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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1030.62544" ID-GBIF-Dataset="91fec855-7fa9-4ad6-84d7-9b9d35e23e8b" ID-PMC="PMC8060247" ID-Pensoft-Pub="1313-2970-1030-1" ID-Pensoft-UUID="875199397EEE5F7898EA1DB25DA62D25" ID-PubMed="33958904" ModsDocID="1313-2970-1030-1" checkinTime="1618371561543" checkinUser="pensoft" docAuthor="Means, Jackson C., Hennen, Derek A. &amp; Marek, Paul E." docDate="2021" docId="834069AA15D95AD5BD10A6938565EB9F" docLanguage="en" docName="ZooKeys 1030: 1-180" docOrigin="ZooKeys 1030" docPubDate="2021-04-13" docSource="http://dx.doi.org/10.3897/zookeys.1030.62544" docTitle="Nannaria fritzae Means &amp; Hennen &amp; Marek 2021, sp. nov." docType="treatment" docVersion="5" id="875199397EEE5F7898EA1DB25DA62D25" lastPageNumber="1" masterDocId="875199397EEE5F7898EA1DB25DA62D25" masterDocTitle="A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae)" masterLastPageNumber="180" masterPageNumber="1" pageNumber="1" updateTime="1673961214978" updateUser="julia">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Means, Jackson C.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7377-0696</mods:nameIdentifier>
<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
<mods:nameIdentifier type="email">mjacks4@vt.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hennen, Derek A.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7005-1151</mods:nameIdentifier>
<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Marek, Paul E.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7048-2514</mods:nameIdentifier>
<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-04-13</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1030</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>180</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1030.62544</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1030.62544</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1030-1</mods:identifier>
<mods:identifier type="Pensoft-UUID">875199397EEE5F7898EA1DB25DA62D25</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="181004731" LSID="urn:lsid:zoobank.org:act:735CBEDE-9F71-4971-9601-BCE0ADF3F53B" httpUri="http://treatment.plazi.org/id/834069AA15D95AD5BD10A6938565EB9F" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName ID-CoL="8RVTP" LSID="http://zoobank.org/735CBEDE-9F71-4971-9601-BCE0ADF3F53B" authority="Means &amp; Hennen &amp; Marek, 2021" authorityName="Means &amp; Hennen &amp; Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nannaria fritzae" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="fritzae" status="sp. nov.">Nannaria fritzae</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Nannaria fritzae sp. nov. ♂ holotype (VTEC, MPE 02359) left gonopod A anterior view; red arrow indicates tooth-like medial flange B medial view; red triangle indicates small, thorn-like prefemoral spine C posterior view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure55" httpUri="https://binary.pensoft.net/fig/531158" pageId="0" pageNumber="1">Figs 55</figureCitation>
<figureCitation captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Nannaria fritzae sp. nov. ♂ holotype (VTEC, MPE 02359) coloration. Scale bar: 4.0 mm." figureDoi="10.3897/zookeys.1030.62544.figure56" httpUri="https://binary.pensoft.net/fig/531159" pageId="0" pageNumber="1">
, 56 Vernacular name: &quot;Kathlyn
<normalizedToken originalValue="Fritzs">Fritz's</normalizedToken>
Twisted-Claw Millipede&quot;
</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation ID-GBIF-Occurrence="3071355512" accessionNumber="MPE02359" collectingDate="2017-03-05" collectorName="J. Means, K. Means" country="Georgia" county="Chattooga County" elevation="385" latitude="34.5628" location="United States" longLatPrecision="6" longitude="-85.0681" municipality="Lake Marvin Rd." specimenCount="♂" typeStatus="Holotype">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">
<typeStatus>Holotype</typeStatus>
</emphasis>
:
</emphasis>
<location LSID="urn:lsid:plazi:treatment:834069AA15D95AD5BD10A6938565EB9F:9B3EC108295A31457BA5697F45B613B7" country="Georgia" county="Chattooga County" latitude="34.5628" longLatPrecision="6" longitude="-85.0681" municipality="Lake Marvin Rd." name="United States">United States</location>
-
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingCountry name="Georgia">Georgia</collectingCountry>
</emphasis>
<specimenCount></specimenCount>
<collectingCounty>Chattooga County</collectingCounty>
, hillside by
<collectingMunicipality>Lake Marvin Rd.</collectingMunicipality>
;
<geoCoordinate degrees="34.5628" direction="north" orientation="latitude" precision="5" value="34.5628">34.5628°N</geoCoordinate>
, -
<geoCoordinate degrees="85.0681" direction="west" orientation="longitude" precision="5" value="-85.0681">85.0681°W</geoCoordinate>
; elev.
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.85" unit="m" value="385.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="3.85" unit="m" value="385.0">385 m</elevation>
</quantity>
;
<collectingDate value="2017-03-05">5 Mar. 2017</collectingDate>
; hand collected;
<collectorName>J. Means</collectorName>
,
<collectorName>K. Means</collectorName>
leg.; VTEC
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE02359">MPE02359</accessionNumber>
.
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation ID-GBIF-Occurrence="3071355746" accessionNumber="MPE02377" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="J. Means &amp; K. Means" country="Georgia" county="Chattooga County" elevation="385" latitude="34.5628" location="United States" longLatPrecision="6" longitude="-85.0681" municipality="Lake Marvin Rd." specimenCount="1" specimenCount-male="1" typeStatus="Paratypes">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">
<typeStatus>Paratypes</typeStatus>
</emphasis>
:
</emphasis>
<location LSID="urn:lsid:plazi:treatment:834069AA15D95AD5BD10A6938565EB9F:80D31B204B8E440E4FA9C1C72C189C02" country="Georgia" county="Chattooga County" latitude="34.5628" longLatPrecision="6" longitude="-85.0681" name="United States">United States</location>
-
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingCountry name="Georgia">Georgia</collectingCountry>
</emphasis>
<specimenCount type="male">1 ♂</specimenCount>
; same collection data as holotype; VTEC
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE02377">MPE02377</accessionNumber>
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355381" accessionNumber="MPE02378" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="J. Means &amp; K. Means" country="Georgia" county="Chattooga County" elevation="385" latitude="34.5628" location="United States" longLatPrecision="6" longitude="-85.0681" municipality="Lake Marvin Rd." specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount type="male">1 ♂</specimenCount>
; same collection data as holotype; VMNH
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE02378">MPE02378</accessionNumber>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
Adult males of
<taxonomicName authorityName="Means &amp; Hennen &amp; Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria fritzae" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="fritzae">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria fritzae</emphasis>
</taxonomicName>
sp. nov. are distinct from other
<taxonomicName authorityName="Chamberlin" authorityYear="1918" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria</emphasis>
</taxonomicName>
and the nearby
<taxonomicName authorityName="Chamberlin" authorityYear="1918" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria</emphasis>
</taxonomicName>
sp. nov.
<normalizedToken originalValue="Amicolola">'Amicolola'</normalizedToken>
(
<taxonomicName authorityName="Hoffman" authorityYear="1949" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria wilsoni" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="wilsoni">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria wilsoni</emphasis>
</taxonomicName>
species group) based on the following combination of characters.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Gonopods</emphasis>
.
</emphasis>
Gonopodal acropodite gently curving medially throughout, not straight with abrupt medial bend at apex as in
<taxonomicName authorityName="Chamberlin" authorityYear="1918" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria</emphasis>
</taxonomicName>
sp. nov.
<normalizedToken originalValue="Amicolola.">'Amicolola.'</normalizedToken>
Distal zone short, rounded, not serpentine as in
<taxonomicName authorityName="Chamberlin" authorityYear="1918" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria</emphasis>
</taxonomicName>
sp. nov.
<normalizedToken originalValue="Amicolola.">'Amicolola.'</normalizedToken>
Acropodite with small, tooth-like medial flange near apex (Fig.
<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Nannaria fritzae sp. nov. ♂ holotype (VTEC, MPE 02359) left gonopod A anterior view; red arrow indicates tooth-like medial flange B medial view; red triangle indicates small, thorn-like prefemoral spine C posterior view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure55" httpUri="https://binary.pensoft.net/fig/531158" pageId="0" pageNumber="1">55A</figureCitation>
, red arrow). Prefemur with long acicular prefemoral process bent at base and directed medially-not curving laterally, crossing acropodite, as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. minor" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="minor">
<emphasis italics="true" pageId="0" pageNumber="1">N. minor</emphasis>
</taxonomicName>
or not large, laminate as in
<taxonomicName authorityName="Chamberlin" authorityYear="1918" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria</emphasis>
</taxonomicName>
sp. nov.
<normalizedToken originalValue="Amicolola.">'Amicolola.'</normalizedToken>
Prefemoral spine reduced to thorn-like structure, arising from prefemoral process, directed ventrally (Fig.
<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Nannaria fritzae sp. nov. ♂ holotype (VTEC, MPE 02359) left gonopod A anterior view; red arrow indicates tooth-like medial flange B medial view; red triangle indicates small, thorn-like prefemoral spine C posterior view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure55" httpUri="https://binary.pensoft.net/fig/531158" pageId="0" pageNumber="1">55B</figureCitation>
, red triangle)-not curving cephalically as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. minor" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="minor">
<emphasis italics="true" pageId="0" pageNumber="1">N. minor</emphasis>
</taxonomicName>
. Telopodite basal zone ca. 1/2 length of acropodite, not &lt;1/2 as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. minor" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="minor">
<emphasis italics="true" pageId="0" pageNumber="1">N. minor</emphasis>
</taxonomicName>
or&gt; 1/4 as in
<taxonomicName authorityName="Chamberlin" authorityYear="1918" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria</emphasis>
</taxonomicName>
sp. nov.
<normalizedToken originalValue="Amicolola.">'Amicolola.'</normalizedToken>
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Color</emphasis>
.
</emphasis>
Tergites with orange paranotal spots (Fig.
<figureCitation captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Nannaria fritzae sp. nov. ♂ holotype (VTEC, MPE 02359) coloration. Scale bar: 4.0 mm." figureDoi="10.3897/zookeys.1030.62544.figure56" httpUri="https://binary.pensoft.net/fig/531159" pageId="0" pageNumber="1">56</figureCitation>
), reduced pigmentation of paranotal spots of terminal segments. Light brown background. Dorsum of collum smooth with orange and white margin.
</paragraph>
<caption doi="10.3897/zookeys.1030.62544.figure56" httpUri="https://binary.pensoft.net/fig/531159" pageId="0" pageNumber="1" start="Figure 56" startId="F56">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 56.</emphasis>
<taxonomicName authorityName="Means &amp; Hennen &amp; Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria fritzae" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="fritzae">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria fritzae</emphasis>
</taxonomicName>
sp. nov. ♂ holotype (VTEC, MPE02359) coloration. Scale bar: 4.0 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Measurements.</paragraph>
<paragraph pageId="0" pageNumber="1">♂ holotype (VTEC, MPE02359): BL = 28.2, CW = 3.8, IW = 2.3, ISW = 0.8, B11W = 4.6, B11H = 3.0.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="variation">
<paragraph pageId="0" pageNumber="1">Variation.</paragraph>
<paragraph pageId="0" pageNumber="1">No known variation.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
Known only from the type locality (Georgia: Chattooga County; Suppl. material 7; Fig.
<figureCitation captionStart="Figure 126" captionStartId="F126" captionText="Figure 126. Distribution map of Nannaria minor species group collection localities in the central Appalachians." figureDoi="10.3897/zookeys.1030.62544.figure126" httpUri="https://binary.pensoft.net/fig/531229" pageId="0" pageNumber="1">126</figureCitation>
). Distribution area: N/A; status: MRE.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="ecology">
<paragraph pageId="0" pageNumber="1">Ecology.</paragraph>
<paragraph pageId="0" pageNumber="1">
Individuals of
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. fritzae" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="fritzae">
<emphasis italics="true" pageId="0" pageNumber="1">N. fritzae</emphasis>
</taxonomicName>
sp. nov. were collected on a hillside in a mesic hardwood forest dominated by oak and maple. Specimens were found under leaf litter in a dry creek bed near the base of an oak tree.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">This species is named for its co-collector, Kathlyn Fritz Means. The name is a genitive noun derived as a matronym.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">
United States, Georgia, Chattooga County, hillside by Lake Marvin Rd.;
<geoCoordinate degrees="34.5628" direction="north" orientation="latitude" precision="5" value="34.5628">34.5628°N</geoCoordinate>
, -
<geoCoordinate degrees="85.0681" direction="west" orientation="longitude" precision="5" value="-85.0681">85.0681°W</geoCoordinate>
.
</paragraph>
</subSubSection>
</treatment>
</document>