://binary.pensoft.net/fig/793039" pageId="0" pageNumber="33">1A</figureCitation>
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,
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<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Wulibei Cave, type locality of Spelaeochthonius wulibeiensis sp. nov. A cave location (red arrow) B entrance C inside the cave entrance D live male of S. wulibeiensis sp. nov. in its natural environment." figureDoi="10.3897/zookeys.1139.96639.figure18" httpUri="https://binary.pensoft.net/fig/793056" pageId="0" pageNumber="33">18</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="33">Paratypes</emphasis>
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: • 2♂; the same data as the holotype; Ps.-MHBU-GZC19080702-GZC19080703 • 1♀; the same location as the holotype; 19 May. 2017, Zhisheng Zhang, Huiming Chen and Luyu Wang leg.; Ps.-MSWU-CZCH-17-06.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="33" type="diagnosis">
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<paragraph pageId="0" pageNumber="33">Diagnosis</paragraph>
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<paragraph pageId="0" pageNumber="33">
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<emphasis bold="true" pageId="0" pageNumber="33">(♂♀).</emphasis>
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The new species can be recognized by the following combination of characters: surfaces mostly with fine reticulations; carapace without eyes or eyespots but eye region bulging and convex in dorsal view; anterior margin without protuberances; cheliceral palm with five setae; rallum with 11 blades (each with fine pinnate, the basal-most blade shorter than the others); coxal spines present on coxa I only, comprising a transverse, contiguous series of seven or eight tridentate blades, which arise from a lightly sclerotized or translucent hillock, the central ramus of each blade (except the basal two) sharply acumino-spatulate and extending beyond the lateral rami; pedipalps slender, femur 7.24 (♂), 6.40 (♀), chela 6.21-6.22 (♂), 5.68 (♀)
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<normalizedToken originalValue="×">x</normalizedToken>
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longer than broad, both chelal fingers with a row of teeth (fixed chelal finger with 22 or 24 teeth; movable chelal finger with 16-19 teeth), slightly retrorse and pointed; chela fingers straight in dorsal view.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="33" type="etymology">
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<paragraph pageId="0" pageNumber="33">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="33">Named after the type locality, Wulibei Cave.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="33" type="description">
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<paragraph pageId="0" pageNumber="33">Description.</paragraph>
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<paragraph pageId="0" pageNumber="33">
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<emphasis bold="true" pageId="0" pageNumber="33">Adult males</emphasis>
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(Figs
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<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Wulibei Cave, type locality of Spelaeochthonius wulibeiensis sp. nov. A cave location (red arrow) B entrance C inside the cave entrance D live male of S. wulibeiensis sp. nov. in its natural environment." figureDoi="10.3897/zookeys.1139.96639.figure18" httpUri="https://binary.pensoft.net/fig/793056" pageId="0" pageNumber="33">18D</figureCitation>
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,
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<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Spelaeochthonius wulibeiensis sp. nov. A holotype male, habitus (minus left chelicera, pedipalp, legs I and IV, dorsal view) B paratype female, habitus (dorsal view). Scale bars: 0.50 mm." figureDoi="10.3897/zookeys.1139.96639.figure19" httpUri="https://binary.pensoft.net/fig/793057" pageId="0" pageNumber="33">19A</figureCitation>
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,
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<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Spelaeochthonius wulibeiensis sp. nov., holotype male A left chela (lateral view) B carapace (dorsal view) C left chelicera (dorsal view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure20" httpUri="https://binary.pensoft.net/fig/793058" pageId="0" pageNumber="33">20</figureCitation>
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,
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Spelaeochthonius wulibeiensis sp. nov., holotype male (A, B), paratype female (C) A coxae (ventral view) B male genital area (ventral view) C female genital area (ventral view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure21" httpUri="https://binary.pensoft.net/fig/793059" pageId="0" pageNumber="33">21A, B</figureCitation>
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,
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<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Spelaeochthonius wulibeiensis sp. nov., holotype male A carapace (dorsal view), with a detail of anterior margin B left chelicera (dorsal view), with details of teeth C coxal spines on coxae I (ventral view) D left pedipalp (minus chela, dorsal view) E rallum. Scale bars: 0.20 mm (A, B, D); 0.10 mm (C, E)." figureDoi="10.3897/zookeys.1139.96639.figure22" httpUri="https://binary.pensoft.net/fig/793060" pageId="0" pageNumber="33">22</figureCitation>
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,
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<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Spelaeochthonius wulibeiensis sp. nov., holotype male A left chela (lateral view), with details of teeth and trichobothrial pattern B left chela (dorsal view) C leg I (lateral view) D leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1139.96639.figure23" httpUri="https://binary.pensoft.net/fig/793061" pageId="0" pageNumber="33">23</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="33">Color</emphasis>
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(Figs
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<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Wulibei Cave, type locality of Spelaeochthonius wulibeiensis sp. nov. A cave location (red arrow) B entrance C inside the cave entrance D live male of S. wulibeiensis sp. nov. in its natural environment." figureDoi="10.3897/zookeys.1139.96639.figure18" httpUri="https://binary.pensoft.net/fig/793056" pageId="0" pageNumber="33">18D</figureCitation>
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,
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<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Spelaeochthonius wulibeiensis sp. nov. A holotype male, habitus (minus left chelicera, pedipalp, legs I and IV, dorsal view) B paratype female, habitus (dorsal view). Scale bars: 0.50 mm." figureDoi="10.3897/zookeys.1139.96639.figure19" httpUri="https://binary.pensoft.net/fig/793057" pageId="0" pageNumber="33">19A</figureCitation>
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,
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<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Spelaeochthonius wulibeiensis sp. nov., holotype male A left chela (lateral view) B carapace (dorsal view) C left chelicera (dorsal view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure20" httpUri="https://binary.pensoft.net/fig/793058" pageId="0" pageNumber="33">20</figureCitation>
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,
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Spelaeochthonius wulibeiensis sp. nov., holotype male (A, B), paratype female (C) A coxae (ventral view) B male genital area (ventral view) C female genital area (ventral view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure21" httpUri="https://binary.pensoft.net/fig/793059" pageId="0" pageNumber="33">21A, B</figureCitation>
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): generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="33">Cephalothorax</emphasis>
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(Figs
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<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Spelaeochthonius wulibeiensis sp. nov., holotype male A left chela (lateral view) B carapace (dorsal view) C left chelicera (dorsal view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure20" httpUri="https://binary.pensoft.net/fig/793058" pageId="0" pageNumber="33">20B</figureCitation>
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,
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Spelaeochthonius wulibeiensis sp. nov., holotype male (A, B), paratype female (C) A coxae (ventral view) B male genital area (ventral view) C female genital area (ventral view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure21" httpUri="https://binary.pensoft.net/fig/793059" pageId="0" pageNumber="33">21A</figureCitation>
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,
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<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Spelaeochthonius wulibeiensis sp. nov., holotype male A carapace (dorsal view), with a detail of anterior margin B left chelicera (dorsal view), with details of teeth C coxal spines on coxae I (ventral view) D left pedipalp (minus chela, dorsal view) E rallum. Scale bars: 0.20 mm (A, B, D); 0.10 mm (C, E)." figureDoi="10.3897/zookeys.1139.96639.figure22" httpUri="https://binary.pensoft.net/fig/793060" pageId="0" pageNumber="33">22A, C</figureCitation>
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): carapace subquadrate, 1.02-1.03
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<normalizedToken originalValue="×">x</normalizedToken>
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longer than broad, gently narrowed posteriorly; surface mostly with fine reticulations, without furrows but with seven or eight lyrifissures; no traces of eyes but eye region bulging and convex in dorsal view; epistome present and with some tiny spinules; with 16 setae arranged s4s: 4: 2: 2: 2, most setae heavy, long, and gently curved. Chaetotaxy of coxae: P 3, I 6-7, II 4-5, III 4, IV 4; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; coxal spines present on coxa I only, comprising a transverse, contiguous series of seven or eight tridentate blades, which arise from a lightly sclerotized or translucent hillock, the central ramus of each blade (except the basal two) sharply acumino-spatulate and extending beyond the lateral rami (Figs
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Spelaeochthonius wulibeiensis sp. nov., holotype male (A, B), paratype female (C) A coxae (ventral view) B male genital area (ventral view) C female genital area (ventral view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure21" httpUri="https://binary.pensoft.net/fig/793059" pageId="0" pageNumber="33">21A</figureCitation>
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,
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<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Spelaeochthonius wulibeiensis sp. nov., holotype male A carapace (dorsal view), with a detail of anterior margin B left chelicera (dorsal view), with details of teeth C coxal spines on coxae I (ventral view) D left pedipalp (minus chela, dorsal view) E rallum. Scale bars: 0.20 mm (A, B, D); 0.10 mm (C, E)." figureDoi="10.3897/zookeys.1139.96639.figure22" httpUri="https://binary.pensoft.net/fig/793060" pageId="0" pageNumber="33">22C</figureCitation>
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); bisetose intercoxal tubercle present between coxae III and IV, tear drop-shaped (Fig.
|
||
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Spelaeochthonius wulibeiensis sp. nov., holotype male (A, B), paratype female (C) A coxae (ventral view) B male genital area (ventral view) C female genital area (ventral view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure21" httpUri="https://binary.pensoft.net/fig/793059" pageId="0" pageNumber="33">21A</figureCitation>
|
||
).
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="33">Chelicera</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Spelaeochthonius wulibeiensis sp. nov., holotype male A left chela (lateral view) B carapace (dorsal view) C left chelicera (dorsal view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure20" httpUri="https://binary.pensoft.net/fig/793058" pageId="0" pageNumber="33">20C</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Spelaeochthonius wulibeiensis sp. nov., holotype male A carapace (dorsal view), with a detail of anterior margin B left chelicera (dorsal view), with details of teeth C coxal spines on coxae I (ventral view) D left pedipalp (minus chela, dorsal view) E rallum. Scale bars: 0.20 mm (A, B, D); 0.10 mm (C, E)." figureDoi="10.3897/zookeys.1139.96639.figure22" httpUri="https://binary.pensoft.net/fig/793060" pageId="0" pageNumber="33">22B, E</figureCitation>
|
||
): large, approximately as long as carapace, 2.37-2.41
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than broad; five setae present on hand, movable finger with a medial seta, all setae acuminate, ventrobasal seta shorter than others; exterior condylar lyrifissure and exterior lyrifissure exist, palm with one extra (between sub-basal seta and an accessory seta). Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 13-15 acute teeth, distal one largest; movable finger with 12 retrorse contiguous teeth of equal length, plus three or four round proximal teeth, 15 or 16 in total; galea represented by a very slight bump on movable finger. Serrula exterior with 21 blades and serrula interior with 17-20 blades. Rallum in two rows and composed of 11 blades with fine pinnate, of which the basal-most blade shorter than the others (Fig.
|
||
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Spelaeochthonius wulibeiensis sp. nov., holotype male A carapace (dorsal view), with a detail of anterior margin B left chelicera (dorsal view), with details of teeth C coxal spines on coxae I (ventral view) D left pedipalp (minus chela, dorsal view) E rallum. Scale bars: 0.20 mm (A, B, D); 0.10 mm (C, E)." figureDoi="10.3897/zookeys.1139.96639.figure22" httpUri="https://binary.pensoft.net/fig/793060" pageId="0" pageNumber="33">22E</figureCitation>
|
||
).
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="33">Pedipalp</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Spelaeochthonius wulibeiensis sp. nov., holotype male A left chela (lateral view) B carapace (dorsal view) C left chelicera (dorsal view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure20" httpUri="https://binary.pensoft.net/fig/793058" pageId="0" pageNumber="33">20A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Spelaeochthonius wulibeiensis sp. nov., holotype male A carapace (dorsal view), with a detail of anterior margin B left chelicera (dorsal view), with details of teeth C coxal spines on coxae I (ventral view) D left pedipalp (minus chela, dorsal view) E rallum. Scale bars: 0.20 mm (A, B, D); 0.10 mm (C, E)." figureDoi="10.3897/zookeys.1139.96639.figure22" httpUri="https://binary.pensoft.net/fig/793060" pageId="0" pageNumber="33">22D</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Spelaeochthonius wulibeiensis sp. nov., holotype male A left chela (lateral view), with details of teeth and trichobothrial pattern B left chela (dorsal view) C leg I (lateral view) D leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1139.96639.figure23" httpUri="https://binary.pensoft.net/fig/793061" pageId="0" pageNumber="33">23A, B</figureCitation>
|
||
): surfaces mostly with fine reticulations; long and slender, trochanter 1.78-2.00, femur 7.24, patella 2.44-2.47, chela 6.21-6.22, hand 2.26-2.36
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than broad; femur 2.62-2.80
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than patella; movable chelal finger 1.61-1.74
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than hand and 0.61-0.63
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than chela. Setae generally long and acuminate; one distal lyrifissure present on patella and femur, respectively (Fig.
|
||
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Spelaeochthonius wulibeiensis sp. nov., holotype male A carapace (dorsal view), with a detail of anterior margin B left chelicera (dorsal view), with details of teeth C coxal spines on coxae I (ventral view) D left pedipalp (minus chela, dorsal view) E rallum. Scale bars: 0.20 mm (A, B, D); 0.10 mm (C, E)." figureDoi="10.3897/zookeys.1139.96639.figure22" httpUri="https://binary.pensoft.net/fig/793060" pageId="0" pageNumber="33">22D</figureCitation>
|
||
). Chelal palm robust and slightly constricted towards fingers. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria,
|
||
<emphasis italics="true" pageId="0" pageNumber="33">ib</emphasis>
|
||
,
|
||
<emphasis italics="true" pageId="0" pageNumber="33">isb</emphasis>
|
||
,
|
||
<emphasis italics="true" pageId="0" pageNumber="33">eb</emphasis>
|
||
,
|
||
<emphasis italics="true" pageId="0" pageNumber="33">esb</emphasis>
|
||
, and
|
||
<emphasis italics="true" pageId="0" pageNumber="33">ist</emphasis>
|
||
clustered at the base of fixed finger,
|
||
<emphasis italics="true" pageId="0" pageNumber="33">ist</emphasis>
|
||
slightly distal to
|
||
<emphasis italics="true" pageId="0" pageNumber="33">esb</emphasis>
|
||
,
|
||
<emphasis italics="true" pageId="0" pageNumber="33">esb</emphasis>
|
||
close to
|
||
<emphasis italics="true" pageId="0" pageNumber="33">ist</emphasis>
|
||
than to
|
||
<emphasis italics="true" pageId="0" pageNumber="33">eb</emphasis>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="33">it</emphasis>
|
||
slightly distal to
|
||
<emphasis italics="true" pageId="0" pageNumber="33">est</emphasis>
|
||
, situated subdistally and forming a pair;
|
||
<emphasis italics="true" pageId="0" pageNumber="33">et</emphasis>
|
||
situated subdistally, very close to chelal teeth;
|
||
<emphasis italics="true" pageId="0" pageNumber="33">dx</emphasis>
|
||
situated distal to
|
||
<emphasis italics="true" pageId="0" pageNumber="33">et</emphasis>
|
||
, near the tip of fixed finger;
|
||
<emphasis italics="true" pageId="0" pageNumber="33">sb</emphasis>
|
||
distinctly closer to
|
||
<emphasis italics="true" pageId="0" pageNumber="33">b</emphasis>
|
||
than to
|
||
<emphasis italics="true" pageId="0" pageNumber="33">st</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Spelaeochthonius wulibeiensis sp. nov., holotype male A left chela (lateral view), with details of teeth and trichobothrial pattern B left chela (dorsal view) C leg I (lateral view) D leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1139.96639.figure23" httpUri="https://binary.pensoft.net/fig/793061" pageId="0" pageNumber="33">23A</figureCitation>
|
||
). Microsetae (chemosensory setae) absent on hand and both palpal fingers. Sensilla absent. Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 22-24 teeth, slightly retrorse and pointed; movable chelal finger with 16-19 teeth (slightly smaller than teeth on fixed chelal finger); a small tubercle between the seventh and eighth teeth present (near trichobothrium
|
||
<emphasis italics="true" pageId="0" pageNumber="33">t</emphasis>
|
||
) (Fig.
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Spelaeochthonius wulibeiensis sp. nov., holotype male A left chela (lateral view), with details of teeth and trichobothrial pattern B left chela (dorsal view) C leg I (lateral view) D leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1139.96639.figure23" httpUri="https://binary.pensoft.net/fig/793061" pageId="0" pageNumber="33">23A</figureCitation>
|
||
). Chelal fingers straight in dorsal view (Fig.
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Spelaeochthonius wulibeiensis sp. nov., holotype male A left chela (lateral view), with details of teeth and trichobothrial pattern B left chela (dorsal view) C leg I (lateral view) D leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1139.96639.figure23" httpUri="https://binary.pensoft.net/fig/793061" pageId="0" pageNumber="33">23B</figureCitation>
|
||
).
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="33">Opisthosoma</emphasis>
|
||
: generally typical, ovate, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I-XII: 2: 4: 6: 6: 6: 7-8: 8: 7: 6: 4: TT: 0, tergites VII-IX each with an unpaired median seta, one lyrifissure present on each side of tergites IV-IX. Sternal chaetotaxy III-XII: 6-9: 10-14: 13-14: 12: 12-13: 12-13: 10-11: 8-9: 0: 2, one lyrifissure present on each side of tergite III. Anterior genital operculum with 10-12 setae, genital opening pit-like, with seven marginal setae on each side, 24-26 in total, with a pair of lyrifissures present anterolateral and posteriolateral to genital opening, respectively (Fig.
|
||
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Spelaeochthonius wulibeiensis sp. nov., holotype male (A, B), paratype female (C) A coxae (ventral view) B male genital area (ventral view) C female genital area (ventral view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure21" httpUri="https://binary.pensoft.net/fig/793059" pageId="0" pageNumber="33">21B</figureCitation>
|
||
).
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="33">Legs</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Spelaeochthonius wulibeiensis sp. nov., holotype male A left chela (lateral view), with details of teeth and trichobothrial pattern B left chela (dorsal view) C leg I (lateral view) D leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1139.96639.figure23" httpUri="https://binary.pensoft.net/fig/793061" pageId="0" pageNumber="33">23C, D</figureCitation>
|
||
): generally typical, long, and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.74
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than patella and with one lyrifissure at the base of femur; tarsus 2.11-2.26
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than tibia. Femoropatella of leg IV 3.70-3.88
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than deep and with one lyrifissure at the base of femur; tibia 5.92-6.00
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than deep; with a long tactile seta on both tarsal segments: basitarsus 4.00
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than deep (TS = 0.22-0.28), telotarsus 11.13-12.43
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than deep and 2.42-2.47
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than basitarsus (TS = 0.26-0.27). Setae of leg I (trochanter to tibia) 1: 13: 12-15: 14-17, setae of leg IV (trochanter to basitarsus) 2: 2: 6: 16: 14. Arolium slightly shorter than the claws, not divided; claws simple.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="33">Dimensions of adult males</emphasis>
|
||
(length/breadth or, in the case of the legs, length/depth in mm). Males: body length 2.50. Pedipalps: trochanter 0.32/0.16-0.18, femur 1.23/0.17, patella 0.44-0.47/0.18-0.19, chela 1.68-1.74/0.27-0.28, hand 0.61-0.66/0.27-0.28, movable finger length 1.06. Chelicera 0.64-0.65/0.27, movable finger length 0.34. Carapace 0.63/0.61-0.62. Leg I: trochanter 0.24-0.25/0.18, femur 0.73-0.75/0.10, patella 0.42-0.43/0.09, tibia 0.34-0.37/0.07, tarsus 0.77-0.78/0.06-0.07. Leg IV: trochanter 0.33-0.36/0.19-0.22, femoropatella 0.97-1.00/0.25-0.27, tibia 0.77-0.78/0.13, basitarsus 0.36/0.09, telotarsus 0.87-0.89/0.07-0.08.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1139.96639.figure18" httpUri="https://binary.pensoft.net/fig/793056" pageId="0" pageNumber="33" start="Figure 18" startId="F18">
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Figure 18.</emphasis>
|
||
Wulibei Cave, type locality of
|
||
<taxonomicName authorityName="Gao & Hou & Zhang" authorityYear="2023" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" kingdom="Animalia" lsidName="Spelaeochthonius wulibeiensis" order="Pseudoscorpiones" pageId="0" pageNumber="33" rank="species" species="wulibeiensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Spelaeochthonius wulibeiensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="33">A</emphasis>
|
||
cave location (red arrow)
|
||
<emphasis bold="true" pageId="0" pageNumber="33">B</emphasis>
|
||
entrance
|
||
<emphasis bold="true" pageId="0" pageNumber="33">C</emphasis>
|
||
inside the cave entrance
|
||
<emphasis bold="true" pageId="0" pageNumber="33">D</emphasis>
|
||
live male of
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. wulibeiensis" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="wulibeiensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">S. wulibeiensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. in its natural environment.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1139.96639.figure19" httpUri="https://binary.pensoft.net/fig/793057" pageId="0" pageNumber="33" start="Figure 19" startId="F19">
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Figure 19.</emphasis>
|
||
<taxonomicName authorityName="Gao & Hou & Zhang" authorityYear="2023" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" kingdom="Animalia" lsidName="Spelaeochthonius wulibeiensis" order="Pseudoscorpiones" pageId="0" pageNumber="33" rank="species" species="wulibeiensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Spelaeochthonius wulibeiensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="33">A</emphasis>
|
||
holotype male, habitus (minus left chelicera, pedipalp, legs I and IV, dorsal view)
|
||
<emphasis bold="true" pageId="0" pageNumber="33">B</emphasis>
|
||
paratype female, habitus (dorsal view). Scale bars: 0.50 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1139.96639.figure20" httpUri="https://binary.pensoft.net/fig/793058" pageId="0" pageNumber="33" start="Figure 20" startId="F20">
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Figure 20.</emphasis>
|
||
<taxonomicName authorityName="Gao & Hou & Zhang" authorityYear="2023" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" kingdom="Animalia" lsidName="Spelaeochthonius wulibeiensis" order="Pseudoscorpiones" pageId="0" pageNumber="33" rank="species" species="wulibeiensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Spelaeochthonius wulibeiensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male
|
||
<emphasis bold="true" pageId="0" pageNumber="33">A</emphasis>
|
||
left chela (lateral view)
|
||
<emphasis bold="true" pageId="0" pageNumber="33">B</emphasis>
|
||
carapace (dorsal view)
|
||
<emphasis bold="true" pageId="0" pageNumber="33">C</emphasis>
|
||
left chelicera (dorsal view). Scale bars: 0.20 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="33">A</emphasis>
|
||
); 0.10 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="33">B, C</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1139.96639.figure21" httpUri="https://binary.pensoft.net/fig/793059" pageId="0" pageNumber="33" start="Figure 21" startId="F21">
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Figure 21.</emphasis>
|
||
<taxonomicName authorityName="Gao & Hou & Zhang" authorityYear="2023" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" kingdom="Animalia" lsidName="Spelaeochthonius wulibeiensis" order="Pseudoscorpiones" pageId="0" pageNumber="33" rank="species" species="wulibeiensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Spelaeochthonius wulibeiensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male (
|
||
<emphasis bold="true" pageId="0" pageNumber="33">A, B</emphasis>
|
||
), paratype female (
|
||
<emphasis bold="true" pageId="0" pageNumber="33">C</emphasis>
|
||
)
|
||
<emphasis bold="true" pageId="0" pageNumber="33">A</emphasis>
|
||
coxae (ventral view)
|
||
<emphasis bold="true" pageId="0" pageNumber="33">B</emphasis>
|
||
male genital area (ventral view)
|
||
<emphasis bold="true" pageId="0" pageNumber="33">C</emphasis>
|
||
female genital area (ventral view). Scale bars: 0.20 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="33">A</emphasis>
|
||
); 0.10 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="33">B, C</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1139.96639.figure22" httpUri="https://binary.pensoft.net/fig/793060" pageId="0" pageNumber="33" start="Figure 22" startId="F22">
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Figure 22.</emphasis>
|
||
<taxonomicName authorityName="Gao & Hou & Zhang" authorityYear="2023" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" kingdom="Animalia" lsidName="Spelaeochthonius wulibeiensis" order="Pseudoscorpiones" pageId="0" pageNumber="33" rank="species" species="wulibeiensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Spelaeochthonius wulibeiensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male
|
||
<emphasis bold="true" pageId="0" pageNumber="33">A</emphasis>
|
||
carapace (dorsal view), with a detail of anterior margin
|
||
<emphasis bold="true" pageId="0" pageNumber="33">B</emphasis>
|
||
left chelicera (dorsal view), with details of teeth
|
||
<emphasis bold="true" pageId="0" pageNumber="33">C</emphasis>
|
||
coxal spines on coxae I (ventral view)
|
||
<emphasis bold="true" pageId="0" pageNumber="33">D</emphasis>
|
||
left pedipalp (minus chela, dorsal view)
|
||
<emphasis bold="true" pageId="0" pageNumber="33">E</emphasis>
|
||
rallum. Scale bars: 0.20 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="33">A, B, D</emphasis>
|
||
); 0.10 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="33">C, E</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1139.96639.figure23" httpUri="https://binary.pensoft.net/fig/793061" pageId="0" pageNumber="33" start="Figure 23" startId="F23">
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Figure 23.</emphasis>
|
||
<taxonomicName authorityName="Gao & Hou & Zhang" authorityYear="2023" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" kingdom="Animalia" lsidName="Spelaeochthonius wulibeiensis" order="Pseudoscorpiones" pageId="0" pageNumber="33" rank="species" species="wulibeiensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Spelaeochthonius wulibeiensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male
|
||
<emphasis bold="true" pageId="0" pageNumber="33">A</emphasis>
|
||
left chela (lateral view), with details of teeth and trichobothrial pattern
|
||
<emphasis bold="true" pageId="0" pageNumber="33">B</emphasis>
|
||
left chela (dorsal view)
|
||
<emphasis bold="true" pageId="0" pageNumber="33">C</emphasis>
|
||
leg I (lateral view)
|
||
<emphasis bold="true" pageId="0" pageNumber="33">D</emphasis>
|
||
leg IV (lateral view). Scale bars: 0.20 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Adult female</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Spelaeochthonius wulibeiensis sp. nov. A holotype male, habitus (minus left chelicera, pedipalp, legs I and IV, dorsal view) B paratype female, habitus (dorsal view). Scale bars: 0.50 mm." figureDoi="10.3897/zookeys.1139.96639.figure19" httpUri="https://binary.pensoft.net/fig/793057" pageId="0" pageNumber="33">19B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Spelaeochthonius wulibeiensis sp. nov., holotype male (A, B), paratype female (C) A coxae (ventral view) B male genital area (ventral view) C female genital area (ventral view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure21" httpUri="https://binary.pensoft.net/fig/793059" pageId="0" pageNumber="33">21C</figureCitation>
|
||
). Mostly same as males; tergal chaetotaxy I-XII: 2: 4: 6: 6: 6: 7: 8: 9: 7: 4: TT: 0; sternal chaetotaxy IV-XII: 10: 13: 11: 12: 12: 11: 8: 0: 2; anterior genital operculum with seven setae, posterior margin with 11 marginal setae, 18 in total; leg IV with a long tactile seta on both tarsal segments: basitarsus 3.60
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than deep (TS = 0.25), telotarsus 11.63
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than deep and 2.58
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than basitarsus (TS = 0.24). Body length 1.93. Pedipalps: trochanter 0.32/0.19 (1.68
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), femur 1.28/0.20 (6.40
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), patella 0.51/0.21 (2.43
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), chela 1.76/0.31 (5.68
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), hand 0.66/0.31 (2.13
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), movable chelal finger length 1.11. Chelicera 0.70/0.30 (2.33
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), movable finger length 0.38. Carapace 0.68/0.68 (1.00
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
). Leg I: trochanter 0.24/0.19 (1.26
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), femur 0.77/0.12 (6.42
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), patella 0.45/0.11 (4.09
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), tibia 0.36/0.08 (4.50
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), tarsus 0.83/0.08 (10.38
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
). Leg IV: trochanter 0.35/0.22 (1.59
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), femoropatella 1.05/0.27 (3.89
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), tibia 0.80/0.14 (5.71
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), basitarsus 0.36/0.10 (3.60
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), telotarsus 0.93/0.08 (11.63
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="33" type="remarks">
|
||
<paragraph pageId="0" pageNumber="33">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
The new species shares similar characters with most species of
|
||
<taxonomicName authorityName="Beier" authorityYear="1931" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Centrochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Centrochthonius" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Centrochthonius</emphasis>
|
||
</taxonomicName>
|
||
Beier, 1931,
|
||
<taxonomicName authorityName="Morikawa" authorityYear="1954" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" kingdom="Animalia" lsidName="Spelaeochthonius" order="Pseudoscorpiones" pageId="0" pageNumber="33" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Spelaeochthonius</emphasis>
|
||
</taxonomicName>
|
||
and all species of "
|
||
<taxonomicName authorityName="Beier" authorityYear="1930" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Pseudotyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudotyrannochthonius" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Pseudotyrannochthonius</emphasis>
|
||
</taxonomicName>
|
||
" Beier, 1930 from the western US by the presence of only 16 setae on the carapace.
|
||
<bibRefCitation DOI="https://doi.org/10.1007/s12542-021-00565-8" author="Schwarze, D" journalOrPublisher="PalZ" pageId="0" pageNumber="33" pagination="11 - 27" refId="B29" refString="Schwarze, D, Harms, D, Hammel, J, Kotthoff, U, 2021. The first fossils of the most basal pseudoscorpion family (Arachnida: Pseudoscorpiones: Pseudotyrannochthoniidae): Evidence for major biogeographical shifts in the European paleofauna. PalZ 96 (1): 11 - 27, DOI: https://doi.org/10.1007/s12542-021-00565-8" title="The first fossils of the most basal pseudoscorpion family (Arachnida: Pseudoscorpiones: Pseudotyrannochthoniidae): Evidence for major biogeographical shifts in the European paleofauna." url="https://doi.org/10.1007/s12542-021-00565-8" volume="96" year="2021">Schwarze et al. (2021)</bibRefCitation>
|
||
emphasized the importance of the number of carapaceal setae in Holarctic pseudotyrannochthoniids, thus, it indicates that the three "
|
||
<taxonomicName authorityName="Beier" authorityYear="1930" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Pseudotyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudotyrannochthonius" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Pseudotyrannochthonius</emphasis>
|
||
</taxonomicName>
|
||
" species in the western US were misclassified in comparison with the twelve species from Australia and the three species from Chile (including the type species, the number of carapaceal setae is more than 18). It can be said that the genus
|
||
<taxonomicName authorityName="Beier" authorityYear="1930" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Pseudotyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudotyrannochthonius" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Pseudotyrannochthonius</emphasis>
|
||
</taxonomicName>
|
||
is endemic to the southern hemisphere (
|
||
<bibRefCitation DOI="https://doi.org/10.1636/JoA-S-21-033" author="Harvey, MS" journalOrPublisher="The Journal of Arachnology" pageId="0" pageNumber="33" pagination="158 - 174" refId="B9" refString="Harvey, MS, Harms, D, 2022. The pseudoscorpion genus Centrochthonius (Pseudoscorpiones: Pseudotyrannochthoniidae) from central Asia and description of a new species from Nepal. The Journal of Arachnology 50 (2): 158 - 174, DOI: https://doi.org/10.1636/JoA-S-21-033" title="The pseudoscorpion genus Centrochthonius (Pseudoscorpiones: Pseudotyrannochthoniidae) from central Asia and description of a new species from Nepal." url="https://doi.org/10.1636/JoA-S-21-033" volume="50" year="2022">Harvey and Harms 2022</bibRefCitation>
|
||
). Thus, it is inappropriate to place this new species in
|
||
<taxonomicName authorityName="Beier" authorityYear="1930" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Pseudotyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudotyrannochthonius" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Pseudotyrannochthonius</emphasis>
|
||
</taxonomicName>
|
||
, even though the shape of this new species of coxal spines is similar to that of the three "
|
||
<taxonomicName authorityName="Beier" authorityYear="1930" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Pseudotyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudotyrannochthonius" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Pseudotyrannochthonius</emphasis>
|
||
</taxonomicName>
|
||
" species.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
The shape and number of the coxal spines are important distinguishing features between
|
||
<taxonomicName authorityName="Beier" authorityYear="1931" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Centrochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Centrochthonius" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Centrochthonius</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Morikawa" authorityYear="1954" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" kingdom="Animalia" lsidName="Spelaeochthonius" order="Pseudoscorpiones" pageId="0" pageNumber="33" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Spelaeochthonius</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation DOI="https://doi.org/10.1636/JoA-S-21-025" author="You, J" journalOrPublisher="The Journal of Arachnology" pageId="0" pageNumber="33" pagination="135 - 157" refId="B33" refString="You, J, Yoo, JS, Harvey, MS, Harms, D, 2022. Some cryptic Korean karst creatures: revalidation of the pseudoscorpion genus Spelaeochthonius (Pseudoscorpiones: Pseudotyrannochthoniidae) and description of two new species from Korea. The Journal of Arachnology 50 (2): 135 - 157, DOI: https://doi.org/10.1636/JoA-S-21-025" title="Some cryptic Korean karst creatures: revalidation of the pseudoscorpion genus Spelaeochthonius (Pseudoscorpiones: Pseudotyrannochthoniidae) and description of two new species from Korea." url="https://doi.org/10.1636/JoA-S-21-025" volume="50" year="2022">You et al. 2022</bibRefCitation>
|
||
). In our opinion, it is appropriate to place this new species to
|
||
<taxonomicName authorityName="Morikawa" authorityYear="1954" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" kingdom="Animalia" lsidName="Spelaeochthonius" order="Pseudoscorpiones" pageId="0" pageNumber="33" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Spelaeochthonius</emphasis>
|
||
</taxonomicName>
|
||
rather than
|
||
<taxonomicName authorityName="Beier" authorityYear="1931" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Centrochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Centrochthonius" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Centrochthonius</emphasis>
|
||
</taxonomicName>
|
||
, the reasons are as follows: for
|
||
<taxonomicName authorityName="Beier" authorityYear="1931" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Centrochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Centrochthonius" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Centrochthonius</emphasis>
|
||
</taxonomicName>
|
||
, the number of carapaceal setae is not fixed (e.g., occasionally 18 are present in
|
||
<taxonomicName lsidName="C. anatonus" pageId="0" pageNumber="33" rank="species" species="anatonus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">C. anatonus</emphasis>
|
||
</taxonomicName>
|
||
Harvey & Harms, 2022) and only four or five coxal spines blades; for
|
||
<taxonomicName authorityName="Morikawa" authorityYear="1954" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" kingdom="Animalia" lsidName="Spelaeochthonius" order="Pseudoscorpiones" pageId="0" pageNumber="33" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Spelaeochthonius</emphasis>
|
||
</taxonomicName>
|
||
, the character of coxal spines is diverse (e.g., in
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. undecimclavatus" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="undecimclavatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">S. undecimclavatus</emphasis>
|
||
</taxonomicName>
|
||
Morikawa, 1956, which is club-shaped, not distally plumose).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<taxonomicName authorityName="Gao & Hou & Zhang" authorityYear="2023" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" kingdom="Animalia" lsidName="Spelaeochthonius wulibeiensis" order="Pseudoscorpiones" pageId="0" pageNumber="33" rank="species" species="wulibeiensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Spelaeochthonius wulibeiensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is similar to
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. cheonsooi" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="cheonsooi">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">S. cheonsooi</emphasis>
|
||
</taxonomicName>
|
||
You, Yoo, Harvey & Harms, 2022, but differs by the number of setae on tergite I (2 vs. 4) and larger body size (body length 1.93 (♀) mm vs. 1.70 (♀) mm; chela 5.68 (♀)
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
vs. 5.32 (♀)
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than board, length 1.76 (♀) mm vs. 1.49 (♀) mm).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<taxonomicName authorityName="Gao & Hou & Zhang" authorityYear="2023" family="Pseudotyrannochthoniidae" genus="Spelaeochthonius" kingdom="Animalia" lsidName="Spelaeochthonius wulibeiensis" order="Pseudoscorpiones" pageId="0" pageNumber="33" rank="species" species="wulibeiensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Spelaeochthonius wulibeiensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be distinguished from
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. seungsookae" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="seungsookae">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">S. seungsookae</emphasis>
|
||
</taxonomicName>
|
||
You, Yoo, Harvey & Harms, 2022 by the number of setae on tergite I (2 vs. 4) and smaller body size (body length 1.93 (♀) mm vs. 2.05-2.36 (♀) mm; chela length 1.68-1.74 (♂), 1.76 (♀) mm vs. 1.90 (♂), 1.92 (♀) mm); from
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. undecimclavatus" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="undecimclavatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">S. undecimclavatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. dorogawaensis" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="dorogawaensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">S. dorogawaensis</emphasis>
|
||
</taxonomicName>
|
||
by the number of setae on chelicera (6 vs. 7), a slender palp (palpal femur 7.24 (♂)
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
vs. 4.80-5.40 (♂)
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than board; chela 6.21-6.22 (♂)
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
vs. 5.50-6.13 (♂)
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than board) and lower number of blades of coxal spines (7-8 vs. 10-11); from
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. akiyoshiensis" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="akiyoshiensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">S. akiyoshiensis</emphasis>
|
||
</taxonomicName>
|
||
Morikawa, 1956 and
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. kobayashii" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="kobayashii">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">S. kobayashii</emphasis>
|
||
</taxonomicName>
|
||
Morikawa, 1956 by the number of setae on chelicera (6 vs. 7), lower number of movable chelal finger teeth (16-19 teeth vs. min. 26 teeth) and a slender palp (palpal femur 7.24 (♂)
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
vs. 5.00-5.60 (♂)
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than board; chela 6.21-6.22 (♂)
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
vs. 5.13-5.74 (♂)
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than board); from
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. dentifer" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="dentifer">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">S. dentifer</emphasis>
|
||
</taxonomicName>
|
||
(Morikawa, 1970) by the number of setae on chelicera (6 vs. 7), lower number of movable chelal finger teeth (16-19 teeth vs. min. 36 teeth) and a shorter chela (chela 6.21-6.22 (♂), 5.68 (♀)
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
vs. 6.85 (♂), 7.12 (♀)
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than board, length 1.68-1.74 (♂), 1.76 (♀) mm vs. 1.85 (♂♀) mm); from
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. kubotai" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="kubotai">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">S. kubotai</emphasis>
|
||
</taxonomicName>
|
||
by the slightly smaller body size (body length 1.93 (♀) mm vs. 2.03 (♀) mm; chela 5.68 (♀)
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
vs. 5.70 (♀)
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than board; movable chelal finger 1.68 (♀)
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
vs. 1.87 (♀)
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than) and the number of setae of coxal spines (7 or 8 vs. 11); from
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="S. kishidai" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="kishidai">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">S. kishidai</emphasis>
|
||
</taxonomicName>
|
||
(Morikawa, 1960) by a slender palp (palpal femur 7.24
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
vs. 4.90
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than board; movable chelal finger 0.61-0.63
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
vs. 0.67-0.69
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than board) (
|
||
<bibRefCitation author="Morikawa, K" journalOrPublisher="Revue Suisse de Zoologie" pageId="0" pageNumber="33" refId="B24" refString="Morikawa, K, 1954. On some pseudoscorpions in Japanese lime-grottoes. Memoirs of Ehime University 2(2B): 79-87." title="On some pseudoscorpions in Japanese lime-grottoes. Memoirs of Ehime University 2 (2 B): 79 - 87." year="1954">Morikawa 1954</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Morikawa, K" journalOrPublisher="Revue Suisse de Zoologie" pageId="0" pageNumber="33" refId="B25" refString="Morikawa, K, 1956. Cave pseudoscorpions of Japan (I). Memoirs of Ehime University 2(2B): 271-282." title="Cave pseudoscorpions of Japan (I). Memoirs of Ehime University 2 (2 B): 271 - 282." year="1956">1956</bibRefCitation>
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,
|
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<bibRefCitation author="Morikawa, K" journalOrPublisher="Revue Suisse de Zoologie" pageId="0" pageNumber="33" refId="B26" refString="Morikawa, K, 1960. Systematics studies of Japanese pseudoscorpions. Memoirs of Ehime University 2(2B): 85-172." title="Systematics studies of Japanese pseudoscorpions. Memoirs of Ehime University 2 (2 B): 85 - 172." year="1960">1960</bibRefCitation>
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,
|
||
<bibRefCitation author="Morikawa, K" journalOrPublisher="Bulletin of the National Science Museum, Tokyo" pageId="0" pageNumber="33" pagination="141 - 148" refId="B27" refString="Morikawa, K, 1970. Results of the speleological survey in South Korea 1966. XX. New pseudoscorpions from South Korea. Bulletin of the National Science Museum, Tokyo 13: 141 - 148" title="Results of the speleological survey in South Korea 1966. XX. New pseudoscorpions from South Korea." volume="13" year="1970">1970</bibRefCitation>
|
||
;
|
||
<bibRefCitation DOI="https://doi.org/10.1636/JoA-S-21-025" author="You, J" journalOrPublisher="The Journal of Arachnology" pageId="0" pageNumber="33" pagination="135 - 157" refId="B33" refString="You, J, Yoo, JS, Harvey, MS, Harms, D, 2022. Some cryptic Korean karst creatures: revalidation of the pseudoscorpion genus Spelaeochthonius (Pseudoscorpiones: Pseudotyrannochthoniidae) and description of two new species from Korea. The Journal of Arachnology 50 (2): 135 - 157, DOI: https://doi.org/10.1636/JoA-S-21-025" title="Some cryptic Korean karst creatures: revalidation of the pseudoscorpion genus Spelaeochthonius (Pseudoscorpiones: Pseudotyrannochthoniidae) and description of two new species from Korea." url="https://doi.org/10.1636/JoA-S-21-025" volume="50" year="2022">You et al. 2022</bibRefCitation>
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).
|
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</paragraph>
|
||
</subSubSection>
|
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<subSubSection pageId="0" pageNumber="33" type="distribution">
|
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<paragraph pageId="0" pageNumber="33">Distribution and habitat.</paragraph>
|
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<paragraph pageId="0" pageNumber="33">
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This species is known only from the type locality, Wulibei Cave (Figs
|
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Study area with indication of cave locations representing the type localities. Each color represents an administrative region (red: Guizhou Province; purple: Weining County; green: Xishui County; blue: Fenggang County) A Wulibei Cave (Spelaeochthonius wulibeiensis sp. nov.) B Liangfeng Cave (Allochthonius bainiensis sp. nov.) C Daozuo Cave (A. pandus sp. nov.) D Sanjie Cave (A. xinqiaoensis sp. nov.)." figureDoi="10.3897/zookeys.1139.96639.figure1" httpUri="https://binary.pensoft.net/fig/793039" pageId="0" pageNumber="33">1A</figureCitation>
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,
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<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Wulibei Cave, type locality of Spelaeochthonius wulibeiensis sp. nov. A cave location (red arrow) B entrance C inside the cave entrance D live male of S. wulibeiensis sp. nov. in its natural environment." figureDoi="10.3897/zookeys.1139.96639.figure18" httpUri="https://binary.pensoft.net/fig/793056" pageId="0" pageNumber="33">18A-C</figureCitation>
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), which is located ~ 1.2 km east of Yangguan Village (Weining County). This limestone cave has an elongated entrance (~ 2.5 m high and 8 m wide) with some corn stalks scattered nearby. Entrance of the cave has a large muddy cave hall, connected to a small hall through a narrow tunnel, which is a more enclosed, completely dark space, covered with gravel, with temperatures ~ 10 °C and humidity ~ 90%. The specimen was collected under a stone in a small cave hall.
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</paragraph>
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</subSubSection>
|
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</treatment>
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</subSection>
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</document> |