288 lines
24 KiB
XML
288 lines
24 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/asp.80.e73829" ID-Pensoft-Pub="1864-8312-80-309" ID-Pensoft-UUID="11907D1B6F665D96B5FD2F9751431366" ID-ZooBank="D5C0468B99A14EF39237D9BC51A8BDA3" ModsDocID="1864-8312-80-309" checkinTime="1660711556436" checkinUser="pensoft" docAuthor="Benedetti, Alipio Rezende & Pinto-da-Rocha, Ricardo" docDate="2022" docId="A257B9E5ECE8555CBEA039657D25FB10" docLanguage="en" docName="ArthropodSystPhyl 80: 309-388" docOrigin="Arthropod Systematics & amp; Phylogeny 80" docPubDate="2022-08-16" docSource="http://dx.doi.org/10.3897/asp.80.e73829" docTitle="Ayacucho insignitus Benedetti & Pinto-da-Rocha 2022, comb. nov." docType="treatment" docVersion="1" id="11907D1B6F665D96B5FD2F9751431366" lastPageNumber="309" masterDocId="11907D1B6F665D96B5FD2F9751431366" masterDocTitle="Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species" masterLastPageNumber="388" masterPageNumber="309" pageNumber="309" updateTime="1660711556436" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Benedetti, Alipio Rezende</mods:namePart>
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<mods:affiliation>Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil & Instituto Federal de Minas Gerais, campus Sao Joao Evangelista, Av. Primeiro de Junho, 1043, Sao Joao Evangelista, MG, Brazil</mods:affiliation>
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<mods:nameIdentifier type="email">alipiobenedetti@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pinto-da-Rocha, Ricardo</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3959-2205</mods:nameIdentifier>
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<mods:affiliation>Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Arthropod Systematics & amp; Phylogeny</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-08-16</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>80</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>309</mods:start>
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<mods:end>388</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/asp.80.e73829</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.80.e73829</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1864-8312-80-309</mods:identifier>
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<mods:identifier type="ZooBank">D5C0468B99A14EF39237D9BC51A8BDA3</mods:identifier>
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<mods:identifier type="Pensoft-UUID">11907D1B6F665D96B5FD2F9751431366</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:A257B9E5ECE8555CBEA039657D25FB10" httpUri="http://treatment.plazi.org/id/A257B9E5ECE8555CBEA039657D25FB10" lastPageNumber="309" pageId="0" pageNumber="309">
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<subSubSection pageId="0" pageNumber="309" type="nomenclature">
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<paragraph pageId="0" pageNumber="309">
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3.14.
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<taxonomicName LSID="A257B9E5-ECE8-555C-BEA0-39657D25FB10" authority="(Roewer, 1956)" authorityName="Benedetti & Pinto-da-Rocha" authorityYear="2022" baseAuthorityName="Roewer" baseAuthorityYear="1956" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho insignitus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="insignitus" status="comb. nov.">Ayacucho insignitus (Roewer, 1956)</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="309">comb. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="309" type="description">
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<paragraph pageId="0" pageNumber="309">
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus, dorsal of Ayacucho. A A. bambamarca (Roewer, 1957) comb. nov., male; B A. glauberrochai sp. nov., male; C A. inermis (Roewer, 1957) comb. nov., male; D A. insignitus (Roewer, 1956) comb. nov., female; E A. pasolinii sp. nov., male; F A. pomacocha sp. nov., male; G A. querococha sp. nov., male; H A. roeweri nom. nov., female; I A. silvae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure2" httpUri="https://binary.pensoft.net/fig/729876" pageId="0" pageNumber="309">Figs 2D</figureCitation>
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<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Distribution of Ayacucho spp. in Peru" figureDoi="10.3897/asp.80.e73829.figure28" httpUri="https://binary.pensoft.net/fig/729902" pageId="0" pageNumber="309">, 28</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="309" type="reference_group">
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<paragraph pageId="0" pageNumber="309">
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<taxonomicName authorityName="Benedetti & Pinto-da-Rocha" authorityYear="2022" baseAuthorityName="Benedetti & Pinto-da-Rocha" baseAuthorityYear="2022" class="Arachnida" family="Metasarcidae" genus="Cargaruaya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cargaruaya insignita" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="insignita" status="comb. nov.">Cargaruaya insignita</taxonomicName>
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Roewer, 1956: 439 (desc.), figs. 11-12 (ocularium), 13 (female pedipalp).
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<bibRefCitation author="Kury, AB" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="309" refId="B51" refString="Kury, AB, 2003. Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol.7: 1-337." title="Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol. 7: 1 - 337." year="2003">Kury 2003</bibRefCitation>
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: 144 (cat.).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="309" type="redescription">
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<paragraph pageId="0" pageNumber="309">Redescription.</paragraph>
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<paragraph pageId="0" pageNumber="309">
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<emphasis bold="true" pageId="0" pageNumber="309">
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FEMALE:
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<emphasis italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
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</emphasis>
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(
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<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
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=1) DSW: 4.0; DWL: 4.5; CL: 1.5. FIVL: 4.2. ChL: 1.5.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Coloration</emphasis>
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(in ethanol): Predominantly yellowish with dark spots at carapace, areas, lateral and posterior margin of the dorsal scutum and free tergites.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Dorsum</emphasis>
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: (Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male trochanter-tibia IV of Ayacucho, dorsal and ventral view, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D A. glauberrochai sp. nov.; E - F A. inermis (Roewer, 1957) comb. nov.; G - H A. pasolinii sp. nov.; I - J A. pomacocha sp. nov.; K - L A. querococha sp. nov.; M - N A. silvae sp. nov.; O - P A. spielbergi sp. nov.; Q - R A. spiniger (Roewer, 1957) comb. nov.; S - T: A. tapacocha nom. nov.; U - V A. titschacki Roewer, 1949; W - X: A. triarmatus nom. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure7" httpUri="https://binary.pensoft.net/fig/729881" pageId="0" pageNumber="309">7D</figureCitation>
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) Alpha-type DSS, with long and wide coda. Anterior margin with median elevation with granules sparsely distributed. Ocularium with a pair of large spines, with few scattered granules. Carapace with granules distributed on lateral and posterior regions. Areas I-IV with granules sparsely distributed, with a pair of small tubercles. Lateral margins of dorsal scutum with granules irregularly distributed throughout its length. Posterior margin of dorsal scutum and free tergite I with a row of tiny tubercles with base wider than its height, interspersed with small granules. Free tergites II-III with a row of acuminate tubercles, larger than those of tergite I and with smaller granules interspersed. DSS: alpha, with coda larger than mid-bulge.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Chelicerae</emphasis>
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: (Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male trochanter-tibia IV of Ayacucho, dorsal and ventral view, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D A. glauberrochai sp. nov.; E - F A. inermis (Roewer, 1957) comb. nov.; G - H A. pasolinii sp. nov.; I - J A. pomacocha sp. nov.; K - L A. querococha sp. nov.; M - N A. silvae sp. nov.; O - P A. spielbergi sp. nov.; Q - R A. spiniger (Roewer, 1957) comb. nov.; S - T: A. tapacocha nom. nov.; U - V A. titschacki Roewer, 1949; W - X: A. triarmatus nom. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure7" httpUri="https://binary.pensoft.net/fig/729881" pageId="0" pageNumber="309">7D</figureCitation>
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) Not swollen. Segment I densely granulate; segment II smooth; four teeth on finger. Segment III with four teeth.
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<taxonomicName genus="Pedipalpus" lsidName="Pedipalpus" pageId="0" pageNumber="309" rank="genus">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Pedipalpus</emphasis>
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</taxonomicName>
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: Small granules distributed on dorsal surface of femur. Trochanter with a ventroapical setiferous tubercle. Femur with a ventrobasal setiferous tubercle; a ventral row of six small setiferous tubercles, except at apex and base, smaller than the ventrobasal tubercle; one tall proapical tubercle. Tibia: prolateral IIi, retrolateral iIIii. Tarsus: prolateral II, retrolateral iIii.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Venter</emphasis>
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: Coxa I with four tubercles. Coxae II-IV with granules sparsely distributed; with two small tubercles at the apical region between the coxae III-IV. Genital area smooth. Free sternites with a row of small granules. Anal operculum with granules sparsely distributed throughout its surface.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Legs</emphasis>
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: (Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male trochanter-tibia IV of Ayacucho, dorsal and ventral view, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D A. glauberrochai sp. nov.; E - F A. inermis (Roewer, 1957) comb. nov.; G - H A. pasolinii sp. nov.; I - J A. pomacocha sp. nov.; K - L A. querococha sp. nov.; M - N A. silvae sp. nov.; O - P A. spielbergi sp. nov.; Q - R A. spiniger (Roewer, 1957) comb. nov.; S - T: A. tapacocha nom. nov.; U - V A. titschacki Roewer, 1949; W - X: A. triarmatus nom. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure7" httpUri="https://binary.pensoft.net/fig/729881" pageId="0" pageNumber="309">7D</figureCitation>
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) Coxae I-III with a prolateral and one retrolateral apophyses; IV with few granules distributed throughout its surface; an apical spiniform apophysis. Trochanters I-IV with few granules; IV with a small blunt retroapical tubercle. Femora I-IV with granules throughout their length, without prominent tubercles. Patellae I-IV with sparse granulation. Tarsal segmentation: (
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<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
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=1): 7, 12, 8, 8. -
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<emphasis bold="true" pageId="0" pageNumber="309">MALE</emphasis>
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: unknown.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="309" type="diagnosis">
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<paragraph pageId="0" pageNumber="309">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="309">
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It differs from females of
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<taxonomicName authorityName="Benedetti & Pinto-da-Rocha" authorityYear="2022" baseAuthorityName="Roewer" baseAuthorityYear="1957" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho uniseriatus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="uniseriatus">
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<emphasis italics="true" pageId="0" pageNumber="309">Ayacucho uniseriatus</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="309">comb. nov.</emphasis>
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and
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<taxonomicName genus="A." lsidName="A. weyrauchi" pageId="0" pageNumber="309" rank="species" species="weyrauchi">
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<emphasis italics="true" pageId="0" pageNumber="309">A. weyrauchi</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="309">comb. nov.</emphasis>
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because it has one pair of median tubercles in areas I-IV; from
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<taxonomicName genus="A." lsidName="A. inermis" pageId="0" pageNumber="309" rank="species" species="inermis">
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<emphasis italics="true" pageId="0" pageNumber="309">A. inermis</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="309">comb. nov.</emphasis>
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,
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<taxonomicName genus="A." lsidName="A. spiniger" pageId="0" pageNumber="309" rank="species" species="spiniger">
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<emphasis italics="true" pageId="0" pageNumber="309">A. spiniger</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="309">comb. nov.</emphasis>
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and
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<taxonomicName genus="A." lsidName="A. querococha" pageId="0" pageNumber="309" rank="species" species="querococha">
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<emphasis italics="true" pageId="0" pageNumber="309">A. querococha</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
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because it has sparser granulation on carapace, and it is larger than
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<taxonomicName genus="A." lsidName="A. inermis" pageId="0" pageNumber="309" rank="species" species="inermis">
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<emphasis italics="true" pageId="0" pageNumber="309">A. inermis</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="309">comb. nov.</emphasis>
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;
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<taxonomicName genus="A." lsidName="A. bambamarca" pageId="0" pageNumber="309" rank="species" species="bambamarca">
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<emphasis italics="true" pageId="0" pageNumber="309">A. bambamarca</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="309">comb. nov.</emphasis>
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and
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<taxonomicName genus="A." lsidName="A. tapacocha" pageId="0" pageNumber="309" rank="species" species="tapacocha">
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<emphasis italics="true" pageId="0" pageNumber="309">A. tapacocha</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="309">nom. nov.</emphasis>
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because it has smaller tubercles present in the posterior margin of dorsal scutum and free tergite I-II; and
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<taxonomicName genus="A." lsidName="A. roeweri" pageId="0" pageNumber="309" rank="species" species="roeweri">
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<emphasis italics="true" pageId="0" pageNumber="309">A. roeweri</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="309">nom. nov.</emphasis>
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because it does not present a huge median spine in free tergites I-II. (Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male trochanter-tibia IV of Ayacucho, dorsal and ventral view, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D A. glauberrochai sp. nov.; E - F A. inermis (Roewer, 1957) comb. nov.; G - H A. pasolinii sp. nov.; I - J A. pomacocha sp. nov.; K - L A. querococha sp. nov.; M - N A. silvae sp. nov.; O - P A. spielbergi sp. nov.; Q - R A. spiniger (Roewer, 1957) comb. nov.; S - T: A. tapacocha nom. nov.; U - V A. titschacki Roewer, 1949; W - X: A. triarmatus nom. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure7" httpUri="https://binary.pensoft.net/fig/729881" pageId="0" pageNumber="309">7D</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="309" type="remarks">
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<paragraph pageId="0" pageNumber="309">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="309">
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This species is known only by the female holotype, which ends up being a taxonomic problem, since it is not uncommon for females of
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<taxonomicName authorityName="Roewer" authorityYear="1949" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="309">Ayacucho</emphasis>
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</taxonomicName>
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, whose males are clearly morphologically distinct, to be very similar to each other. This similarity between the females justifies the not so much informative diagnosis (see above). Geographically,
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<taxonomicName genus="A." lsidName="A. insignitus" pageId="0" pageNumber="309" rank="species" species="insignitus">
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<emphasis italics="true" pageId="0" pageNumber="309">A. insignitus</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="309">comb. nov.</emphasis>
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occurs near to three other species of the genus:
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<taxonomicName genus="A." lsidName="A. inermis" pageId="0" pageNumber="309" rank="species" species="inermis">
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<emphasis italics="true" pageId="0" pageNumber="309">A. inermis</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="309">comb. nov.</emphasis>
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,
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<taxonomicName genus="A." lsidName="A. triarmatus" pageId="0" pageNumber="309" rank="species" species="triarmatus">
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<emphasis italics="true" pageId="0" pageNumber="309">A. triarmatus</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="309">nom. nov.</emphasis>
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and
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<taxonomicName genus="A." lsidName="A. weyrauchi" pageId="0" pageNumber="309" rank="species" species="weyrauchi">
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<emphasis italics="true" pageId="0" pageNumber="309">A. weyrauchi</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="309">comb. nov.</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Distribution of Ayacucho spp. in Peru" figureDoi="10.3897/asp.80.e73829.figure28" httpUri="https://binary.pensoft.net/fig/729902" pageId="0" pageNumber="309">28</figureCitation>
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). The female of
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<taxonomicName genus="A." lsidName="A. triarmatus" pageId="0" pageNumber="309" rank="species" species="triarmatus">
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<emphasis italics="true" pageId="0" pageNumber="309">A. triarmatus</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="309">nom. nov.</emphasis>
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is unknown and the type locality of this species is closest to the type locality of
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<taxonomicName genus="A." lsidName="A. insignitus" pageId="0" pageNumber="309" rank="species" species="insignitus">
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<emphasis italics="true" pageId="0" pageNumber="309">A. insignitus</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="309">comb. nov.</emphasis>
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It would be possible that
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<taxonomicName genus="A." lsidName="A. insignitus" pageId="0" pageNumber="309" rank="species" species="insignitus">
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<emphasis italics="true" pageId="0" pageNumber="309">A. insignitus</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="309">comb. nov.</emphasis>
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was tentatively pointed as the female of
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<taxonomicName genus="A." lsidName="A. triarmatus" pageId="0" pageNumber="309" rank="species" species="triarmatus">
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<emphasis italics="true" pageId="0" pageNumber="309">A. triarmatus</emphasis>
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</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="309">nom. nov.</emphasis>
|
|
, although the male of
|
|
<taxonomicName genus="A." lsidName="A. triarmatus" pageId="0" pageNumber="309" rank="species" species="triarmatus">
|
|
<emphasis italics="true" pageId="0" pageNumber="309">A. triarmatus</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="309">nom. nov.</emphasis>
|
|
has very minute armature in areas I-IV of the DS (compared to
|
|
<taxonomicName genus="A." lsidName="A. insignitus" pageId="0" pageNumber="309" rank="species" species="insignitus">
|
|
<emphasis italics="true" pageId="0" pageNumber="309">A. insignitus</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="309">comb. nov.</emphasis>
|
|
). Furthermore,
|
|
<taxonomicName genus="A." lsidName="A. insignitus" pageId="0" pageNumber="309" rank="species" species="insignitus">
|
|
<emphasis italics="true" pageId="0" pageNumber="309">A. insignitus</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="309">comb. nov.</emphasis>
|
|
has subtle differences in relation to females of
|
|
<taxonomicName genus="A." lsidName="A. inermis" pageId="0" pageNumber="309" rank="species" species="inermis">
|
|
<emphasis italics="true" pageId="0" pageNumber="309">A. inermis</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="309">comb. nov.</emphasis>
|
|
and
|
|
<taxonomicName genus="A." lsidName="A. weyrauchi" pageId="0" pageNumber="309" rank="species" species="weyrauchi">
|
|
<emphasis italics="true" pageId="0" pageNumber="309">A. weyrauchi</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="309">comb. nov.</emphasis>
|
|
Therefore, it is not possible to point out with sufficient certainty that
|
|
<taxonomicName genus="A." lsidName="A. insignitus" pageId="0" pageNumber="309" rank="species" species="insignitus">
|
|
<emphasis italics="true" pageId="0" pageNumber="309">A. insignitus</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="309">comb. nov.</emphasis>
|
|
is synonymous with one of these three species and, therefore, we prefer the more conservative option of keeping the species valid, until further evidence (e.g. collection of males from the type locality) can clarify this issue.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="309" type="distribution">
|
|
<paragraph pageId="0" pageNumber="309">Distribution.</paragraph>
|
|
<paragraph pageId="0" pageNumber="309">
|
|
(Fig.
|
|
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Distribution of Ayacucho spp. in Peru" figureDoi="10.3897/asp.80.e73829.figure28" httpUri="https://binary.pensoft.net/fig/729902" pageId="0" pageNumber="309">28</figureCitation>
|
|
) PERU. La Libertad. Hacienda
|
|
<normalizedToken originalValue="Llaguén">Llaguen</normalizedToken>
|
|
. Rejo Cargaruay forest.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="309" type="materials_examined">
|
|
<paragraph pageId="0" pageNumber="309">Material examined.</paragraph>
|
|
<paragraph pageId="0" pageNumber="309">
|
|
<emphasis bold="true" pageId="0" pageNumber="309">
|
|
<emphasis italics="true" pageId="0" pageNumber="309">Type material</emphasis>
|
|
:
|
|
<materialsCitation collectingDate="1952-12-14" collectorName="Koepcke" country="PERU" elevation="2650" location="Rejo Cargaruay forest" municipality="Hacienda Llaguen" specimenCount="♀" stateProvince="La Libertad" typeStatus="Holotype">
|
|
<typeStatus>Holotype</typeStatus>
|
|
<specimenCount>♀</specimenCount>
|
|
, '
|
|
<collectingCountry name="Peru">PERU</collectingCountry>
|
|
,
|
|
<collectingRegion country="Peru" name="La Libertad">La Libertad</collectingRegion>
|
|
.
|
|
<collectingMunicipality>
|
|
Hacienda
|
|
<normalizedToken originalValue="Llaguén">Llaguen</normalizedToken>
|
|
</collectingMunicipality>
|
|
,
|
|
<location LSID="urn:lsid:plazi:treatment:A257B9E5ECE8555CBEA039657D25FB10:FAE76BD5BA15F6C07825AB60824F4CBC" country="PERU" municipality="Hacienda Llaguen" name="Rejo Cargaruay forest" stateProvince="La Libertad">Rejo Cargaruay forest</location>
|
|
,
|
|
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.65" unit="m" value="2650.0">
|
|
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.65" unit="m" value="2650.0">2,650m</quantity>
|
|
a.s.l.
|
|
</elevation>
|
|
,
|
|
<collectingDate value="1952-12-14">14/XII/1952</collectingDate>
|
|
,
|
|
<collectorName>Koepcke</collectorName>
|
|
leg. (SMF RII 9706)
|
|
</materialsCitation>
|
|
</emphasis>
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |