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<mods:title id="2CEB129F1D1EC72E7EB8BBCAD6268BFF">Composition of the genus Loricella (Mollusca: Polyplacophora: Loricidae) and the description of two new species</mods:title>
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<treatment id="82770F72FF9428792BE1FAA0746C8D34" ID-DOI="http://doi.org/10.5281/zenodo.4923612" ID-GBIF-Taxon="183227978" ID-Zenodo-Dep="4923612" LSID="urn:lsid:plazi:treatment:82770F72FF9428792BE1FAA0746C8D34" httpUri="http://treatment.plazi.org/id/82770F72FF9428792BE1FAA0746C8D34" lastPageId="2" lastPageNumber="276" pageId="1" pageNumber="276">
<subSubSection id="42C4EDEFFF94287A2BE1FAA0751E8929" box="[151,517,1374,1401]" pageId="1" pageNumber="276" type="nomenclature">
<paragraph id="0A61BE64FF94287A2BE1FAA0751E8929" blockId="1.[151,517,1374,1401]" box="[151,517,1374,1401]" pageId="1" pageNumber="276">
<heading id="51290908FF94287A2BE1FAA0751E8929" bold="true" box="[151,517,1374,1401]" fontSize="11" level="1" pageId="1" pageNumber="276" reason="1">
<emphasis id="38AA6276FF94287A2BE1FAA0751E8929" bold="true" box="[151,517,1374,1401]" pageId="1" pageNumber="276">
Genus
<taxonomicName id="CDDEC5E7FF94287A2B98FAA0751E8929" ID-CoL="7P7VF" authority="Pilsbry, 1893" authorityName="Pilsbry" authorityYear="1893" box="[238,517,1374,1401]" class="Polyplacophora" family="Loricidae" genus="Loricella" kingdom="Animalia" order="Chitonida" pageId="1" pageNumber="276" phylum="Mollusca" rank="genus">
<emphasis id="38AA6276FF94287A2B98FAA076408928" bold="true" box="[238,347,1374,1401]" italics="true" pageId="1" pageNumber="276">Loricella</emphasis>
<bibRefCitation id="6E4FC395FF94287A2A15FAA0751E8929" author="Pilsbry, H. A." box="[355,517,1374,1400]" pageId="1" pageNumber="276" pagination="129 - 350" refId="ref9572" refString="Pilsbry, H. A. (1893) Polyplacophora. In: Tryon, G. W. (Ed.), Manual of Conchology. Vol. 14. Academy of Natural Sciences, Philadelphia, Pennsylvania, pp. 129 - 350, pls. 31 - 68." type="book chapter" year="1893">Pilsbry, 1893</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="42C4EDEFFF94287A2BE1FA5973778AEC" pageId="1" pageNumber="276" type="type_taxon">
<paragraph id="0A61BE64FF94287A2BE1FA5973228990" blockId="1.[151,1437,1446,2014]" box="[151,1081,1446,1473]" pageId="1" pageNumber="276">
<emphasis id="38AA6276FF94287A2BE1FA5976298991" bold="true" box="[151,306,1446,1473]" pageId="1" pageNumber="276">
<typeStatus id="D56500C6FF94287A2BE1FA5977C98990" box="[151,210,1447,1473]" pageId="1" pageNumber="276">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="CDDEC5E7FF94287A2A4FFA5975578990" ID-CoL="3W75Z" authority="H. Adams" authorityName="H. Adams" box="[313,588,1447,1473]" class="Polyplacophora" family="Loricidae" genus="Lorica" kingdom="Animalia" order="Chitonida" pageId="1" pageNumber="276" phylum="Mollusca" rank="species" species="angasi">
<emphasis id="38AA6276FF94287A2A4FFA5976CD8991" box="[313,470,1447,1472]" italics="true" pageId="1" pageNumber="276">Lorica angasi</emphasis>
H. Adams
</taxonomicName>
in H.
<bibRefCitation id="6E4FC395FF94287A29E7FA5974888990" author="Adams, H. &amp; Angas, G. F." box="[657,915,1447,1473]" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="1" pageNumber="276" pagination="192 - 194" part="1864" refId="ref8578" refString="Adams, H. &amp; Angas, G. F. (1864) Descriptions of new genera and species of Chitonidae from the Australian Seas, in the collection of George French Angas. Proceedings of the Zoological Society of London, 1864, 192 - 194." title="Descriptions of new genera and species of Chitonidae from the Australian Seas" type="journal article" volumeTitle="the collection of George French Angas" year="1864">Adams &amp; Angas, 1864</bibRefCitation>
, by monotypy.
</paragraph>
<paragraph id="0A61BE64FF94287A2BB1FA34726789B4" blockId="1.[151,1437,1446,2014]" box="[199,1404,1482,1509]" pageId="1" pageNumber="276">
<emphasis id="38AA6276FF94287A2BB1FA34765289B5" bold="true" box="[199,329,1482,1508]" pageId="1" pageNumber="276">Synonyms.</emphasis>
<taxonomicName id="CDDEC5E7FF94287A2A26FA3575CB89B4" authority="Ashby, 1925: 198" authorityName="Ashby" authorityPageNumber="198" authorityYear="1925" box="[336,720,1482,1509]" class="Polyplacophora" family="Loricidae" genus="Pseudoloricella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chitonida" pageId="1" pageNumber="276" phylum="Mollusca" rank="genus">
<emphasis id="38AA6276FF94287A2A26FA35751A89B4" box="[336,513,1483,1509]" italics="true" pageId="1" pageNumber="276">Pseudoloricella</emphasis>
<bibRefCitation id="6E4FC395FF94287A2971FA3575CB89B4" author="Ashby, E." box="[519,720,1482,1509]" pageId="1" pageNumber="276" pagination="170 - 205" refId="ref8671" refString="Ashby, E. (1925) Monograph of Australian fossil Polyplacophora (chitons). Proceedings of the Royal Society of Victoria, 37 (2), 170 - 205." type="journal article" year="1925">Ashby, 1925: 198</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="CDDEC5E7FF94287A29AAFA35735789B4" authority="Milne, 1963: 25" authorityName="Milne" authorityPageNumber="25" authorityYear="1963" box="[732,1100,1482,1509]" class="Polyplacophora" family="Loricidae" genus="Componochiton" higherTaxonomySource="CoL" kingdom="Animalia" order="Chitonida" pageId="1" pageNumber="276" phylum="Mollusca" rank="genus">
<emphasis id="38AA6276FF94287A29AAFA35749489B4" box="[732,911,1483,1509]" italics="true" pageId="1" pageNumber="276">Componochiton</emphasis>
<bibRefCitation id="6E4FC395FF94287A28E0FA35735789B4" author="Milne, K. L." box="[918,1100,1482,1509]" pageId="1" pageNumber="276" pagination="25 - 27" refId="ref9437" refString="Milne, K. L. (1963) A new deep water chiton (Polyplacophora; Chitonidae) from eastern Australia. Journal of the Malacological Society of Australia, 7, 25 - 27, figs. 1 - 5. https: // doi. org / 10.1080 / 00852988.1963.10673796" type="journal article" year="1963">Milne, 1963: 25</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="CDDEC5E7FF94287A2F2EFA35726389B4" authority="Xu, 1990: 376" authorityName="Xu" authorityPageNumber="376" authorityYear="1990" box="[1112,1400,1483,1509]" class="Polyplacophora" family="Loricidae" genus="Sinolorica" higherTaxonomySource="CoL" kingdom="Animalia" order="Chitonida" pageId="1" pageNumber="276" phylum="Mollusca" rank="genus">
<emphasis id="38AA6276FF94287A2F2EFA3573D689B4" box="[1112,1229,1483,1509]" italics="true" pageId="1" pageNumber="276">Sinolorica</emphasis>
Xu, 1990: 376
</taxonomicName>
.
</paragraph>
<paragraph id="0A61BE64FF94287A2BB1FA1075E38A00" blockId="1.[151,1437,1446,2014]" pageId="1" pageNumber="276">
<emphasis id="38AA6276FF94287A2BB1FA1076B58A59" bold="true" box="[199,430,1518,1544]" pageId="1" pageNumber="276">Genus distribution.</emphasis>
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<collectingCounty id="E300C6E8FF94287A2AC5FA1175218A58" box="[435,570,1519,1545]" pageId="1" pageNumber="276">Sagami Bay</collectingCounty>
and
<collectingMunicipality id="EA05241EFF94287A2918FA1174228A58" box="[622,825,1519,1545]" pageId="1" pageNumber="276">Omurodashi Bank</collectingMunicipality>
(
<collectingCountry id="72C9FEF4FF94287A2830FA1174908A58" box="[838,907,1519,1545]" name="Japan" pageId="1" pageNumber="276">Japan</collectingCountry>
),
<location id="0F01E8BFFF94287A28EBFA11732A8A58" LSID="urn:lsid:plazi:treatment:82770F72FF9428792BE1FAA0746C8D34:0F01E8BFFF94287A28EBFA11732A8A58" box="[925,1073,1519,1545]" country="Japan" county="Sagami Bay" municipality="Omurodashi Bank" name="South of East" pageId="1" pageNumber="276" stateProvince="Tasmania">South of East</location>
<collectingCountry id="72C9FEF4FF94287A2F41FA1173618A58" box="[1079,1146,1519,1545]" name="China" pageId="1" pageNumber="276">China</collectingCountry>
<location id="0F01E8BFFF94287A2FF6FA1173B08A58" LSID="urn:lsid:plazi:treatment:82770F72FF9428792BE1FAA0746C8D34:0F01E8BFFF94287A2FF6FA1173B08A58" box="[1152,1195,1519,1545]" country="Japan" county="Sagami Bay" municipality="Omurodashi Bank" name="Sea" pageId="1" pageNumber="276" stateProvince="Tasmania">Sea</location>
, north of
<location id="0F01E8BFFF94287A2E64FA1172488A58" LSID="urn:lsid:plazi:treatment:82770F72FF9428792BE1FAA0746C8D34:0F01E8BFFF94287A2E64FA1172488A58" box="[1298,1363,1519,1545]" country="Japan" county="Sagami Bay" municipality="Omurodashi Bank" name="South" pageId="1" pageNumber="276" stateProvince="Tasmania">South</location>
<collectingCountry id="72C9FEF4FF94287A2E2FFA1172878A58" box="[1369,1436,1519,1545]" name="China" pageId="1" pageNumber="276">China</collectingCountry>
<location id="0F01E8BFFF94287A2BE1F9ED77D98A7C" LSID="urn:lsid:plazi:treatment:82770F72FF9428792BE1FAA0746C8D34:0F01E8BFFF94287A2BE1F9ED77D98A7C" box="[151,194,1555,1581]" country="Japan" county="Sagami Bay" municipality="Omurodashi Bank" name="Sea" pageId="1" pageNumber="276" stateProvince="Tasmania">Sea</location>
,
<collectingCountry id="72C9FEF4FF94287A2BB8F9ED76258A7C" box="[206,318,1555,1581]" name="Indonesia" pageId="1" pageNumber="276">Indonesia</collectingCountry>
,
<location id="0F01E8BFFF94287A2A3EF9ED76668A7C" LSID="urn:lsid:plazi:treatment:82770F72FF9428792BE1FAA0746C8D34:0F01E8BFFF94287A2A3EF9ED76668A7C" box="[328,381,1555,1581]" country="Japan" county="Sagami Bay" municipality="Omurodashi Bank" name="New" pageId="1" pageNumber="276" stateProvince="Tasmania">New</location>
<collectingCountry id="72C9FEF4FF94287A2AF2F9ED75BA8A7C" box="[388,673,1555,1581]" isEnumeration="true" pageId="1" pageNumber="276">Guinea, Solomon Islands</collectingCountry>
,
<collectingCountry id="72C9FEF4FF94287A29DDF9ED75EA8A7C" box="[683,753,1555,1581]" name="Tonga" pageId="1" pageNumber="276">Tonga</collectingCountry>
<location id="0F01E8BFFF94287A298EF9ED74518A7C" LSID="urn:lsid:plazi:treatment:82770F72FF9428792BE1FAA0746C8D34:0F01E8BFFF94287A298EF9ED74518A7C" box="[760,842,1555,1581]" country="Japan" county="Sagami Bay" municipality="Omurodashi Bank" name="Islands" pageId="1" pageNumber="276" stateProvince="Tasmania">Islands</location>
,
<collectingCountry id="72C9FEF4FF94287A2823F9ED731E8A7C" box="[853,1029,1555,1581]" name="New Caledonia" pageId="1" pageNumber="276">New Caledonia</collectingCountry>
,
<location id="0F01E8BFFF94287A2F66F9ED734A8A7C" LSID="urn:lsid:plazi:treatment:82770F72FF9428792BE1FAA0746C8D34:0F01E8BFFF94287A2F66F9ED734A8A7C" box="[1040,1105,1555,1581]" country="Japan" county="Sagami Bay" municipality="Omurodashi Bank" name="South" pageId="1" pageNumber="276" stateProvince="Tasmania">South</location>
and
<location id="0F01E8BFFF94287A2FFEF9ED73A38A7C" LSID="urn:lsid:plazi:treatment:82770F72FF9428792BE1FAA0746C8D34:0F01E8BFFF94287A2FFEF9ED73A38A7C" box="[1160,1208,1555,1581]" country="Japan" county="Sagami Bay" municipality="Omurodashi Bank" name="East" pageId="1" pageNumber="276" stateProvince="Tasmania">East</location>
<collectingCountry id="72C9FEF4FF94287A2FCBF9ED723C8A7C" box="[1213,1319,1555,1581]" name="Australia" pageId="1" pageNumber="276">Australia</collectingCountry>
,
<collectingRegion id="C81A7086FF94287A2E47F9ED72878A7C" box="[1329,1436,1555,1581]" country="Australia" name="Tasmania" pageId="1" pageNumber="276">Tasmania</collectingRegion>
and
<collectingCountry id="72C9FEF4FF94287A2BB1F9C976448A00" box="[199,351,1591,1617]" name="New Zealand" pageId="1" pageNumber="276">New Zealand</collectingCountry>
, intertidal–
<specimenCount id="1CD875EDFF94287A2AA8F9C975378A00" box="[478,556,1591,1617]" pageId="1" pageNumber="276" type="generic">
<quantity id="CD261381FF94287A2AA8F9C975338A00" box="[478,552,1591,1617]" metricMagnitude="2" metricUnit="m" metricValue="7.72" pageId="1" pageNumber="276" unit="m" value="772.0">
<elevation id="81F35957FF94287A2AA8F9C975338A00" box="[478,552,1591,1617]" metricMagnitude="2" metricUnit="m" metricValue="7.72" pageId="1" pageNumber="276" unit="m" value="772.0">772 m</elevation>
</quantity>
.
</specimenCount>
Miocene–Recent
</materialsCitation>
.
</paragraph>
<paragraph id="0A61BE64FF94287A2BB1F9A573778AEC" blockId="1.[151,1437,1446,2014]" pageId="1" pageNumber="276">
The inclusion of the genus
<taxonomicName id="CDDEC5E7FF94287A2975F9A5747F8A24" authority="Nierstrasz, 1905" authorityName="Nierstrasz" authorityYear="1905" box="[515,868,1626,1653]" class="Polyplacophora" family="Loricidae" genus="Squamophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Chitonida" pageId="1" pageNumber="276" phylum="Mollusca" rank="genus">
<emphasis id="38AA6276FF94287A2975F9A575BB8A24" box="[515,672,1627,1653]" italics="true" pageId="1" pageNumber="276">Squamophora</emphasis>
<bibRefCitation id="6E4FC395FF94287A29DCF9A5747F8A24" author="Nierstrasz, H. F." box="[682,868,1626,1653]" pageId="1" pageNumber="276" pagination="1 - 112" refId="ref9490" refString="Nierstrasz, H. F. (1905) Die Chitonen der Siboga-Expedition. Siboga Expeditie, 48, 1 - 112, pls 1 - 8." type="journal article" year="1905">Nierstrasz, 1905</bibRefCitation>
</taxonomicName>
as a junior synonym of the genus
<taxonomicName id="CDDEC5E7FF94287A2F81F9A572458A24" ID-CoL="7P7VF" authorityName="Pilsbry" authorityYear="1893" box="[1271,1374,1627,1653]" class="Polyplacophora" family="Loricidae" genus="Loricella" kingdom="Animalia" order="Chitonida" pageId="1" pageNumber="276" phylum="Mollusca" rank="genus">
<emphasis id="38AA6276FF94287A2F81F9A572458A24" box="[1271,1374,1627,1653]" italics="true" pageId="1" pageNumber="276">Loricella</emphasis>
</taxonomicName>
(
<bibRefCitation id="6E4FC395FF94287A2E19F9A5760E8AC8" author="Van Belle, R. A." pageId="1" pageNumber="276" pagination="1 - 18" refId="ref10341" refString="Van Belle, R. A. (1978) Sur la classification des Polyplacophora. IV. Classification systematique des Mopaliidae (Neoloricata: Chitonina), avec la description de Heterochitoninae nov. subfam. Informations de la Societe Belge de Malacologie, 6, 1 - 18, pls. VI + VII." type="journal article" year="1978">Van Belle 1978</bibRefCitation>
,
<bibRefCitation id="6E4FC395FF94287A2A56F98176428AC8" author="Van Belle, R. A." box="[288,345,1663,1689]" pageId="1" pageNumber="276" pagination="1 - 179" refId="ref10398" refString="Van Belle, R. A. (1983) The systematic classification of the chitons (Mollusca: Polyplacophora). Informations de la Societe Belge de Malacologie, 11, 1 - 179." type="journal article" year="1983">1983</bibRefCitation>
) led to a confusion of generic characters of
<taxonomicName id="CDDEC5E7FF94287A283FF98174A98AC8" ID-CoL="7P7VF" authorityName="Pilsbry" authorityYear="1893" box="[841,946,1663,1689]" class="Polyplacophora" family="Loricidae" genus="Loricella" kingdom="Animalia" order="Chitonida" pageId="1" pageNumber="276" phylum="Mollusca" rank="genus">Loricella</taxonomicName>
. The restoration of the genus
<taxonomicName id="CDDEC5E7FF94287A2F89F981765B8AEC" authority="(Sirenko 2021)" baseAuthorityName="Sirenko" baseAuthorityYear="2021" class="Polyplacophora" family="Loricidae" genus="Squamophora" higherTaxonomySource="CoL" kingdom="Animalia" order="Chitonida" pageId="1" pageNumber="276" phylum="Mollusca" rank="genus">
<emphasis id="38AA6276FF94287A2F89F98172878AC8" box="[1279,1436,1663,1689]" italics="true" pageId="1" pageNumber="276">Squamophora</emphasis>
(
<bibRefCitation id="6E4FC395FF94287A2BE9F95D76238AEC" author="Sirenko, B. I." box="[159,312,1699,1725]" pageId="1" pageNumber="276" pagination="51 - 58" refId="ref10156" refString="Sirenko, B. I. (2021) Restoration of the genus Squamophora (Mollusca: Polyplacophora: Loricidae. Ruthenica, 31 (2), 51 - 58." type="journal article" year="2021">Sirenko 2021</bibRefCitation>
)
</taxonomicName>
makes an emendation of the diagnosis of the genus
<taxonomicName id="CDDEC5E7FF94287A28FFF95D74EB8AEC" ID-CoL="7P7VF" authorityName="Pilsbry" authorityYear="1893" box="[905,1008,1699,1725]" class="Polyplacophora" family="Loricidae" genus="Loricella" kingdom="Animalia" order="Chitonida" pageId="1" pageNumber="276" phylum="Mollusca" rank="genus">
<emphasis id="38AA6276FF94287A28FFF95D74EB8AEC" box="[905,1008,1699,1725]" italics="true" pageId="1" pageNumber="276">Loricella</emphasis>
</taxonomicName>
necessary:
</paragraph>
</subSubSection>
<subSubSection id="42C4EDEFFF9428792BB1F938746C8D34" lastPageId="2" lastPageNumber="277" pageId="1" pageNumber="276" type="diagnosis">
<paragraph id="0A61BE64FF9428792BB1F938746C8D34" blockId="1.[151,1437,1446,2014]" lastBlockId="2.[151,1437,151,357]" lastPageId="2" lastPageNumber="277" pageId="1" pageNumber="276">
<emphasis id="38AA6276FF94287A2BB1F93876598AB1" bold="true" box="[199,322,1734,1760]" pageId="1" pageNumber="276">Diagnosis.</emphasis>
Small to large sized, ovoid and broadened anteriorly; head valve large about more than twice as wide as tail valve, intermediate valves short and wide, tail valve with sinus and usually several indistinct slits, mucro terminal. Tegmentum of all species except for old specimens of
<taxonomicName id="CDDEC5E7FF94287A2805F8F174C58B79" box="[883,990,1807,1832]" class="Polyplacophora" family="Loricidae" genus="Lorica" kingdom="Animalia" order="Chitonida" pageId="1" pageNumber="276" phylum="Mollusca" rank="species" species="angasi">
<emphasis id="38AA6276FF94287A2805F8F174C58B79" box="[883,990,1807,1832]" italics="true" pageId="1" pageNumber="276">L. angasi</emphasis>
</taxonomicName>
has numerous pits, which are arranged mainly in radial rows on the head valve and lateral areas and in longitudinal rows on central areas. There is a row of larger and deeper pits on each diagonal between central and lateral areas of the tegmentum in deep-water species. Pores of aesthetes are numerous on surface of tegmentum between the pits. The jugal plate is usually well developed and often separated from the apophyses by a notch, on the intermediate valves it is wide, and on the tale valve it is narrow, tooth-shaped. Slit formula 8–20/1–(rare 2)/sinus +2–8 obsolete. Narrow mantle fold and wide head. Posterior slit in girdle is present or absent. Girdle, dorsally covered with small scales with tubular hollow and rare tufts of bristles. The tufts are located on the perinotum closer to the outer margin. There are rare single scattered bristles. The hyponotum scales are elongate, rectangular and smooth. There are 14–27 gills on each side arranged from valve II or III to valve VII or VIII. Radula
<quantity id="CD261381FF9728792945FEFD75A18D4C" box="[563,698,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="2.8499999999999996" metricValueMax="4.1" metricValueMin="1.6" pageId="2" pageNumber="277" unit="mm" value="2.8499999999999996" valueMax="4.1" valueMin="1.6">1.6–4.1 mm</quantity>
long with 28–45 transverse rows of mature teeth in chitons of BL 9.0–26.0 mm. Central tooth of radula short, bulbous, first lateral tooth slender, with small cusp at top, head of major lateral tooth bicuspid, major uncinal tooth with rather wide blade.
</paragraph>
</subSubSection>
</treatment>
</document>
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