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<document id="82FE63957CAF8E1A298F3489600EC2AD" ID-CLB-Dataset="8481" ID-DOI="10.11646/zootaxa.4853.4.7" ID-GBIF-Dataset="a416516e-a6f0-4fab-9e3c-1b50abe53f1e" ID-ISSN="1175-5326" ID-Zenodo-Dep="4506498" ID-ZooBank="C3FFB4EE-B46B-41D2-99CB-9A5B0E8B3F68" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609606137281" checkinUser="plazi" docAuthor="Martínez, Leonel, Brescovit, Antonio Domingos &amp; Oliveira, Luiz Fernando M." docDate="2020" docId="AA067B1DFFF7FF95FF67F891AA293EA0" docLanguage="en" docName="zootaxa.4853.4.7.pdf" docOrigin="Zootaxa 4853 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macrophyes sanzi Martínez &amp; Brescovit &amp; Oliveira 2020, new species" docType="treatment" docVersion="8" lastPageNumber="584" masterDocId="563F0365FFF6FF96FFF0FFB6A9643B3D" masterDocTitle="Two new species of the ghost spider genus Macrophyes O. Pickard-Cambridge 1893 from Colombia and description of the first male of Mesilla Simon, 1903 (Araneae: Anyphaenidae: Anyphaeninae)" masterLastPageNumber="590" masterPageNumber="581" pageNumber="582" updateTime="1698851393780" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="D9A68838CAF45859EDE6A6E2C6FF0FED" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="91388235218007B26CFF1A71AEEE6FB8">
<mods:title id="28B58E3F3A03FA5E4A43A4C9FCD599BA">Two new species of the ghost spider genus Macrophyes O. Pickard-Cambridge 1893 from Colombia and description of the first male of Mesilla Simon, 1903 (Araneae: Anyphaenidae: Anyphaeninae)</mods:title>
</mods:titleInfo>
<mods:name id="92132A493FE8B7A901AC7EFA29A075C9" type="personal">
<mods:role id="9E19167CA10F048D2C4E53A376FDE944">
<mods:roleTerm id="8BD7891A705142A19731F46AC38E2E77">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="27FD2CD2A82C5197D368C4829860E14C">Martínez, Leonel</mods:namePart>
<mods:nameIdentifier id="67A22F504D3823F8199678188EE02915" type="ORCID">0000-0002-4166-0561</mods:nameIdentifier>
<mods:affiliation id="913DDBF9092AF7D3B64822DB1BE3EB55">Grupo de Investigación Biodiversidad del Caribe Colombiano, Semillero de Investigación Sistemática de Artrópodos Neotropicales, Departamento de Biología, Universidad del Atlántico, Barranquilla, Colombia. &amp; leonelmarbio @ gmail. com; https: // orcid. org / 0000 - 0002 - 4166 - 0561</mods:affiliation>
<mods:nameIdentifier id="D5E7FD2A5899AC8EB7C6F1EB2157BFBA" type="email">leonelmarbio@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="B4208467CECAFF0F415257DF65B071C6" type="personal">
<mods:role id="9CABC85444268DDFBAF9E3C7354638C6">
<mods:roleTerm id="194DFD5EFD3B6720105B7638FF147319">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="246E6C6CF9508A4F541B09E0B39F9DF2">Brescovit, Antonio Domingos</mods:namePart>
<mods:nameIdentifier id="77F35117A0F7383D68E5CF2A09F77709" type="ORCID">0000-0002-1511-5324</mods:nameIdentifier>
<mods:affiliation id="58CE371CFE2C23C6C41FECE8940CB1E5">Laboratório de Coleções Zoológicas, Instituto Butantan, Av. Vital Brasil, 1500, 05503 - 900, São Paulo, SP, Brazil. &amp; antonio. brescovit @ butantan. gov. br; https: // orcid. org / 000.002 - 1511 - 5324 &amp; luiz. moura. lfm @ gmail. com; https: // orcid. org / 0000 - 0002 - 4054 - 4681</mods:affiliation>
<mods:nameIdentifier id="CC1E251D80E79614F9D1D218B8DD6EC6" type="email">antonio.brescovit@butantan.gov.br</mods:nameIdentifier>
</mods:name>
<mods:name id="31D8E0E61427618D8447A7294023AD43" type="personal">
<mods:role id="F84C1FC0F0E80EAE257E176C188B492C">
<mods:roleTerm id="7B14838C2D8C067053635D7F66ABAC4B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2729DBB281999E0B5B6537E7B82E4401">Oliveira, Luiz Fernando M.</mods:namePart>
<mods:nameIdentifier id="AD3E3F1D16E4D01D4D002D366BF6F5F3" type="ORCID">0000-0002-4054-4681</mods:nameIdentifier>
<mods:affiliation id="628B3ACA8A851F9046205150FD254330">Laboratório de Coleções Zoológicas, Instituto Butantan, Av. Vital Brasil, 1500, 05503 - 900, São Paulo, SP, Brazil. &amp; antonio. brescovit @ butantan. gov. br; https: // orcid. org / 000.002 - 1511 - 5324 &amp; luiz. moura. lfm @ gmail. com; https: // orcid. org / 0000 - 0002 - 4054 - 4681 &amp; Programa de Pós-Graduação em Zoologia, Instituto de Biociências, Rua do Matão, 14, 05508 - 090, Universidade de São Paulo, São Paulo, SP, Brazil. &amp; luiz. moura. lfm @ gmail. com; https: // orcid. org / 0000 - 0002 - 4054 - 4681</mods:affiliation>
<mods:nameIdentifier id="EFB87EAA712798D26F928FCFE452858B" type="email">antonio.brescovit@butantan.gov.br</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="149528E2CB9D262CD2EC59882F56B712">text</mods:typeOfResource>
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<mods:titleInfo id="681F1FE5A305744A579BA1068B1ADAB1">
<mods:title id="B2F66E9D5D6020E72D5F98843C627CE9">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="BD40FDF9943A8C815BFC25B1E5BD5314">
<mods:date id="8885255F92B7662DA09BC6FB5382CEF1">2020</mods:date>
<mods:detail id="A719FECE1D86140BCBB79723B1937894" type="pubDate">
<mods:number id="00CDE3B2F9EB397AD1C429CED7EB304A">2020-09-24</mods:number>
</mods:detail>
<mods:detail id="27CCE91418CE0003959A2D4F6E9CD329" type="volume">
<mods:number id="6D9DCB827DF1C1FF534DC41D28F38085">4853</mods:number>
</mods:detail>
<mods:detail id="1FADC11124ABC65DFD5FB286D4C792C3" type="issue">
<mods:number id="31D1365A34B295027171AF85E01F7A27">4</mods:number>
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<mods:classification id="C4F02E32C9A6916B8D34244E89D2307B">journal article</mods:classification>
<mods:identifier id="070A7C50C14F1297B4AFACFF4B22DA54" type="CLB-Dataset">8481</mods:identifier>
<mods:identifier id="90906B3EDF86F1C9CD5088B6CE7F97C8" type="DOI">10.11646/zootaxa.4853.4.7</mods:identifier>
<mods:identifier id="C577245DD5842905B6A5B64685EFB5BE" type="GBIF-Dataset">a416516e-a6f0-4fab-9e3c-1b50abe53f1e</mods:identifier>
<mods:identifier id="163D51FD1F9D1CEF2B651FB8A5F144BD" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="AB239BFA46493CF8124BE07332E1888C" type="Zenodo-Dep">4506498</mods:identifier>
<mods:identifier id="9586AEA2BE05DDA79CE86F167BD10E9C" type="ZooBank">C3FFB4EE-B46B-41D2-99CB-9A5B0E8B3F68</mods:identifier>
</mods:mods>
<treatment id="AA067B1DFFF7FF95FF67F891AA293EA0" ID-DOI="http://doi.org/10.5281/zenodo.4506504" ID-GBIF-Taxon="177209789" ID-Zenodo-Dep="4506504" LSID="urn:lsid:plazi:treatment:AA067B1DFFF7FF95FF67F891AA293EA0" httpUri="http://treatment.plazi.org/id/AA067B1DFFF7FF95FF67F891AA293EA0" lastPageId="3" lastPageNumber="584" pageId="1" pageNumber="582">
<subSubSection id="6AB59980FFF7FF97FF67F891AB6C3C7F" box="[151,520,1831,1858]" pageId="1" pageNumber="582" type="nomenclature">
<paragraph id="2210CA0BFFF7FF97FF67F891AB6C3C7F" blockId="1.[151,520,1831,1894]" box="[151,520,1831,1858]" pageId="1" pageNumber="582">
<heading id="79587D67FFF7FF97FF67F891AB6C3C7F" bold="true" box="[151,520,1831,1858]" fontSize="11" level="1" pageId="1" pageNumber="582" reason="1">
<emphasis id="10DB1619FFF7FF97FF67F891AB6C3C7F" bold="true" box="[151,520,1831,1858]" pageId="1" pageNumber="582">
<taxonomicName id="E5AFB188FFF7FF97FF67F891A8153C7F" ID-CoL="98ZBV" authority="Martínez &amp; Brescovit &amp; Oliveira, 2020" authorityName="Martínez &amp; Brescovit &amp; Oliveira" authorityYear="2020" box="[151,369,1831,1858]" class="Arachnida" family="Anyphaenidae" genus="Macrophyes" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="582" phylum="Arthropoda" rank="species" species="sanzi" status="sp. nov.">
<emphasis id="10DB1619FFF7FF97FF67F891A8153C7F" bold="true" box="[151,369,1831,1858]" italics="true" pageId="1" pageNumber="582">Macrophyes sanzi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0BE8AB62FFF7FF97FE88F89EAB6C3C7F" box="[376,520,1832,1858]" pageId="1" pageNumber="582" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6AB59980FFF7FF97FF67F8FAA8073C5B" box="[151,355,1868,1894]" pageId="1" pageNumber="582" type="description">
<paragraph id="2210CA0BFFF7FF97FF67F8FAA8073C5B" blockId="1.[151,520,1831,1894]" box="[151,355,1868,1894]" pageId="1" pageNumber="582">
<heading id="79587D67FFF7FF97FF67F8FAA8073C5B" bold="true" box="[151,355,1868,1894]" fontSize="11" level="1" pageId="1" pageNumber="582" reason="1">
<emphasis id="10DB1619FFF7FF97FF67F8FAA8073C5B" bold="true" box="[151,355,1868,1894]" pageId="1" pageNumber="582">
<figureCitation id="BA94D68EFFF7FF97FF67F8FAA8673C5B" box="[151,259,1868,1894]" captionStart="FIGURE" captionStartId="2.[151,168,1741,1767]" captionTargetBox="[189,1398,306,1717]" captionTargetId="figure-76@2.[189,1398,306,1717]" captionTargetPageId="2" captionText="FIGURE 1A−G. Macrophyes sanzi n. sp. Male (MPUJ_ENT 0061936): A habitus, dorsal view; C left palp, retroventral view; D same, retrolateral view; E same, ventral view. Female (MPUJ_ENT 0061923): B habitus, dorsal view; F epigynum, ventral view; G epigynum, dorsal view.Abbreviations:A, atrium; CD, copulatory ducts; E, embolus; FD, fertilization ducts; LB, lateral borders; MA, median apophysis; RTA, retrolateral tibial apophysis; S, spermathecae; SD, spermatic ducts; SR, seminal receptacles; T, tegulum; VTP, ventral tegular process. Scale bars: A, B, F: 2mm; C: 1mm; D−E: 0.5mm; F, G: 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.4506500" httpUri="https://zenodo.org/record/4506500/files/figure.png" pageId="1" pageNumber="582">Figures 1</figureCitation>
A−G, 4.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6AB59980FFF7FF94FF67F822AA0C3BE8" lastPageId="2" lastPageNumber="583" pageId="1" pageNumber="582" type="materials_examined">
<paragraph id="2210CA0BFFF7FF94FF67F822AA0C3BE8" blockId="1.[151,1436,1940,2039]" lastBlockId="2.[151,1436,150,285]" lastPageId="2" lastPageNumber="583" pageId="1" pageNumber="582">
<emphasis id="10DB1619FFF7FF97FF67F822A8233C93" bold="true" box="[151,327,1940,1966]" pageId="1" pageNumber="582">Type material.</emphasis>
<materialsCitation id="92C7C056FFF7FF97FEBFF822AD8E3CEE" ID-GBIF-Occurrence="3029721303" collectingDate="2009-03-28" collectionCode="MPUJ" collectorName="M. Vargas &amp; C. Zornasa" country="Colombia" elevation="90" latitude="8.631653" location="Jardin Botanico del Darien" longLatPrecision="1" longitude="-77.35651" pageId="1" pageNumber="582" specimenCode="ENT 0061936" specimenCount="1" typeStatus="holotype">
<typeStatus id="FD1474A9FFF7FF97FEBFF822A8D83C93" box="[335,444,1940,1966]" pageId="1" pageNumber="582" type="holotype">Holotype</typeStatus>
: male from
<location id="27709CD0FFF7FF97FDB6F822AA1C3C93" LSID="urn:lsid:plazi:treatment:AA067B1DFFF7FF95FF67F891AA293EA0:27709CD0FFF7FF97FDB6F822AA1C3C93" box="[582,888,1940,1966]" country="Colombia" latitude="8.631653" longLatPrecision="1" longitude="-77.35651" name="Jardin Botanico del Darien" pageId="1" pageNumber="582">Jardin Botánico del Darién</location>
(
<geoCoordinate id="479BACCCFFF7FF97FC79F823AD433C8D" box="[905,1063,1941,1968]" degrees="8" direction="north" minutes="37" orientation="latitude" pageId="1" pageNumber="582" precision="1" seconds="53.95" value="8.631653">8°37′53.95″N</geoCoordinate>
;
<geoCoordinate id="479BACCCFFF7FF97FBC4F822AD943C8D" box="[1076,1264,1940,1968]" degrees="77" direction="west" minutes="21" orientation="longitude" pageId="1" pageNumber="582" precision="1" seconds="23.43" value="-77.35651">77° 21′23.43″W</geoCoordinate>
),
<quantity id="E55767EEFFF7FF97FAF4F822AC5D3C93" box="[1284,1337,1940,1966]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="582" unit="m" value="90.0">
<elevation id="A9822D38FFF7FF97FAF4F822AC5D3C93" box="[1284,1337,1940,1966]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="582" unit="m" value="90.0">90m</elevation>
</quantity>
, Acandí, Capurgana, Chocó,
<collectingCountry id="5AB88A9BFFF7FF97FE8AF80EA88F3CEF" box="[378,491,1976,2002]" name="Colombia" pageId="1" pageNumber="582">Colombia</collectingCountry>
,
<date id="5611ECCBFFF7FF97FE08F80EAB1D3CEF" box="[504,633,1976,2002]" pageId="1" pageNumber="582" value="2009-03-28">
<collectingDate id="46551523FFF7FF97FE08F80EAB1D3CEF" box="[504,633,1976,2002]" pageId="1" pageNumber="582" value="2009-03-28">28.III.2009</collectingDate>
</date>
,
<collectorName id="8F5AAFDDFFF7FF97FD77F80FAB9F3CEE" box="[647,763,1977,2003]" pageId="1" pageNumber="582">M. Vargas</collectorName>
&amp;
<collectorName id="8F5AAFDDFFF7FF97FCD3F80EAAC43CEE" box="[803,928,1976,2003]" pageId="1" pageNumber="582">C. Zornasa</collectorName>
leg. (
<collectionCode id="44BE52CEFFF7FF97FC13F80EAD4F3CEF" box="[995,1067,1976,2002]" pageId="1" pageNumber="582">MPUJ</collectionCode>
_
<specimenCode id="72096270FFF7FF97FBCAF80EAD853CEE" box="[1082,1249,1976,2003]" collectionCode="MPUJ" pageId="1" pageNumber="582">ENT 0061936</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="92C7C056FFF7FF94FB06F80FAA003BE8" ID-GBIF-Occurrence="3029721304" collectingDate="2008-11-03" collectingDateMax="2009-10-07" collectingDateMin="2008-11-03" collectionCode="MPUJ" collectorName="One &amp; J. Aristizabal" country="Colombia" elevation="250" lastPageId="2" lastPageNumber="583" latitude="8.631653" location="Jardin Botanico del Darien" longLatPrecision="1" longitude="-77.35651" pageId="1" pageNumber="582" specimenCode="ENT 0061923, ENT 0061910, ENT 0061884, ENT 0061897, IBSP 244556" specimenCount="1" typeStatus="paratype">
<typeStatus id="FD1474A9FFF7FF97FB06F80FAC033CEE" box="[1270,1383,1977,2003]" pageId="1" pageNumber="582" type="paratype">Paratypes</typeStatus>
: one male and female same data as holotype (
<collectionCode id="44BE52CEFFF7FF97FD93F86AABCF3CCB" box="[611,683,2012,2038]" pageId="1" pageNumber="582">MPUJ</collectionCode>
_
<specimenCode id="72096270FFF7FF97FD4AF86AAA3E3CCB" box="[698,858,2012,2038]" collectionCode="MPUJ" pageId="1" pageNumber="582">ENT 0061923</specimenCode>
[m#1];
<collectionCode id="44BE52CEFFF7FF97FC46F86BAA993CCA" box="[950,1021,2013,2039]" pageId="1" pageNumber="582">MPUJ</collectionCode>
_
<specimenCode id="72096270FFF7FF97FBFCF86BADC83CCB" box="[1036,1196,2012,2039]" collectionCode="MPUJ" pageId="1" pageNumber="582">ENT 0061910</specimenCode>
[f#1]); one male and female same locality data,
<quantity id="E55767EEFFF4FF94FE37FF20AB6D3B8C" box="[455,521,150,177]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="583" unit="m" value="250.0">
<elevation id="A9822D38FFF4FF94FE37FF20AB6D3B8C" box="[455,521,150,177]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="583" unit="m" value="250.0">250m</elevation>
</quantity>
,
<date id="5611ECCBFFF4FF94FDE6FF21AB193B8C" box="[534,637,151,177]" pageId="2" pageNumber="583" value="2009-10-07">
<collectingDate id="46551523FFF4FF94FDE6FF21AB193B8C" box="[534,637,151,177]" pageId="2" pageNumber="583" value="2009-10-07">7.X.2009</collectingDate>
</date>
(
<collectionCode id="44BE52CEFFF4FF94FD7DFF21ABB13B8C" box="[653,725,151,177]" pageId="2" pageNumber="583">MPUJ</collectionCode>
_
<specimenCode id="72096270FFF4FF94FD14FF21AAE13B8C" box="[740,901,151,177]" collectionCode="MPUJ" pageId="2" pageNumber="583">ENT 0061884</specimenCode>
[m#1];
<collectionCode id="44BE52CEFFF4FF94FC11FF21AD4C3B8C" box="[993,1064,151,177]" pageId="2" pageNumber="583">MPUJ</collectionCode>
_
<specimenCode id="72096270FFF4FF94FBC7FF21ADB33B8C" box="[1079,1239,151,177]" collectionCode="MPUJ" pageId="2" pageNumber="583">ENT 0061897</specimenCode>
[f#1]).
<collectorName id="8F5AAFDDFFF4FF94FADFFF21AC3A3B8C" box="[1327,1374,151,177]" pageId="2" pageNumber="583">One</collectorName>
male same locality data,
<date id="5611ECCBFFF4FF94FE9FFF0DA8803BE8" box="[367,484,187,213]" pageId="2" pageNumber="583" value="2008-11-03">
<collectingDate id="46551523FFF4FF94FE9FFF0DA8803BE8" box="[367,484,187,213]" pageId="2" pageNumber="583" value="2008-11-03">3.XI.2008</collectingDate>
</date>
,
<collectorName id="8F5AAFDDFFF4FF94FE1FFF0DABE53BE8" box="[495,641,187,213]" pageId="2" pageNumber="583">J. Aristizabal</collectorName>
leg. (
<specimenCode id="72096270FFF4FF94FD30FF0DAA383BE8" box="[704,860,186,213]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="2" pageNumber="583">IBSP 244556</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="6AB59980FFF4FF94FF37FF69AB5D3A20" pageId="2" pageNumber="583" type="etymology">
<paragraph id="2210CA0BFFF4FF94FF37FF69AB5D3A20" blockId="2.[151,1436,150,285]" pageId="2" pageNumber="583">
<emphasis id="10DB1619FFF4FF94FF37FF69A82C3BC4" bold="true" box="[199,328,223,249]" pageId="2" pageNumber="583">Etymology</emphasis>
. The specific epithet is a patronymic given in honor of the Spanish singer Alejandro Sanz, who has greatly contributed to Spanish music.
</paragraph>
</subSubSection>
<caption id="76D09A83FFF4FF94FF67F97BAD933C4A" ID-DOI="http://doi.org/10.5281/zenodo.4506500" ID-Zenodo-Dep="4506500" httpUri="https://zenodo.org/record/4506500/files/figure.png" pageId="2" pageNumber="583" startId="2.[151,168,1741,1767]" targetBox="[189,1398,306,1717]" targetPageId="2">
<paragraph id="2210CA0BFFF4FF94FF67F97BAD933C4A" blockId="2.[151,1437,1741,1911]" pageId="2" pageNumber="583">
<emphasis id="10DB1619FFF4FF94FF67F97BA82E3DDA" bold="true" box="[151,330,1741,1767]" pageId="2" pageNumber="583">FIGURE 1A−G.</emphasis>
<taxonomicName id="E5AFB188FFF4FF94FEBFF979AB623DDA" authority="Martínez &amp; Brescovit &amp; Oliveira, 2020" authorityName="Martínez &amp; Brescovit &amp; Oliveira" authorityYear="2020" box="[335,518,1743,1767]" class="Arachnida" family="Anyphaenidae" genus="Macrophyes" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="583" phylum="Arthropoda" rank="species" species="sanzi" status="sp. nov.">
<emphasis id="10DB1619FFF4FF94FEBFF979AB623DDA" box="[335,518,1743,1767]" italics="true" pageId="2" pageNumber="583">Macrophyes sanzi</emphasis>
</taxonomicName>
<emphasis id="10DB1619FFF4FF94FDFCF979AB223DDA" bold="true" box="[524,582,1743,1767]" pageId="2" pageNumber="583">
<taxonomicNameLabel id="0BE8AB62FFF4FF94FDFCF979AB223DDA" box="[524,582,1743,1767]" pageId="2" pageNumber="583" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Male (MPUJ_ENT 0061936): A habitus, dorsal view; C left palp, retroventral view; D same, retrolateral view; E same, ventral view. Female (MPUJ_ENT 0061923): B habitus, dorsal view; F epigynum, ventral view; G epigynum, dorsal view. Abbreviations: A, atrium; CD, copulatory ducts; E, embolus; FD, fertilization ducts; LB, lateral borders; MA, median apophysis; RTA, retrolateral tibial apophysis; S, spermathecae; SD, spermatic ducts; SR, seminal receptacles; T, tegulum; VTP, ventral tegular process. Scale bars: A, B, F: 2mm; C: 1mm; D−E: 0.5mm; F, G: 0.2mm.
</paragraph>
</caption>
<subSubSection id="6AB59980FFF4FF95FF37F813AA153A20" lastPageId="3" lastPageNumber="584" pageId="2" pageNumber="583" type="diagnosis">
<paragraph id="2210CA0BFFF4FF95FF37F813AA153A20" blockId="2.[151,1436,1957,2020]" lastBlockId="3.[151,1437,151,1437]" lastPageId="3" lastPageNumber="584" pageId="2" pageNumber="583">
<emphasis id="10DB1619FFF4FF94FF37F813A8263C82" bold="true" box="[199,322,1957,1983]" pageId="2" pageNumber="583">Diagnosis.</emphasis>
Males of
<taxonomicName id="E5AFB188FFF4FF94FE44F813AB1B3C82" authority="Martínez &amp; Brescovit &amp; Oliveira, 2020" authorityName="Martínez &amp; Brescovit &amp; Oliveira" authorityYear="2020" box="[436,639,1957,1983]" class="Arachnida" family="Anyphaenidae" genus="Macrophyes" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="583" phylum="Arthropoda" rank="species" species="sanzi" status="sp. nov.">
<emphasis id="10DB1619FFF4FF94FE44F813AB1B3C82" box="[436,639,1957,1983]" italics="true" pageId="2" pageNumber="583">Macrophyes sanzi</emphasis>
</taxonomicName>
<emphasis id="10DB1619FFF4FF94FD76F813ABA23C82" bold="true" box="[646,710,1957,1983]" pageId="2" pageNumber="583">
<taxonomicNameLabel id="0BE8AB62FFF4FF94FD76F813ABA23C82" box="[646,710,1957,1983]" pageId="2" pageNumber="583" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, resemble those of
<taxonomicName id="E5AFB188FFF4FF94FC53F810AD923C82" authority="Chickering, 1937" authorityName="Chickering" authorityYear="1937" box="[931,1270,1957,1983]" class="Arachnida" family="Anyphaenidae" genus="Macrophyes" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="583" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="10DB1619FFF4FF94FC53F810AD4F3C82" box="[931,1067,1957,1983]" italics="true" pageId="2" pageNumber="583">M. elongata</emphasis>
Chickering, 1937
</taxonomicName>
by the conical ventral tegular process and a retrolateral tibial apophysis with a small posterior projection (see
<bibRefCitation id="463EB7FAFFF4FF94FB4BF87FAC0C3CDE" author="Brescovit, A. D." box="[1211,1384,1993,2019]" pageId="2" pageNumber="583" pagination="101 - 106" refId="ref5208" refString="Brescovit, A. D. (1992) Revisao das aranhas do genero Macrophyes O. Pickard-Cambridge, da regiao Neotropical (Araneae, Anyphaenidae). Revista Brasileira de Entomologia, 36, 101 - 106." type="journal article" year="1992">Brescovit 1992</bibRefCitation>
, figs 6, 7), but can be distinguished by the quadrangular, straight retrolateral tibial apophysis and wider ventral tegular process (
<figureCitation id="BA94D68EFFF5FF95FF0CFF0DA8243BE8" box="[252,320,187,213]" captionStart="FIGURE" captionStartId="2.[151,168,1741,1767]" captionTargetBox="[189,1398,306,1717]" captionTargetId="figure-76@2.[189,1398,306,1717]" captionTargetPageId="2" captionText="FIGURE 1A−G. Macrophyes sanzi n. sp. Male (MPUJ_ENT 0061936): A habitus, dorsal view; C left palp, retroventral view; D same, retrolateral view; E same, ventral view. Female (MPUJ_ENT 0061923): B habitus, dorsal view; F epigynum, ventral view; G epigynum, dorsal view.Abbreviations:A, atrium; CD, copulatory ducts; E, embolus; FD, fertilization ducts; LB, lateral borders; MA, median apophysis; RTA, retrolateral tibial apophysis; S, spermathecae; SD, spermatic ducts; SR, seminal receptacles; T, tegulum; VTP, ventral tegular process. Scale bars: A, B, F: 2mm; C: 1mm; D−E: 0.5mm; F, G: 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.4506500" httpUri="https://zenodo.org/record/4506500/files/figure.png" pageId="3" pageNumber="584">Figs 1</figureCitation>
C−E). Females are similar to those of
<taxonomicName id="E5AFB188FFF5FF95FD0DFF0DAAE23BE8" authorityName="Chickering" authorityYear="1937" box="[765,902,187,213]" class="Arachnida" family="Anyphaenidae" genus="Macrophyes" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="584" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="10DB1619FFF5FF95FD0DFF0DAAE23BE8" box="[765,902,187,213]" italics="true" pageId="3" pageNumber="584">M. elongata</emphasis>
</taxonomicName>
by having an oval atrium, differing mainly by their rounded and large spermathecae. Also, resembles to
<taxonomicName id="E5AFB188FFF5FF95FCEEFF69AD1B3BC4" authority="Brescovit et al., 2019" authorityName="Brescovit" authorityYear="2019" box="[798,1151,223,249]" class="Arachnida" family="Anyphaenidae" genus="Macrophyes" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="584" phylum="Arthropoda" rank="species" species="pacoti">
<emphasis id="10DB1619FFF5FF95FCEEFF69AAED3BC5" box="[798,905,223,248]" italics="true" pageId="3" pageNumber="584">M. pacoti</emphasis>
<bibRefCitation id="463EB7FAFFF5FF95FC60FF69AD1B3BC4" author="Brescovit, A. D. &amp; Villanueva-Bonilla, G. A. &amp; Sobczak, J. C. M. S. M. &amp; Nobrega, F. A. de &amp; Sousa &amp; Oliveira, L. F. M. &amp; Arruda, I. D. P. &amp; Sobczak, J. F." box="[912,1151,223,249]" pageId="3" pageNumber="584" pagination="294 - 300" refId="ref5336" refString="Brescovit, A. D., Villanueva-Bonilla, G. A., Sobczak, J. C. M. S. M., Nobrega, F. A. de Sousa, Oliveira, L. F. M., Arruda, I. D. P. &amp; Sobczak, J. F. (2019) Macrophyes pacoti n. sp. (Araneae: Anyphaenidae) from Brazilian Atlantic Forest, with notes on an araneopathogenic fungus. Zootaxa, 4629 (2), 294 - 300. https: // doi. org / 10.11646 / zootaxa. 4629.2.12" type="journal article" year="2019">
Brescovit
<emphasis id="10DB1619FFF5FF95FBF3FF69AD733BC5" box="[1027,1047,223,248]" italics="true" pageId="3" pageNumber="584">et</emphasis>
al., 2019
</bibRefCitation>
</taxonomicName>
by the large and rounded spermathecae, separating by the shape of the atrium (
<figureCitation id="BA94D68EFFF5FF95FD1AFEB5AA003A20" box="[746,868,259,285]" captionStart="FIGURE" captionStartId="2.[151,168,1741,1767]" captionTargetBox="[189,1398,306,1717]" captionTargetId="figure-76@2.[189,1398,306,1717]" captionTargetPageId="2" captionText="FIGURE 1A−G. Macrophyes sanzi n. sp. Male (MPUJ_ENT 0061936): A habitus, dorsal view; C left palp, retroventral view; D same, retrolateral view; E same, ventral view. Female (MPUJ_ENT 0061923): B habitus, dorsal view; F epigynum, ventral view; G epigynum, dorsal view.Abbreviations:A, atrium; CD, copulatory ducts; E, embolus; FD, fertilization ducts; LB, lateral borders; MA, median apophysis; RTA, retrolateral tibial apophysis; S, spermathecae; SD, spermatic ducts; SR, seminal receptacles; T, tegulum; VTP, ventral tegular process. Scale bars: A, B, F: 2mm; C: 1mm; D−E: 0.5mm; F, G: 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.4506500" httpUri="https://zenodo.org/record/4506500/files/figure.png" pageId="3" pageNumber="584">Figs 1F, G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="6AB59980FFF5FF95FF37FE90ABFA3E44" pageId="3" pageNumber="584" type="description">
<paragraph id="2210CA0BFFF5FF95FF37FE90AD113828" blockId="3.[151,1437,151,1437]" pageId="3" pageNumber="584">
<emphasis id="10DB1619FFF5FF95FF37FE90A83D3A7D" bold="true" box="[199,345,294,320]" pageId="3" pageNumber="584">Description.</emphasis>
<emphasis id="10DB1619FFF5FF95FEAFFE91A8C43A7C" box="[351,416,295,321]" italics="true" pageId="3" pageNumber="584">Male.</emphasis>
(
<typeStatus id="FD1474A9FFF5FF95FE5EFE91AB7F3A7C" box="[430,539,295,321]" pageId="3" pageNumber="584" type="holotype">Holotype</typeStatus>
, MPUJ_ENT 0061936). Carapace yellowish dark in the cephalic region. Chelicerae brown, labium and endites brown. Sternum yellowish-brown. Legs yellow.Abdomen dorsally pale yellow, with two yellow lateral stripes; ventrally pale yellow. Spinnerets pale yellow (
<figureCitation id="BA94D68EFFF5FF95FC4BFED9AD773AB4" box="[955,1043,367,393]" captionStart="FIGURE" captionStartId="2.[151,168,1741,1767]" captionTargetBox="[189,1398,306,1717]" captionTargetId="figure-76@2.[189,1398,306,1717]" captionTargetPageId="2" captionText="FIGURE 1A−G. Macrophyes sanzi n. sp. Male (MPUJ_ENT 0061936): A habitus, dorsal view; C left palp, retroventral view; D same, retrolateral view; E same, ventral view. Female (MPUJ_ENT 0061923): B habitus, dorsal view; F epigynum, ventral view; G epigynum, dorsal view.Abbreviations:A, atrium; CD, copulatory ducts; E, embolus; FD, fertilization ducts; LB, lateral borders; MA, median apophysis; RTA, retrolateral tibial apophysis; S, spermathecae; SD, spermatic ducts; SR, seminal receptacles; T, tegulum; VTP, ventral tegular process. Scale bars: A, B, F: 2mm; C: 1mm; D−E: 0.5mm; F, G: 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.4506500" httpUri="https://zenodo.org/record/4506500/files/figure.png" pageId="3" pageNumber="584">Fig. 1A</figureCitation>
). Total length 7.12, carapace length 2.49, width 1.99, high 0.68. Clypeus height 0.15. Eye diameters and interdistances: AME 0.10, ALE 0.18, PME 0.2, PLE 0.21; AME–AME 0.23, AME–ALE 0.35, PME–PME 0.48, PME–PLE 0.38, ALE–PLE 0.43. Chelicerae 1.72 long, distal large retromarginal teeth, six promarginal teeth, six retromarginal denticle Leg I—femur 6.13/ patella 0.86/ tibia 7.08/ metatarsus 6.82/ tarsus 1.87/ total 22.76; II—5.77/ 0.81/ 6.68/ 6.59/ 1.85/ 21.7; III—3.3/ 0.81/ 2.63/ 3.54/ 1.17/ 11.45; IV—4.42/ 0.73/ 4.77/ 6.31/ 1.55/ 17.78. Leg spination: III—tibia v2-0-2-0, p0-1-0-1, r1-0-0-1, metatarsus p1-1-0-0, r=I; IV—tibia v2-0-2-2-0, p0-1-0-1-0, r1-0-0-0-1, metatarsus v0-1-0-0-1, p1-0-1-1-0, r1-0- 1-0-0. Abdomen, twice the length of carapace, length 4.61, epigastric furrow 0.68 from tracheal spiracle, spiracle 2.93 from base of spinnerets. Palp: retrolateral tibial apophysis small and apically straight, with a small projection on posterior side; cymbium shorter than tibia; subtegulum longer than wide; tegulum longer than wide, with ventral tegular projection wide, laminar; median apophysis long, curved, apically inserted on the tegulum; embolus short, apically filiform, with narrow base and proximally inserted on the tegulum (
<figureCitation id="BA94D68EFFF5FF95FC1CFD4DAD4A3828" box="[1004,1070,763,789]" captionStart="FIGURE" captionStartId="2.[151,168,1741,1767]" captionTargetBox="[189,1398,306,1717]" captionTargetId="figure-76@2.[189,1398,306,1717]" captionTargetPageId="2" captionText="FIGURE 1A−G. Macrophyes sanzi n. sp. Male (MPUJ_ENT 0061936): A habitus, dorsal view; C left palp, retroventral view; D same, retrolateral view; E same, ventral view. Female (MPUJ_ENT 0061923): B habitus, dorsal view; F epigynum, ventral view; G epigynum, dorsal view.Abbreviations:A, atrium; CD, copulatory ducts; E, embolus; FD, fertilization ducts; LB, lateral borders; MA, median apophysis; RTA, retrolateral tibial apophysis; S, spermathecae; SD, spermatic ducts; SR, seminal receptacles; T, tegulum; VTP, ventral tegular process. Scale bars: A, B, F: 2mm; C: 1mm; D−E: 0.5mm; F, G: 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.4506500" httpUri="https://zenodo.org/record/4506500/files/figure.png" pageId="3" pageNumber="584">Figs 1</figureCitation>
C–E).
</paragraph>
<paragraph id="2210CA0BFFF5FF95FF37FCA9A8423FD4" blockId="3.[151,1437,151,1437]" pageId="3" pageNumber="584">
<emphasis id="10DB1619FFF5FF95FF37FCA9A87F3804" box="[199,283,799,825]" italics="true" pageId="3" pageNumber="584">Female</emphasis>
(
<typeStatus id="FD1474A9FFF5FF95FED9FCA9A8F43804" box="[297,400,799,825]" pageId="3" pageNumber="584" type="paratype">Paratype</typeStatus>
, MPUJ_ENT 0061923). Carapace yellowish. Chelicerae, labium and endites yellow. Sternum yellowish-brown. Legs pale yellow. Abdomen and spinnerets pale yellow (
<figureCitation id="BA94D68EFFF5FF95FC04FCF5AD2A3860" box="[1012,1102,835,861]" captionStart="FIGURE" captionStartId="2.[151,168,1741,1767]" captionTargetBox="[189,1398,306,1717]" captionTargetId="figure-76@2.[189,1398,306,1717]" captionTargetPageId="2" captionText="FIGURE 1A−G. Macrophyes sanzi n. sp. Male (MPUJ_ENT 0061936): A habitus, dorsal view; C left palp, retroventral view; D same, retrolateral view; E same, ventral view. Female (MPUJ_ENT 0061923): B habitus, dorsal view; F epigynum, ventral view; G epigynum, dorsal view.Abbreviations:A, atrium; CD, copulatory ducts; E, embolus; FD, fertilization ducts; LB, lateral borders; MA, median apophysis; RTA, retrolateral tibial apophysis; S, spermathecae; SD, spermatic ducts; SR, seminal receptacles; T, tegulum; VTP, ventral tegular process. Scale bars: A, B, F: 2mm; C: 1mm; D−E: 0.5mm; F, G: 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.4506500" httpUri="https://zenodo.org/record/4506500/files/figure.png" pageId="3" pageNumber="584">Fig. 1B</figureCitation>
). Total length 7.02, carapace length 2.06, width 1.90, high 0.77. Clypeus height 0.09. Eye diameters and interdistances: AME 0.07, ALE 0.18, PME 0.15, PLE 0.17; AME–AME 0.22, AME–ALE 0.30, PME–PME 0.44, PME–PLE 0.42, ALE–PLE 0.37. Chelicerae 1.41 long, six promarginal teeth, six retromarginal teeth, sixth largest. Leg measurements: leg I—femur 6.84/ patella 1.12/ tibia 7.86/ metatarsus 6.98/ tarsus 2.01/ total 24.81; II—5.88/ 1.02 / 5.56/ 5.94/ 1.81/ 20.21; III—3.99/ 0.91/ 3.49/ 4.38/ 1.22/ 13.99; IV—5.86/ 1.06/ 5.8/ 7.07/ 1.81/ 21.6. Leg spination: III—tibia v2-2-2-2, p1-1-0-0-1, r1-1-0-0-1, metatarsus v2-1-1-2; IV—tibia v1-2-1-1-1, p0-1-1-0-0, r1-1-0-0-1, metatarsus v2-0-1-1-0, p1-0-1-0-1, r1-0-1-0-0-1-1. Abdomen: length 4.86, epigastric furrow 0.83 from tracheal spiracle, spiracle 3.07 from base of spinnerets. Epigynum: atrium longer than wide, oval-shaped lateral borders wide, sclerotized and laminar; internally with long, thin, curved copulatory ducts; seminal receptacles small, rounded, medially situated on the copulatory ducts; spermathecae large, rounded and posteriorly positioned; fertilization ducts as long as the spermathecae width (
<figureCitation id="BA94D68EFFF5FF95FF6FFB79A87D3FD4" box="[159,281,1231,1257]" captionStart="FIGURE" captionStartId="2.[151,168,1741,1767]" captionTargetBox="[189,1398,306,1717]" captionTargetId="figure-76@2.[189,1398,306,1717]" captionTargetPageId="2" captionText="FIGURE 1A−G. Macrophyes sanzi n. sp. Male (MPUJ_ENT 0061936): A habitus, dorsal view; C left palp, retroventral view; D same, retrolateral view; E same, ventral view. Female (MPUJ_ENT 0061923): B habitus, dorsal view; F epigynum, ventral view; G epigynum, dorsal view.Abbreviations:A, atrium; CD, copulatory ducts; E, embolus; FD, fertilization ducts; LB, lateral borders; MA, median apophysis; RTA, retrolateral tibial apophysis; S, spermathecae; SD, spermatic ducts; SR, seminal receptacles; T, tegulum; VTP, ventral tegular process. Scale bars: A, B, F: 2mm; C: 1mm; D−E: 0.5mm; F, G: 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.4506500" httpUri="https://zenodo.org/record/4506500/files/figure.png" pageId="3" pageNumber="584">Figs 1F, G</figureCitation>
).
</paragraph>
<paragraph id="2210CA0BFFF5FF95FF37FB44AA0A3E0C" blockId="3.[151,1437,151,1437]" pageId="3" pageNumber="584">
<emphasis id="10DB1619FFF5FF95FF37FB44A85B3E31" bold="true" box="[199,319,1266,1292]" pageId="3" pageNumber="584">Variation.</emphasis>
Males (n=3): total length: 7.03–8.20; carapace length: 2.05–2.71; femur I length: 6.13–6.66. Females (n=6): total length: 1.02–1.17; carapace length: 0.41–0.51.
</paragraph>
<paragraph id="2210CA0BFFF5FF95FF37FA8CABFA3E44" blockId="3.[151,1437,151,1437]" pageId="3" pageNumber="584">
<emphasis id="10DB1619FFF5FF95FF37FA8CA84E3E69" bold="true" box="[199,298,1338,1364]" pageId="3" pageNumber="584">Habitat.</emphasis>
The specimens were caught manually and beating foliage of shrubs and low vegetation in a fragment of rain forest in
<collectingRegion id="E06B04E9FFF5FF95FEBCFAE9AB793E44" box="[332,541,1375,1401]" country="Colombia" name="Choco" pageId="3" pageNumber="584">Chocó department</collectingRegion>
,
<collectingCountry id="5AB88A9BFFF5FF95FDD8FAE9ABFE3E44" box="[552,666,1375,1401]" name="Colombia" pageId="3" pageNumber="584">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="6AB59980FFF5FF95FF37FA34AA293EA0" box="[199,845,1410,1437]" pageId="3" pageNumber="584" type="distribution">
<paragraph id="2210CA0BFFF5FF95FF37FA34AA293EA0" blockId="3.[151,1437,151,1437]" box="[199,845,1410,1437]" pageId="3" pageNumber="584">
<emphasis id="10DB1619FFF5FF95FF37FA34A83D3EA1" bold="true" box="[199,345,1410,1436]" pageId="3" pageNumber="584">Distribution</emphasis>
. Only known from the
<typeStatus id="FD1474A9FFF5FF95FD90FA35ABF43EA0" box="[608,656,1411,1437]" pageId="3" pageNumber="584">type</typeStatus>
locality (
<figureCitation id="BA94D68EFFF5FF95FD0BFA35AA243EA0" box="[763,832,1411,1437]" captionStart="FIGURE 4" captionStartId="7.[151,250,1529,1553]" captionTargetBox="[156,1431,186,1501]" captionTargetId="figure-18@7.[151,1436,181,1505]" captionTargetPageId="7" captionText="FIGURE 4. Distribution maps of Macrophyes and Mesilla species from Colombia, including the records of Mesilla vittiventris Simon, 1903 in Brescovit (1997)." figureDoi="http://doi.org/10.5281/zenodo.4506508" httpUri="https://zenodo.org/record/4506508/files/figure.png" pageId="3" pageNumber="584">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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