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<document id="4533D710A669D57B26D20F9BEFFDF330" ID-CLB-Dataset="9534" ID-DOI="10.11646/zootaxa.4881.2.1" ID-GBIF-Dataset="120e0bf4-0fd9-47c7-85fd-4dd9ff878245" ID-ISSN="1175-5326" ID-Zenodo-Dep="4283530" ID-ZooBank="DA5F7402-0D1E-42D7-9DBE-26DF54671F0C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1605973600690" checkinUser="plazi" docAuthor="Moravec, Jiří &amp; Vybíral, Jan" docDate="2020" docId="824987FADA5BFFCCFF0EFEB5FDDCFE2C" docLanguage="en" docName="zootaxa.4881.2.1.pdf" docOrigin="Zootaxa 4881 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pogonostoma (Pogonostoma) ondravybirali Moravec &amp; Vybíral 2020, sp. nov." docType="treatment" docVersion="7" lastPageNumber="211" masterDocId="7E70FF82DA5FFFC6FF99FF85FFF2FFA5" masterDocTitle="New or rare Madagascar tiger beetles- 18. Pogonostoma (Pogonostoma) ondravybirali sp. nov., elaborated redescriptions of P. (P.) atrorotundatum, P. (P.) densisculptum and P. (Microstenocera) fabiocassolai (Coleoptera: Cicindelidae)" masterLastPageNumber="230" masterPageNumber="201" pageNumber="205" updateTime="1698846399937" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="168F215DD79FEB3976FAEE6CB021DB82" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A8F4F35E29F23E21CED9CFCFCB884A93">
<mods:title id="7C01DB767CA4A6CA2D4D70C80FF5C2BB">New or rare Madagascar tiger beetles- 18. Pogonostoma (Pogonostoma) ondravybirali sp. nov., elaborated redescriptions of P. (P.) atrorotundatum, P. (P.) densisculptum and P. (Microstenocera) fabiocassolai (Coleoptera: Cicindelidae)</mods:title>
</mods:titleInfo>
<mods:name id="B53F1E400C608F6AD2382BAC5864E355" type="personal">
<mods:role id="1EEB6F99B7973210574D9444F9FCF039">
<mods:roleTerm id="5E6DC9A317533F9196FAC589538F459B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="78158990A61DD1AE058302E178F4F213">Moravec, Jiří</mods:namePart>
<mods:affiliation id="1CD6703907CE8B53BBC6E8F520B974AC">Mendel University in Brno, Department of Forest Ecology, Zemědělská 3, 613 00 Brno, Czech Republic.</mods:affiliation>
</mods:name>
<mods:name id="4E1922B3466591F0364B0C938FD4A947" type="personal">
<mods:role id="9AE64D78813A9B50B68990243C6141A3">
<mods:roleTerm id="98F09AD61D2FA36F4791EABA459AC5DA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="435A43B2FDB2FE2FA285D5B6AC85FB2F">Vybíral, Jan</mods:namePart>
<mods:affiliation id="98261FD4AFAB60F3928118816E1EBC6F">Biosférická rezervace Dolní Morava, o. p. s., Národních hrdinů 23, 690 02 Břeclav, Czech Republic.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="24B8C153B8B3307F02E6B3ED3CE2C34E">text</mods:typeOfResource>
<mods:relatedItem id="30D254C36E2AD79C962EC1229D069E6F" type="host">
<mods:titleInfo id="DEFBB64BD10D2B677EB7324093CF1416">
<mods:title id="E83669C9EC9EEA4D5629850CEFC37DAA">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="906FEA921F393311DB6062556CAB3C7A">
<mods:date id="1017159485F96963D21016ED9BAA4482">2020</mods:date>
<mods:detail id="F6C346D27CFA5DE1BA82CD5B84D8DE13" type="pubDate">
<mods:number id="F9CE217FC759B79FBB1171A56D1D2B99">2020-11-19</mods:number>
</mods:detail>
<mods:detail id="3953E03E7B508D66E05D32D20850B4F2" type="volume">
<mods:number id="4502794A4002D569CC5AA814A4C63602">4881</mods:number>
</mods:detail>
<mods:detail id="119DD87FC3CAD2190798B9A78AD03277" type="issue">
<mods:number id="9F22EF8B933CEB466B878FD6DFED5CF7">2</mods:number>
</mods:detail>
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<mods:start id="888827ADE8F6AC9B7600C7DAF8DB5413">201</mods:start>
<mods:end id="B2607F39731D112851E35BFA39975F6F">230</mods:end>
</mods:extent>
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</mods:relatedItem>
<mods:classification id="F45B56A81A462BB16044DBE30C570F0D">journal article</mods:classification>
<mods:identifier id="3FB33CEFF600A3460CF31A54CA98EBC0" type="CLB-Dataset">9534</mods:identifier>
<mods:identifier id="D81ACB4D36B5F7FE0C3785EB112F71FD" type="DOI">10.11646/zootaxa.4881.2.1</mods:identifier>
<mods:identifier id="A0D894D8BCFC3CE24F2D52C83E71E332" type="GBIF-Dataset">120e0bf4-0fd9-47c7-85fd-4dd9ff878245</mods:identifier>
<mods:identifier id="FFE46F3C3F6B9FB0FB544E6AAEDDD520" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A84B2D353A31DFB9414B3C570F33238E" type="Zenodo-Dep">4283530</mods:identifier>
<mods:identifier id="BC6DB1843521ED81C1DE0C81403D4520" type="ZooBank">DA5F7402-0D1E-42D7-9DBE-26DF54671F0C</mods:identifier>
</mods:mods>
<treatment id="824987FADA5BFFCCFF0EFEB5FDDCFE2C" ID-DOI="http://doi.org/10.5281/zenodo.4328711" ID-GBIF-Taxon="170510467" ID-Zenodo-Dep="4328711" LSID="urn:lsid:plazi:treatment:824987FADA5BFFCCFF0EFEB5FDDCFE2C" httpUri="http://treatment.plazi.org/id/824987FADA5BFFCCFF0EFEB5FDDCFE2C" lastPageId="10" lastPageNumber="211" pageId="4" pageNumber="205">
<subSubSection id="42FA6567DA5BFFC2FF0EFEB5FCEEFEEF" box="[151,796,303,330]" pageId="4" pageNumber="205" type="nomenclature">
<paragraph id="0A5F36ECDA5BFFC2FF0EFEB5FCEEFEEF" blockId="4.[151,796,303,366]" box="[151,796,303,330]" pageId="4" pageNumber="205">
<heading id="51178180DA5BFFC2FF0EFEB5FCEEFEEF" bold="true" box="[151,796,303,330]" fontSize="11" level="1" pageId="4" pageNumber="205" reason="1">
<emphasis id="3894EAFEDA5BFFC2FF0EFEB5FCEEFEEF" bold="true" box="[151,796,303,330]" pageId="4" pageNumber="205">
<taxonomicName id="CDE04D6FDA5BFFC2FF0EFEB5FD44FEEF" authority="Moravec &amp; Vybíral, 2020" authorityName="Moravec &amp; Vybíral" authorityYear="2020" box="[151,694,303,330]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="ondravybirali" status="sp. nov." subGenus="Pogonostoma">
<emphasis id="3894EAFEDA5BFFC2FF0EFEB5FECCFEEF" bold="true" box="[151,318,304,330]" italics="true" pageId="4" pageNumber="205">Pogonostoma</emphasis>
(
<emphasis id="3894EAFEDA5BFFC2FEC9FEB5FE05FEEF" bold="true" box="[336,503,304,330]" italics="true" pageId="4" pageNumber="205">Pogonostoma</emphasis>
)
<emphasis id="3894EAFEDA5BFFC2FD90FEAAFD44FEEF" bold="true" box="[521,694,303,330]" italics="true" pageId="4" pageNumber="205">ondravybirali</emphasis>
</taxonomicName>
<taxonomicNameLabel id="23A75785DA5BFFC2FD24FEB5FCEEFEEF" box="[701,796,304,330]" pageId="4" pageNumber="205" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="42FA6567DA5BFFC2FF0EFED1FEA3FECB" box="[151,337,340,366]" pageId="4" pageNumber="205" type="description">
<paragraph id="0A5F36ECDA5BFFC2FF0EFED1FEA3FECB" blockId="4.[151,796,303,366]" box="[151,337,340,366]" pageId="4" pageNumber="205">
(
<figureCitation id="92DB2A69DA5BFFC2FF06FED1FEE0FECB" box="[159,274,340,366]" captionStart-0="FIGURES 1–10" captionStart-1="FIGURES 11–16" captionStart-2="FIGURES 17–20" captionStart-3="FIGURES 21–29" captionStartId-0="5.[151,264,1867,1891]" captionStartId-1="6.[151,264,1844,1868]" captionStartId-2="7.[151,264,1841,1865]" captionStartId-3="9.[151,264,1943,1967]" captionTargetBox-0="[160,1427,185,1831]" captionTargetBox-1="[165,1425,193,1810]" captionTargetBox-2="[163,1423,183,1804]" captionTargetBox-3="[163,1424,1213,1911]" captionTargetId-0="figure-17@5.[151,1435,181,1844]" captionTargetId-1="figure-20@6.[151,1435,181,1821]" captionTargetId-2="figure-17@7.[152,1435,181,1817]" captionTargetId-3="figure-484@9.[158,1429,1209,1919]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="9" captionText-0="FIGURES 1–10. Pogonostoma (P.) ondravybirali sp. nov. (from type locality). 1–2—habitus: 1—♁, 10.5 mm, HT (SDEI); 2—♀, 10.9 mm, AT (CJVB); 3–4—mandibles: 3—♀, AT; 4—♁, HT; 5—galea and lacinia, ♁, PT (CCJM); 6–8—labrum: 6—♁, HT; 7—♁, PT (CJVB); 8—♀, AT; 9—antennal scape and pedicel, ♀, AT; 10—head, ♁, HT. Bars = 1 mm." captionText-1="FIGURES 11–16. Pogonostoma (P.) ondravybirali sp. nov. (from type locality). 11–12—elytral apices: 11—♁, HT (SDEI); 12—♀, AT (CJVB); 13–14—elytron: 13—♁, HT; 14—♀, AT; 15—sculpture on elytral disc, ♁, HT; 16—ditto (higher magnifi- cation and on larger area of both elytra). Bars = 1 mm." captionText-2="FIGURES 17–20. Pogonostoma (P.) ondravybirali sp. nov. (from type locality), pronotum. 17—♁, HT (SDEI); 18—♀, AT (CJVB); 19—♁, PT (CJVB) 20—ditto, lateral view. Bars = 1 mm." captionText-3="FIGURES 21–29. Pogonostoma (P.) ondravybirali sp. nov. (from type locality), male aedeagus or its apex. 21—HT (SDEI); 22—ditto, ventral view; 23—PT (CJVB); 24—ditto, dorsal view; 25—ditto, ventral view; 26—PT (CCJM); 27—ditto, ventral view; 28—PT (CJVB); 29—ditto, ventral view. Bar = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4283532" figureDoi-1="http://doi.org/10.5281/zenodo.4283534" figureDoi-2="http://doi.org/10.5281/zenodo.4283536" figureDoi-3="http://doi.org/10.5281/zenodo.4283538" httpUri-0="https://zenodo.org/record/4283532/files/figure.png" httpUri-1="https://zenodo.org/record/4283534/files/figure.png" httpUri-2="https://zenodo.org/record/4283536/files/figure.png" httpUri-3="https://zenodo.org/record/4283538/files/figure.png" pageId="4" pageNumber="205">Figs 1–29</figureCitation>
,
<figureCitation id="92DB2A69DA5BFFC2FE87FED1FEBBFECB" box="[286,329,340,366]" captionStart="FIGURES 104–107" captionStartId="27.[151,264,1724,1748]" captionTargetBox="[164,1423,167,1692]" captionTargetId="figure-17@27.[151,1436,151,1707]" captionTargetPageId="27" captionText="FIGURES 104–107. Maps of distribution. 104—Pogonostoma (Pogonostoma) ondravybirali sp. nov.; 105—Pogonostoma (P.) atrorotundatum W. Horn; 106—Pogonostoma (P.) densisculptum Moravec; 107—Pogonostoma (Microstenocera) fabiocassolai Moravec." figureDoi="http://doi.org/10.5281/zenodo.4283560" httpUri="https://zenodo.org/record/4283560/files/figure.png" pageId="4" pageNumber="205">104</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="42FA6567DA5BFFC2FF0EFE12FF0CFC24" pageId="4" pageNumber="205" type="materials_examined">
<paragraph id="0A5F36ECDA5BFFC2FF0EFE12FCDAFE71" blockId="4.[151,1437,406,2042]" pageId="4" pageNumber="205">
<materialsCitation id="BA883CB1DA5BFFC2FF0EFE12FCD6FE71" ID-GBIF-Occurrence="2985230318" country="Madagascar" county="Ranohira" location="Southwestern" municipality="Isalo National Park" pageId="4" pageNumber="205" specimenCount="1" typeStatus="holotype">
<emphasis id="3894EAFEDA5BFFC2FF0EFE12FEC3FE15" bold="true" box="[151,305,406,433]" pageId="4" pageNumber="205">
<typeStatus id="D55B884EDA5BFFC2FF0EFE12FF20FE14" box="[151,210,407,433]" pageId="4" pageNumber="205">Type</typeStatus>
locality
</emphasis>
.
<location id="0F3F6037DA5BFFC2FED9FE13FE2AFE15" LSID="urn:lsid:plazi:treatment:824987FADA5BFFCCFF0EFEB5FDDCFE2C:0F3F6037DA5BFFC2FED9FE13FE2AFE15" box="[320,472,406,432]" country="Madagascar" county="Ranohira" municipality="Isalo National Park" name="Southwestern" pageId="4" pageNumber="205">Southwestern</location>
<collectingCountry id="72F7767CDA5BFFC2FE78FE13FD9AFE15" box="[481,616,406,432]" name="Madagascar" pageId="4" pageNumber="205">Madagascar</collectingCountry>
(
<location id="0F3F6037DA5BFFC2FDE1FE13FCB4FE14" LSID="urn:lsid:plazi:treatment:824987FADA5BFFCCFF0EFEB5FDDCFE2C:0F3F6037DA5BFFC2FDE1FE13FCB4FE14" box="[632,838,406,433]" country="Madagascar" county="Ranohira" municipality="Isalo National Park" name="Ihorombe Region" pageId="4" pageNumber="205">Ihorombe Region</location>
): sclerophylous woodland area with
<location id="0F3F6037DA5BFFC2FB7DFE13FAD0FE15" LSID="urn:lsid:plazi:treatment:824987FADA5BFFCCFF0EFEB5FDDCFE2C:0F3F6037DA5BFFC2FB7DFE13FAD0FE15" box="[1252,1314,406,432]" country="Madagascar" county="Ranohira" municipality="Isalo National Park" name="Tapia" pageId="4" pageNumber="205">Tapia</location>
trees (
<taxonomicName id="CDE04D6FDA5BFFC2FAE8FE12FEBDFE70" authority="Baill" authorityName="Baill" class="Magnoliopsida" family="Phyllanthaceae" genus="Uapaca" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="205" phylum="Tracheophyta" rank="species" species="boeri">
<emphasis id="3894EAFEDA5BFFC2FAE8FE12FEE2FE71" italics="true" pageId="4" pageNumber="205">Uapaca boeri</emphasis>
Baill
</taxonomicName>
) in the
<collectingMunicipality id="EA3BAC96DA5BFFC2FE3AFE3FFD89FE71" box="[419,635,442,468]" pageId="4" pageNumber="205">Isalo National Park</collectingMunicipality>
near
<collectingCounty id="E33E4E60DA5BFFC2FD23FE3FFCD6FE71" box="[698,804,442,468]" pageId="4" pageNumber="205">Ranohira</collectingCounty>
</materialsCitation>
.
</paragraph>
<paragraph id="0A5F36ECDA5BFFC2FF5EFE5BFD46FC9F" blockId="4.[151,1437,406,2042]" pageId="4" pageNumber="205">
<emphasis id="3894EAFEDA5BFFC2FF5EFE5BFE9DFE5D" bold="true" box="[199,367,478,504]" pageId="4" pageNumber="205">Type material</emphasis>
.
<materialsCitation id="BA883CB1DA5BFFC2FEE6FE5BFD31FDB9" ID-GBIF-Occurrence="2985230315" collectionCode="SDEI" collectorName="Jan Vybiral" country="Madagascar" elevation="825" latitude="-22.551943" location="Madagascar" longLatPrecision="15" longitude="45.41375" pageId="4" pageNumber="205" specimenCount="1" typeStatus="holotype">
<typeStatus id="D55B884EDA5BFFC2FEE6FE5BFE15FE5D" box="[383,487,478,504]" pageId="4" pageNumber="205" type="holotype">Holotype</typeStatus>
♁ in
<collectionCode id="6CF1AE29DA5BFFC2FDB5FE5BFD9FFE5D" box="[556,621,478,504]" pageId="4" pageNumber="205">SDEI</collectionCode>
, labelled: “
<collectingCountry id="72F7767CDA5BFFC2FD75FE5BFC89FE5D" box="[748,891,478,504]" name="Madagascar" pageId="4" pageNumber="205">Madagascar</collectingCountry>
SW: Isalo N.P /
<geoCoordinate id="6FD4502BDA5BFFC2FBDCFE5BFB2BFE5D" box="[1093,1241,478,504]" degrees="22" direction="south" minutes="33" orientation="latitude" pageId="4" pageNumber="205" precision="15" seconds="07" value="-22.551943">S22°33´07´´</geoCoordinate>
;
<geoCoordinate id="6FD4502BDA5BFFC2FB7FFE5BFA79FE5C" box="[1254,1419,478,505]" degrees="45" direction="east" minutes="24" orientation="longitude" pageId="4" pageNumber="205" precision="1" seconds="49.5" value="45.41375">E45°24´49.5´´</geoCoordinate>
/
<quantity id="CD189B09DA5BFFC2FF0EFD87FF15FDB8" box="[151,231,514,541]" metricMagnitude="2" metricUnit="m" metricValue="8.25" pageId="4" pageNumber="205" unit="m" value="825.0">
<elevation id="81CDD1DFDA5BFFC2FF0EFD87FF15FDB8" box="[151,231,514,541]" metricMagnitude="2" metricUnit="m" metricValue="8.25" pageId="4" pageNumber="205" unit="m" value="825.0">825 m</elevation>
</quantity>
,
<date id="7E5E102CDA5BFFC2FF6DFD87FE9DFDB9" box="[244,367,514,540]" pageId="4" pageNumber="205" value="2010-01-11">11-I. 2010</date>
/ leg.
<collectorName id="A715533ADA5BFFC2FE27FD86FDB4FDB9" box="[446,582,514,541]" pageId="4" pageNumber="205">Jan Vybíral</collectorName>
” [printed]
</materialsCitation>
.
<materialsCitation id="BA883CB1DA5BFFC2FD57FD87FDABFDE5" ID-GBIF-Occurrence="2985230301" collectingDate="2017-01-04" collectionCode="CJVB" collectorName="Jan" country="Madagascar" elevation="825" latitude="-22.551943" location="Madagascar" longLatPrecision="15" longitude="45.41375" pageId="4" pageNumber="205" specimenCount="1" specimenCount-female="1" typeStatus="allotype">
<typeStatus id="D55B884EDA5BFFC2FD57FD87FCC6FDB9" box="[718,820,514,540]" pageId="4" pageNumber="205" type="allotype">Allotype</typeStatus>
<specimenCount id="1CE6FD65DA5BFFC2FCA4FD86FCA0FDB8" box="[829,850,515,541]" pageId="4" pageNumber="205" type="female">♀</specimenCount>
in
<collectionCode id="6CF1AE29DA5BFFC2FCE5FD87FC30FDB9" box="[892,962,514,540]" pageId="4" pageNumber="205">CJVB</collectionCode>
with the same locality label except for: “
<date id="7E5E102CDA5BFFC2FF3BFDA3FEF7FDE5" box="[162,261,550,576]" pageId="4" pageNumber="205" value="2017-01-04">
<collectingDate id="6E1AE9C4DA5BFFC2FF3BFDA3FEF7FDE5" box="[162,261,550,576]" pageId="4" pageNumber="205" value="2017-01-04">4.I.2017</collectingDate>
</date>
/ leg.
<collectorName id="A715533ADA5BFFC2FED6FDA3FE84FDE5" box="[335,374,550,576]" pageId="4" pageNumber="205">Jan</collectorName>
&amp; Ondřej Vybíral”
</materialsCitation>
.
<materialsCitation id="BA883CB1DA5BFFC2FDFDFDA3FC3EFDE5" ID-GBIF-Occurrence="2985230305" box="[612,972,550,578]" collectionCode="CJVB" country="Madagascar" pageId="4" pageNumber="205" specimenCount="6" specimenCount-female="6" typeStatus="paratype">
<typeStatus id="D55B884EDA5BFFC2FDFDFDA3FD2BFDE5" box="[612,729,550,576]" pageId="4" pageNumber="205" type="paratype">Paratypes</typeStatus>
. 1 ♁,
<specimenCount id="1CE6FD65DA5BFFC2FC86FDA3FC92FDE5" box="[799,864,550,576]" pageId="4" pageNumber="205" type="female">6 ♀♀</specimenCount>
in
<collectionCode id="6CF1AE29DA5BFFC2FC1FFDA3FC3EFDE5" box="[902,972,550,576]" pageId="4" pageNumber="205">CJVB</collectionCode>
</materialsCitation>
<materialsCitation id="BA883CB1DA5BFFC2FC4DFDA3FF2BFDC3" ID-GBIF-Occurrence="2985230313" collectionCode="CJVB" country="Madagascar" pageId="4" pageNumber="205" specimenCount="1" typeStatus="holotype">
with same locality label as in
<typeStatus id="D55B884EDA5BFFC2FAA9FDA3FA6BFDE5" box="[1328,1433,550,576]" pageId="4" pageNumber="205" type="holotype">holotype</typeStatus>
. 7 ♁♁
</materialsCitation>
,
<materialsCitation id="BA883CB1DA5BFFC2FF7FFDCFFE56FDC1" ID-GBIF-Occurrence="2985230329" box="[230,420,585,612]" collectionCode="CJVB" country="Madagascar" pageId="4" pageNumber="205" specimenCount="22" specimenCount-female="22" typeStatus="paratype">
<specimenCount id="1CE6FD65DA5BFFC2FF7FFDCFFEC4FDC1" box="[230,310,586,612]" pageId="4" pageNumber="205" type="female">22 ♀♀</specimenCount>
in
<collectionCode id="6CF1AE29DA5BFFC2FEC2FDCFFE56FDC1" box="[347,420,586,612]" pageId="4" pageNumber="205">CJVB</collectionCode>
</materialsCitation>
, 1 ♁ in
<collectionCode id="6CF1AE29DA5BFFC2FD99FDCFFDA9FDC1" box="[512,603,586,612]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="205">MNHN</collectionCode>
,
<materialsCitation id="BA883CB1DA5BFFC2FDFFFDCFFD0BFDC1" ID-GBIF-Occurrence="2985230319" box="[614,761,585,612]" collectionCode="SDEI" country="Madagascar" pageId="4" pageNumber="205" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="1CE6FD65DA5BFFC2FDFFFDCFFD62FDC1" box="[614,656,586,612]" pageId="4" pageNumber="205" type="female">1 ♀</specimenCount>
in
<collectionCode id="6CF1AE29DA5BFFC2FD2FFDCFFD0BFDC1" box="[694,761,586,612]" pageId="4" pageNumber="205">SDEI</collectionCode>
</materialsCitation>
,
<materialsCitation id="BA883CB1DA5BFFC2FC9CFDCFFC5BFDC1" ID-GBIF-Occurrence="2985230332" box="[773,937,585,612]" collectionCode="MFNB" country="Madagascar" pageId="4" pageNumber="205" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="1CE6FD65DA5BFFC2FC9CFDCFFCDDFDC1" box="[773,815,586,612]" pageId="4" pageNumber="205" type="female">1 ♀</specimenCount>
in
<collectionCode id="6CF1AE29DA5BFFC2FCCCFDCFFC5BFDC1" box="[853,937,586,612]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="4" pageNumber="205">MFNB</collectionCode>
</materialsCitation>
,
<materialsCitation id="BA883CB1DA5BFFC2FC2CFDCFFBA5FDC1" ID-GBIF-Occurrence="2985230311" box="[949,1111,585,612]" collectionCode="BMNH" country="Madagascar" pageId="4" pageNumber="205" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="1CE6FD65DA5BFFC2FC2CFDCFFC28FDC6" box="[949,986,585,612]" pageId="4" pageNumber="205" type="female">1 ♀</specimenCount>
in
<collectionCode id="6CF1AE29DA5BFFC2FC67FDCFFBA5FDC1" box="[1022,1111,586,612]" pageId="4" pageNumber="205">BMNH</collectionCode>
</materialsCitation>
, 2 ♁♁,
<materialsCitation id="BA883CB1DA5BFFC2FB28FDCCFA90FDC1" ID-GBIF-Occurrence="2985230309" box="[1201,1378,585,612]" collectionCode="CCJM" country="Madagascar" pageId="4" pageNumber="205" specimenCount="5" specimenCount-female="5" typeStatus="paratype">
<specimenCount id="1CE6FD65DA5BFFC2FB28FDCCFB00FDC1" box="[1201,1266,585,612]" pageId="4" pageNumber="205" type="female">5 ♀♀</specimenCount>
in
<collectionCode id="6CF1AE29DA5BFFC2FA8EFDCFFA90FDC1" box="[1303,1378,586,612]" pageId="4" pageNumber="205">CCJM</collectionCode>
</materialsCitation>
<materialsCitation id="BA883CB1DA5BFFC2FAF0FDCCFD90FD2F" ID-GBIF-Occurrence="2985230324" collectionCode="CCJM" country="Madagascar" pageId="4" pageNumber="205" specimenCount="1" typeStatus="allotype">
with same locality label as in
<typeStatus id="D55B884EDA5BFFC2FE2CFDE8FDE4FD22" box="[437,534,621,647]" pageId="4" pageNumber="205" type="allotype">allotype</typeStatus>
. 4 ♁♁
</materialsCitation>
,
<materialsCitation id="BA883CB1DA5BFFC2FDE9FDE8FCD1FD22" ID-GBIF-Occurrence="2985230325" box="[624,803,621,648]" collectionCode="MSCB" country="Madagascar" pageId="4" pageNumber="205" specimenCount="4" specimenCount-female="4" typeStatus="paratype">
<specimenCount id="1CE6FD65DA5BFFC2FDE9FDE8FD42FD2D" box="[624,688,621,648]" pageId="4" pageNumber="205" type="female">4 ♀♀</specimenCount>
in
<collectionCode id="6CF1AE29DA5BFFC2FD4CFDE8FCD1FD22" box="[725,803,621,647]" pageId="4" pageNumber="205">MSCB</collectionCode>
</materialsCitation>
with same labels as in
<typeStatus id="D55B884EDA5BFFC2FBBAFDE8FB8FFD22" box="[1059,1149,621,647]" pageId="4" pageNumber="205" type="allotype">allotype</typeStatus>
except for “leg.
<collectorName id="A715533ADA5BFFC2FAAEFDE8FF0DFD0E" pageId="4" pageNumber="205">Miroslav Svoboda</collectorName>
”.
<materialsCitation id="BA883CB1DA5BFFC2FE8AFD14FE4AFD0E" ID-GBIF-Occurrence="2985230322" box="[275,440,657,683]" collectionCode="CMTD" country="Madagascar" pageId="4" pageNumber="205" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="1CE6FD65DA5BFFC2FE8AFD14FECFFD0E" box="[275,317,657,683]" pageId="4" pageNumber="205" type="female">1 ♀</specimenCount>
in
<collectionCode id="6CF1AE29DA5BFFC2FEFBFD14FE4AFD0E" box="[354,440,657,683]" pageId="4" pageNumber="205">CMTD</collectionCode>
</materialsCitation>
, 2 ♁♁ in
<collectionCode id="6CF1AE29DA5BFFC2FDBFFD14FD83FD0E" box="[550,625,657,683]" pageId="4" pageNumber="205">CJVB</collectionCode>
:
<materialsCitation id="BA883CB1DA5BFFC2FDE5FD14FD23FD6A" ID-GBIF-Occurrence="2985230326" collectionCode="ZUAC" collectorName="M. Tryzna" country="Madagascar" county="Ranohira" location="Isalo Nat. Park" municipality="Namaza" pageId="4" pageNumber="205" specimenCount="1" typeStatus="paratype">
“
<collectingCountry id="72F7767CDA5BFFC2FD1CFD14FCE4FD0E" box="[645,790,657,683]" name="Madagascar" pageId="4" pageNumber="205">Madagascar</collectingCountry>
, 2007 /
<location id="0F3F6037DA5BFFC2FCE9FD14FBD2FD0E" LSID="urn:lsid:plazi:treatment:824987FADA5BFFCCFF0EFEB5FDDCFE2C:0F3F6037DA5BFFC2FCE9FD14FBD2FD0E" box="[880,1056,657,683]" country="Madagascar" county="Ranohira" municipality="Namaza" name="Isalo Nat. Park" pageId="4" pageNumber="205">Isalo Nat. Park</location>
, camp /
<collectingMunicipality id="EA3BAC96DA5BFFC2FBE6FD14FB13FD0E" box="[1151,1249,657,683]" pageId="4" pageNumber="205">Namaza</collectingMunicipality>
, near
<collectingCounty id="E33E4E60DA5BFFC2FABDFD14FA7CFD0E" box="[1316,1422,657,683]" pageId="4" pageNumber="205">Ranohira</collectingCounty>
/
<collectorName id="A715533ADA5BFFC2FF0EFD30FEFDFD6A" box="[151,271,693,719]" pageId="4" pageNumber="205">M. Trýzna</collectorName>
leg., 17-18.I.” [printed]. 1 ♁ in
<collectionCode id="6CF1AE29DA5BFFC2FD19FD30FD23FD6A" box="[640,721,693,719]" country="Madagascar" httpUri="http://grbio.org/cool/dprk-p82z" name="University of Antananarivo" pageId="4" pageNumber="205">ZUAC</collectionCode>
</materialsCitation>
, 2 ♁♁,
<materialsCitation id="BA883CB1DA5BFFC2FCB0FD30FC25FD6A" ID-GBIF-Occurrence="2985230306" box="[809,983,693,719]" collectionCode="MHCW" country="Madagascar" pageId="4" pageNumber="205" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="1CE6FD65DA5BFFC2FCB0FD30FCA1FD6A" box="[809,851,693,719]" pageId="4" pageNumber="205" type="female">1 ♀</specimenCount>
in
<collectionCode id="6CF1AE29DA5BFFC2FCEEFD30FC25FD6A" box="[887,983,693,719]" pageId="4" pageNumber="205">MHCW</collectionCode>
</materialsCitation>
:
<materialsCitation id="BA883CB1DA5BFFC2FC78FD30FD43FC9F" ID-GBIF-Occurrence="2985230310" collectingDate="2017-12-09" collectorName="Elyse Razanajaonarivalona" country="Madagascar" determinerName="Jiri Moravec" location="Madagascar" pageId="4" pageNumber="205" specimenCount="3" typeStatus="holotype">
“near Ranohira, Isalo N. P., / Iforombe Reg. /
<collectingCountry id="72F7767CDA5BFFC2FF7BFD5DFE9CFD57" box="[226,366,728,754]" name="Madagascar" pageId="4" pageNumber="205">Madagascar</collectingCountry>
/
<date id="7E5E102CDA5BFFC2FE1AFD5CFDEBFD56" box="[387,537,729,755]" pageId="4" pageNumber="205" value="2017-12-09">
<collectingDate id="6E1AE9C4DA5BFFC2FE1AFD5CFDEBFD56" box="[387,537,729,755]" pageId="4" pageNumber="205" value="2017-12-09">Dec. 9, 2017</collectingDate>
</date>
, Michio Hori &amp; /
<collectorName id="A715533ADA5BFFC2FD7DFD5DFBE1FD57" box="[740,1043,728,754]" pageId="4" pageNumber="205">Elysé Razanajaonarivalona</collectorName>
leg.” [printed]. All type specimens labelled: “
<typeStatus id="D55B884EDA5BFFC2FE95FD79FE89FCB3" box="[268,379,764,790]" pageId="4" pageNumber="205" type="holotype">Holotype</typeStatus>
(
<typeStatus id="D55B884EDA5BFFC2FE11FD79FE02FCB3" box="[392,496,764,790]" pageId="4" pageNumber="205" type="allotype">Allotype</typeStatus>
or
<typeStatus id="D55B884EDA5BFFC2FD8DFD78FD88FCB2" box="[532,634,765,791]" pageId="4" pageNumber="205" type="paratype">Paratype</typeStatus>
respectively) /
<taxonomicName id="CDE04D6FDA5BFFC2FCB2FD78FC3AFCB2" authorityName="Klug" authorityYear="1835" box="[811,968,765,791]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="genus">Pogonostoma</taxonomicName>
(s. str.) / ondravybirali
<emphasis id="3894EAFEDA5BFFC2FB4FFD79FAC3FCB3" bold="true" box="[1238,1329,764,790]" pageId="4" pageNumber="205">
<taxonomicNameLabel id="23A75785DA5BFFC2FB4FFD79FAC3FCB3" box="[1238,1329,764,790]" pageId="4" pageNumber="205" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
/ det.
<determinerName id="FF83DA28DA5BFFC2FAEEFD79FF0CFC9F" pageId="4" pageNumber="205">Jiří Moravec</determinerName>
/ &amp; Jan Vybíral 2020”. [red, printed]
</materialsCitation>
.
</paragraph>
<paragraph id="0A5F36ECDA5BFFC2FF5EFCC6FF0CFC24" blockId="4.[151,1437,406,2042]" pageId="4" pageNumber="205">
<emphasis id="3894EAFEDA5BFFC2FF5EFCC6FE0FFCF8" bold="true" box="[199,509,835,861]" pageId="4" pageNumber="205">Other material examined</emphasis>
.
<materialsCitation id="BA883CB1DA5BFFC2FD94FCC1FF08FC24" ID-GBIF-Occurrence="2985230328" collectingDate="2017-01-06" collectionCode="CJVB" collectorName="Jan" country="Madagascar" location="Madagascar" pageId="4" pageNumber="205" specimenCount="1">
2 ♁♁ in
<collectionCode id="6CF1AE29DA5BFFC2FDEBFCC1FD4FFCFB" box="[626,701,836,862]" pageId="4" pageNumber="205">CJVB</collectionCode>
: “
<collectingCountry id="72F7767CDA5BFFC2FD48FCC1FC90FCFB" box="[721,866,836,862]" name="Madagascar" pageId="4" pageNumber="205">Madagascar</collectingCountry>
SW / Zombitse N.P.,
<date id="7E5E102CDA5BFFC2FBFDFCC6FB33FCF8" box="[1124,1217,835,861]" pageId="4" pageNumber="205" value="2017-01-06">
<collectingDate id="6E1AE9C4DA5BFFC2FBFDFCC6FB33FCF8" box="[1124,1217,835,861]" pageId="4" pageNumber="205" value="2017-01-06">6.I.2017</collectingDate>
</date>
/ leg.
<collectorName id="A715533ADA5BFFC2FA9FFCC1FAD9FCFB" box="[1286,1323,836,862]" pageId="4" pageNumber="205">Jan</collectorName>
&amp; Ondřej Vybíral”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="42FA6567DA5BFFC2FF5EFC0EFE13F9D4" pageId="4" pageNumber="205" type="diagnosis">
<paragraph id="0A5F36ECDA5BFFC2FF5EFC0EFB06FC6C" blockId="4.[151,1437,406,2042]" pageId="4" pageNumber="205">
<emphasis id="3894EAFEDA5BFFC2FF5EFC0EFE23FC00" bold="true" box="[199,465,907,933]" pageId="4" pageNumber="205">Differential diagnosis</emphasis>
.
<taxonomicName id="CDE04D6FDA5BFFC2FE7BFC09FC26FC00" authority="Moravec &amp; Vybíral, 2020" authorityName="Moravec &amp; Vybíral" authorityYear="2020" box="[482,980,907,934]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="ondravybirali" status="sp. nov." subGenus="Pogonostoma">
<emphasis id="3894EAFEDA5BFFC2FE7BFC09FD8EFC00" box="[482,636,908,933]" italics="true" pageId="4" pageNumber="205">Pogonostoma</emphasis>
(
<emphasis id="3894EAFEDA5BFFC2FD17FC09FCDAFC00" box="[654,808,908,933]" italics="true" pageId="4" pageNumber="205">Pogonostoma</emphasis>
)
<emphasis id="3894EAFEDA5BFFC2FCA3FC0EFC26FC00" box="[826,980,907,933]" italics="true" pageId="4" pageNumber="205">ondravybirali</emphasis>
</taxonomicName>
<emphasis id="3894EAFEDA5BFFC2FC45FC0EFBC5FC00" bold="true" box="[988,1079,907,933]" pageId="4" pageNumber="205">
<taxonomicNameLabel id="23A75785DA5BFFC2FC45FC0EFBC5FC00" box="[988,1079,907,933]" pageId="4" pageNumber="205" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is placed here in the
<taxonomicName id="CDE04D6FDA5BFFC2FAA9FC09FF2DFC6C" authorityName="W. Horn" authorityYear="1898" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="alluaudi" subGenus="Pogonostoma">
<emphasis id="3894EAFEDA5BFFC2FAA9FC09FAB4FC00" box="[1328,1350,908,933]" italics="true" pageId="4" pageNumber="205">P.</emphasis>
(
<emphasis id="3894EAFEDA5BFFC2FACEFC09FA9EFC00" box="[1367,1388,908,933]" italics="true" pageId="4" pageNumber="205">P.</emphasis>
)
<emphasis id="3894EAFEDA5BFFC2FAE4FC0EFF2DFC6C" italics="true" pageId="4" pageNumber="205">alluaudi</emphasis>
</taxonomicName>
species-complex within the rather large
<taxonomicName id="CDE04D6FDA5BFFC2FD3FFC35FCBBFC6C" authorityName="Brulle in Audouin &amp; Brulle" authorityYear="1834" box="[678,841,943,969]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="219" phylum="Arthropoda" rank="species" species="elegans" subGenus="Pogonostoma">
<emphasis id="3894EAFEDA5BFFC2FD3FFC35FD49FC6C" box="[678,699,944,969]" italics="true" pageId="4" pageNumber="205">P.</emphasis>
(
<emphasis id="3894EAFEDA5BFFC2FD52FC35FD2EFC6C" box="[715,732,944,969]" italics="true" pageId="4" pageNumber="205">P</emphasis>
.)
<emphasis id="3894EAFEDA5BFFC2FD6AFC2AFCBBFC6C" box="[755,841,943,969]" italics="true" pageId="4" pageNumber="205">elegans</emphasis>
</taxonomicName>
species-group (sensu
<bibRefCitation id="6E714B1DDA5BFFC2FBD8FC2AFB1AFC6C" author="MORAVEC" box="[1089,1256,943,969]" pageId="4" pageNumber="205" refId="ref17625" refString="MORAVEC, J. (2007) Tiger beetles of Madagascar 1. A monograph of the genus Pogonostoma (Coleoptera: Cicindelidae). Nakladatelstvi Kabourek, Zlin, 499 pp." type="book" year="2007">Moravec 2007</bibRefCitation>
).
</paragraph>
<paragraph id="0A5F36ECDA5BFFC2FF5EFC56FC09F9A1" blockId="4.[151,1437,406,2042]" pageId="4" pageNumber="205">
<taxonomicName id="CDE04D6FDA5BFFC2FF5EFC56FD94FC48" authority="W. Horn, 1934" authorityName="W. Horn" authorityYear="1934" box="[199,614,978,1005]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="atrorotundatum" subGenus="Pogonostoma">
<emphasis id="3894EAFEDA5BFFC2FF5EFC56FF2FFC49" box="[199,221,979,1004]" italics="true" pageId="4" pageNumber="205">P.</emphasis>
(
<emphasis id="3894EAFEDA5BFFC2FF72FC56FEF3FC49" box="[235,257,979,1004]" italics="true" pageId="4" pageNumber="205">P.</emphasis>
)
<emphasis id="3894EAFEDA5BFFC2FE96FC57FE4CFC49" box="[271,446,978,1004]" italics="true" pageId="4" pageNumber="205">atrorotundatum</emphasis>
W. Horn, 1934
</taxonomicName>
(treated below) shares the punctate-setose posterior pronotal lobe with the new species but is clearly differentiated by notably larger punctures on its elytral surface (
<figureCitation id="92DB2A69DA5BFFC2FB1FFC73FAFBFBB5" box="[1158,1289,1014,1040]" captionStart="FIGURES 41–49" captionStartId="14.[151,264,1850,1874]" captionTargetBox="[153,1430,189,1816]" captionTargetId="figure-20@14.[151,1435,181,1826]" captionTargetPageId="14" captionText="FIGURES 41–49. Pogonostoma (P.) atrorotundatum W. Horn. 41–44—elytral apices: 41—♁, “C. W. Madagascar”, LT (SDEI); 42—♁, Bongolava, HT of syn. P. (P.) rufidens Rivalier (MNHN); 43—♀, “Tananarivo”, PLT (SDEI); 44—♀, Miandrivazo, PLT (MNHN); 45–49—elytron or its detail: 45—♁, LT (SDEI); 46—ditto, sculpture on elytral disc; 47—♁, Antsalova (CCJM ex APCA); 48—♁, “Tananarivo”, as syn. P. (P.) rufidens Rivalier (CCJM ex APCA); 49—♀, Antsalova (CCJM). Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4283542" httpUri="https://zenodo.org/record/4283542/files/figure.png" pageId="4" pageNumber="205">Figs 41–49</figureCitation>
), and its pronotal disc is covered with more anastomosing tubercles forming short or more elongate and continuous transverse rugae on wide median discal area (
<figureCitation id="92DB2A69DA5BFFC2FD8DFBBBFD64FBFC" box="[532,662,1086,1113]" captionStart="FIGURES 50–54" captionStartId="15.[151,264,1867,1891]" captionTargetBox="[152,1431,189,1827]" captionTargetId="figure-17@15.[151,1435,181,1844]" captionTargetPageId="15" captionText="FIGURES 50–54. Pogonostoma (P.) atrorotundatum W. Horn, pronotum. 50—♁, “C. W. Madagascar”, LT (SDEI); 51—♁, ibid., PLT (SDEI); 52—♁, Antsalova (CCJM ex APCA); 53—♁, Bongolava, HT of syn. P. (P.) rufidens Rivalier (MNHN); 54—♀, Antsalova (CCJM). Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4283546" httpUri="https://zenodo.org/record/4283546/files/figure.png" pageId="4" pageNumber="205">Figs 50–54</figureCitation>
); moreover the aedeagus in
<taxonomicName id="CDE04D6FDA5BFFC2FC50FBBBFB57FBFD" box="[969,1189,1086,1112]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="atrorotundatum" subGenus="Pogonostoma">
(
<emphasis id="3894EAFEDA5BFFC2FC4BFBBAFC15FBFD" box="[978,999,1087,1112]" italics="true" pageId="4" pageNumber="205">P.</emphasis>
)
<emphasis id="3894EAFEDA5BFFC2FC6FFBBBFB57FBFD" box="[1014,1189,1086,1112]" italics="true" pageId="4" pageNumber="205">atrorotundatum</emphasis>
</taxonomicName>
has notably elongated and ventrad-bent apex (in its left lateral aspect,
<figureCitation id="92DB2A69DA5BFFC2FD20FBE4FCCFFBD9" box="[697,829,1121,1148]" captionStart="FIGURES 55–66" captionStartId="16.[151,264,1818,1842]" captionTargetBox="[163,1426,193,1782]" captionTargetId="figure-20@16.[151,1435,181,1794]" captionTargetPageId="16" captionText="FIGURES 55–66. Pogonostoma (P.) atrorotundatum W. Horn, male aedeagus or its apex. 55—“C.W. Madagascar”, LT (SDEI); 56—ditto, dorsal view; 57—ibid., PLT (SDEI); 58—Bongolava, HT of syn. P. (P.) rufidens Rivalier (MNHN); 59—ibid., PT of syn. P. (P.) rufidens Rivalier (MNHN); 60—Antsalova (CCJM ex APCA); 61—ditto, ventral view; 62—ditto, dorsal view; 63—ibid. (CCJM) 64—ditto dorsal view; 65—Ambalatomby (CCJM); 66—labels of LT (SDEI). Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4283548" httpUri="https://zenodo.org/record/4283548/files/figure.png" pageId="4" pageNumber="205">Figs 55–66</figureCitation>
). Other two rather similar species of the complex are
<taxonomicName id="CDE04D6FDA5BFFC2FF0EFB03FCCEFB3A" authority="Moravec, Razanajaonarivalona &amp; Hori 2020" authorityName="Moravec, Razanajaonarivalona &amp; Hori" authorityYear="2020" box="[151,828,1157,1184]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="natsuae" subGenus="Pogonostoma">
<emphasis id="3894EAFEDA5BFFC2FF0EFB03FF5FFB3A" box="[151,173,1158,1183]" italics="true" pageId="4" pageNumber="205">P.</emphasis>
(
<emphasis id="3894EAFEDA5BFFC2FF24FB03FF21FB3A" box="[189,211,1158,1183]" italics="true" pageId="4" pageNumber="205">P.</emphasis>
)
<emphasis id="3894EAFEDA5BFFC2FF7AFB03FEC8FB3A" box="[227,314,1158,1183]" italics="true" pageId="4" pageNumber="205">natsuae</emphasis>
<bibRefCitation id="6E714B1DDA5BFFC2FEDBFB03FCCEFB3A" author="Moravec &amp; Razanajaonarivalona, E. H. &amp; HORI, M." box="[322,828,1157,1184]" pageId="4" pageNumber="205" pagination="261 - 276" refId="ref17781" refString="Moravec, J., Razanajaonarivalona, E. H. &amp; HORI, M. (2020) New or rare Madagascar tiger beetles- 19. Description of Pogonostoma (Pogonostoma) natsuae sp. nov. and elaborated redescription of P. (M.) meridionale Fleutiaux (Coleoptera: Cicindelidae). Zootaxa, 4790 (2), 261 - 276. https: // doi. org / 10.11646 / zootaxa. 4790.2.4" type="journal article" year="2020">Moravec, Razanajaonarivalona &amp; Hori 2020</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="CDE04D6FDA5BFFC2FCEDFB03FAF1FB3A" authority="Fleutiaux, 1899" authorityName="Fleutiaux" authorityYear="1899" box="[884,1283,1157,1184]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="meridionale" subGenus="Pogonostoma">
<emphasis id="3894EAFEDA5BFFC2FCEDFB03FC78FB3A" box="[884,906,1158,1183]" italics="true" pageId="4" pageNumber="205">P.</emphasis>
(
<emphasis id="3894EAFEDA5BFFC2FC03FB03FC42FB3A" box="[922,944,1158,1183]" italics="true" pageId="4" pageNumber="205">P.</emphasis>
)
<emphasis id="3894EAFEDA5BFFC2FC59FB00FBB5FB3A" box="[960,1095,1157,1183]" italics="true" pageId="4" pageNumber="205">meridionale</emphasis>
Fleutiaux, 1899
</taxonomicName>
. Both share a comparatively fine elytral punctation with the new species, but differ in having their posterior pronotal lobe glabrous and aedeagus apex of a different shape (see
<bibRefCitation id="6E714B1DDA5BFFC2FD52FB48FC31FB42" author="Moravec &amp; Razanajaonarivalona, E. H. &amp; HORI, M." box="[715,963,1229,1255]" pageId="4" pageNumber="205" pagination="261 - 276" refId="ref17781" refString="Moravec, J., Razanajaonarivalona, E. H. &amp; HORI, M. (2020) New or rare Madagascar tiger beetles- 19. Description of Pogonostoma (Pogonostoma) natsuae sp. nov. and elaborated redescription of P. (M.) meridionale Fleutiaux (Coleoptera: Cicindelidae). Zootaxa, 4790 (2), 261 - 276. https: // doi. org / 10.11646 / zootaxa. 4790.2.4" type="journal article" year="2020">
Moravec
<emphasis id="3894EAFEDA5BFFC2FCAFFB48FC95FB42" box="[822,871,1229,1255]" italics="true" pageId="4" pageNumber="205">et al</emphasis>
. (2020)
</bibRefCitation>
. In addition,
<taxonomicName id="CDE04D6FDA5BFFC2FBCEFB48FADBFB42" authorityName="Fleutiaux" authorityYear="1899" box="[1111,1321,1229,1255]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="meridionale" subGenus="Pogonostoma">
<emphasis id="3894EAFEDA5BFFC2FBCEFB48FB9FFB43" box="[1111,1133,1229,1254]" italics="true" pageId="4" pageNumber="205">P.</emphasis>
(
<emphasis id="3894EAFEDA5BFFC2FBE4FB48FB60FB43" box="[1149,1170,1229,1254]" italics="true" pageId="4" pageNumber="205">P.</emphasis>
)
<emphasis id="3894EAFEDA5BFFC2FB3BFB48FADBFB42" box="[1186,1321,1229,1255]" italics="true" pageId="4" pageNumber="205">meridionale</emphasis>
</taxonomicName>
possesses notably more irregular elytral punctation and conspicuously ferrugineous (rusty) setosity.
<taxonomicName id="CDE04D6FDA5BFFC2FB1DFB74FF3DFA8A" authority="Moravec, 2005" authorityName="Moravec" authorityYear="2005" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="rivalieri" subGenus="Pogonostoma">
<emphasis id="3894EAFEDA5BFFC2FB1DFB74FB68FAAF" box="[1156,1178,1265,1290]" italics="true" pageId="4" pageNumber="205">P.</emphasis>
(
<emphasis id="3894EAFEDA5BFFC2FB33FB74FB4DFAAF" box="[1194,1215,1265,1290]" italics="true" pageId="4" pageNumber="205">P.</emphasis>
)
<emphasis id="3894EAFEDA5BFFC2FB56FB75FAD9FAAF" box="[1231,1323,1264,1290]" italics="true" pageId="4" pageNumber="205">rivalieri</emphasis>
<bibRefCitation id="6E714B1DDA5BFFC2FAABFB74FF3DFA8A" author="MORAVEC, J." pageId="4" pageNumber="205" pagination="49 - 69" refId="ref17582" refString="MORAVEC, J. (2005) New or rare Madagascar tiger beetles- 10. Species closely related to Pogonostoma (Pogonostoma) alluaudi. (Coleoptera: Cicindelidae). Folia Heyrovskyana, Series A, 13, 49 - 69." type="journal article" year="2005">Moravec, 2005</bibRefCitation>
</taxonomicName>
possesses a similar (though rather straighter) aedeagus apex, but principally differs from the new species in having outer pronotal margins subparallel to parallel (particularly in male) and posterior pronotal lobe glabrous.
<taxonomicName id="CDE04D6FDA5BFFC2FF0EFAD9FDDBFAD3" authority="Moravec, 2005" authorityName="Moravec" authorityYear="2005" box="[151,553,1371,1398]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="praetervisum" subGenus="Pogonostoma">
<emphasis id="3894EAFEDA5BFFC2FF0EFAD9FF5FFAD0" box="[151,173,1372,1397]" italics="true" pageId="4" pageNumber="205">P.</emphasis>
(
<emphasis id="3894EAFEDA5BFFC2FF24FAD9FF21FAD0" box="[189,211,1372,1397]" italics="true" pageId="4" pageNumber="205">P.</emphasis>
)
<emphasis id="3894EAFEDA5BFFC2FF7AFAD9FE85FAD0" box="[227,375,1372,1397]" italics="true" pageId="4" pageNumber="205">praetervisum</emphasis>
<bibRefCitation id="6E714B1DDA5BFFC2FEE6FAD9FDDBFAD3" author="MORAVEC, J." box="[383,553,1371,1398]" pageId="4" pageNumber="205" pagination="49 - 69" refId="ref17582" refString="MORAVEC, J. (2005) New or rare Madagascar tiger beetles- 10. Species closely related to Pogonostoma (Pogonostoma) alluaudi. (Coleoptera: Cicindelidae). Folia Heyrovskyana, Series A, 13, 49 - 69." type="journal article" year="2005">Moravec, 2005</bibRefCitation>
</taxonomicName>
has its aedeagus (in its lateral view) somewhat similar to the new species, but clearly differs in having its elytral punctation much coarser (similar to that of
<taxonomicName id="CDE04D6FDA5BFFC2FC60FA05FB03FA3C" box="[1017,1265,1407,1434]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="atrorotundatum" subGenus="Pogonostoma">
<emphasis id="3894EAFEDA5BFFC2FC60FA05FBFCFA3C" box="[1017,1038,1408,1433]" italics="true" pageId="4" pageNumber="205">P.</emphasis>
(
<emphasis id="3894EAFEDA5BFFC2FB87FA05FBC1FA3C" box="[1054,1075,1408,1433]" italics="true" pageId="4" pageNumber="205">P.</emphasis>
)
<emphasis id="3894EAFEDA5BFFC2FBDBFAFAFB03FA3C" box="[1090,1265,1407,1433]" italics="true" pageId="4" pageNumber="205">atrorotundatum</emphasis>
</taxonomicName>
) and its pronotal disc covered with rather fine and shallow transverse rugae (see
<bibRefCitation id="6E714B1DDA5BFFC2FCE1FA21FBEFFA1B" author="MORAVEC, J." box="[888,1053,1443,1470]" pageId="4" pageNumber="205" pagination="49 - 69" refId="ref17582" refString="MORAVEC, J. (2005) New or rare Madagascar tiger beetles- 10. Species closely related to Pogonostoma (Pogonostoma) alluaudi. (Coleoptera: Cicindelidae). Folia Heyrovskyana, Series A, 13, 49 - 69." type="journal article" year="2005">Moravec 2005</bibRefCitation>
,
<bibRefCitation id="6E714B1DDA5BFFC2FBBEFA26FB96FA18" author="MORAVEC" box="[1063,1124,1443,1469]" pageId="4" pageNumber="205" refId="ref17625" refString="MORAVEC, J. (2007) Tiger beetles of Madagascar 1. A monograph of the genus Pogonostoma (Coleoptera: Cicindelidae). Nakladatelstvi Kabourek, Zlin, 499 pp." type="book" year="2007">2007</bibRefCitation>
). The leading representative of the
<taxonomicName id="CDE04D6FDA5BFFC2FF79FA4DFE7BFA44" authorityName="W. Horn" authorityYear="1898" box="[224,393,1479,1505]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="alluaudi" subGenus="Pogonostoma">
<emphasis id="3894EAFEDA5BFFC2FF79FA4DFF04FA44" box="[224,246,1480,1505]" italics="true" pageId="4" pageNumber="205">P.</emphasis>
(
<emphasis id="3894EAFEDA5BFFC2FE9FFA4DFEE9FA44" box="[262,283,1480,1505]" italics="true" pageId="4" pageNumber="205">P.</emphasis>
)
<emphasis id="3894EAFEDA5BFFC2FEB5FA42FE7BFA44" box="[300,393,1479,1505]" italics="true" pageId="4" pageNumber="205">alluaudi</emphasis>
</taxonomicName>
species-complex,
<taxonomicName id="CDE04D6FDA5BFFC2FDC2FA4DFC74FA44" authority="W. Horn, 1898" authorityName="W. Horn" authorityYear="1898" box="[603,902,1479,1505]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis id="3894EAFEDA5BFFC2FDC2FA4DFD26FA44" box="[603,724,1479,1505]" italics="true" pageId="4" pageNumber="205">P. alluaudi</emphasis>
W. Horn, 1898
</taxonomicName>
, differs from the new species in having coarser elytral punctation, and immediately in its black-blue to violaceous-blue body.
</paragraph>
<paragraph id="0A5F36ECDA5BFFC2FF5EF98BFE13F9D4" blockId="4.[151,1437,406,2042]" pageId="4" pageNumber="205">
The labrum possesses a rather similar shape (including usual variability) in all species of the
<taxonomicName id="CDE04D6FDA5BFFC2FB6EF98AFA6EF98D" authorityName="W. Horn" authorityYear="1898" box="[1271,1436,1550,1577]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="alluaudi" subGenus="Pogonostoma">
<emphasis id="3894EAFEDA5BFFC2FB6EF98AFAFEF98D" box="[1271,1292,1551,1576]" italics="true" pageId="4" pageNumber="205">P.</emphasis>
(
<emphasis id="3894EAFEDA5BFFC2FA82F98AFAC2F98D" box="[1307,1328,1551,1576]" italics="true" pageId="4" pageNumber="205">P.</emphasis>
)
<emphasis id="3894EAFEDA5BFFC2FAA6F98BFA6EF98D" box="[1343,1436,1550,1576]" italics="true" pageId="4" pageNumber="205">alluaudi</emphasis>
</taxonomicName>
species-complex, except for the number of anterior setae (but their number may vary in some specimens and the setae may be easily abraded).
</paragraph>
</subSubSection>
<subSubSection id="42FA6567DA5BFFCFFF5EF9FCFDB6FB04" lastPageId="9" lastPageNumber="210" pageId="4" pageNumber="205" type="description">
<paragraph id="0A5F36ECDA5BFFC2FF5EF9FCFDEBF912" blockId="4.[151,1437,406,2042]" pageId="4" pageNumber="205">
<emphasis id="3894EAFEDA5BFFC2FF5EF9FCFEA4F936" bold="true" box="[199,342,1657,1683]" pageId="4" pageNumber="205">Description</emphasis>
. Body (
<figureCitation id="92DB2A69DA5BFFC2FE2AF9FCFDE8F931" box="[435,538,1657,1684]" captionStart="FIGURES 1–10" captionStartId="5.[151,264,1867,1891]" captionTargetBox="[160,1427,185,1831]" captionTargetId="figure-17@5.[151,1435,181,1844]" captionTargetPageId="5" captionText="FIGURES 1–10. Pogonostoma (P.) ondravybirali sp. nov. (from type locality). 1–2—habitus: 1—♁, 10.5 mm, HT (SDEI); 2—♀, 10.9 mm, AT (CJVB); 3–4—mandibles: 3—♀, AT; 4—♁, HT; 5—galea and lacinia, ♁, PT (CCJM); 6–8—labrum: 6—♁, HT; 7—♁, PT (CJVB); 8—♀, AT; 9—antennal scape and pedicel, ♀, AT; 10—head, ♁, HT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4283532" httpUri="https://zenodo.org/record/4283532/files/figure.png" pageId="4" pageNumber="205">Figs 1–2</figureCitation>
) medium-sized, 9.50–11.0 (HT 10.5, AT 10.9) mm long, 2.40–2.90 (HT 2.65, AT 2.90) mm wide, pitchy-black.
</paragraph>
<paragraph id="0A5F36ECDA5BFFC2FF5EF944FE94F95A" blockId="4.[151,1437,406,2042]" pageId="4" pageNumber="205">
Head (
<figureCitation id="92DB2A69DA5BFFC2FE8BF944FEACF97E" box="[274,350,1729,1755]" captionStart="FIGURES 1–10" captionStartId="5.[151,264,1867,1891]" captionTargetBox="[160,1427,185,1831]" captionTargetId="figure-17@5.[151,1435,181,1844]" captionTargetPageId="5" captionText="FIGURES 1–10. Pogonostoma (P.) ondravybirali sp. nov. (from type locality). 1–2—habitus: 1—♁, 10.5 mm, HT (SDEI); 2—♀, 10.9 mm, AT (CJVB); 3–4—mandibles: 3—♀, AT; 4—♁, HT; 5—galea and lacinia, ♁, PT (CCJM); 6–8—labrum: 6—♁, HT; 7—♁, PT (CJVB); 8—♀, AT; 9—antennal scape and pedicel, ♀, AT; 10—head, ♁, HT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4283532" httpUri="https://zenodo.org/record/4283532/files/figure.png" pageId="4" pageNumber="205">Fig 10</figureCitation>
) notably narrower than body, width 2.00–
<quantity id="CD189B09DA5BFFC2FCA4F944FC57F97E" box="[829,933,1729,1756]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" pageId="4" pageNumber="205" unit="mm" value="2.15">2.15 mm</quantity>
; temples short (approximately 2.6–2.8 times shorter than eyes).
</paragraph>
<paragraph id="0A5F36ECDA5BFFC2FF5EF88CFBC5F8CF" blockId="4.[151,1437,406,2042]" pageId="4" pageNumber="205">Frons merging with clypeus in middle and not differentiated from vertex, flat or moderately convex; supraantennal keels sometimes merging with surface sculpture, usually consisting of moderately elevated posterior crest forming semi-ovoid apex of supraantennal plate; surface very irregularly scabrous.</paragraph>
<paragraph id="0A5F36ECDA5BFFC2FF5EF8F1FA77F85F" blockId="4.[151,1437,406,2042]" pageId="4" pageNumber="205">Vertex moderately convex in middle with faint posteromedian impression, surface rather coarsely and very ir-regularly scabriculous-cristulate, posterior and occipital area covered with somewhat finer, irregularly vermicular or wavy, short crests, passing to irregular and coarsely scabrous sculpture on temples; dorsal surface of head covered with erect, very short, whitish, or partly pale straw-yellow hairlike setae which are better obvious in lateral view.</paragraph>
<caption id="5E9F6664DA5AFFC3FF0EF8CEFB39F80E" ID-DOI="http://doi.org/10.5281/zenodo.4283532" ID-Zenodo-Dep="4283532" httpUri="https://zenodo.org/record/4283532/files/figure.png" pageId="5" pageNumber="206" startId="5.[151,264,1867,1891]" targetBox="[160,1427,185,1831]" targetPageId="5">
<paragraph id="0A5F36ECDA5AFFC3FF0EF8CEFB39F80E" blockId="5.[151,1436,1867,1966]" pageId="5" pageNumber="206">
<emphasis id="3894EAFEDA5AFFC3FF0EF8CEFEBBF8C1" bold="true" box="[151,329,1867,1892]" pageId="5" pageNumber="206">FIGURES 1–10.</emphasis>
<taxonomicName id="CDE04D6FDA5AFFC3FEC8F8C9FE2EF8C6" authorityName="Klug" authorityYear="1835" box="[337,476,1868,1891]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="genus">
<emphasis id="3894EAFEDA5AFFC3FEC8F8C9FE2EF8C6" box="[337,476,1868,1891]" italics="true" pageId="5" pageNumber="206">Pogonostoma</emphasis>
</taxonomicName>
<taxonomicName id="CDE04D6FDA5AFFC3FE7DF8C9FD6CF8C1" authority="Moravec &amp; Vybíral, 2020" authorityName="Moravec &amp; Vybíral" authorityYear="2020" box="[484,670,1868,1892]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="ondravybirali" status="sp. nov." subGenus="Pogonostoma">
(
<emphasis id="3894EAFEDA5AFFC3FE74F8C9FE0EF8C6" box="[493,508,1868,1891]" italics="true" pageId="5" pageNumber="206">P</emphasis>
.)
<emphasis id="3894EAFEDA5AFFC3FD8AF8C9FD6CF8C1" box="[531,670,1868,1892]" italics="true" pageId="5" pageNumber="206">ondravybirali</emphasis>
</taxonomicName>
<emphasis id="3894EAFEDA5AFFC3FD3FF8C9FD0BF8C1" bold="true" box="[678,761,1868,1892]" pageId="5" pageNumber="206">
<taxonomicNameLabel id="23A75785DA5AFFC3FD3FF8C9FD0BF8C1" box="[678,761,1868,1892]" pageId="5" pageNumber="206" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(from type locality). 1–2—habitus: 1—♁, 10.5 mm, HT (SDEI); 2—♀, 10.9 mm, AT (CJVB); 3–4—mandibles: 3—♀, AT; 4—♁, HT; 5—galea and lacinia, ♁, PT (CCJM); 6–8—labrum: 6—♁, HT; 7—♁, PT (CJVB); 8—♀, AT; 9—antennal scape and pedicel, ♀, AT; 10—head, ♁, HT. Bars = 1 mm.
</paragraph>
</caption>
<caption id="5E9F6664DA59FFC0FF0EF8B1FD32F831" ID-DOI="http://doi.org/10.5281/zenodo.4283534" ID-Zenodo-Dep="4283534" httpUri="https://zenodo.org/record/4283534/files/figure.png" pageId="6" pageNumber="207" startId="6.[151,264,1844,1868]" targetBox="[165,1425,193,1810]" targetPageId="6">
<paragraph id="0A5F36ECDA59FFC0FF0EF8B1FD32F831" blockId="6.[151,1436,1844,1941]" pageId="6" pageNumber="207">
<emphasis id="3894EAFEDA59FFC0FF0EF8B1FEA6F8E8" bold="true" box="[151,340,1844,1869]" pageId="6" pageNumber="207">FIGURES 11–16.</emphasis>
<taxonomicName id="CDE04D6FDA59FFC0FEC5F8B0FE15F8E9" authorityName="Klug" authorityYear="1835" box="[348,487,1845,1868]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="genus">
<emphasis id="3894EAFEDA59FFC0FEC5F8B0FE15F8E9" box="[348,487,1845,1868]" italics="true" pageId="6" pageNumber="207">Pogonostoma</emphasis>
</taxonomicName>
<taxonomicName id="CDE04D6FDA59FFC0FE76F8B0FD5AF8E8" authority="Moravec &amp; Vybíral, 2020" authorityName="Moravec &amp; Vybíral" authorityYear="2020" box="[495,680,1845,1869]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="ondravybirali" status="sp. nov." subGenus="Pogonostoma">
(
<emphasis id="3894EAFEDA59FFC0FE6EF8B0FDF4F8E9" box="[503,518,1845,1868]" italics="true" pageId="6" pageNumber="207">P</emphasis>
.)
<emphasis id="3894EAFEDA59FFC0FD84F8B0FD5AF8E8" box="[541,680,1845,1869]" italics="true" pageId="6" pageNumber="207">ondravybirali</emphasis>
</taxonomicName>
<emphasis id="3894EAFEDA59FFC0FD29F8B0FCF0F8E8" bold="true" box="[688,770,1845,1869]" pageId="6" pageNumber="207">
<taxonomicNameLabel id="23A75785DA59FFC0FD29F8B0FCF0F8E8" box="[688,770,1845,1869]" pageId="6" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(from type locality). 11–12—elytral apices: 11—♁, HT (SDEI); 12—♀, AT (CJVB); 13–14—elytron: 13—♁, HT; 14—♀, AT; 15—sculpture on elytral disc, ♁, HT; 16—ditto (higher magnifi- cation and on larger area of both elytra). Bars = 1 mm.
</paragraph>
</caption>
<caption id="5E9F6664DA58FFC1FF0EF8B4FCCBF8C9" ID-DOI="http://doi.org/10.5281/zenodo.4283536" ID-Zenodo-Dep="4283536" httpUri="https://zenodo.org/record/4283536/files/figure.png" pageId="7" pageNumber="208" startId="7.[151,264,1841,1865]" targetBox="[163,1423,183,1804]" targetPageId="7">
<paragraph id="0A5F36ECDA58FFC1FF0EF8B4FCCBF8C9" blockId="7.[151,1437,1841,1903]" pageId="7" pageNumber="208">
<emphasis id="3894EAFEDA58FFC1FF0EF8B4FEA4F8EC" bold="true" box="[151,342,1841,1865]" pageId="7" pageNumber="208">FIGURES 17–20.</emphasis>
<taxonomicName id="CDE04D6FDA58FFC1FEC4F8B7FE1AF8EC" authorityName="Klug" authorityYear="1835" box="[349,488,1842,1865]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="genus">
<emphasis id="3894EAFEDA58FFC1FEC4F8B7FE1AF8EC" box="[349,488,1842,1865]" italics="true" pageId="7" pageNumber="208">Pogonostoma</emphasis>
</taxonomicName>
<taxonomicName id="CDE04D6FDA58FFC1FE68F8B4FD58F8EC" authority="Moravec &amp; Vybíral, 2020" authorityName="Moravec &amp; Vybíral" authorityYear="2020" box="[497,682,1841,1865]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="ondravybirali" status="sp. nov." subGenus="Pogonostoma">
(
<emphasis id="3894EAFEDA58FFC1FE60F8B7FDFAF8EC" box="[505,520,1842,1865]" italics="true" pageId="7" pageNumber="208">P</emphasis>
.)
<emphasis id="3894EAFEDA58FFC1FD86F8B4FD58F8EC" box="[543,682,1841,1865]" italics="true" pageId="7" pageNumber="208">ondravybirali</emphasis>
</taxonomicName>
<emphasis id="3894EAFEDA58FFC1FD2AF8B4FCF7F8EC" bold="true" box="[691,773,1841,1865]" pageId="7" pageNumber="208">
<taxonomicNameLabel id="23A75785DA58FFC1FD2AF8B4FCF7F8EC" box="[691,773,1841,1865]" pageId="7" pageNumber="208" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(from type locality), pronotum. 17—♁, HT (SDEI); 18—♀, AT (CJVB); 19—♁, PT (CJVB) 20—ditto, lateral view. Bars = 1 mm.
</paragraph>
</caption>
<paragraph id="0A5F36ECDA58FFC1FF5EF81EFBA3F87F" blockId="7.[151,1437,1947,2010]" pageId="7" pageNumber="208">Genae rather finely and shallowly striate, striae coarser and more irregular on postgenal and juxtaorbital area while basal area almost smooth and with few semierect, easily abraded whitish setae.</paragraph>
<paragraph id="0A5F36ECDA57FFCEFF5EFF12FEE4FF70" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="209">Clypeus pitchy-black, rough coriaceous to irregularly scabriculous, covered with scattered whitish, mostly short setae.</paragraph>
<paragraph id="0A5F36ECDA57FFCEFF5EFF5AFAB9FE50" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="209">
Labrum primarily 8-setose, rarely 7-setose (with 5–6 anterior and 2 lateral, ochre to ochre-brown setae); surface black, with only moderate central convexity, rough-coriaceous, glabrous (except for occasional setae passing from clypeus); lateral margins moderately arcuate towards indistinct lateral indentations with deep setigerous puncture, sexually dimorphic; male labrum (
<figureCitation id="92DB2A69DA57FFCEFD8DFECEFD85FEC3" box="[532,631,331,358]" captionStart="FIGURES 1–10" captionStartId="5.[151,264,1867,1891]" captionTargetBox="[160,1427,185,1831]" captionTargetId="figure-17@5.[151,1435,181,1844]" captionTargetPageId="5" captionText="FIGURES 1–10. Pogonostoma (P.) ondravybirali sp. nov. (from type locality). 1–2—habitus: 1—♁, 10.5 mm, HT (SDEI); 2—♀, 10.9 mm, AT (CJVB); 3–4—mandibles: 3—♀, AT; 4—♁, HT; 5—galea and lacinia, ♁, PT (CCJM); 6–8—labrum: 6—♁, HT; 7—♁, PT (CJVB); 8—♀, AT; 9—antennal scape and pedicel, ♀, AT; 10—head, ♁, HT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4283532" httpUri="https://zenodo.org/record/4283532/files/figure.png" pageId="8" pageNumber="209">Figs 6–7</figureCitation>
) short,
<quantity id="CD189B09DA57FFCEFD50FECFFC99FEC0" box="[713,875,330,357]" metricMagnitude="-4" metricUnit="m" metricValue="6.25" metricValueMax="6.5" metricValueMin="6.0" pageId="8" pageNumber="209" unit="mm" value="0.625" valueMax="0.65" valueMin="0.6">0.60–0.65 mm</quantity>
long,
<quantity id="CD189B09DA57FFCEFC36FECFFBA3FEC0" box="[943,1105,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.25" metricValueMin="1.05" pageId="8" pageNumber="209" unit="mm" value="1.15" valueMax="1.25" valueMin="1.05">1.05–1.25 mm</quantity>
wide, anterolateral teeth variably rounded or subacute or else flattened, anterior margin truncate or shallowly emarginate, irregularly dentate, the teeth usually partly or entirely effaced; female labrum (
<figureCitation id="92DB2A69DA57FFCEFC91FE16FCB9FE08" box="[776,843,403,429]" captionStart="FIGURES 1–10" captionStartId="5.[151,264,1867,1891]" captionTargetBox="[160,1427,185,1831]" captionTargetId="figure-17@5.[151,1435,181,1844]" captionTargetPageId="5" captionText="FIGURES 1–10. Pogonostoma (P.) ondravybirali sp. nov. (from type locality). 1–2—habitus: 1—♁, 10.5 mm, HT (SDEI); 2—♀, 10.9 mm, AT (CJVB); 3–4—mandibles: 3—♀, AT; 4—♁, HT; 5—galea and lacinia, ♁, PT (CCJM); 6–8—labrum: 6—♁, HT; 7—♁, PT (CJVB); 8—♀, AT; 9—antennal scape and pedicel, ♀, AT; 10—head, ♁, HT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4283532" httpUri="https://zenodo.org/record/4283532/files/figure.png" pageId="8" pageNumber="209">Fig. 8</figureCitation>
) notably prolonged anteriad, mostly almost semicircular due to almost arcuate anterolateral margins towards small almost right-angled subacute anterolateral teeth and rather narrow, truncate and irregularly dentate anterior margin; length
<quantity id="CD189B09DA57FFCEFC30FE5FFBBCFE50" box="[937,1102,474,501]" metricMagnitude="-4" metricUnit="m" metricValue="7.75" metricValueMax="8.0" metricValueMin="7.5" pageId="8" pageNumber="209" unit="mm" value="0.775" valueMax="0.8" valueMin="0.75">0.75–0.80 mm</quantity>
, width
<quantity id="CD189B09DA57FFCEFB38FE5EFAB5FE50" box="[1185,1351,475,501]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="8" pageNumber="209" unit="mm" value="1.2" valueMax="1.3" valueMin="1.1">1.10–1.30 mm</quantity>
.
</paragraph>
<paragraph id="0A5F36ECDA57FFCEFF5EFE7AFDE0FD20" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="209">
Maxillae (
<figureCitation id="92DB2A69DA57FFCEFEA5FE7AFE73FDBC" box="[316,385,510,537]" captionStart="FIGURES 1–10" captionStartId="5.[151,264,1867,1891]" captionTargetBox="[160,1427,185,1831]" captionTargetId="figure-17@5.[151,1435,181,1844]" captionTargetPageId="5" captionText="FIGURES 1–10. Pogonostoma (P.) ondravybirali sp. nov. (from type locality). 1–2—habitus: 1—♁, 10.5 mm, HT (SDEI); 2—♀, 10.9 mm, AT (CJVB); 3–4—mandibles: 3—♀, AT; 4—♁, HT; 5—galea and lacinia, ♁, PT (CCJM); 6–8—labrum: 6—♁, HT; 7—♁, PT (CJVB); 8—♀, AT; 9—antennal scape and pedicel, ♀, AT; 10—head, ♁, HT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4283532" httpUri="https://zenodo.org/record/4283532/files/figure.png" pageId="8" pageNumber="209">Fig. 5</figureCitation>
): galea black, usually with depigmented median orifice and tip of basal (longest) galeomere and with brownish tip of the terminal palpomere; lacinia with apical portion moderately to more distinctly spatulatedilated,
<quantity id="CD189B09DA57FFCEFF69FDC2FE61FDC4" box="[240,403,583,609]" metricMagnitude="-4" metricUnit="m" metricValue="3.15" metricValueMax="3.3" metricValueMin="3.0" pageId="8" pageNumber="209" unit="mm" value="0.315" valueMax="0.33" valueMin="0.3">0.30–0.33 mm</quantity>
wide, black with brownish, rarely reddish-brown faint tinge and with brownish or rusty setae mixed with whitish hairlike setae.
</paragraph>
<paragraph id="0A5F36ECDA57FFCEFF5EFD0AFD63FCB0" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="209">
Palpi (
<figureCitation id="92DB2A69DA57FFCEFE88FD0AFE94FD0C" box="[273,358,655,681]" captionStart="FIGURES 1–10" captionStartId="5.[151,264,1867,1891]" captionTargetBox="[160,1427,185,1831]" captionTargetId="figure-17@5.[151,1435,181,1844]" captionTargetPageId="5" captionText="FIGURES 1–10. Pogonostoma (P.) ondravybirali sp. nov. (from type locality). 1–2—habitus: 1—♁, 10.5 mm, HT (SDEI); 2—♀, 10.9 mm, AT (CJVB); 3–4—mandibles: 3—♀, AT; 4—♁, HT; 5—galea and lacinia, ♁, PT (CCJM); 6–8—labrum: 6—♁, HT; 7—♁, PT (CJVB); 8—♀, AT; 9—antennal scape and pedicel, ♀, AT; 10—head, ♁, HT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4283532" httpUri="https://zenodo.org/record/4283532/files/figure.png" pageId="8" pageNumber="209">Fig. 10</figureCitation>
). Both maxillary and labial palpi elongate (of usual length and shape), black, except for cinnamon-brown apical half of terminal palpomeres, with long, black-brown setae with mahogany to violet tinge; surface of maxillary palpi covered with scattered, short whitish or greyish microtrichia; longest palpomeres of labial palpi with bumpy surface and depigmented apices.
</paragraph>
<paragraph id="0A5F36ECDA57FFCEFF5EFC9AFE29FC6C" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="209">
Mandibles (
<figureCitation id="92DB2A69DA57FFCEFED5FC9AFE53FC9C" box="[332,417,799,825]" captionStart="FIGURES 30–40" captionStartId="13.[151,264,1852,1876]" captionTargetBox="[161,1427,189,1819]" captionTargetId="figure-17@13.[151,1435,181,1828]" captionTargetPageId="13" captionText="FIGURES 30–40. Pogonostoma (P.) atrorotundatum W. Horn. 30–31—habitus: 30—♁, 9.9 mm, “C. W. Madagascar”, LT (SDEI); 31—♁, 9.7 mm, Bongolava, HT of syn. P. (P.) rufidens Rivalier (MNHN); 32—mandibles and labrum, ♁, Antsalova (CCJM ex APCA); 33—mandibles, ♀, ibid.; 34—galea and lacinia, ♁, ibid.; 35—lacinia, ♁, “C. W. Madagascar”, PLT (SDEI); 36–40—labrum: 36—♁, LT; 37—♁, “C. W. Madagascar”, PLT (SDEI); 38—♁, Antsalova (CCJM); 39—♁, HT of syn. P. (P.) rufidens; 40—♀, Miandrivazo, PLT (MNHN). Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4283540" httpUri="https://zenodo.org/record/4283540/files/figure.png" pageId="8" pageNumber="209">Figs 34</figureCitation>
) mahogany-brown, teeth usually paler, with rather short terminal teeth; sexually dimorphic in shape: male mandibles (
<figureCitation id="92DB2A69DA57FFCEFE51FCC6FDFFFCF8" box="[456,525,835,861]" captionStart="FIGURES 1–10" captionStartId="5.[151,264,1867,1891]" captionTargetBox="[160,1427,185,1831]" captionTargetId="figure-17@5.[151,1435,181,1844]" captionTargetPageId="5" captionText="FIGURES 1–10. Pogonostoma (P.) ondravybirali sp. nov. (from type locality). 1–2—habitus: 1—♁, 10.5 mm, HT (SDEI); 2—♀, 10.9 mm, AT (CJVB); 3–4—mandibles: 3—♀, AT; 4—♁, HT; 5—galea and lacinia, ♁, PT (CCJM); 6–8—labrum: 6—♁, HT; 7—♁, PT (CJVB); 8—♀, AT; 9—antennal scape and pedicel, ♀, AT; 10—head, ♁, HT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4283532" httpUri="https://zenodo.org/record/4283532/files/figure.png" pageId="8" pageNumber="209">Fig. 4</figureCitation>
) subsymmetrical (apart from the universally longer terminal tooth in right mandible) with second tooth in both mandibles notably smaller than third tooth; female mandibles (
<figureCitation id="92DB2A69DA57FFCEFB4DFCE2FAE5FC24" box="[1236,1303,871,897]" captionStart="FIGURES 1–10" captionStartId="5.[151,264,1867,1891]" captionTargetBox="[160,1427,185,1831]" captionTargetId="figure-17@5.[151,1435,181,1844]" captionTargetPageId="5" captionText="FIGURES 1–10. Pogonostoma (P.) ondravybirali sp. nov. (from type locality). 1–2—habitus: 1—♁, 10.5 mm, HT (SDEI); 2—♀, 10.9 mm, AT (CJVB); 3–4—mandibles: 3—♀, AT; 4—♁, HT; 5—galea and lacinia, ♁, PT (CCJM); 6–8—labrum: 6—♁, HT; 7—♁, PT (CJVB); 8—♀, AT; 9—antennal scape and pedicel, ♀, AT; 10—head, ♁, HT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4283532" httpUri="https://zenodo.org/record/4283532/files/figure.png" pageId="8" pageNumber="209">Fig. 3</figureCitation>
) with inner teeth of right mandible longer and almost of the same size; convex basolateral portions black-brown with short, scattered greyish-white setae
</paragraph>
<paragraph id="0A5F36ECDA57FFCEFF5EFC56FB30FB90" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="209">
Antennae (
<figureCitation id="92DB2A69DA57FFCEFEDBFC56FE20FC48" box="[322,466,979,1005]" captionStart="FIGURES 1–10" captionStartId="5.[151,264,1867,1891]" captionTargetBox="[160,1427,185,1831]" captionTargetId="figure-17@5.[151,1435,181,1844]" captionTargetPageId="5" captionText="FIGURES 1–10. Pogonostoma (P.) ondravybirali sp. nov. (from type locality). 1–2—habitus: 1—♁, 10.5 mm, HT (SDEI); 2—♀, 10.9 mm, AT (CJVB); 3–4—mandibles: 3—♀, AT; 4—♁, HT; 5—galea and lacinia, ♁, PT (CCJM); 6–8—labrum: 6—♁, HT; 7—♁, PT (CJVB); 8—♀, AT; 9—antennal scape and pedicel, ♀, AT; 10—head, ♁, HT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4283532" httpUri="https://zenodo.org/record/4283532/files/figure.png" pageId="8" pageNumber="209">Figs 1–2, 10</figureCitation>
) black, in male as long as body or slightly longer, in female somewhat shorter; scape with one or two long, yellow-brown subapical (easily abraded) setae, surface covered with indistinct whitish microsetae; antennomeres 2–4 with scattered whitish microsetae, 5–11 with usual micro-pubescence.
</paragraph>
<paragraph id="0A5F36ECDA57FFCEFF5EFBBAFEFBF944" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="209">
Thorax: pronotum (
<figureCitation id="92DB2A69DA57FFCEFE3EFBBAFDDBFBFC" box="[423,553,1087,1113]" captionStart="FIGURES 17–20" captionStartId="7.[151,264,1841,1865]" captionTargetBox="[163,1423,183,1804]" captionTargetId="figure-17@7.[152,1435,181,1817]" captionTargetPageId="7" captionText="FIGURES 17–20. Pogonostoma (P.) ondravybirali sp. nov. (from type locality), pronotum. 17—♁, HT (SDEI); 18—♀, AT (CJVB); 19—♁, PT (CJVB) 20—ditto, lateral view. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4283536" httpUri="https://zenodo.org/record/4283536/files/figure.png" pageId="8" pageNumber="209">Figs 17–20</figureCitation>
) elongate,
<quantity id="CD189B09DA57FFCEFD30FBBBFCBCFBFC" box="[681,846,1086,1113]" metricMagnitude="-3" metricUnit="m" metricValue="2.525" metricValueMax="2.7" metricValueMin="2.35" pageId="8" pageNumber="209" unit="mm" value="2.525" valueMax="2.7" valueMin="2.35">2.35–2.70 mm</quantity>
long,
<quantity id="CD189B09DA57FFCEFC00FBBBFBCCFBFC" box="[921,1086,1086,1113]" metricMagnitude="-3" metricUnit="m" metricValue="1.675" metricValueMax="1.8" metricValueMin="1.55" pageId="8" pageNumber="209" unit="mm" value="1.675" valueMax="1.8" valueMin="1.55">1.55–1.80 mm</quantity>
wide; anterior lobe slightly or more notably narrower than posterior lobe, its surface rather coarsely and irregularly scabrous-rugulose with scattered umbilicate tubercles bearing whitish hairlike setae; pronotal disc in male (
<figureCitation id="92DB2A69DA57FFCEFB83FB02FB69FB04" box="[1050,1179,1159,1185]" captionStart="FIGURES 17–20" captionStartId="7.[151,264,1841,1865]" captionTargetBox="[163,1423,183,1804]" captionTargetId="figure-17@7.[152,1435,181,1817]" captionTargetPageId="7" captionText="FIGURES 17–20. Pogonostoma (P.) ondravybirali sp. nov. (from type locality), pronotum. 17—♁, HT (SDEI); 18—♀, AT (CJVB); 19—♁, PT (CJVB) 20—ditto, lateral view. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4283536" httpUri="https://zenodo.org/record/4283536/files/figure.png" pageId="8" pageNumber="209">Figs 17, 19</figureCitation>
) with moderately convex, lateral margins (ellipsoidal); in lateral view subgibbose (
<figureCitation id="92DB2A69DA57FFCEFCD6FB2EFC53FB60" box="[847,929,1195,1221]" captionStart="FIGURES 17–20" captionStartId="7.[151,264,1841,1865]" captionTargetBox="[163,1423,183,1804]" captionTargetId="figure-17@7.[152,1435,181,1817]" captionTargetPageId="7" captionText="FIGURES 17–20. Pogonostoma (P.) ondravybirali sp. nov. (from type locality), pronotum. 17—♁, HT (SDEI); 18—♀, AT (CJVB); 19—♁, PT (CJVB) 20—ditto, lateral view. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4283536" httpUri="https://zenodo.org/record/4283536/files/figure.png" pageId="8" pageNumber="209">Fig. 20</figureCitation>
) in female subglobose (
<figureCitation id="92DB2A69DA57FFCEFB2BFB2EFAF4FB60" box="[1202,1286,1195,1221]" captionStart="FIGURES 17–20" captionStartId="7.[151,264,1841,1865]" captionTargetBox="[163,1423,183,1804]" captionTargetId="figure-17@7.[152,1435,181,1817]" captionTargetPageId="7" captionText="FIGURES 17–20. Pogonostoma (P.) ondravybirali sp. nov. (from type locality), pronotum. 17—♁, HT (SDEI); 18—♀, AT (CJVB); 19—♁, PT (CJVB) 20—ditto, lateral view. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4283536" httpUri="https://zenodo.org/record/4283536/files/figure.png" pageId="8" pageNumber="209">Fig. 18</figureCitation>
); notopleural sutures invisible from above; median line indistinct, partly merging with irregular surface sculpture consisting of rather coarse tubercles which are mostly isolated and umbilicate, particularly on lateral areas, while those on median area are occasionally or more commonly transversely connected into very short, transverse crests; shiny posteromedian area adjacent to posterior lobe covered with fine and shallow, mostly transverse stria-like rugae; whole discal surface covered with scattered, short, decumbent or erect, whitish to pale straw-yellow hairlike setae mostly arising from setigerous pits of the umbilicate tubercles (setae easily abraded and usually barely obvious on median area in dorsal view, but better recognizable in lateral view (
<figureCitation id="92DB2A69DA57FFCEFD40FA22FCDFFA64" box="[729,813,1447,1473]" captionStart="FIGURES 17–20" captionStartId="7.[151,264,1841,1865]" captionTargetBox="[163,1423,183,1804]" captionTargetId="figure-17@7.[152,1435,181,1817]" captionTargetPageId="7" captionText="FIGURES 17–20. Pogonostoma (P.) ondravybirali sp. nov. (from type locality), pronotum. 17—♁, HT (SDEI); 18—♀, AT (CJVB); 19—♁, PT (CJVB) 20—ditto, lateral view. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4283536" httpUri="https://zenodo.org/record/4283536/files/figure.png" pageId="8" pageNumber="209">Fig. 20</figureCitation>
); posterior lobe shiny, sparsely irregularly wrinkled and sparsely or rather densely punctate-setose with whitish setae which may appear blackened, particularly on lateral areas; proepisterna (
<figureCitation id="92DB2A69DA57FFCEFEE7FA6AFE23F9AC" box="[382,465,1519,1545]" captionStart="FIGURES 17–20" captionStartId="7.[151,264,1841,1865]" captionTargetBox="[163,1423,183,1804]" captionTargetId="figure-17@7.[152,1435,181,1817]" captionTargetPageId="7" captionText="FIGURES 17–20. Pogonostoma (P.) ondravybirali sp. nov. (from type locality), pronotum. 17—♁, HT (SDEI); 18—♀, AT (CJVB); 19—♁, PT (CJVB) 20—ditto, lateral view. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4283536" httpUri="https://zenodo.org/record/4283536/files/figure.png" pageId="8" pageNumber="209">Fig. 20</figureCitation>
) large, almost smooth and shiny, with only sparse and fine setigerous punctures and short parallel-wrinkles on juxtanotopleural area; mesepisterna shiny, with occasional setae on ventral area adjacent to mesosternum; mesepimeron deeply impressed; metepisterna notably long, deeply longitudinally impressed, smooth, with very sparse, barely visible whitish microtrichia; prosternum smooth, except for very fine transverse wrinkles, together with mesosternum rather densely covered with long and erect, whitish hairlike setae arising from barely visible setigerous micropunctures; metasternum almost smooth, indistinctly sparsely whitish setose, glabrous in middle.
</paragraph>
<paragraph id="0A5F36ECDA57FFCFFF5EF96EFBB7FE74" blockId="8.[151,1437,151,2013]" lastBlockId="9.[151,1437,151,1185]" lastPageId="9" lastPageNumber="210" pageId="8" pageNumber="209">
Elytra (
<figureCitation id="92DB2A69DA57FFCEFE83F96EFE6BF8A3" box="[282,409,1771,1798]" captionStart="FIGURES 11–16" captionStartId="6.[151,264,1844,1868]" captionTargetBox="[165,1425,193,1810]" captionTargetId="figure-20@6.[151,1435,181,1821]" captionTargetPageId="6" captionText="FIGURES 11–16. Pogonostoma (P.) ondravybirali sp. nov. (from type locality). 11–12—elytral apices: 11—♁, HT (SDEI); 12—♀, AT (CJVB); 13–14—elytron: 13—♁, HT; 14—♀, AT; 15—sculpture on elytral disc, ♁, HT; 16—ditto (higher magnifi- cation and on larger area of both elytra). Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4283534" httpUri="https://zenodo.org/record/4283534/files/figure.png" pageId="8" pageNumber="209">Figs 11–16</figureCitation>
) elongate,
<quantity id="CD189B09DA57FFCEFD8DF96FFD45F8A0" box="[532,695,1770,1797]" metricMagnitude="-3" metricUnit="m" metricValue="5.95" metricValueMax="6.4" metricValueMin="5.5" pageId="8" pageNumber="209" unit="mm" value="5.95" valueMax="6.4" valueMin="5.5">5.50–6.40 mm</quantity>
long, outer margins of elytral base obliquely sloped towards arcu-ate-rounded humeri; lateral margins almost parallel, with only slightly arched bulge in anterior third of the margin in male, and only indistinctly enlarged towards rounded lateral anteapical angles; elytral apices sexually dimorphic: male apex (
<figureCitation id="92DB2A69DA57FFCEFE87F8D2FE50F8D4" box="[286,418,1879,1905]" captionStart="FIGURES 11–16" captionStartId="6.[151,264,1844,1868]" captionTargetBox="[165,1425,193,1810]" captionTargetId="figure-20@6.[151,1435,181,1821]" captionTargetPageId="6" captionText="FIGURES 11–16. Pogonostoma (P.) ondravybirali sp. nov. (from type locality). 11–12—elytral apices: 11—♁, HT (SDEI); 12—♀, AT (CJVB); 13–14—elytron: 13—♁, HT; 14—♀, AT; 15—sculpture on elytral disc, ♁, HT; 16—ditto (higher magnifi- cation and on larger area of both elytra). Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4283534" httpUri="https://zenodo.org/record/4283534/files/figure.png" pageId="8" pageNumber="209">Figs 11, 13</figureCitation>
) with mostly acute, almost thorn-like outer tooth, then faintly obliquely sloped towards small, right-angled inner tooth, and with faint sutural emargination towards very small or indistinct sutural spine; female apex (
<figureCitation id="92DB2A69DA57FFCEFEA8F81AFE46F81C" box="[305,436,1951,1977]" captionStart="FIGURES 11–16" captionStartId="6.[151,264,1844,1868]" captionTargetBox="[165,1425,193,1810]" captionTargetId="figure-20@6.[151,1435,181,1821]" captionTargetPageId="6" captionText="FIGURES 11–16. Pogonostoma (P.) ondravybirali sp. nov. (from type locality). 11–12—elytral apices: 11—♁, HT (SDEI); 12—♀, AT (CJVB); 13–14—elytron: 13—♁, HT; 14—♀, AT; 15—sculpture on elytral disc, ♁, HT; 16—ditto (higher magnifi- cation and on larger area of both elytra). Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4283534" httpUri="https://zenodo.org/record/4283534/files/figure.png" pageId="8" pageNumber="209">Figs 12, 14</figureCitation>
) with blunter and right-angled outer tooth and with right-angled but notably large inner tooth and deep (but narrow) excision towards indistinct sutural spine; elytral surface convex with moderate basodis-cal convexity delineated by deep (mutually V-shaped) discal impression, punctate throughout, except for effaced narrow basal and anterior basohumeral area; punctures larger and deep on two elytral thirds, large and mostly isolated also on lateral areas, deepest, largest and irregularly anastomosing by lowered lateral intervals on basodiscal convexity and median part of elytral disc, but reduced or partly (never entirely) effaced within the deep discal impression; limited area along sutures is covered with smaller and very irregular punctures; posterior declivity and whole anteapical area covered with smaller and more spaced punctures of a carinate shape, partly with almost aciculate posterior margins of intervals; surface of intervals shiny; setal vesture consisting of nearly erect, copious, moderately long, pale to straw-yellow ornamental setae which are mixed with very sparse, long and erect, whitish hairlike sensory setae, which are longest and numerous on lateral portion of humeri.
</paragraph>
<paragraph id="0A5F36ECDA56FFCFFF5EFE5EFE6DFDBC" blockId="9.[151,1437,151,1185]" pageId="9" pageNumber="210">Abdomen. Ventrites pitchy-black, covered with scattered, short and mostly appressed whitish setae; female ventrite 8 usually paler.</paragraph>
<paragraph id="0A5F36ECDA56FFCFFF5EFDA6FBCDFCB0" blockId="9.[151,1437,151,1185]" pageId="9" pageNumber="210">Legs pitchy-black; coxae rather densely whitish-setose; profemora covered with rather dense medium-long whitish and darker setae; mesofemora with dense whitish setae on their dorsal surface, much sparser setae on ventral area; metafemora with sparser, brownish and black mostly stiffer setae; protibiae with dense whitish appressed setae and a few longer erect brown setae on ventral area; mesotibiae with blackened setae except for dark ferrugineous, dense setae on their apical area; metatibiae densely covered with mostly black setae; protarsi covered with whitish and darkened setae (and as in all species, first three protarsomeres in male dilated and with dense setose pad); meso-and metatarsi covered with brown and blackened setae; claws rusty or rusty-brown.</paragraph>
<paragraph id="0A5F36ECDA56FFCFFF5EFC9AFD27FC00" blockId="9.[151,1437,151,1185]" pageId="9" pageNumber="210">
Aedeagus of almost uniform size,
<quantity id="CD189B09DA56FFCFFDCAFC9BFD0AFC9C" box="[595,760,798,825]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.6" metricValueMin="2.5" pageId="9" pageNumber="210" unit="mm" value="2.55" valueMax="2.6" valueMin="2.5">2.50–2.60 mm</quantity>
long,
<quantity id="CD189B09DA56FFCFFCDDFC9AFC58FC9C" box="[836,938,799,825]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="210" unit="mm" value="0.4">0.40 mm</quantity>
wide, in its left lateral aspect (
<figureCitation id="92DB2A69DA56FFCFFA8BFC9AFF12FCF8" captionStart="FIGURES 21–29" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[163,1424,1213,1911]" captionTargetId="figure-484@9.[158,1429,1209,1919]" captionTargetPageId="9" captionText="FIGURES 21–29. Pogonostoma (P.) ondravybirali sp. nov. (from type locality), male aedeagus or its apex. 21—HT (SDEI); 22—ditto, ventral view; 23—PT (CJVB); 24—ditto, dorsal view; 25—ditto, ventral view; 26—PT (CCJM); 27—ditto, ventral view; 28—PT (CJVB); 29—ditto, ventral view. Bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4283538" httpUri="https://zenodo.org/record/4283538/files/figure.png" pageId="9" pageNumber="210">Figs 21, 23, 26, 28</figureCitation>
) rather distinctly bent in middle, apical portion conically tapered towards rather small, rounded, only very slightly dorsad-bent apex; the aedeagus in its ventral (and dorsal) aspect (
<figureCitation id="92DB2A69DA56FFCFFC5BFCE2FB49FC24" box="[962,1211,870,897]" captionStart="FIGURES 21–29" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[163,1424,1213,1911]" captionTargetId="figure-484@9.[158,1429,1209,1919]" captionTargetPageId="9" captionText="FIGURES 21–29. Pogonostoma (P.) ondravybirali sp. nov. (from type locality), male aedeagus or its apex. 21—HT (SDEI); 22—ditto, ventral view; 23—PT (CJVB); 24—ditto, dorsal view; 25—ditto, ventral view; 26—PT (CCJM); 27—ditto, ventral view; 28—PT (CJVB); 29—ditto, ventral view. Bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4283538" httpUri="https://zenodo.org/record/4283538/files/figure.png" pageId="9" pageNumber="210">Figs 22, 24, 25, 27, 29</figureCitation>
) is conically attenuated and narrowed into moderately elongated apex.
</paragraph>
<paragraph id="0A5F36ECDA56FFCFFF5EFC2BFDB6FB04" blockId="9.[151,1437,151,1185]" pageId="9" pageNumber="210">
<emphasis id="3894EAFEDA56FFCFFF5EFC2BFEA0FC6D" bold="true" box="[199,338,942,968]" pageId="9" pageNumber="210">Variability.</emphasis>
Apart from the somewhat variable shape of the labrum (particularly in male as obvious in
<figureCitation id="92DB2A69DA56FFCFFAF2FC2AFF34FC4B" captionStart="FIGURES 1–10" captionStartId="5.[151,264,1867,1891]" captionTargetBox="[160,1427,185,1831]" captionTargetId="figure-17@5.[151,1435,181,1844]" captionTargetPageId="5" captionText="FIGURES 1–10. Pogonostoma (P.) ondravybirali sp. nov. (from type locality). 1–2—habitus: 1—♁, 10.5 mm, HT (SDEI); 2—♀, 10.9 mm, AT (CJVB); 3–4—mandibles: 3—♀, AT; 4—♁, HT; 5—galea and lacinia, ♁, PT (CCJM); 6–8—labrum: 6—♁, HT; 7—♁, PT (CJVB); 8—♀, AT; 9—antennal scape and pedicel, ♀, AT; 10—head, ♁, HT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4283532" httpUri="https://zenodo.org/record/4283532/files/figure.png" pageId="9" pageNumber="210">Figs 6–7</figureCitation>
), the size of the elytral punctures in the new species may slightly vary (but the punctation never consist of so extremely large punctures as in
<taxonomicName id="CDE04D6FDA56FFCFFDACFC72FCC4FBB4" box="[565,822,1015,1041]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="atrorotundatum" subGenus="Pogonostoma">
<emphasis id="3894EAFEDA56FFCFFDACFC72FDB4FBB5" box="[565,582,1015,1040]" italics="true" pageId="9" pageNumber="210">P</emphasis>
. (
<emphasis id="3894EAFEDA56FFCFFDF9FC72FD87FBB5" box="[608,629,1015,1040]" italics="true" pageId="9" pageNumber="210">P.</emphasis>
)
<emphasis id="3894EAFEDA56FFCFFD1EFC72FCC4FBB4" box="[647,822,1015,1041]" italics="true" pageId="9" pageNumber="210">atrorotundatum</emphasis>
</taxonomicName>
,
<taxonomicName id="CDE04D6FDA56FFCFFCDFFC72FBD9FBB5" authorityName="Moravec" authorityYear="2005" box="[838,1067,1015,1041]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="praetervisum" subGenus="Pogonostoma">
<emphasis id="3894EAFEDA56FFCFFCDFFC72FCA5FBB5" box="[838,855,1015,1040]" italics="true" pageId="9" pageNumber="210">P</emphasis>
. (
<emphasis id="3894EAFEDA56FFCFFCE9FC72FC74FBB5" box="[880,902,1015,1040]" italics="true" pageId="9" pageNumber="210">P.</emphasis>
)
<emphasis id="3894EAFEDA56FFCFFC0EFC72FBD9FBB5" box="[919,1067,1015,1040]" italics="true" pageId="9" pageNumber="210">praetervisum</emphasis>
</taxonomicName>
, and
<taxonomicName id="CDE04D6FDA56FFCFFBF5FC72FAE9FBB4" authorityName="W. Horn" authorityYear="1898" box="[1132,1307,1015,1041]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="alluaudi" subGenus="Pogonostoma">
<emphasis id="3894EAFEDA56FFCFFBF5FC72FB8FFBB5" box="[1132,1149,1015,1040]" italics="true" pageId="9" pageNumber="210">P</emphasis>
. (
<emphasis id="3894EAFEDA56FFCFFB0EFC72FB5EFBB5" box="[1175,1196,1015,1040]" italics="true" pageId="9" pageNumber="210">P.</emphasis>
)
<emphasis id="3894EAFEDA56FFCFFB27FC72FAE9FBB4" box="[1214,1307,1015,1041]" italics="true" pageId="9" pageNumber="210">alluaudi</emphasis>
</taxonomicName>
. Likewise, the tubercles on the pronotal surface may be somewhat more anastomosing on the discal median area, but never consisting of transverse rugae as in the above-mentioned species. The aedeagus apex, which is predominantly consistent in shape, may appear shorter due to the state of its apical orifice (as for instance with partly prolapsed internal sac in HT shown in
<figureCitation id="92DB2A69DA56FFCFFE78FB02FDC4FB04" box="[481,566,1159,1185]" captionStart="FIGURES 21–29" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[163,1424,1213,1911]" captionTargetId="figure-484@9.[158,1429,1209,1919]" captionTargetPageId="9" captionText="FIGURES 21–29. Pogonostoma (P.) ondravybirali sp. nov. (from type locality), male aedeagus or its apex. 21—HT (SDEI); 22—ditto, ventral view; 23—PT (CJVB); 24—ditto, dorsal view; 25—ditto, ventral view; 26—PT (CCJM); 27—ditto, ventral view; 28—PT (CJVB); 29—ditto, ventral view. Bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4283538" httpUri="https://zenodo.org/record/4283538/files/figure.png" pageId="9" pageNumber="210">Fig. 21</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="5E9F6664DA56FFCFFF0EF812FCF2F853" ID-DOI="http://doi.org/10.5281/zenodo.4283538" ID-Zenodo-Dep="4283538" httpUri="https://zenodo.org/record/4283538/files/figure.png" pageId="9" pageNumber="210" startId="9.[151,264,1943,1967]" targetBox="[163,1424,1213,1911]" targetPageId="9">
<paragraph id="0A5F36ECDA56FFCFFF0EF812FCF2F853" blockId="9.[151,1436,1943,2039]" pageId="9" pageNumber="210">
<emphasis id="3894EAFEDA56FFCFFF0EF812FEA7F80A" bold="true" box="[151,341,1943,1967]" pageId="9" pageNumber="210">FIGURES 21–29.</emphasis>
<taxonomicName id="CDE04D6FDA56FFCFFEC5F81DFE15F80A" authorityName="Klug" authorityYear="1835" box="[348,487,1944,1967]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="genus">
<emphasis id="3894EAFEDA56FFCFFEC5F81DFE15F80A" box="[348,487,1944,1967]" italics="true" pageId="9" pageNumber="210">Pogonostoma</emphasis>
</taxonomicName>
<taxonomicName id="CDE04D6FDA56FFCFFE77F812FD55F80A" authority="Moravec &amp; Vybíral, 2020" authorityName="Moravec &amp; Vybíral" authorityYear="2020" box="[494,679,1943,1967]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="ondravybirali" status="sp. nov." subGenus="Pogonostoma">
(
<emphasis id="3894EAFEDA56FFCFFE6FF81DFDF7F80A" box="[502,517,1944,1967]" italics="true" pageId="9" pageNumber="210">P</emphasis>
.)
<emphasis id="3894EAFEDA56FFCFFD85F812FD55F80A" box="[540,679,1943,1967]" italics="true" pageId="9" pageNumber="210">ondravybirali</emphasis>
</taxonomicName>
<emphasis id="3894EAFEDA56FFCFFD37F812FD0DF80A" bold="true" box="[686,767,1943,1967]" pageId="9" pageNumber="210">
<taxonomicNameLabel id="23A75785DA56FFCFFD37F812FD0DF80A" box="[686,767,1943,1967]" pageId="9" pageNumber="210" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(from type locality), male aedeagus or its apex. 21—HT (SDEI); 22—ditto, ventral view; 23—PT (CJVB); 24—ditto, dorsal view; 25—ditto, ventral view; 26—PT (CCJM); 27—ditto, ventral view; 28—PT (CJVB); 29—ditto, ventral view. Bar = 1 mm.
</paragraph>
</caption>
<subSubSection id="42FA6567DA55FFCCFF5EFF12FF1BFF70" pageId="10" pageNumber="211" type="etymology">
<paragraph id="0A5F36ECDA55FFCCFF5EFF12FF1BFF70" blockId="10.[151,1437,151,393]" pageId="10" pageNumber="211">
<emphasis id="3894EAFEDA55FFCCFF5EFF12FEBEFF14" bold="true" box="[199,332,151,177]" pageId="10" pageNumber="211">Etymology</emphasis>
. Named after one of the collectors, Ondřej Vybíral (son of the second author of the present paper).
</paragraph>
</subSubSection>
<subSubSection id="42FA6567DA55FFCCFF5EFF5BFDDCFE2C" pageId="10" pageNumber="211" type="distribution">
<paragraph id="0A5F36ECDA55FFCCFF5EFF5BFDDCFE2C" blockId="10.[151,1437,151,393]" pageId="10" pageNumber="211">
<emphasis id="3894EAFEDA55FFCCFF5EFF5BFE19FF5D" bold="true" box="[199,491,222,249]" pageId="10" pageNumber="211">Distribution and biology</emphasis>
.
<taxonomicName id="CDE04D6FDA55FFCCFE60FF5AFD61FF5D" authorityName="Klug" authorityYear="1835" box="[505,659,223,248]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="genus">
<emphasis id="3894EAFEDA55FFCCFE60FF5AFD61FF5D" box="[505,659,223,248]" italics="true" pageId="10" pageNumber="211">Pogonostoma</emphasis>
</taxonomicName>
<taxonomicName id="CDE04D6FDA55FFCCFD03FF5AFC97FF5C" authority="Moravec &amp; Vybíral, 2020" authorityName="Moravec &amp; Vybíral" authorityYear="2020" box="[666,869,223,249]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="ondravybirali" status="sp. nov." subGenus="Pogonostoma">
(
<emphasis id="3894EAFEDA55FFCCFD3AFF5AFD46FF5D" box="[675,692,223,248]" italics="true" pageId="10" pageNumber="211">P</emphasis>
.)
<emphasis id="3894EAFEDA55FFCCFD52FF5AFC97FF5C" box="[715,869,223,249]" italics="true" pageId="10" pageNumber="211">ondravybirali</emphasis>
</taxonomicName>
<taxonomicNameLabel id="23A75785DA55FFCCFCF5FF5AFC35FF5C" box="[876,967,223,249]" pageId="10" pageNumber="211" rank="species">
<emphasis id="3894EAFEDA55FFCCFCF5FF5AFC32FF5C" bold="true" box="[876,960,223,249]" pageId="10" pageNumber="211">sp. nov</emphasis>
.
</taxonomicNameLabel>
inhabits the sclerophylous woodland area of the Isalo National Park near Ranohira (Southwestern
<collectingCountry id="72F7767CDA55FFCCFC92FE86FC66FEB8" box="[779,916,259,285]" name="Madagascar" pageId="10" pageNumber="211">Madagascar</collectingCountry>
, region of Ihorombe). Adults from the
<typeStatus id="D55B884EDA55FFCCFADFFE86FA84FEB8" box="[1350,1398,259,285]" pageId="10" pageNumber="211">type</typeStatus>
lo-cality were taken from the coarse bark of the fire-resistant trees called Tapia (=
<taxonomicName id="CDE04D6FDA55FFCCFB9DFEA2FB15FEE7" authority="Baill" authorityName="Baill" authorityYear="1874" box="[1028,1255,295,322]" class="Magnoliopsida" family="Phyllanthaceae" genus="Uapaca" kingdom="Plantae" order="Malpighiales" pageId="10" pageNumber="211" phylum="Tracheophyta" rank="species" species="bojeri">
<emphasis id="3894EAFEDA55FFCCFB9DFEA2FB54FEE4" box="[1028,1190,295,321]" italics="true" pageId="10" pageNumber="211">Uapaca bojeri</emphasis>
Baill
</taxonomicName>
). Two examined males were caught on a different kind of tree in the Zombitse-Vohibasia National Park near Sakaraha, situated
<quantity id="CD189B09DA55FFCCFA19FECEFF49FE2C" metricMagnitude="4" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="211" unit="km" value="90.0">90 km</quantity>
southwest from the Isalo massif.
</paragraph>
</subSubSection>
</treatment>
</document>
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