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<mods:title id="5F118FFB27EE60469AA3365B7CAC172D">Cirsium paludigenum and C. zhejiangense (Asteraceae) are synonymous with C. sieboldii and C. yezoense</mods:title>
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2. 
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(Maximowicz) 
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— 
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s.n.
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(
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Z.H.Chen &amp; X.F. 
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X. 
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. 
<materialsCitation id="8084F971FF95FFE84E6D834B5405805A" ID-GBIF-Occurrence="3750509337" box="[1357,1371,1642,1663]" collectionCode="T" country="China" isTailEnumeration="true" location="Jin" pageId="9" pageNumber="60" specimenCount="1" stateProvince="Zhejiang">
<collectionCode id="56FD6BE9FF95FFE84E6D834B5405805A" box="[1357,1371,1642,1663]" name="Tavera, Department of Geology and Geophysics" pageId="9" pageNumber="60">T</collectionCode>
</materialsCitation>
.
</emphasis>
<materialsCitation id="8084F971FF95FFE84E7E834B53FA8086" ID-GBIF-Occurrence="3750509386" collectionCode="ZM, HTC, PE, A, B" country="China" location="Yu" pageId="9" pageNumber="60" specimenCount="1" stateProvince="Zhejiang" typeStatus="holotype">
<emphasis id="02982F3EFF95FFE84E7E834B54F5805A" box="[1374,1451,1642,1663]" italics="true" pageId="9" pageNumber="60">
<location id="3533A5F7FF95FFE84E7E834B5428805A" LSID="urn:lsid:plazi:treatment:B845423AFF95FFEA4BA880F752968725:3533A5F7FF95FFE84E7E834B5428805A" box="[1374,1398,1642,1663]" country="China" name="Yu" pageId="9" pageNumber="60" stateProvince="Zhejiang">Yu</location>
3202
</emphasis>
(
<typeStatus id="EF574D8EFF95FFE84BE483AC50428086" box="[196,284,1677,1699]" pageId="9" pageNumber="60" type="holotype">holotype</typeStatus>
, 
<collectionCode id="56FD6BE9FF95FFE84A0583AF500E8086" box="[293,336,1678,1699]" country="China" lsid="urn:lsid:biocol.org:col:14833" name="Zhejiang Museum of Natural History" pageId="9" pageNumber="60" type="Herbarium">ZM</collectionCode>
!; isotypes, 
<collectionCode id="56FD6BE9FF95FFE84A9983AF50B58086" box="[441,491,1678,1699]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13187" lsid="urn:lsid:biocol.org:col:13187" name="Hangzhou Normal College" pageId="9" pageNumber="60" type="Herbarium">HTC</collectionCode>
, 
<collectionCode id="56FD6BE9FF95FFE84AD583AF53468081" box="[501,536,1678,1700]" country="China" lsid="urn:lsid:biocol.org:col:15858" name="Institute of Botany, Chinese Academy of Sciences" pageId="9" pageNumber="60" type="Herbarium">PE</collectionCode>
!). 
<figureCitation id="A8D7EFA9FF95FFE8490D83AC533B8086" box="[557,613,1677,1699]" captionStart="FIGURE 6" captionStartId="7.[136,229,1945,1967]" captionTargetBox="[183,1405,186,1925]" captionTargetId="figure-17@7.[183,1405,186,1925]" captionTargetPageId="7" captionText="FIGURE 6. Specimens of Cirsium yezoense. A, B, D. China, Zhejiang, Pan’an, Weixin, Lingjiangyuan, by stream under forest, X.F.Jin &amp; T.T.Yu 3202 (holotype of Cirsium zhejiangense, ZM; isotypes of Cirsium zhejiangense, PE; D. capitulum); C, E. Japan, Hakodate, Mohidzi, C.J.Maximowicz s.n. (type of Cirsium yezoense, K000778217; E. capitulum)." figureDoi="http://doi.org/10.5281/zenodo.6424362" httpUri="https://zenodo.org/record/6424362/files/figure.png" pageId="9" pageNumber="60">Fig. 6</figureCitation>
(
<collectionCode id="56FD6BE9FF95FFE8494C83AC53DE8086" box="[620,640,1677,1699]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="9" pageNumber="60" type="Herbarium">A</collectionCode>
, 
<collectionCode id="56FD6BE9FF95FFE849AA83AF53C28086" box="[650,668,1678,1699]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="9" pageNumber="60" type="Herbarium">B</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="78F6A0A7FF95FFEB4BA883F3534F8791" lastPageId="10" lastPageNumber="61" pageId="9" pageNumber="60" type="description">
<paragraph id="3053F32CFF95FFEB4BA883F3534F8791" blockId="9.[136,1452,1746,2061]" lastBlockId="10.[136,1452,158,2093]" lastPageId="10" lastPageNumber="61" pageId="9" pageNumber="60">
<emphasis id="02982F3EFF95FFE84BA883F3504280C9" bold="true" box="[136,284,1746,1772]" pageId="9" pageNumber="60">Description:</emphasis>
—Herbs 
<quantity id="F7145EC9FF95FFE84AA483F3535C80C9" box="[388,514,1746,1772]" metricMagnitude="-1" metricUnit="m" metricValue="7.0" metricValueMax="11.0" metricValueMin="3.0" pageId="9" pageNumber="60" unit="cm" value="70.0" valueMax="110.0" valueMin="30.0">30–110 cm</quantity>
tall, perennial. Roots fleshy. Stems, erect, unbranched or branched above, ribbed, unwinged, with sparse brown hairs and dense tomentose under capitula. Leaves concolorous, green. Basal leaves present at anthesis, rosulate, with petiole, 
<quantity id="F7145EC9FF95FFE84951823B53878111" box="[625,729,1818,1844]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" metricValueMax="10.0" metricValueMin="4.0" pageId="9" pageNumber="60" unit="cm" value="7.0" valueMax="10.0" valueMin="4.0">4–10 cm</quantity>
; leaf blade ovate or elliptic, 25–50 × 
<quantity id="F7145EC9FF95FFE84FB5823B54558111" box="[1173,1291,1818,1844]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="9" pageNumber="60" unit="cm" value="20.0" valueMax="30.0" valueMin="10.0">10–30 cm</quantity>
, pinnatilobate or pinnatifid, adaxailly green, with short brown hairs or glabrous, abaxially light green, glabrous or with sparsely arachnoid; segments 4–7 pairs, ovate-lanceolate, elliptic or triangular, toothed, teeth with marginal and 
<quantity id="F7145EC9FF95FFE84E268243543D8159" box="[1286,1379,1890,1916]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="9" pageNumber="60" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">1–3 mm</quantity>
apical spines; terminal segment largest, triangular. Cauline leaves, sessile, ovate, elliptic or oblong-elliptic, 5–20 × 
<quantity id="F7145EC9FF95FFE84E6382A754F98185" box="[1347,1447,1926,1952]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="12.0" metricValueMin="3.0" pageId="9" pageNumber="60" unit="cm" value="7.5" valueMax="12.0" valueMin="3.0">3–12 cm</quantity>
, pinnatilobate or pinnatifid; segments 4–6 pairs, semielliptic, teeth with a ca. 
<quantity id="F7145EC9FF95FFE848C4828B557B81E1" box="[996,1061,1962,1988]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="60" unit="mm" value="3.0">3 mm</quantity>
apical spine, adaxailly green, with short brown hairs or glabrous, abaxially light green, glabrous or with sparsely arachnoid; gradually smaller upward. Capitula solitary to many, terminal, nodding in flower, erect in fruit. Involucre campanulate, 
<quantity id="F7145EC9FF95FFE84F9782D354528E29" box="[1207,1292,2034,2060]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="9" pageNumber="60" unit="cm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 cm</quantity>
(in vivo) and 
<quantity id="F7145EC9FF96FFEB4BA885BF51AA869C" box="[136,244,158,185]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" metricValueMax="4.5" metricValueMin="3.0" pageId="10" pageNumber="61" unit="cm" value="3.75" valueMax="4.5" valueMin="3.0">3–4.5 cm</quantity>
(in sicco) in diam., 
<quantity id="F7145EC9FF96FFEB4AC785BF5360869C" box="[487,574,158,185]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="10" pageNumber="61" unit="cm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 cm</quantity>
high, glabrous or with sparsely arachnoid. Peduncles 
<quantity id="F7145EC9FF96FFEB4F9985BF547E869C" box="[1209,1312,158,185]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="10" pageNumber="61" unit="cm" value="7.5" valueMax="10.0" valueMin="5.0">5–10 cm</quantity>
, with dense tomentose. Phyllaries imbricate, patent, in 5–10 rows, lacking marginal spinules, wings, and scarious appendage; outer and middle phyllaries, ovate or ovate-lanceolate, with acuminate tips, patent, 5–13 × 
<quantity id="F7145EC9FF96FFEB4F5D85C755AF8724" box="[1149,1265,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="10" pageNumber="61" unit="mm" value="3.25" valueMax="3.5" valueMin="3.0">3–3.5 mm</quantity>
, narrowed into a 
<quantity id="F7145EC9FF96FFEB4BA8842B51BB8700" box="[136,229,266,293]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="10" pageNumber="61" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
spine; inner phyllaries lanceolate to linear-lanceolate, 1.5–2 × 
<quantity id="F7145EC9FF96FFEB4882842B557F8700" box="[930,1057,266,293]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="10" pageNumber="61" unit="cm" value="0.25" valueMax="0.3" valueMin="0.2">0.2–0.3 cm</quantity>
, apex purplish pink, acuminate and softly spiny. Florets bisexual. Corolla purple, ca. 
<quantity id="F7145EC9FF96FFEB4986840F53AA876C" box="[678,756,302,329]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="10" pageNumber="61" unit="cm" value="2.8">2.8 cm</quantity>
, tube ca. 
<quantity id="F7145EC9FF96FFEB487A840F52F9876C" box="[858,935,302,329]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="10" pageNumber="61" unit="cm" value="1.3">1.3 cm</quantity>
, limb ca. 
<quantity id="F7145EC9FF96FFEB4F30840F5505876C" box="[1040,1115,302,329]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="61" unit="cm" value="1.5">1.5 cm</quantity>
long, lobes 5, linear, ca. 
<quantity id="F7145EC9FF96FFEB4E4C840F54F2876C" box="[1388,1452,302,329]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="61" unit="mm" value="6.0">6 mm</quantity>
long. Anther purple, ca. 
<quantity id="F7145EC9FF96FFEB4AB8847350878748" box="[408,473,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="61" unit="mm" value="9.0">9 mm</quantity>
long including appendage. Achenes cylindric, laterally compressed, ca. 
<quantity id="F7145EC9FF96FFEB4E248473541B8748" box="[1284,1349,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="61" unit="mm" value="5.0">5 mm</quantity>
long, ca. 
<quantity id="F7145EC9FF96FFEB4BA88457518087B4" box="[136,222,374,401]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="10" pageNumber="61" unit="mm" value="1.7">1.7 mm</quantity>
wide, smooth, indistinctly longitudinally ribbed, apical rim forming a crown. Pappus of 3–4 rows of plumose bristles, pale brown, ca. 
<quantity id="F7145EC9FF96FFEB4AB984BB508E8790" box="[409,464,410,437]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="61" unit="cm" value="2.0">2 cm</quantity>
long.
</paragraph>
</subSubSection>
<subSubSection id="78F6A0A7FF96FFEB4B9D849F52E787FD" box="[189,953,446,473]" pageId="10" pageNumber="61" type="biology_ecology">
<paragraph id="3053F32CFF96FFEB4B9D849F52E787FD" blockId="10.[136,1452,158,2093]" box="[189,953,446,473]" pageId="10" pageNumber="61">
<emphasis id="02982F3EFF96FFEB4B9D849F501D87FD" bold="true" box="[189,323,446,472]" pageId="10" pageNumber="61">Phenology:</emphasis>
—Flowering and fruiting from September to November.
</paragraph>
</subSubSection>
<subSubSection id="78F6A0A7FF96FFEB4B9D84C3521E84A8" pageId="10" pageNumber="61" type="distribution">
<paragraph id="3053F32CFF96FFEB4B9D84C3521E84A8" blockId="10.[136,1452,158,2093]" pageId="10" pageNumber="61">
<emphasis id="02982F3EFF96FFEB4B9D84C350AA87D9" bold="true" box="[189,500,482,508]" pageId="10" pageNumber="61">Distribution and habitat:</emphasis>
— 
<taxonomicName id="F7EC88AFFF96FFEB493084C2538587D9" authorityName="Makino" authorityYear="1905" baseAuthorityName="Maximowicz" baseAuthorityYear="1874" box="[528,731,483,509]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="61" phylum="Tracheophyta" rank="species" species="yezoense">
<emphasis id="02982F3EFF96FFEB493084C2538587D9" box="[528,731,483,509]" italics="true" pageId="10" pageNumber="61">Cirsium yezoense</emphasis>
</taxonomicName>
is distributed in 
<collectingRegion id="F2283DCEFF96FFEB488A84C352AD87D9" box="[938,1011,482,508]" country="China" name="Anhui" pageId="10" pageNumber="61">Anhui</collectingRegion>
, 
<collectingRegion id="F2283DCEFF96FFEB4F2484C3553987D9" box="[1028,1127,482,508]" country="China" name="Guizhou" pageId="10" pageNumber="61">Guizhou</collectingRegion>
, Hunnan, 
<collectingRegion id="F2283DCEFF96FFEB4FC484C3546687D9" box="[1252,1336,482,508]" country="China" name="Jiangxi" pageId="10" pageNumber="61">Jiangxi</collectingRegion>
, 
<collectingRegion id="F2283DCEFF96FFEB4E6884C351B48405" country="China" name="Zhejiang" pageId="10" pageNumber="61">Zhejiang province</collectingRegion>
of 
<collectingCountry id="48FBB3BCFF96FFEB4A0B8727502C8405" box="[299,370,518,544]" name="China" pageId="10" pageNumber="61">China</collectingCountry>
. In the website of Chinese Field Herbarium (https://www.cfh.ac.cn/album/ShowPhoto. aspx?photoid=6b60814b-3301-4dc1-b2d3-70550ec479d4) and Plant Photo Bank of 
<collectingCountry id="48FBB3BCFF96FFEB4F47870B55F48461" box="[1127,1194,554,580]" name="China" pageId="10" pageNumber="61">China</collectingCountry>
(http://ppbc.iplant.cn/ tu/7001749), we also find a similar plant from Yuexi county and Chizhou county, 
<collectingRegion id="F2283DCEFF96FFEB4F07876F5587844D" box="[1063,1241,590,616]" country="China" name="Anhui" pageId="10" pageNumber="61">Anhui Province</collectingRegion>
(
<figureCitation id="A8D7EFA9FF96FFEB4FC9876F5470844D" box="[1257,1326,590,616]" captionStart="FIGURE 8" captionStartId="9.[136,229,1209,1231]" captionTargetBox="[163,1425,197,1177]" captionTargetId="figure-17@9.[151,1436,186,1189]" captionTargetPageId="9" captionText="FIGURE 8. Distribution map of Cirsium sieboldii (red circles) and Cirsium yezoense (blue diamonds) in China." figureDoi="http://doi.org/10.5281/zenodo.6424366" httpUri="https://zenodo.org/record/6424366/files/figure.png" pageId="10" pageNumber="61">Fig. 8</figureCitation>
). It mainly grows on the forest margins at altitudes of about 
<quantity id="F7145EC9FF96FFEB498C8753526284A8" box="[684,828,626,653]" metricMagnitude="2" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="10" pageNumber="61" unit="m" value="750.0" valueMax="1000.0" valueMin="500.0">500–1000 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="78F6A0A7FF96FFEB4B9D87B7541F828C" pageId="10" pageNumber="61" type="materials_examined">
<paragraph id="3053F32CFF96FFEB4B9D87B7541F828C" blockId="10.[136,1452,158,2093]" pageId="10" pageNumber="61">
<emphasis id="02982F3EFF96FFEB4B9D87B753098495" bold="true" box="[189,599,662,688]" pageId="10" pageNumber="61">Additional specimens examined:</emphasis>
— 
<materialsCitation id="8084F971FF96FFEB495487B7530184F0" ID-GBIF-Occurrence="3750509378" collectionCode="PE" collectorName="Sino-American Guizhou Botanical Expedition" country="China" location="Guizhou" pageId="10" pageNumber="61" specimenCode="PE00456363" specimenCount="1" stateProvince="Guizhou">
<collectingCountry id="48FBB3BCFF96FFEB495487B753918495" box="[628,719,662,688]" name="China" pageId="10" pageNumber="61">CHINA</collectingCountry>
. 
<emphasis id="02982F3EFF96FFEB49C487B752158495" bold="true" box="[740,843,662,688]" pageId="10" pageNumber="61">
<collectingRegion id="F2283DCEFF96FFEB49C487B752158495" box="[740,843,662,688]" country="China" name="Guizhou" pageId="10" pageNumber="61">Guizhou</collectingRegion>
</emphasis>
: 
<collectorName id="9D1996FAFF96FFEB484587B752938495" box="[869,973,662,688]" pageId="10" pageNumber="61">Jiangkou</collectorName>
, 
<collectorName id="9D1996FAFF96FFEB48C387B755238495" box="[995,1149,662,688]" pageId="10" pageNumber="61">Fangjingshan</collectorName>
, 
<emphasis id="02982F3EFF96FFEB4FB387B650EC84F1" italics="true" pageId="10" pageNumber="61">
<collectorName id="9D1996FAFF96FFEB4FB387B654658494" box="[1171,1339,663,689]" pageId="10" pageNumber="61">Sino-American</collectorName>
<collectingRegion id="F2283DCEFF96FFEB4E6C87B754F58495" box="[1356,1451,662,688]" country="China" name="Guizhou" pageId="10" pageNumber="61">Guizhou</collectingRegion>
<collectorName id="9D1996FAFF96FFEB4BA8879A502A84F1" box="[136,372,698,724]" pageId="10" pageNumber="61">Botanicai Expedition</collectorName>
1040
</emphasis>
(
<specimenCode id="604A5B57FF96FFEB4AE0879B530684F0" box="[448,600,698,725]" collectionCode="PE" country="China" lsid="urn:lsid:biocol.org:col:15858" name="Institute of Botany, Chinese Academy of Sciences" pageId="10" pageNumber="61" type="Herbarium">PE00456363</specimenCode>
)
</materialsCitation>
. 
<materialsCitation id="8084F971FF96FFEB4948879B55FA84F1" ID-GBIF-Occurrence="3750509302" box="[616,1188,698,725]" collectionCode="Z, H" collectorName="Yang" country="China" location="Hengshan" pageId="10" pageNumber="61" specimenCode="PEY0060419" specimenCount="1" stateProvince="Hunan">
<emphasis id="02982F3EFF96FFEB4948879B53E484F1" bold="true" box="[616,698,698,724]" pageId="10" pageNumber="61">
<collectingRegion id="F2283DCEFF96FFEB4948879B53E484F1" box="[616,698,698,724]" country="China" name="Hunan" pageId="10" pageNumber="61">Hunan</collectingRegion>
</emphasis>
: 
<location id="3533A5F7FF96FFEB49E9879B526284F1" LSID="urn:lsid:plazi:treatment:B845423AFF95FFEA4BA880F752968725:3533A5F7FF96FFEB49E9879B526284F1" box="[713,828,698,724]" country="China" name="Hengshan" pageId="10" pageNumber="61" stateProvince="Hunan">Hengshan</location>
, 
<emphasis id="02982F3EFF96FFEB4866879A52E584F1" box="[838,955,698,724]" italics="true" pageId="10" pageNumber="61">
<collectorName id="9D1996FAFF96FFEB4866879A522384F1" box="[838,893,699,724]" pageId="10" pageNumber="61">Yang</collectorName>
5861
</emphasis>
(
<specimenCode id="604A5B57FF96FFEB48E8879B553B84F0" box="[968,1125,698,725]" collectionCode="PEY" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13846" lsid="urn:lsid:biocol.org:col:13846" name="Peking University" pageId="10" pageNumber="61" type="Herbarium">PEY0060419</specimenCode>
), 
<emphasis id="02982F3EFF96FFEB4F57879A55FA84F1" box="[1143,1188,699,724]" italics="true" pageId="10" pageNumber="61">
<collectionCode id="56FD6BE9FF96FFEB4F57879A55D484F1" box="[1143,1162,699,724]" country="Switzerland" name="Universität Zürich" pageId="10" pageNumber="61" type="Herbarium">Z</collectionCode>
. 
<collectionCode id="56FD6BE9FF96FFEB4FAD879A55FA84F1" box="[1165,1188,699,724]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="10" pageNumber="61" type="Herbarium">H</collectionCode>
</emphasis>
</materialsCitation>
<emphasis id="02982F3EFF96FFEB4F84879A55F684F1" box="[1188,1192,699,724]" italics="true" pageId="10" pageNumber="61">.</emphasis>
<materialsCitation id="8084F971FF96FFEB4F88879A51EB84DD" ID-GBIF-Occurrence="3750509385" collectionCode="Z, H" country="China" location="Hu" pageId="10" pageNumber="61" specimenCode="PE01834991" specimenCount="1">
<emphasis id="02982F3EFF96FFEB4F88879A55A584F1" box="[1192,1275,698,724]" italics="true" pageId="10" pageNumber="61">
<location id="3533A5F7FF96FFEB4F88879A559584F1" LSID="urn:lsid:plazi:treatment:B845423AFF95FFEA4BA880F752968725:3533A5F7FF96FFEB4F88879A559584F1" box="[1192,1227,699,724]" country="China" name="Hu" pageId="10" pageNumber="61">Hu</location>
512
</emphasis>
(
<specimenCode id="604A5B57FF96FFEB4E29879B54C184F1" box="[1289,1439,698,724]" collectionCode="PE" country="China" lsid="urn:lsid:biocol.org:col:15858" name="Institute of Botany, Chinese Academy of Sciences" pageId="10" pageNumber="61" type="Herbarium">PE01834991</specimenCode>
), 
<emphasis id="02982F3EFF96FFEB4BA887FE51EB84DD" box="[136,181,735,760]" italics="true" pageId="10" pageNumber="61">
<collectionCode id="56FD6BE9FF96FFEB4BA887FE51C584DD" box="[136,155,735,760]" country="Switzerland" name="Universität Zürich" pageId="10" pageNumber="61" type="Herbarium">Z</collectionCode>
. 
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513
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(
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), 
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. 
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<emphasis id="02982F3EFF96FFEB4AC987FE50B384DC" box="[489,493,735,761]" italics="true" pageId="10" pageNumber="61">.</emphasis>
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HS2021021
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(
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)
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. 
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<emphasis id="02982F3EFF96FFEB49DB87FF520C84DD" bold="true" box="[763,850,734,760]" pageId="10" pageNumber="61">
<collectingRegion id="F2283DCEFF96FFEB49DB87FF520C84DD" box="[763,850,734,760]" country="China" name="Jiangxi" pageId="10" pageNumber="61">Jiangxi</collectingRegion>
</emphasis>
: 
<collectorName id="9D1996FAFF96FFEB487F87FF52E084DD" box="[863,958,734,760]" pageId="10" pageNumber="61">Fengxin</collectorName>
, 
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<emphasis id="02982F3EFF96FFEB48E687FE52A584DC" box="[966,1019,735,761]" italics="true" pageId="10" pageNumber="61">LBG</emphasis>
</collectionCode>
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<emphasis id="02982F3EFF96FFEB4F2087FF55C984DD" box="[1024,1175,734,760]" italics="true" pageId="10" pageNumber="61">
<location id="3533A5F7FF96FFEB4F2087FF551384DD" LSID="urn:lsid:plazi:treatment:B845423AFF95FFEA4BA880F752968725:3533A5F7FF96FFEB4F2087FF551384DD" box="[1024,1101,734,760]" country="China" name="Exped." pageId="10" pageNumber="61">Exped.</location>
00416
</emphasis>
(
<specimenCode id="604A5B57FF96FFEB4F8487FF546584DC" box="[1188,1339,734,761]" collectionCode="PE" country="China" lsid="urn:lsid:biocol.org:col:15858" name="Institute of Botany, Chinese Academy of Sciences" pageId="10" pageNumber="61" type="Herbarium">PE01776765</specimenCode>
); 
<collectorName id="9D1996FAFF96FFEB4E6B87FF54F984DD" box="[1355,1447,734,760]" pageId="10" pageNumber="61">Wuning</collectorName>
, 
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<emphasis id="02982F3EFF96FFEB4BA8862251E38538" box="[136,189,771,797]" italics="true" pageId="10" pageNumber="61">LBG</emphasis>
</collectionCode>
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<emphasis id="02982F3EFF96FFEB4BE68623500D8539" box="[198,339,770,796]" italics="true" pageId="10" pageNumber="61">
<location id="3533A5F7FF96FFEB4BE68623504D8539" LSID="urn:lsid:plazi:treatment:B845423AFF95FFEA4BA880F752968725:3533A5F7FF96FFEB4BE68623504D8539" box="[198,275,770,796]" country="China" name="Exped." pageId="10" pageNumber="61">Exped.</location>
0047
</emphasis>
(
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), 
<emphasis id="02982F3EFF96FFEB492F8622531F8538" box="[527,577,771,797]" italics="true" pageId="10" pageNumber="61">
<collectionCode id="56FD6BE9FF96FFEB492F8622537C8538" box="[527,546,771,797]" country="Denmark" name="University of Copenhagen" pageId="10" pageNumber="61" type="Herbarium">C</collectionCode>
. 
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</emphasis>
</materialsCitation>
<emphasis id="02982F3EFF96FFEB49618622531B8538" box="[577,581,771,797]" italics="true" pageId="10" pageNumber="61">.</emphasis>
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<emphasis id="02982F3EFF96FFEB4964862253958539" box="[580,715,770,797]" italics="true" pageId="10" pageNumber="61">
<location id="3533A5F7FF96FFEB49648622532E8538" LSID="urn:lsid:plazi:treatment:B845423AFF95FFEA4BA880F752968725:3533A5F7FF96FFEB49648622532E8538" box="[580,624,771,797]" country="China" name="Tan" pageId="10" pageNumber="61">Tan</location>
941188
</emphasis>
(
<specimenCode id="604A5B57FF96FFEB49FC8623522D8539" box="[732,883,770,796]" collectionCode="PE" country="China" lsid="urn:lsid:biocol.org:col:15858" name="Institute of Botany, Chinese Academy of Sciences" pageId="10" pageNumber="61" type="Herbarium">PE01837338</specimenCode>
); 
<collectorName id="9D1996FAFF96FFEB48A7862252818538" box="[903,991,771,797]" pageId="10" pageNumber="61">Tonggu</collectorName>
, 
<emphasis id="02982F3EFF96FFEB48CC8622554B8538" box="[1004,1045,771,797]" italics="true" pageId="10" pageNumber="61">
<collectionCode id="56FD6BE9FF96FFEB48CC862252A58538" box="[1004,1019,771,797]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="10" pageNumber="61" type="Herbarium">S</collectionCode>
. 
<collectionCode id="56FD6BE9FF96FFEB48DF8622554B8538" box="[1023,1045,771,797]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="10" pageNumber="61" type="Herbarium">K</collectionCode>
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</materialsCitation>
<emphasis id="02982F3EFF96FFEB4F34862255468538" box="[1044,1048,771,797]" italics="true" pageId="10" pageNumber="61">.</emphasis>
<materialsCitation id="8084F971FF96FFEB4F38862251EF8564" ID-GBIF-Occurrence="3750509366" collectionCode="Y, Q" collectorName="Xiushui" country="China" location="Lai" pageId="10" pageNumber="61" specimenCode="PE00455899" specimenCount="1">
<emphasis id="02982F3EFF96FFEB4F38862255D18539" box="[1048,1167,770,797]" italics="true" pageId="10" pageNumber="61">
<location id="3533A5F7FF96FFEB4F388622551E8538" LSID="urn:lsid:plazi:treatment:B845423AFF95FFEA4BA880F752968725:3533A5F7FF96FFEB4F388622551E8538" box="[1048,1088,771,797]" country="China" name="Lai" pageId="10" pageNumber="61">Lai</location>
03673
</emphasis>
(
<specimenCode id="604A5B57FF96FFEB4FBF862354698538" box="[1183,1335,770,797]" collectionCode="PE" country="China" lsid="urn:lsid:biocol.org:col:15858" name="Institute of Botany, Chinese Academy of Sciences" pageId="10" pageNumber="61" type="Herbarium">PE00455899</specimenCode>
); 
<collectorName id="9D1996FAFF96FFEB4E6C862354F68539" box="[1356,1448,770,796]" pageId="10" pageNumber="61">Xiushui</collectorName>
, 
<emphasis id="02982F3EFF96FFEB4BA8860651EF8564" box="[136,177,807,833]" italics="true" pageId="10" pageNumber="61">
<collectionCode id="56FD6BE9FF96FFEB4BA8860651C78564" box="[136,153,807,833]" country="USA" lsid="urn:lsid:biocol.org:col:14792" name="Yale University" pageId="10" pageNumber="61" type="Herbarium">Y</collectionCode>
. 
<collectionCode id="56FD6BE9FF96FFEB4BBD860651EF8564" box="[157,177,807,833]" country="Ecuador" lsid="urn:lsid:biocol.org:col:14273" name="Universidad Central" pageId="10" pageNumber="61" type="Herbarium">Q</collectionCode>
</emphasis>
</materialsCitation>
<emphasis id="02982F3EFF96FFEB4B91860651EB8564" box="[177,181,807,833]" italics="true" pageId="10" pageNumber="61">.</emphasis>
<materialsCitation id="8084F971FF96FFEB4B94860653AE8564" ID-GBIF-Occurrence="3750509353" box="[180,752,806,833]" collectionCode="JJF" collectorName="Tancm" country="China" location="Miu" pageId="10" pageNumber="61" specimenCode="JJF00028632, JJF00028633" specimenCount="1">
<emphasis id="02982F3EFF96FFEB4B94860650FF8565" box="[180,417,806,833]" italics="true" pageId="10" pageNumber="61">
<location id="3533A5F7FF96FFEB4B94860651BA8564" LSID="urn:lsid:plazi:treatment:B845423AFF95FFEA4BA880F752968725:3533A5F7FF96FFEB4B94860651BA8564" box="[180,228,807,833]" country="China" name="Miu" pageId="10" pageNumber="61">Miu</location>
et al. 
<collectorName id="9D1996FAFF96FFEB4A3F860750348565" box="[287,362,806,832]" pageId="10" pageNumber="61">Tancm</collectorName>
2451
</emphasis>
(
<specimenCode id="604A5B57FF96FFEB4A8C860753188565" box="[428,582,806,832]" collectionCode="JJF" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13403" lsid="urn:lsid:biocol.org:col:13403" name="Jiujiang Forestry Institute" pageId="10" pageNumber="61" type="Herbarium">JJF00028632</specimenCode>
, 
<specimenCode id="604A5B57FF96FFEB496E860753B68564" box="[590,744,806,833]" collectionCode="JJF" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13403" lsid="urn:lsid:biocol.org:col:13403" name="Jiujiang Forestry Institute" pageId="10" pageNumber="61" type="Herbarium">JJF00028633</specimenCode>
)
</materialsCitation>
. 
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<emphasis id="02982F3EFF96FFEB49D78607523C8565" bold="true" box="[759,866,806,832]" pageId="10" pageNumber="61">
<collectingRegion id="F2283DCEFF96FFEB49D78607523C8565" box="[759,866,806,832]" country="China" name="Zhejiang" pageId="10" pageNumber="61">Zhejiang</collectingRegion>
</emphasis>
: 
<collectorName id="9D1996FAFF96FFEB484D860752E48565" box="[877,954,806,832]" pageId="10" pageNumber="61">Lin’an</collectorName>
, 
<collectorName id="9D1996FAFF96FFEB48E28607553C8565" box="[962,1122,806,832]" pageId="10" pageNumber="61">Xitianmushan</collectorName>
, 
<collectionCode id="56FD6BE9FF96FFEB4F4A860655DF8565" box="[1130,1153,807,832]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="10" pageNumber="61" type="Herbarium">
<emphasis id="02982F3EFF96FFEB4F4A860655DF8565" box="[1130,1153,807,832]" italics="true" pageId="10" pageNumber="61">H</emphasis>
</collectionCode>
</materialsCitation>
<emphasis id="02982F3EFF96FFEB4FA1860655DB8565" box="[1153,1157,807,832]" italics="true" pageId="10" pageNumber="61">.</emphasis>
<materialsCitation id="8084F971FF96FFEB4FA48606500D8541" ID-GBIF-Occurrence="3750509373" accessionNumber="HZ047017" collectionCode="HZ, K" country="China" location="He" pageId="10" pageNumber="61" specimenCode="PE01837442" specimenCount="1">
<emphasis id="02982F3EFF96FFEB4FA4860654598565" box="[1156,1287,806,832]" italics="true" pageId="10" pageNumber="61">
X. 
<location id="3533A5F7FF96FFEB4FBC860655E38565" LSID="urn:lsid:plazi:treatment:B845423AFF95FFEA4BA880F752968725:3533A5F7FF96FFEB4FBC860655E38565" box="[1180,1213,807,832]" country="China" name="He" pageId="10" pageNumber="61">He</location>
25779
</emphasis>
(
<specimenCode id="604A5B57FF96FFEB4E32860754F68565" box="[1298,1448,806,832]" collectionCode="PE" country="China" lsid="urn:lsid:biocol.org:col:15858" name="Institute of Botany, Chinese Academy of Sciences" pageId="10" pageNumber="61" type="Herbarium">PE01837442</specimenCode>
, 
<collectionCode id="56FD6BE9FF96FFEB4BA8866B51F18541" box="[136,175,842,868]" pageId="10" pageNumber="61">HZ</collectionCode>
047017), 
<emphasis id="02982F3EFF96FFEB4A3A866A500D8541" box="[282,339,843,868]" italics="true" pageId="10" pageNumber="61">
<collectionCode id="56FD6BE9FF96FFEB4A3A866A506E8541" box="[282,304,843,868]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="10" pageNumber="61" type="Herbarium">K</collectionCode>
. 
<collectionCode id="56FD6BE9FF96FFEB4A1D866A500D8541" box="[317,339,843,868]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="10" pageNumber="61" type="Herbarium">K</collectionCode>
</emphasis>
</materialsCitation>
<emphasis id="02982F3EFF96FFEB4A73866A50098541" box="[339,343,843,868]" italics="true" pageId="10" pageNumber="61">.</emphasis>
<materialsCitation id="8084F971FF96FFEB4A7F866A50FD85AC" ID-GBIF-Occurrence="3750509395" collectionCode="S, Y" collectorName="Zhejaing Agricultural University D" country="China" location="Tsoong" pageId="10" pageNumber="61" specimenCode="PE00455997, PE00455874, PE00455875, PE00455879" specimenCount="1">
<emphasis id="02982F3EFF96FFEB4A7F866A50BC8541" box="[351,482,842,868]" italics="true" pageId="10" pageNumber="61">
<location id="3533A5F7FF96FFEB4A7F866A50F18541" LSID="urn:lsid:plazi:treatment:B845423AFF95FFEA4BA880F752968725:3533A5F7FF96FFEB4A7F866A50F18541" box="[351,431,843,868]" country="China" name="Tsoong" pageId="10" pageNumber="61">Tsoong</location>
754
</emphasis>
(
<specimenCode id="604A5B57FF96FFEB4AD2866B53D98540" box="[498,647,842,869]" collectionCode="PE" country="China" lsid="urn:lsid:biocol.org:col:15858" name="Institute of Botany, Chinese Academy of Sciences" pageId="10" pageNumber="61" type="Herbarium">PE00455997</specimenCode>
, 
<specimenCode id="604A5B57FF96FFEB49B4866B52778540" box="[660,809,842,869]" collectionCode="PE" country="China" lsid="urn:lsid:biocol.org:col:15858" name="Institute of Botany, Chinese Academy of Sciences" pageId="10" pageNumber="61" type="Herbarium">PE00455874</specimenCode>
), 
<emphasis id="02982F3EFF96FFEB481E866B55A68541" box="[830,1272,842,868]" italics="true" pageId="10" pageNumber="61">
<collectorName id="9D1996FAFF96FFEB481E866B55908541" box="[830,1230,842,868]" pageId="10" pageNumber="61">Zhejaing Agricultural University D</collectorName>
330
</emphasis>
(
<specimenCode id="604A5B57FF96FFEB4E29866B54C18540" box="[1289,1439,842,869]" collectionCode="PE" country="China" lsid="urn:lsid:biocol.org:col:15858" name="Institute of Botany, Chinese Academy of Sciences" pageId="10" pageNumber="61" type="Herbarium">PE00455875</specimenCode>
), 
<emphasis id="02982F3EFF96FFEB4BA8864F519085AD" box="[136,206,878,904]" italics="true" pageId="10" pageNumber="61">10343</emphasis>
(
<specimenCode id="604A5B57FF96FFEB4BF9864F502F85AC" box="[217,369,878,905]" collectionCode="PE" country="China" lsid="urn:lsid:biocol.org:col:15858" name="Institute of Botany, Chinese Academy of Sciences" pageId="10" pageNumber="61" type="Herbarium">PE00455879</specimenCode>
), 
<emphasis id="02982F3EFF96FFEB4A5F864E50FD85AC" box="[383,419,879,905]" italics="true" pageId="10" pageNumber="61">
<collectionCode id="56FD6BE9FF96FFEB4A5F864E50D085AC" box="[383,398,879,905]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="10" pageNumber="61" type="Herbarium">S</collectionCode>
. 
<collectionCode id="56FD6BE9FF96FFEB4AB2864E50FD85AC" box="[402,419,879,905]" country="USA" lsid="urn:lsid:biocol.org:col:14792" name="Yale University" pageId="10" pageNumber="61" type="Herbarium">Y</collectionCode>
</emphasis>
</materialsCitation>
<emphasis id="02982F3EFF96FFEB4A83864E50F985AC" box="[419,423,879,905]" italics="true" pageId="10" pageNumber="61">.</emphasis>
<materialsCitation id="8084F971FF96FFEB4A86864E539885AC" ID-GBIF-Occurrence="3750509303" box="[422,710,878,905]" collectionCode="Z, C" country="China" location="Hu" pageId="10" pageNumber="61" specimenCode="PE00457428" specimenCount="1">
<emphasis id="02982F3EFF96FFEB4A86864E50B485AD" box="[422,490,878,905]" italics="true" pageId="10" pageNumber="61">
<location id="3533A5F7FF96FFEB4A86864E509485AC" LSID="urn:lsid:plazi:treatment:B845423AFF95FFEA4BA880F752968725:3533A5F7FF96FFEB4A86864E509485AC" box="[422,458,879,905]" country="China" name="Hu" pageId="10" pageNumber="61">Hu</location>
31
</emphasis>
(
<specimenCode id="604A5B57FF96FFEB4AD5864F53D285AC" box="[501,652,878,905]" collectionCode="PE" country="China" lsid="urn:lsid:biocol.org:col:15858" name="Institute of Botany, Chinese Academy of Sciences" pageId="10" pageNumber="61" type="Herbarium">PE00457428</specimenCode>
), 
<emphasis id="02982F3EFF96FFEB49BB864E539885AC" box="[667,710,879,905]" italics="true" pageId="10" pageNumber="61">
<collectionCode id="56FD6BE9FF96FFEB49BB864E53F085AC" box="[667,686,879,905]" country="Switzerland" name="Universität Zürich" pageId="10" pageNumber="61" type="Herbarium">Z</collectionCode>
. 
<collectionCode id="56FD6BE9FF96FFEB4992864E539885AC" box="[690,710,879,905]" country="Denmark" name="University of Copenhagen" pageId="10" pageNumber="61" type="Herbarium">C</collectionCode>
</emphasis>
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<emphasis id="02982F3EFF96FFEB49E6864E539485AC" box="[710,714,879,905]" italics="true" pageId="10" pageNumber="61">.</emphasis>
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ZJ2021009
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(
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), 
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. 
<collectionCode id="56FD6BE9FF96FFEB48C5864E52A785AC" box="[997,1017,879,905]" country="Denmark" name="University of Copenhagen" pageId="10" pageNumber="61" type="Herbarium">C</collectionCode>
</emphasis>
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<emphasis id="02982F3EFF96FFEB48D9864E52A385AC" box="[1017,1021,879,905]" italics="true" pageId="10" pageNumber="61">.</emphasis>
<materialsCitation id="8084F971FF96FFEB48DC864E51E88589" ID-GBIF-Occurrence="3750509347" collectionCode="IBSC, K" collectorName="Jin ZJ &amp; Yandangshan" country="China" pageId="10" pageNumber="61" specimenCount="1">
<emphasis id="02982F3EFF96FFEB48DC864E55EF85AD" box="[1020,1201,878,905]" italics="true" pageId="10" pageNumber="61">
<collectorName id="9D1996FAFF96FFEB48DC864E551F85AD" box="[1020,1089,878,905]" pageId="10" pageNumber="61">Jin ZJ</collectorName>
20210010
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(
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); 
<collectorName id="9D1996FAFF96FFEB4E2F864F54F685AD" box="[1295,1448,878,904]" pageId="10" pageNumber="61">Yandangshan</collectorName>
, 
<emphasis id="02982F3EFF96FFEB4BA886B251E88589" box="[136,182,915,940]" italics="true" pageId="10" pageNumber="61">
<collectionCode id="56FD6BE9FF96FFEB4BA886B251C38589" box="[136,157,915,940]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="10" pageNumber="61" type="Herbarium">K</collectionCode>
. 
<collectionCode id="56FD6BE9FF96FFEB4B8186B251E88589" box="[161,182,915,940]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="10" pageNumber="61" type="Herbarium">K</collectionCode>
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<emphasis id="02982F3EFF96FFEB4B9686B251E48589" box="[182,186,915,940]" italics="true" pageId="10" pageNumber="61">.</emphasis>
<materialsCitation id="8084F971FF96FFEB4B9A86B253FE8589" ID-GBIF-Occurrence="3750509368" box="[186,672,914,941]" collectionCode="K" collectorName="Tiantaishan" country="China" location="Tsoong" pageId="10" pageNumber="61" specimenCode="PE0455871" specimenCount="1">
<emphasis id="02982F3EFF96FFEB4B9A86B250678589" box="[186,313,915,940]" italics="true" pageId="10" pageNumber="61">
<location id="3533A5F7FF96FFEB4B9A86B250528589" LSID="urn:lsid:plazi:treatment:B845423AFF95FFEA4BA880F752968725:3533A5F7FF96FFEB4B9A86B250528589" box="[186,268,915,940]" country="China" name="Tsoong" pageId="10" pageNumber="61">Tsoong</location>
s.n.
</emphasis>
(
<specimenCode id="604A5B57FF96FFEB4A6786B3508F8588" box="[327,465,914,941]" collectionCode="PE" country="China" lsid="urn:lsid:biocol.org:col:15858" name="Institute of Botany, Chinese Academy of Sciences" pageId="10" pageNumber="61" type="Herbarium">PE0455871</specimenCode>
); 
<collectorName id="9D1996FAFF96FFEB4AC386B353368589" box="[483,616,914,940]" pageId="10" pageNumber="61">Tiantaishan</collectorName>
, 
<emphasis id="02982F3EFF96FFEB495286B253FE8589" box="[626,672,915,940]" italics="true" pageId="10" pageNumber="61">
<collectionCode id="56FD6BE9FF96FFEB495286B253D98589" box="[626,647,915,940]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="10" pageNumber="61" type="Herbarium">K</collectionCode>
. 
<collectionCode id="56FD6BE9FF96FFEB49AB86B253FE8589" box="[651,672,915,940]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="10" pageNumber="61" type="Herbarium">K</collectionCode>
</emphasis>
</materialsCitation>
<emphasis id="02982F3EFF96FFEB498186B253FB8589" box="[673,677,915,940]" italics="true" pageId="10" pageNumber="61">.</emphasis>
<materialsCitation id="8084F971FF96FFEB498486B2547C8589" ID-GBIF-Occurrence="3750509322" box="[676,1314,914,941]" collectionCode="H" collectorName="Changhua" country="China" location="Tsoong" pageId="10" pageNumber="61" specimenCode="PE00455873, PE00455882" specimenCount="1">
<emphasis id="02982F3EFF96FFEB498486B2526A8589" box="[676,820,914,940]" italics="true" pageId="10" pageNumber="61">
<location id="3533A5F7FF96FFEB498486B253A88589" LSID="urn:lsid:plazi:treatment:B845423AFF95FFEA4BA880F752968725:3533A5F7FF96FFEB498486B253A88589" box="[676,758,915,940]" country="China" name="Tsoong" pageId="10" pageNumber="61">Tsoong</location>
3712
</emphasis>
(
<specimenCode id="604A5B57FF96FFEB486386B352868588" box="[835,984,914,941]" collectionCode="PE" country="China" lsid="urn:lsid:biocol.org:col:15858" name="Institute of Botany, Chinese Academy of Sciences" pageId="10" pageNumber="61" type="Herbarium">PE00455873</specimenCode>
, 
<specimenCode id="604A5B57FF96FFEB48C386B355278588" box="[995,1145,914,941]" collectionCode="PE" country="China" lsid="urn:lsid:biocol.org:col:15858" name="Institute of Botany, Chinese Academy of Sciences" pageId="10" pageNumber="61" type="Herbarium">PE00455882</specimenCode>
); 
<collectorName id="9D1996FAFF96FFEB4FAB86B355A18589" box="[1163,1279,914,940]" pageId="10" pageNumber="61">Changhua</collectorName>
, 
<collectionCode id="56FD6BE9FF96FFEB4E2B86B2547C8589" box="[1291,1314,915,940]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="10" pageNumber="61" type="Herbarium">
<emphasis id="02982F3EFF96FFEB4E2B86B2547C8589" box="[1291,1314,915,940]" italics="true" pageId="10" pageNumber="61">H</emphasis>
</collectionCode>
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<emphasis id="02982F3EFF96FFEB4E0286B254788589" box="[1314,1318,915,940]" italics="true" pageId="10" pageNumber="61">.</emphasis>
<materialsCitation id="8084F971FF96FFEB4E0586B2504A85F5" ID-GBIF-Occurrence="3750509331" accessionNumber="HZ047043" collectionCode="HZ" collectorName="He" country="China" pageId="10" pageNumber="61" specimenCount="1">
<emphasis id="02982F3EFF96FFEB4E0586B254F58589" box="[1317,1451,914,940]" italics="true" pageId="10" pageNumber="61">
X. 
<collectorName id="9D1996FAFF96FFEB4E1D86B254008589" box="[1341,1374,915,940]" pageId="10" pageNumber="61">He</collectorName>
15314
</emphasis>
(
<collectionCode id="56FD6BE9FF96FFEB4BB0869751E885F5" box="[144,182,950,976]" pageId="10" pageNumber="61">HZ</collectionCode>
047043)
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.— 
<materialsCitation id="8084F971FF96FFEB4A148697535885F5" ID-GBIF-Occurrence="3750509352" box="[308,518,950,976]" collectionCode="M" country="Japan" location="Akita" pageId="10" pageNumber="61" specimenCount="1" stateProvince="Akita">
<collectingCountry id="48FBB3BCFF96FFEB4A14869750D185F5" box="[308,399,950,976]" name="Japan" pageId="10" pageNumber="61">JAPAN</collectingCountry>
. 
<collectingRegion id="F2283DCEFF96FFEB4ABB8697508285F5" box="[411,476,950,976]" country="Japan" name="Akita" pageId="10" pageNumber="61">Akita</collectingRegion>
, 
<collectionCode id="56FD6BE9FF96FFEB4ACB8696535885F5" box="[491,518,951,976]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="10" pageNumber="61" type="Herbarium">
<emphasis id="02982F3EFF96FFEB4ACB8696535885F5" box="[491,518,951,976]" italics="true" pageId="10" pageNumber="61">M</emphasis>
</collectionCode>
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<emphasis id="02982F3EFF96FFEB49268696535485F5" box="[518,522,951,976]" italics="true" pageId="10" pageNumber="61">.</emphasis>
<materialsCitation id="8084F971FF96FFEB49338697524F85F4" ID-GBIF-Occurrence="3750509344" accessionNumber="TNS50829" box="[531,785,950,977]" collectionCode="TNS" country="Japan" location="Matsuda" pageId="10" pageNumber="61" specimenCount="1">
<emphasis id="02982F3EFF96FFEB49338697532B85F5" box="[531,629,950,976]" italics="true" pageId="10" pageNumber="61">
<location id="3533A5F7FF96FFEB49338697532B85F5" LSID="urn:lsid:plazi:treatment:B845423AFF95FFEA4BA880F752968725:3533A5F7FF96FFEB49338697532B85F5" box="[531,629,950,976]" country="Japan" name="Matsuda" pageId="10" pageNumber="61">Matsuda</location>
</emphasis>
(
<collectionCode id="56FD6BE9FF96FFEB49A7869753E085F4" box="[647,702,950,977]" country="Japan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15622" lsid="urn:lsid:biocol.org:col:15622" name="National Museum of Nature and Science" pageId="10" pageNumber="61" type="Herbarium">TNS</collectionCode>
50829)
</materialsCitation>
; 
<materialsCitation id="8084F971FF96FFEB483E869752FF85F5" ID-GBIF-Occurrence="3750509326" box="[798,929,950,976]" collectionCode="M" country="Japan" location="Aomori" pageId="10" pageNumber="61" specimenCount="1" stateProvince="Aomori">
<collectingRegion id="F2283DCEFF96FFEB483E8697522685F5" box="[798,888,950,976]" country="Japan" name="Aomori" pageId="10" pageNumber="61">Aomori</collectingRegion>
, 
<collectionCode id="56FD6BE9FF96FFEB48A6869652FF85F5" box="[902,929,951,976]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="10" pageNumber="61" type="Herbarium">
<emphasis id="02982F3EFF96FFEB48A6869652FF85F5" box="[902,929,951,976]" italics="true" pageId="10" pageNumber="61">M</emphasis>
</collectionCode>
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<emphasis id="02982F3EFF96FFEB4881869652FB85F5" box="[929,933,951,976]" italics="true" pageId="10" pageNumber="61">.</emphasis>
<materialsCitation id="8084F971FF96FFEB48848696507985D0" ID-GBIF-Occurrence="3750509356" collectionCode="PE, Y" country="Japan" location="Furuse" pageId="10" pageNumber="61" specimenCode="TNS753035" specimenCount="1">
<emphasis id="02982F3EFF96FFEB48848696556885F5" box="[932,1078,950,976]" italics="true" pageId="10" pageNumber="61">
<location id="3533A5F7FF96FFEB4884869652AA85F5" LSID="urn:lsid:plazi:treatment:B845423AFF95FFEA4BA880F752968725:3533A5F7FF96FFEB4884869652AA85F5" box="[932,1012,951,976]" country="Japan" name="Furuse" pageId="10" pageNumber="61">Furuse</location>
9652
</emphasis>
(
<collectionCode id="56FD6BE9FF96FFEB4F688697553485F5" box="[1096,1130,950,976]" country="China" lsid="urn:lsid:biocol.org:col:15858" name="Institute of Botany, Chinese Academy of Sciences" pageId="10" pageNumber="61" type="Herbarium">PE</collectionCode>
01292177), 
<emphasis id="02982F3EFF96FFEB4FD5869754F585F5" box="[1269,1451,950,976]" italics="true" pageId="10" pageNumber="61">
<collectionCode id="56FD6BE9FF96FFEB4FD58697545B85F5" box="[1269,1285,950,976]" country="USA" lsid="urn:lsid:biocol.org:col:14792" name="Yale University" pageId="10" pageNumber="61" type="Herbarium">Y</collectionCode>
. 
<location id="3533A5F7FF96FFEB4E288697540585F5" LSID="urn:lsid:plazi:treatment:B845423AFF95FFEA4BA880F752968725:3533A5F7FF96FFEB4E288697540585F5" box="[1288,1371,950,976]" country="Japan" name="Kadota" pageId="10" pageNumber="61">Kadota</location>
55001
</emphasis>
(
<specimenCode id="604A5B57FF96FFEB4BB086FB504185D0" box="[144,287,986,1013]" collectionCode="TNS" country="Japan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15622" lsid="urn:lsid:biocol.org:col:15622" name="National Museum of Nature and Science" pageId="10" pageNumber="61" type="Herbarium">TNS753035</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="8084F971FF96FFEB4A1486FB50C685D1" ID-GBIF-Occurrence="3750509301" box="[308,408,986,1012]" collectionCode="N" country="Japan" location="Fukui" pageId="10" pageNumber="61" specimenCount="1" stateProvince="Fukui">
<collectingRegion id="F2283DCEFF96FFEB4A1486FB502685D1" box="[308,376,986,1012]" country="Japan" name="Fukui" pageId="10" pageNumber="61">Fukui</collectingRegion>
, 
<collectionCode id="56FD6BE9FF96FFEB4AA586FA50C685D1" box="[389,408,987,1012]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="10" pageNumber="61" type="Herbarium">
<emphasis id="02982F3EFF96FFEB4AA586FA50C685D1" box="[389,408,987,1012]" italics="true" pageId="10" pageNumber="61">N</emphasis>
</collectionCode>
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<emphasis id="02982F3EFF96FFEB4AB786FB50C585D1" box="[407,411,986,1012]" italics="true" pageId="10" pageNumber="61">.</emphasis>
<materialsCitation id="8084F971FF96FFEB4ABB86FB525785D0" ID-GBIF-Occurrence="3750509310" box="[411,777,986,1013]" collectionCode="TNS" country="Japan" location="Kurosaki" pageId="10" pageNumber="61" specimenCode="TNS591236" specimenCount="1">
<emphasis id="02982F3EFF96FFEB4ABB86FB533F85D1" box="[411,609,986,1012]" italics="true" pageId="10" pageNumber="61">
<location id="3533A5F7FF96FFEB4ABB86FB535A85D1" LSID="urn:lsid:plazi:treatment:B845423AFF95FFEA4BA880F752968725:3533A5F7FF96FFEB4ABB86FB535A85D1" box="[411,516,986,1012]" country="Japan" name="Kurosaki" pageId="10" pageNumber="61">Kurosaki</location>
177031
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(
<specimenCode id="604A5B57FF96FFEB495186FB525E85D0" box="[625,768,986,1013]" collectionCode="TNS" country="Japan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15622" lsid="urn:lsid:biocol.org:col:15622" name="National Museum of Nature and Science" pageId="10" pageNumber="61" type="Herbarium">TNS591236</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="8084F971FF96FFEB483586FB522D85D1" ID-GBIF-Occurrence="3750509359" box="[789,883,986,1012]" collectionCode="M" country="Japan" location="Gifu" pageId="10" pageNumber="61" specimenCount="1" stateProvince="Gifu">
<collectingRegion id="F2283DCEFF96FFEB483586FB521585D1" box="[789,843,986,1012]" country="Japan" name="Gifu" pageId="10" pageNumber="61">Gifu</collectingRegion>
, 
<collectionCode id="56FD6BE9FF96FFEB487886FA522D85D1" box="[856,883,987,1012]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="10" pageNumber="61" type="Herbarium">
<emphasis id="02982F3EFF96FFEB487886FA522D85D1" box="[856,883,987,1012]" italics="true" pageId="10" pageNumber="61">M</emphasis>
</collectionCode>
</materialsCitation>
<emphasis id="02982F3EFF96FFEB485386FA522985D1" box="[883,887,987,1012]" italics="true" pageId="10" pageNumber="61">.</emphasis>
<materialsCitation id="8084F971FF96FFEB485686FA55BB85D1" ID-GBIF-Occurrence="3750509390" box="[886,1253,986,1012]" collectionCode="PE, H" country="Japan" location="Furuse" pageId="10" pageNumber="61" specimenCount="1">
<emphasis id="02982F3EFF96FFEB485686FA554A85D1" box="[886,1044,986,1012]" italics="true" pageId="10" pageNumber="61">
<location id="3533A5F7FF96FFEB485686FA529885D1" LSID="urn:lsid:plazi:treatment:B845423AFF95FFEA4BA880F752968725:3533A5F7FF96FFEB485686FA529885D1" box="[886,966,987,1012]" country="Japan" name="Furuse" pageId="10" pageNumber="61">Furuse</location>
52183
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(
<collectionCode id="56FD6BE9FF96FFEB4F0486FB551985D1" box="[1060,1095,986,1012]" country="China" lsid="urn:lsid:biocol.org:col:15858" name="Institute of Botany, Chinese Academy of Sciences" pageId="10" pageNumber="61" type="Herbarium">PE</collectionCode>
01292173), 
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<emphasis id="02982F3EFF96FFEB4FEF86FB55BB85D1" box="[1231,1253,986,1012]" italics="true" pageId="10" pageNumber="61">H</emphasis>
</collectionCode>
</materialsCitation>
<emphasis id="02982F3EFF96FFEB4FC686FB55B485D1" box="[1254,1258,986,1012]" italics="true" pageId="10" pageNumber="61">.</emphasis>
<materialsCitation id="8084F971FF96FFEB4FC986FB5079823C" ID-GBIF-Occurrence="3750509398" collectionCode="TNS" country="Japan" location="Takahashi" pageId="10" pageNumber="61" specimenCode="TNS714682" specimenCount="1">
<emphasis id="02982F3EFF96FFEB4FC986FB54F585D1" box="[1257,1451,986,1012]" italics="true" pageId="10" pageNumber="61">
<location id="3533A5F7FF96FFEB4FC986FB540385D1" LSID="urn:lsid:plazi:treatment:B845423AFF95FFEA4BA880F752968725:3533A5F7FF96FFEB4FC986FB540385D1" box="[1257,1373,986,1012]" country="Japan" name="Takahashi" pageId="10" pageNumber="61">Takahashi</location>
20840
</emphasis>
(
<specimenCode id="604A5B57FF96FFEB4BB086DF5041823C" box="[144,287,1022,1049]" collectionCode="TNS" country="Japan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15622" lsid="urn:lsid:biocol.org:col:15622" name="National Museum of Nature and Science" pageId="10" pageNumber="61" type="Herbarium">TNS714682</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="8084F971FF96FFEB4A1686DF50EC823D" ID-GBIF-Occurrence="3750509374" box="[310,434,1022,1048]" collectionCode="H" country="Japan" location="Gunma" pageId="10" pageNumber="61" specimenCount="1" stateProvince="Gunma">
<collectingRegion id="F2283DCEFF96FFEB4A1686DF50D5823D" box="[310,395,1022,1048]" country="Japan" name="Gunma" pageId="10" pageNumber="61">Gunma</collectingRegion>
, 
<collectionCode id="56FD6BE9FF96FFEB4ABA86DE50EC823D" box="[410,434,1023,1048]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="10" pageNumber="61" type="Herbarium">
<emphasis id="02982F3EFF96FFEB4ABA86DE50EC823D" box="[410,434,1023,1048]" italics="true" pageId="10" pageNumber="61">H</emphasis>
</collectionCode>
</materialsCitation>
<emphasis id="02982F3EFF96FFEB4A9186DE50EB823D" box="[433,437,1023,1048]" italics="true" pageId="10" pageNumber="61">.</emphasis>
<materialsCitation id="8084F971FF96FFEB4A9586DE53F1823C" ID-GBIF-Occurrence="3750509336" box="[437,687,1022,1049]" collectionCode="TNS" country="Japan" location="Utsumi" pageId="10" pageNumber="61" specimenCode="TNS142954" specimenCount="1">
<emphasis id="02982F3EFF96FFEB4A9586DE5358823D" box="[437,518,1023,1048]" italics="true" pageId="10" pageNumber="61">
<location id="3533A5F7FF96FFEB4A9586DE5358823D" LSID="urn:lsid:plazi:treatment:B845423AFF95FFEA4BA880F752968725:3533A5F7FF96FFEB4A9586DE5358823D" box="[437,518,1023,1048]" country="Japan" name="Utsumi" pageId="10" pageNumber="61">Utsumi</location>
</emphasis>
(
<specimenCode id="604A5B57FF96FFEB493886DF53F9823C" box="[536,679,1022,1049]" collectionCode="TNS" country="Japan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15622" lsid="urn:lsid:biocol.org:col:15622" name="National Museum of Nature and Science" pageId="10" pageNumber="61" type="Herbarium">TNS142954</specimenCode>
)
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; 
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1195
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(
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), 
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<emphasis id="02982F3EFF96FFEB4F8586DE55E8823C" box="[1189,1206,1023,1049]" italics="true" pageId="10" pageNumber="61">T</emphasis>
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<emphasis id="02982F3EFF96FFEB4F9686DE55E4823C" box="[1206,1210,1023,1049]" italics="true" pageId="10" pageNumber="61">.</emphasis>
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<emphasis id="02982F3EFF96FFEB4F9986DE55A0823C" box="[1209,1278,1023,1049]" italics="true" pageId="10" pageNumber="61">
<location id="3533A5F7FF96FFEB4F9986DE55A0823C" LSID="urn:lsid:plazi:treatment:B845423AFF95FFEA4BA880F752968725:3533A5F7FF96FFEB4F9986DE55A0823C" box="[1209,1278,1023,1049]" country="Japan" name="Satow" pageId="10" pageNumber="61">Satow</location>
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(
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)
</materialsCitation>
; 
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<collectingRegion id="F2283DCEFF96FFEB4BA8810251978218" box="[136,201,1059,1085]" country="Japan" name="Iwate" pageId="10" pageNumber="61">Iwate</collectingRegion>
, 
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<emphasis id="02982F3EFF96FFEB4BFE810251A78219" box="[222,249,1059,1084]" italics="true" pageId="10" pageNumber="61">M</emphasis>
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<emphasis id="02982F3EFF96FFEB4BD9810251A38219" box="[249,253,1059,1084]" italics="true" pageId="10" pageNumber="61">.</emphasis>
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<emphasis id="02982F3EFF96FFEB4BDC810250FD8219" box="[252,419,1058,1084]" italics="true" pageId="10" pageNumber="61">
<location id="3533A5F7FF96FFEB4BDC810250128219" LSID="urn:lsid:plazi:treatment:B845423AFF95FFEA4BA880F752968725:3533A5F7FF96FFEB4BDC810250128219" box="[252,332,1059,1084]" country="Japan" name="Furuse" pageId="10" pageNumber="61">Furuse</location>
41390
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(
<collectionCode id="56FD6BE9FF96FFEB4A9B810350808218" box="[443,478,1058,1085]" country="China" lsid="urn:lsid:biocol.org:col:15858" name="Institute of Botany, Chinese Academy of Sciences" pageId="10" pageNumber="61" type="Herbarium">PE</collectionCode>
01292175), 
<collectionCode id="56FD6BE9FF96FFEB494F810353D78219" box="[623,649,1058,1084]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="10" pageNumber="61" type="Herbarium">
<emphasis id="02982F3EFF96FFEB494F810353D78219" box="[623,649,1058,1084]" italics="true" pageId="10" pageNumber="61">M</emphasis>
</collectionCode>
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<emphasis id="02982F3EFF96FFEB49A9810353D38219" box="[649,653,1058,1084]" italics="true" pageId="10" pageNumber="61">.</emphasis>
<materialsCitation id="8084F971FF96FFEB49AC810352E78219" ID-GBIF-Occurrence="3750509350" box="[652,953,1058,1085]" collectionCode="T" country="Japan" location="Kikuchi" pageId="10" pageNumber="61" specimenCode="TNS176756" specimenCount="1">
<emphasis id="02982F3EFF96FFEB49AC810353BA8219" box="[652,740,1058,1084]" italics="true" pageId="10" pageNumber="61">
<location id="3533A5F7FF96FFEB49AC810353BA8219" LSID="urn:lsid:plazi:treatment:B845423AFF95FFEA4BA880F752968725:3533A5F7FF96FFEB49AC810353BA8219" box="[652,740,1058,1084]" country="Japan" name="Kikuchi" pageId="10" pageNumber="61">Kikuchi</location>
</emphasis>
(
<specimenCode id="604A5B57FF96FFEB49DD810352D28218" box="[765,908,1058,1085]" collectionCode="TNS" country="Japan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15622" lsid="urn:lsid:biocol.org:col:15622" name="National Museum of Nature and Science" pageId="10" pageNumber="61" type="Herbarium">TNS176756</specimenCode>
), 
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<emphasis id="02982F3EFF96FFEB4888810352E78219" box="[936,953,1058,1084]" italics="true" pageId="10" pageNumber="61">T</emphasis>
</collectionCode>
</materialsCitation>
<emphasis id="02982F3EFF96FFEB4899810352E38219" box="[953,957,1058,1084]" italics="true" pageId="10" pageNumber="61">.</emphasis>
<materialsCitation id="8084F971FF96FFEB489C810354748218" ID-GBIF-Occurrence="3750509399" box="[956,1322,1058,1085]" collectionCode="HAST" country="Japan" location="Kawasaki" pageId="10" pageNumber="61" specimenCode="HAST82445" specimenCount="1">
<emphasis id="02982F3EFF96FFEB489C8103552A8219" box="[956,1140,1058,1084]" italics="true" pageId="10" pageNumber="61">
<location id="3533A5F7FF96FFEB489C810355758219" LSID="urn:lsid:plazi:treatment:B845423AFF95FFEA4BA880F752968725:3533A5F7FF96FFEB489C810355758219" box="[956,1067,1058,1084]" country="Japan" name="Kawasaki" pageId="10" pageNumber="61">Kawasaki</location>
7309
</emphasis>
(
<specimenCode id="604A5B57FF96FFEB4FAD8103547C8218" box="[1165,1314,1058,1085]" collectionCode="HAST" country="Taiwan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15074" lsid="urn:lsid:biocol.org:col:15074" name="Research Center for Biodiversity, Academia Sinica" pageId="10" pageNumber="61" type="Herbarium">HAST82445</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="8084F971FF96FFEB4E1F810351C58245" ID-GBIF-Occurrence="3750509394" collectionCode="N" country="Japan" location="Ishikawa" pageId="10" pageNumber="61" specimenCount="1" stateProvince="Ishikawa">
<collectingRegion id="F2283DCEFF96FFEB4E1F810354F98219" box="[1343,1447,1058,1084]" country="Japan" name="Ishikawa" pageId="10" pageNumber="61">Ishikawa</collectingRegion>
, 
<collectionCode id="56FD6BE9FF96FFEB4BA8816651C58245" box="[136,155,1095,1120]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="10" pageNumber="61" type="Herbarium">
<emphasis id="02982F3EFF96FFEB4BA8816651C58245" box="[136,155,1095,1120]" italics="true" pageId="10" pageNumber="61">N</emphasis>
</collectionCode>
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<emphasis id="02982F3EFF96FFEB4BBB816751C18245" box="[155,159,1094,1120]" italics="true" pageId="10" pageNumber="61">.</emphasis>
<materialsCitation id="8084F971FF96FFEB4BBE816753F58245" ID-GBIF-Occurrence="3750509348" box="[158,683,1094,1120]" collectionCode="M" collectorName="Koodzuke" country="Japan" location="Kurosaki" pageId="10" pageNumber="61" specimenCode="TNS310773" specimenCount="1">
<emphasis id="02982F3EFF96FFEB4BBE816750128245" box="[158,332,1094,1120]" italics="true" pageId="10" pageNumber="61">
<location id="3533A5F7FF96FFEB4BBE816750598245" LSID="urn:lsid:plazi:treatment:B845423AFF95FFEA4BA880F752968725:3533A5F7FF96FFEB4BBE816750598245" box="[158,263,1094,1120]" country="Japan" name="Kurosaki" pageId="10" pageNumber="61">Kurosaki</location>
3718
</emphasis>
(
<specimenCode id="604A5B57FF96FFEB4A41816750AE8245" box="[353,496,1094,1120]" collectionCode="TNS" country="Japan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15622" lsid="urn:lsid:biocol.org:col:15622" name="National Museum of Nature and Science" pageId="10" pageNumber="61" type="Herbarium">TNS310773</specimenCode>
); 
<collectorName id="9D1996FAFF96FFEB4929816753218245" box="[521,639,1094,1120]" pageId="10" pageNumber="61">Koodzuke</collectorName>
, 
<collectionCode id="56FD6BE9FF96FFEB49B0816653F58245" box="[656,683,1095,1120]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="10" pageNumber="61" type="Herbarium">
<emphasis id="02982F3EFF96FFEB49B0816653F58245" box="[656,683,1095,1120]" italics="true" pageId="10" pageNumber="61">M</emphasis>
</collectionCode>
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<emphasis id="02982F3EFF96FFEB498B816653F18245" box="[683,687,1095,1120]" italics="true" pageId="10" pageNumber="61">.</emphasis>
<materialsCitation id="8084F971FF96FFEB498E816655F78245" ID-GBIF-Occurrence="3750509400" box="[686,1193,1094,1120]" collectionCode="PE" country="Japan" location="Furuse" pageId="10" pageNumber="61" specimenCount="1">
<emphasis id="02982F3EFF96FFEB498E8166520F8245" box="[686,849,1094,1120]" italics="true" pageId="10" pageNumber="61">
<location id="3533A5F7FF96FFEB498E816653A08245" LSID="urn:lsid:plazi:treatment:B845423AFF95FFEA4BA880F752968725:3533A5F7FF96FFEB498E816653A08245" box="[686,766,1095,1120]" country="Japan" name="Furuse" pageId="10" pageNumber="61">Furuse</location>
47139
</emphasis>
(
<collectionCode id="56FD6BE9FF96FFEB4845816752D98245" box="[869,903,1094,1120]" country="China" lsid="urn:lsid:biocol.org:col:15858" name="Institute of Botany, Chinese Academy of Sciences" pageId="10" pageNumber="61" type="Herbarium">PE</collectionCode>
01292169, 
<collectionCode id="56FD6BE9FF96FFEB4F2B816755738245" box="[1035,1069,1094,1120]" country="China" lsid="urn:lsid:biocol.org:col:15858" name="Institute of Botany, Chinese Academy of Sciences" pageId="10" pageNumber="61" type="Herbarium">PE</collectionCode>
01292170)
</materialsCitation>
; 
<materialsCitation id="8084F971FF96FFEB4F9A816654628245" ID-GBIF-Occurrence="3750509305" box="[1210,1340,1094,1121]" collectionCode="H" country="Japan" location="Nagano" pageId="10" pageNumber="61" specimenCount="1" stateProvince="Nagano">
<collectingRegion id="F2283DCEFF96FFEB4F9A8166544A8244" box="[1210,1300,1095,1121]" country="Japan" name="Nagano" pageId="10" pageNumber="61">Nagano</collectingRegion>
, 
<collectionCode id="56FD6BE9FF96FFEB4E05816754628245" box="[1317,1340,1094,1120]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="10" pageNumber="61" type="Herbarium">
<emphasis id="02982F3EFF96FFEB4E05816754628245" box="[1317,1340,1094,1120]" italics="true" pageId="10" pageNumber="61">H</emphasis>
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<emphasis id="02982F3EFF96FFEB4E1B816754618245" box="[1339,1343,1094,1120]" italics="true" pageId="10" pageNumber="61">.</emphasis>
<materialsCitation id="8084F971FF96FFEB4E1F8167502782A1" ID-GBIF-Occurrence="3750509306" collectionCode="TNS" country="Japan" location="Koidzumi" pageId="10" pageNumber="61" specimenCode="TNS178984" specimenCount="1">
<emphasis id="02982F3EFF96FFEB4E1F8167519082A1" italics="true" pageId="10" pageNumber="61">
<location id="3533A5F7FF96FFEB4E1F816754F28245" LSID="urn:lsid:plazi:treatment:B845423AFF95FFEA4BA880F752968725:3533A5F7FF96FFEB4E1F816754F28245" box="[1343,1452,1094,1120]" country="Japan" name="Koidzumi" pageId="10" pageNumber="61">Koidzumi</location>
34704
</emphasis>
(
<specimenCode id="604A5B57FF96FFEB4BC2814B502F82A1" box="[226,369,1130,1156]" collectionCode="TNS" country="Japan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15622" lsid="urn:lsid:biocol.org:col:15622" name="National Museum of Nature and Science" pageId="10" pageNumber="61" type="Herbarium">TNS178984</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="8084F971FF96FFEB4AA9814B50A382A1" ID-GBIF-Occurrence="3750509342" box="[393,509,1130,1156]" collectionCode="Y" country="Japan" location="Miyagi" pageId="10" pageNumber="61" specimenCount="1" stateProvince="Miyagi">
<collectingRegion id="F2283DCEFF96FFEB4AA9814B508382A1" box="[393,477,1130,1156]" country="Japan" name="Miyagi" pageId="10" pageNumber="61">Miyagi</collectingRegion>
, 
<collectionCode id="56FD6BE9FF96FFEB4ACC814A50A382A1" box="[492,509,1131,1156]" country="USA" lsid="urn:lsid:biocol.org:col:14792" name="Yale University" pageId="10" pageNumber="61" type="Herbarium">
<emphasis id="02982F3EFF96FFEB4ACC814A50A382A1" box="[492,509,1131,1156]" italics="true" pageId="10" pageNumber="61">Y</emphasis>
</collectionCode>
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<emphasis id="02982F3EFF96FFEB4ADE814A535C82A1" box="[510,514,1131,1156]" italics="true" pageId="10" pageNumber="61">.</emphasis>
<materialsCitation id="8084F971FF96FFEB4921814A520182A1" ID-GBIF-Occurrence="3750509304" box="[513,863,1130,1157]" collectionCode="H" country="Japan" location="Ueno" pageId="10" pageNumber="61" specimenCode="TNS692011" specimenCount="1">
<emphasis id="02982F3EFF96FFEB4921814A53D182A1" box="[513,655,1130,1156]" italics="true" pageId="10" pageNumber="61">
<location id="3533A5F7FF96FFEB4921814A536382A1" LSID="urn:lsid:plazi:treatment:B845423AFF95FFEA4BA880F752968725:3533A5F7FF96FFEB4921814A536382A1" box="[513,573,1131,1156]" country="Japan" name="Ueno" pageId="10" pageNumber="61">Ueno</location>
36413
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(
<specimenCode id="604A5B57FF96FFEB4982814B526E82A0" box="[674,816,1130,1157]" collectionCode="TNS" country="Japan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15622" lsid="urn:lsid:biocol.org:col:15622" name="National Museum of Nature and Science" pageId="10" pageNumber="61" type="Herbarium">TNS692011</specimenCode>
), 
<collectionCode id="56FD6BE9FF96FFEB4868814B520182A1" box="[840,863,1130,1156]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="10" pageNumber="61" type="Herbarium">
<emphasis id="02982F3EFF96FFEB4868814B520182A1" box="[840,863,1130,1156]" italics="true" pageId="10" pageNumber="61">H</emphasis>
</collectionCode>
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<emphasis id="02982F3EFF96FFEB487F814B523D82A1" box="[863,867,1130,1156]" italics="true" pageId="10" pageNumber="61">.</emphasis>
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<emphasis id="02982F3EFF96FFEB4842814B555882A1" box="[866,1030,1130,1156]" italics="true" pageId="10" pageNumber="61">
<location id="3533A5F7FF96FFEB4842814B52ED82A1" LSID="urn:lsid:plazi:treatment:B845423AFF95FFEA4BA880F752968725:3533A5F7FF96FFEB4842814B52ED82A1" box="[866,947,1130,1156]" country="Japan" name="Ohashi" pageId="10" pageNumber="61">Ohashi</location>
21836
</emphasis>
(
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)
</materialsCitation>
; 
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<collectingRegion id="F2283DCEFF96FFEB4FF7814A546A82A0" box="[1239,1332,1131,1157]" country="Japan" name="Toyama" pageId="10" pageNumber="61">Toyama</collectingRegion>
, 
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<emphasis id="02982F3EFF96FFEB4E64814B540782A1" box="[1348,1369,1130,1156]" italics="true" pageId="10" pageNumber="61">K</emphasis>
</collectionCode>
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<emphasis id="02982F3EFF96FFEB4E79814B540382A1" box="[1369,1373,1130,1156]" italics="true" pageId="10" pageNumber="61">.</emphasis>
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<emphasis id="02982F3EFF96FFEB4E7C814B54F582A1" box="[1372,1451,1130,1156]" italics="true" pageId="10" pageNumber="61">
<location id="3533A5F7FF96FFEB4E7C814B54F582A1" LSID="urn:lsid:plazi:treatment:B845423AFF95FFEA4BA880F752968725:3533A5F7FF96FFEB4E7C814B54F582A1" box="[1372,1451,1130,1156]" country="Japan" name="Shinno" pageId="10" pageNumber="61">Shinno</location>
</emphasis>
(
<collectionCode id="56FD6BE9FF96FFEB4BB081AF5199828C" box="[144,199,1166,1193]" country="Japan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15622" lsid="urn:lsid:biocol.org:col:15622" name="National Museum of Nature and Science" pageId="10" pageNumber="61" type="Herbarium">TNS</collectionCode>
56383)
</materialsCitation>
; 
<materialsCitation id="8084F971FF96FFEB4A0381AE50F1828D" ID-GBIF-Occurrence="3750509324" box="[291,431,1166,1193]" collectionCode="S" country="Japan" location="Yamagata" pageId="10" pageNumber="61" specimenCount="1" stateProvince="Yamagata">
<collectingRegion id="F2283DCEFF96FFEB4A0381AE50CB828C" box="[291,405,1167,1193]" country="Japan" name="Yamagata" pageId="10" pageNumber="61">Yamagata</collectingRegion>
, 
<collectionCode id="56FD6BE9FF96FFEB4A8081AF50F1828D" box="[416,431,1166,1192]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="10" pageNumber="61" type="Herbarium">
<emphasis id="02982F3EFF96FFEB4A8081AF50F1828D" box="[416,431,1166,1192]" italics="true" pageId="10" pageNumber="61">S</emphasis>
</collectionCode>
</materialsCitation>
<emphasis id="02982F3EFF96FFEB4A8F81AF50ED828D" box="[431,435,1166,1192]" italics="true" pageId="10" pageNumber="61">.</emphasis>
<materialsCitation id="8084F971FF96FFEB4A9281AF5387828D" ID-GBIF-Occurrence="3750509343" accessionNumber="TNS36317" box="[434,729,1166,1193]" collectionCode="TNS, H" country="Japan" location="Okuyama" pageId="10" pageNumber="61" specimenCount="1">
<emphasis id="02982F3EFF96FFEB4A9281AF537E828D" box="[434,544,1166,1192]" italics="true" pageId="10" pageNumber="61">
<location id="3533A5F7FF96FFEB4A9281AF537E828D" LSID="urn:lsid:plazi:treatment:B845423AFF95FFEA4BA880F752968725:3533A5F7FF96FFEB4A9281AF537E828D" box="[434,544,1166,1192]" country="Japan" name="Okuyama" pageId="10" pageNumber="61">Okuyama</location>
</emphasis>
(
<collectionCode id="56FD6BE9FF96FFEB490F81AF5338828C" box="[559,614,1166,1193]" country="Japan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15622" lsid="urn:lsid:biocol.org:col:15622" name="National Museum of Nature and Science" pageId="10" pageNumber="61" type="Herbarium">TNS</collectionCode>
36317), 
<collectionCode id="56FD6BE9FF96FFEB49E381AE5387828D" box="[707,729,1167,1192]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="10" pageNumber="61" type="Herbarium">
<emphasis id="02982F3EFF96FFEB49E381AE5387828D" box="[707,729,1167,1192]" italics="true" pageId="10" pageNumber="61">H</emphasis>
</collectionCode>
</materialsCitation>
<emphasis id="02982F3EFF96FFEB49F981AE5383828D" box="[729,733,1167,1192]" italics="true" pageId="10" pageNumber="61">.</emphasis>
<materialsCitation id="8084F971FF96FFEB49FD81AE5575828C" ID-GBIF-Occurrence="3750509318" box="[733,1067,1166,1193]" country="United States of America" location="Koyama" pageId="10" pageNumber="61" specimenCount="1">
<emphasis id="02982F3EFF96FFEB49FD81AE5224828D" box="[733,890,1166,1192]" italics="true" pageId="10" pageNumber="61">
<location id="3533A5F7FF96FFEB49FD81AE5262828D" LSID="urn:lsid:plazi:treatment:B845423AFF95FFEA4BA880F752968725:3533A5F7FF96FFEB49FD81AE5262828D" box="[733,828,1167,1192]" country="United States of America" name="Koyama" pageId="10" pageNumber="61">Koyama</location>
4054
</emphasis>
(
<collectingCountry id="48FBB3BCFF96FFEB48A981AF52F3828C" box="[905,941,1166,1193]" name="United States of America" pageId="10" pageNumber="61">US</collectingCountry>
02159684)
</materialsCitation>
, 
<emphasis id="02982F3EFF96FFEB4F1681AF5511828D" box="[1078,1103,1166,1192]" italics="true" pageId="10" pageNumber="61">K.</emphasis>
<materialsCitation id="8084F971FF96FFEB4F6E81AF5460828C" ID-GBIF-Occurrence="3750509328" box="[1102,1342,1166,1193]" collectionCode="HAST" country="United States of America" location="Akuta" pageId="10" pageNumber="61" specimenCode="HAST40095" specimenCount="1">
<emphasis id="02982F3EFF96FFEB4F6E81AF55CF828D" box="[1102,1169,1166,1192]" italics="true" pageId="10" pageNumber="61">
<location id="3533A5F7FF96FFEB4F6E81AF55CF828D" LSID="urn:lsid:plazi:treatment:B845423AFF95FFEA4BA880F752968725:3533A5F7FF96FFEB4F6E81AF55CF828D" box="[1102,1169,1166,1192]" country="United States of America" name="Akuta" pageId="10" pageNumber="61">Akuta</location>
</emphasis>
(
<specimenCode id="604A5B57FF96FFEB4F8081AF546B828C" box="[1184,1333,1166,1193]" collectionCode="HAST" country="Taiwan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15074" lsid="urn:lsid:biocol.org:col:15074" name="Research Center for Biodiversity, Academia Sinica" pageId="10" pageNumber="61" type="Herbarium">HAST40095</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="78F6A0A7FF96FFEA4B9D819352968725" lastPageId="11" lastPageNumber="62" pageId="10" pageNumber="61" type="discussion">
<paragraph id="3053F32CFF96FFEB4B9D8193502080E0" blockId="10.[136,1452,158,2093]" pageId="10" pageNumber="61">
<emphasis id="02982F3EFF96FFEB4B9D8193505782E9" bold="true" box="[189,265,1202,1228]" pageId="10" pageNumber="61">Notes:</emphasis>
— 
<taxonomicName id="F7EC88AFFF96FFEB4A068192525B82E9" authority="Z. H. Chen &amp; X. F. Jin" authorityName="Z. H. Chen &amp; X. F. Jin" authorityYear="2021" box="[294,773,1202,1229]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="61" phylum="Tracheophyta" rank="species" species="zhejiangense">
<emphasis id="02982F3EFF96FFEB4A068192534782E9" box="[294,537,1202,1229]" italics="true" pageId="10" pageNumber="61">Cirsium zhejiangense</emphasis>
Z.H.Chen &amp; X.F.Jin
</taxonomicName>
was described on the basis of a collection, 
<emphasis id="02982F3EFF96FFEB4FD38192541C82E9" box="[1267,1346,1203,1228]" italics="true" pageId="10" pageNumber="61">X.F.Jin</emphasis>
&amp; 
<emphasis id="02982F3EFF96FFEB4E478192519E82D5" italics="true" pageId="10" pageNumber="61">T.T.Yu 3202</emphasis>
(
<typeStatus id="EF574D8EFF96FFEB4BEF81F7506B82D5" box="[207,309,1238,1264]" pageId="10" pageNumber="61" type="holotype">holotype</typeStatus>
, ZM; isotypes, HTC, PE), from 
<collectingRegion id="F2283DCEFF96FFEB498381F7525482D5" box="[675,778,1238,1264]" country="China" name="Zhejiang" pageId="10" pageNumber="61">Zhejiang</collectingRegion>
, 
<collectingCountry id="48FBB3BCFF96FFEB483581F7520282D5" box="[789,860,1238,1264]" name="China" pageId="10" pageNumber="61">China</collectingCountry>
. In the protologue, the author stated that this species differs from 
<taxonomicName id="F7EC88AFFF96FFEB4A3B81DA50FC8331" authorityName="Makino" authorityYear="1905" baseAuthorityName="Maximowicz" baseAuthorityYear="1874" box="[283,418,1275,1301]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="61" phylum="Tracheophyta" rank="species" species="yezoense">
<emphasis id="02982F3EFF96FFEB4A3B81DA50FC8331" box="[283,418,1275,1301]" italics="true" pageId="10" pageNumber="61">C. yezoense</emphasis>
</taxonomicName>
, a widespread species in 
<collectingCountry id="48FBB3BCFF96FFEB49EF81DA524C8330" box="[719,786,1275,1301]" name="Japan" pageId="10" pageNumber="61">Japan</collectingCountry>
, by having 5–6 rows of phyllaries (vs. 8–10 rows), outer phyllaries shorter than inner (vs. subequal), leaf blade 15–52 × 
<quantity id="F7145EC9FF96FFEB486D803F52F0831C" box="[845,942,1310,1337]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="10" pageNumber="61" unit="cm" value="10.0" valueMax="15.0" valueMin="5.0">5–15 cm</quantity>
(vs. 30–65 × 
<quantity id="F7145EC9FF96FFEB4F69803F55E4831C" box="[1097,1210,1310,1337]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="10" pageNumber="61" unit="cm" value="20.0" valueMax="30.0" valueMin="10.0">10–30 cm</quantity>
) and leaves adaxailly villose and abaxially pubescent (vs. glabrous). But based on observations on both herbarium specimens (including type material) and living plants, we demonstrate that 
<taxonomicName id="F7EC88AFFF96FFEB49C3804652C883A5" authorityName="Z. H. Chen &amp; X. F. Jin" authorityYear="2021" box="[739,918,1382,1409]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="61" phylum="Tracheophyta" rank="species" species="zhejiangense">
<emphasis id="02982F3EFF96FFEB49C3804652C883A5" box="[739,918,1382,1409]" italics="true" pageId="10" pageNumber="61">C. zhejiangense</emphasis>
</taxonomicName>
is indistinguishable from 
<taxonomicName id="F7EC88AFFF96FFEB4F9D8046541D83A5" authorityName="Makino" authorityYear="1905" baseAuthorityName="Maximowicz" baseAuthorityYear="1874" box="[1213,1347,1383,1409]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="61" phylum="Tracheophyta" rank="species" species="yezoense">
<emphasis id="02982F3EFF96FFEB4F9D8046541D83A5" box="[1213,1347,1383,1409]" italics="true" pageId="10" pageNumber="61">C. yezoense</emphasis>
</taxonomicName>
. 
<taxonomicName id="F7EC88AFFF96FFEB4E71804651B28381" authorityName="Makino" authorityYear="1905" baseAuthorityName="Maximowicz" baseAuthorityYear="1874" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="61" phylum="Tracheophyta" rank="species" species="yezoense">
<emphasis id="02982F3EFF96FFEB4E71804651B28381" italics="true" pageId="10" pageNumber="61">Cirsium yezoense</emphasis>
</taxonomicName>
, described from 
<collectingCountry id="48FBB3BCFF96FFEB4A8B80AA50B08380" box="[427,494,1419,1445]" name="Japan" pageId="10" pageNumber="61">Japan</collectingCountry>
, on the basis of a gathering, 
<emphasis id="02982F3EFF96FFEB481380AA557A8381" box="[819,1060,1419,1445]" italics="true" pageId="10" pageNumber="61">C.J. Maximowicz s.n.</emphasis>
(K000778217; 
<figureCitation id="A8D7EFA9FF96FFEB4FF780AB546B8381" box="[1239,1333,1418,1444]" captionStart="FIGURE 6" captionStartId="7.[136,229,1945,1967]" captionTargetBox="[183,1405,186,1925]" captionTargetId="figure-17@7.[183,1405,186,1925]" captionTargetPageId="7" captionText="FIGURE 6. Specimens of Cirsium yezoense. A, B, D. China, Zhejiang, Pan’an, Weixin, Lingjiangyuan, by stream under forest, X.F.Jin &amp; T.T.Yu 3202 (holotype of Cirsium zhejiangense, ZM; isotypes of Cirsium zhejiangense, PE; D. capitulum); C, E. Japan, Hakodate, Mohidzi, C.J.Maximowicz s.n. (type of Cirsium yezoense, K000778217; E. capitulum)." figureDoi="http://doi.org/10.5281/zenodo.6424362" httpUri="https://zenodo.org/record/6424362/files/figure.png" pageId="10" pageNumber="61">Fig. 6 C</figureCitation>
), is a very widely distributed species. Through extensive herbarium survey, we found that the characters of 
<taxonomicName id="F7EC88AFFF96FFEB4FF0808E540883ED" authorityName="Makino" authorityYear="1905" baseAuthorityName="Maximowicz" baseAuthorityYear="1874" box="[1232,1366,1455,1481]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="61" phylum="Tracheophyta" rank="species" species="yezoense">
<emphasis id="02982F3EFF96FFEB4FF0808E540883ED" box="[1232,1366,1455,1481]" italics="true" pageId="10" pageNumber="61">C. yezoense</emphasis>
</taxonomicName>
are not completely consistent with that stated of 
<bibRefCitation id="547D8EDDFF96FFEB496C80F253A283C9" author="Lu, Y. F. &amp; Chen, Z. H. &amp; Sun, W. Y. &amp; Jin, X. F." box="[588,764,1490,1517]" pageId="10" pageNumber="61" pagination="1 - 8" refId="ref5902" refString="Lu, Y. F., Chen, Z. H., Sun, W. Y. &amp; Jin, X. F. (2021) Two new species in Cirsium (Asteraceae) from Zhejiang. Journal of Plant Resources and Environment 30: 1 - 8." type="journal article" year="2021">
Lu 
<emphasis id="02982F3EFF96FFEB495180F253F783C9" box="[625,681,1490,1516]" italics="true" pageId="10" pageNumber="61">et al.</emphasis>
(2021)
</bibRefCitation>
. It can be clearly seen from the involucre of type and common specimens of 
<taxonomicName id="F7EC88AFFF96FFEB4A0380D650F68035" authorityName="Makino" authorityYear="1905" baseAuthorityName="Maximowicz" baseAuthorityYear="1874" box="[291,424,1527,1553]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="61" phylum="Tracheophyta" rank="species" species="yezoense">
<emphasis id="02982F3EFF96FFEB4A0380D650F68035" box="[291,424,1527,1553]" italics="true" pageId="10" pageNumber="61">C. yezoense</emphasis>
</taxonomicName>
(
<figureCitation id="A8D7EFA9FF96FFEB4A9880D750A08035" box="[440,510,1526,1552]" captionStart="FIGURE 6" captionStartId="7.[136,229,1945,1967]" captionTargetBox="[183,1405,186,1925]" captionTargetId="figure-17@7.[183,1405,186,1925]" captionTargetPageId="7" captionText="FIGURE 6. Specimens of Cirsium yezoense. A, B, D. China, Zhejiang, Pan’an, Weixin, Lingjiangyuan, by stream under forest, X.F.Jin &amp; T.T.Yu 3202 (holotype of Cirsium zhejiangense, ZM; isotypes of Cirsium zhejiangense, PE; D. capitulum); C, E. Japan, Hakodate, Mohidzi, C.J.Maximowicz s.n. (type of Cirsium yezoense, K000778217; E. capitulum)." figureDoi="http://doi.org/10.5281/zenodo.6424362" httpUri="https://zenodo.org/record/6424362/files/figure.png" pageId="10" pageNumber="61">Fig. 6</figureCitation>
: E, 
<figureCitation id="A8D7EFA9FF96FFEB490A80D753338034" box="[554,621,1526,1553]" captionStart="FIGURE 7" captionStartId="8.[136,229,1963,1985]" captionTargetBox="[151,1436,186,1943]" captionTargetId="figure-35@8.[151,1436,186,1943]" captionTargetPageId="8" captionText="FIGURE 7. Selected specimens of Cirsium yezoense. A. Japan, Prov. Koodzuke, Pref. Gumma, M.Furuse 47139 (PE01292170); B. Japan, Prov. Rikuchu, Pref. Iwate, M.Furuse 41390 (PE01292175). C. Japan, Pref. Iwate, T.Kawasaki 7309 (HAST82445); D. Japan, Pref. Yamagata, H.Koyama 4054 (US02159684)." figureDoi="http://doi.org/10.5281/zenodo.6424364" httpUri="https://zenodo.org/record/6424364/files/figure.png" pageId="10" pageNumber="61">Fig. 7</figureCitation>
) that the outer phyllaries is shorter than the inner ones and 
<taxonomicName id="F7EC88AFFF96FFEB4E2C80D654CC8035" authorityName="Makino" authorityYear="1905" baseAuthorityName="Maximowicz" baseAuthorityYear="1874" box="[1292,1426,1527,1553]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="61" phylum="Tracheophyta" rank="species" species="yezoense">
<emphasis id="02982F3EFF96FFEB4E2C80D654CC8035" box="[1292,1426,1527,1553]" italics="true" pageId="10" pageNumber="61">C. yezoense</emphasis>
</taxonomicName>
is itself indeed somewhat variable in the rows of phyllaries. In fact, according to the description of 
<taxonomicName id="F7EC88AFFF96FFEB4F90833A546D8011" authorityName="Makino" authorityYear="1905" baseAuthorityName="Maximowicz" baseAuthorityYear="1874" box="[1200,1331,1563,1589]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="61" phylum="Tracheophyta" rank="species" species="yezoense">
<emphasis id="02982F3EFF96FFEB4F90833A546D8011" box="[1200,1331,1563,1589]" italics="true" pageId="10" pageNumber="61">C. yezoense</emphasis>
</taxonomicName>
by 
<bibRefCitation id="547D8EDDFF96FFEB4E7A833B518D807C" author="Kadota, Y." pageId="10" pageNumber="61" pagination="119 - 151" refId="ref5822" refString="Kadota, Y. (1995) Cirsium Mill. In: Iwatsuki, K., Yamazaki, T., Boufford, D. E. &amp; Ohba, H. (Eds.) Flora of Japan, vol. IIIb. Kodansha, Tokyo, pp. 119 - 151." type="book chapter" year="1995">Kadota (1995)</bibRefCitation>
in Flora of 
<collectingCountry id="48FBB3BCFF96FFEB4A7D831E50FE807C" box="[349,416,1599,1625]" name="Japan" pageId="10" pageNumber="61">Japan</collectingCountry>
, its phyllaries are 6-seriate, outer phyllaries shorter than inner ones, leaves blades glabrous on both surfaces, or covered with short brown hairs on upper surface, or sparingly arachnoid on lower surface. These descriptions of 
<taxonomicName id="F7EC88AFFF96FFEB4A1483A650E78085" authorityName="Makino" authorityYear="1905" baseAuthorityName="Maximowicz" baseAuthorityYear="1874" box="[308,441,1671,1697]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="61" phylum="Tracheophyta" rank="species" species="yezoense">
<emphasis id="02982F3EFF96FFEB4A1483A650E78085" box="[308,441,1671,1697]" italics="true" pageId="10" pageNumber="61">C. yezoense</emphasis>
</taxonomicName>
are contradictory to those given by 
<bibRefCitation id="547D8EDDFF96FFEB486A83A652A58085" author="Lu, Y. F. &amp; Chen, Z. H. &amp; Sun, W. Y. &amp; Jin, X. F." box="[842,1019,1670,1697]" pageId="10" pageNumber="61" pagination="1 - 8" refId="ref5902" refString="Lu, Y. F., Chen, Z. H., Sun, W. Y. &amp; Jin, X. F. (2021) Two new species in Cirsium (Asteraceae) from Zhejiang. Journal of Plant Resources and Environment 30: 1 - 8." type="journal article" year="2021">
Lu 
<emphasis id="02982F3EFF96FFEB485083A652FF8085" box="[880,929,1670,1696]" italics="true" pageId="10" pageNumber="61">et al</emphasis>
. (2021)
</bibRefCitation>
, making 
<taxonomicName id="F7EC88AFFF96FFEB4F4183A655B88085" authorityName="Makino" authorityYear="1905" baseAuthorityName="Maximowicz" baseAuthorityYear="1874" box="[1121,1254,1671,1697]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="61" phylum="Tracheophyta" rank="species" species="yezoense">
<emphasis id="02982F3EFF96FFEB4F4183A655B88085" box="[1121,1254,1671,1697]" italics="true" pageId="10" pageNumber="61">C. yezoense</emphasis>
</taxonomicName>
indistinguishable from 
<taxonomicName id="F7EC88AFFF96FFEB4BE5838A502980E1" authorityName="Z. H. Chen &amp; X. F. Jin" authorityYear="2021" box="[197,375,1706,1733]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="61" phylum="Tracheophyta" rank="species" species="zhejiangense">
<emphasis id="02982F3EFF96FFEB4BE5838A502980E1" box="[197,375,1706,1733]" italics="true" pageId="10" pageNumber="61">C. zhejiangense</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="3053F32CFF96FFEB4B9D83EE50218E2D" blockId="10.[136,1452,158,2093]" pageId="10" pageNumber="61">
In 2021 we visited Tianmushan Mt. in 
<collectingRegion id="F2283DCEFF96FFEB495183EF526080CD" box="[625,830,1742,1768]" country="China" name="Zhejiang" pageId="10" pageNumber="61">Zhejiang province</collectingRegion>
and Hengshan Mt. in 
<collectingRegion id="F2283DCEFF96FFEB4F1883EE55B280CD" box="[1080,1260,1742,1769]" country="China" name="Hunan" pageId="10" pageNumber="61">Hunan province</collectingRegion>
and successfully found two populations of 
<taxonomicName id="F7EC88AFFF96FFEB4A9383D253398129" authorityName="Z. H. Chen &amp; X. F. Jin" authorityYear="2021" box="[435,615,1778,1805]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="61" phylum="Tracheophyta" rank="species" species="zhejiangense">
<emphasis id="02982F3EFF96FFEB4A9383D253398129" box="[435,615,1778,1805]" italics="true" pageId="10" pageNumber="61">C. zhejiangense</emphasis>
</taxonomicName>
. We found that the size of leaves of this species varies greatly (25–50 × 
<quantity id="F7145EC9FF96FFEB4BA8823751A78114" box="[136,249,1814,1841]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="10" pageNumber="61" unit="cm" value="20.0" valueMax="30.0" valueMin="10.0">10–30 cm</quantity>
, 
<figureCitation id="A8D7EFA9FF96FFEB4A23823750188114" box="[259,326,1814,1841]" captionStart="FIGURE 5" captionStartId="6.[136,229,1979,2001]" captionTargetBox="[151,1436,186,1958]" captionTargetId="figure-35@6.[151,1436,186,1958]" captionTargetPageId="6" captionText="FIGURE 5. Living plants of Cirsium yezoense. A. habitat; B. plant; C. plant; D. synflorescence; E. basal leaves; F. cauline leaves; G. capitulum; H. longitudinal section of capitulum; I. florets; J. anther; K. phyllaries; L. pappus; M. seed.All photos by Zi-Chao Jin based on Z.C.Jin ZJ2021009 (IBSC) from Tianmushan, Lin’an, Zhejiang." figureDoi="http://doi.org/10.5281/zenodo.6424360" httpUri="https://zenodo.org/record/6424360/files/figure.png" pageId="10" pageNumber="61">Fig. 5</figureCitation>
: E) and the leaf blade is pinnatilobate or pinnatifid. We found the leaf indumentum in 
<taxonomicName id="F7EC88AFFF96FFEB4FDB823654F28115" authorityName="Z. H. Chen &amp; X. F. Jin" authorityYear="2021" box="[1275,1452,1814,1841]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="61" phylum="Tracheophyta" rank="species" species="zhejiangense">
<emphasis id="02982F3EFF96FFEB4FDB823654F28115" box="[1275,1452,1814,1841]" italics="true" pageId="10" pageNumber="61">C. zhejiangense</emphasis>
</taxonomicName>
is variable in both 
<collectingRegion id="F2283DCEFF96FFEB4A7A821A50FB8170" box="[346,421,1851,1877]" country="China" name="Hunan" pageId="10" pageNumber="61">Hunan</collectingRegion>
and 
<collectingRegion id="F2283DCEFF96FFEB4AFC821B531E8171" box="[476,576,1850,1876]" country="China" name="Zhejiang" pageId="10" pageNumber="61">Zhejiang</collectingRegion>
populations, being sometimes glabrous, sometimes covered with short brown hairs on the upper surface, or sparingly arachnoid on the lower surface. The leaf indumentum in 
<taxonomicName id="F7EC88AFFF96FFEB4FE3827E542B815D" authorityName="Z. H. Chen &amp; X. F. Jin" authorityYear="2021" box="[1219,1397,1886,1913]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="61" phylum="Tracheophyta" rank="species" species="zhejiangense">
<emphasis id="02982F3EFF96FFEB4FE3827E542B815D" box="[1219,1397,1886,1913]" italics="true" pageId="10" pageNumber="61">C. zhejiangense</emphasis>
</taxonomicName>
is in fact consistent with the description of 
<taxonomicName id="F7EC88AFFF96FFEB491D82A2522D81B8" authority="(Kadota 1995)" baseAuthorityName="Kadota" baseAuthorityYear="1995" box="[573,883,1922,1949]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="61" phylum="Tracheophyta" rank="species" species="yezoense">
<emphasis id="02982F3EFF96FFEB491D82A2539A81B9" box="[573,708,1923,1949]" italics="true" pageId="10" pageNumber="61">C. yezoense</emphasis>
(
<bibRefCitation id="547D8EDDFF96FFEB49F382A3523481B8" author="Kadota, Y." box="[723,874,1922,1949]" pageId="10" pageNumber="61" pagination="119 - 151" refId="ref5822" refString="Kadota, Y. (1995) Cirsium Mill. In: Iwatsuki, K., Yamazaki, T., Boufford, D. E. &amp; Ohba, H. (Eds.) Flora of Japan, vol. IIIb. Kodansha, Tokyo, pp. 119 - 151." type="book chapter" year="1995">Kadota 1995</bibRefCitation>
)
</taxonomicName>
. To summarize, the morphological variation of 
<taxonomicName id="F7EC88AFFF96FFEB4EB282A2504781E5" authorityName="Z. H. Chen &amp; X. F. Jin" authorityYear="2021" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="61" phylum="Tracheophyta" rank="species" species="zhejiangense">
<emphasis id="02982F3EFF96FFEB4EB282A2504781E5" italics="true" pageId="10" pageNumber="61">C. zhejiangense</emphasis>
</taxonomicName>
is within the range of 
<taxonomicName id="F7EC88AFFF96FFEB493B828653FF81E5" baseAuthorityName="Kadota" baseAuthorityYear="1995" box="[539,673,1959,1985]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="61" phylum="Tracheophyta" rank="species" species="yezoense">
<emphasis id="02982F3EFF96FFEB493B828653FF81E5" box="[539,673,1959,1985]" italics="true" pageId="10" pageNumber="61">C. yezoense</emphasis>
</taxonomicName>
. Therefore, the morphological characters used by 
<bibRefCitation id="547D8EDDFF96FFEB4FFB828654D381E5" author="Lu, Y. F. &amp; Chen, Z. H. &amp; Sun, W. Y. &amp; Jin, X. F." box="[1243,1421,1958,1985]" pageId="10" pageNumber="61" pagination="1 - 8" refId="ref5902" refString="Lu, Y. F., Chen, Z. H., Sun, W. Y. &amp; Jin, X. F. (2021) Two new species in Cirsium (Asteraceae) from Zhejiang. Journal of Plant Resources and Environment 30: 1 - 8." type="journal article" year="2021">
Lu 
<emphasis id="02982F3EFF96FFEB4E228286546581E5" box="[1282,1339,1958,1984]" italics="true" pageId="10" pageNumber="61">et al.</emphasis>
(2021)
</bibRefCitation>
to distinguish 
<taxonomicName id="F7EC88AFFF96FFEB4A2982EA50E481C1" authorityName="Z. H. Chen &amp; X. F. Jin" authorityYear="2021" box="[265,442,1994,2021]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="61" phylum="Tracheophyta" rank="species" species="zhejiangense">
<emphasis id="02982F3EFF96FFEB4A2982EA50E481C1" box="[265,442,1994,2021]" italics="true" pageId="10" pageNumber="61">C. zhejiangense</emphasis>
</taxonomicName>
from 
<taxonomicName id="F7EC88AFFF96FFEB4ADC82EA532181C1" baseAuthorityName="Kadota" baseAuthorityYear="1995" box="[508,639,1995,2021]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="61" phylum="Tracheophyta" rank="species" species="yezoense">
<emphasis id="02982F3EFF96FFEB4ADC82EA532181C1" box="[508,639,1995,2021]" italics="true" pageId="10" pageNumber="61">C. yezoense</emphasis>
</taxonomicName>
are not diagnostic. We therefore reduce 
<taxonomicName id="F7EC88AFFF96FFEB4F1C82EA55B381C1" authorityName="Z. H. Chen &amp; X. F. Jin" authorityYear="2021" box="[1084,1261,1994,2021]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="61" phylum="Tracheophyta" rank="species" species="zhejiangense">
<emphasis id="02982F3EFF96FFEB4F1C82EA55B381C1" box="[1084,1261,1994,2021]" italics="true" pageId="10" pageNumber="61">C. zhejiangense</emphasis>
</taxonomicName>
to the synonymy of 
<taxonomicName id="F7EC88AFFF96FFEB4B8682CE50758E2D" baseAuthorityName="Kadota" baseAuthorityYear="1995" box="[166,299,2031,2057]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="61" phylum="Tracheophyta" rank="species" species="yezoense">
<emphasis id="02982F3EFF96FFEB4B8682CE50758E2D" box="[166,299,2031,2057]" italics="true" pageId="10" pageNumber="61">C. yezoense</emphasis>
</taxonomicName>
herein.
</paragraph>
<paragraph id="3053F32CFF96FFEA4B9D8D3252968725" blockId="10.[136,1452,158,2093]" lastBlockId="11.[136,1451,158,257]" lastPageId="11" lastPageNumber="62" pageId="10" pageNumber="61">
Because its phyllaries lack marginal spinules, wings, and scarious appendage and leaves abaxially lack spinules, 
<taxonomicName id="F7EC88AFFF97FFEA4BA885BE5011869D" baseAuthorityName="Kadota" baseAuthorityYear="1995" box="[136,335,159,185]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="11" pageNumber="62" phylum="Tracheophyta" rank="species" species="yezoense">
<emphasis id="02982F3EFF97FFEA4BA885BE5011869D" box="[136,335,159,185]" italics="true" pageId="11" pageNumber="62">Cirsium yezoense</emphasis>
</taxonomicName>
should belong to 
<emphasis id="02982F3EFF97FFEA493E85BE5366869C" box="[542,568,159,185]" italics="true" pageId="11" pageNumber="62">C.</emphasis>
sect. 
<taxonomicName id="F7EC88AFFF97FFEA495C85BE5388869C" authorityName="Miller" authorityYear="1754" box="[636,726,159,185]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="11" pageNumber="62" phylum="Tracheophyta" rank="genus">
<emphasis id="02982F3EFF97FFEA495C85BE5388869C" box="[636,726,159,185]" italics="true" pageId="11" pageNumber="62">Cirsium</emphasis>
</taxonomicName>
. 
<taxonomicName id="F7EC88AFFF97FFEA49C685BE52F3869D" baseAuthorityName="Kadota" baseAuthorityYear="1995" box="[742,941,159,185]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="11" pageNumber="62" phylum="Tracheophyta" rank="species" species="yezoense">
<emphasis id="02982F3EFF97FFEA49C685BE52F3869D" box="[742,941,159,185]" italics="true" pageId="11" pageNumber="62">Cirsium yezoense</emphasis>
</taxonomicName>
is mostly similar to 
<taxonomicName id="F7EC88AFFF97FFEA4FBB85BE546D869D" authorityName="Candolle" authorityYear="1838" box="[1179,1331,159,185]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="11" pageNumber="62" phylum="Tracheophyta" rank="species" species="japonicum">
<emphasis id="02982F3EFF97FFEA4FBB85BE546D869D" box="[1179,1331,159,185]" italics="true" pageId="11" pageNumber="62">C. japonicum</emphasis>
</taxonomicName>
and many specimens of 
<taxonomicName id="F7EC88AFFF97FFEA4A0285E250F986F9" baseAuthorityName="Kadota" baseAuthorityYear="1995" box="[290,423,195,221]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="11" pageNumber="62" phylum="Tracheophyta" rank="species" species="yezoense">
<emphasis id="02982F3EFF97FFEA4A0285E250F986F9" box="[290,423,195,221]" italics="true" pageId="11" pageNumber="62">C. yezoense</emphasis>
</taxonomicName>
in Chinese herbaria were usually incorrectly identified as 
<taxonomicName id="F7EC88AFFF97FFEA4F1485E2559486F9" authorityName="Candolle" authorityYear="1838" box="[1076,1226,195,221]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="11" pageNumber="62" phylum="Tracheophyta" rank="species" species="japonicum">
<emphasis id="02982F3EFF97FFEA4F1485E2559486F9" box="[1076,1226,195,221]" italics="true" pageId="11" pageNumber="62">C. japonicum</emphasis>
</taxonomicName>
. The comparison of morphological characters between these two species are shown in 
<tableCitation id="7D6EC697FF97FFEA484E85C7529C8725" box="[878,962,230,256]" captionStart="TABLE 1" captionStartId="11.[136,229,302,328]" captionTargetPageId="11" captionText="TABLE 1. Comparison of morphological characters and habitats between Cirsium sieboldii, C. yezoense and C. japonicum." httpUri="http://table.plazi.org/id/6493A3A4FF97FFEA4BA8840F505B8748" pageId="11" pageNumber="62" tableUuid="6493A3A4FF97FFEA4BA8840F505B8748">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>