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<document ID-DOI="10.7554/eLife.09560" ID-GBIF-Dataset="9f78d18a-b7d7-4b2e-94b3-9a970823b6a9" ID-PMC="PMC4559886" ID-PubMed="26354291" checkinTime="1441993702984" checkinUser="donat" docAuthor="Lee R Berger, John Hawks, Darryl J de Ruiter, Steven E Churchill, Peter Schmid, Lucas K Delezene, Tracy L Kivell, Heather M Garvin, Scott A Williams, Jeremy M DeSilva, Matthew M Skinner, Charles M Musiba, Noel Cameron, Trenton W Holliday, William Harcourt-Smith, Rebecca R Ackermann, Markus Bastir, Barry Bogin, Debra Bolter, Juliet Brophy, Zachary D Cofran, Kimberly A Congdon, Andrew S Deane, Mana Dembo, Michelle Drapeau, Marina C Elliott, , Elen M Feuerriegel, Daniel Garcia-Martinez, David J Green, Alia Gurtov, Joel D Irish, Ashley Kruger, Myra F Laird, Damiano Marchi, Marc R Meyer, Shahed Nalla, Enquye W Negash, Caley M Orr, Davorka Radovcic, Lauren Schroeder, Jill E Scott, Zachary Throckmorton, Matthew W Tocheri, Caroline VanSickle, Christopher S Walker, Pianpian Wei &amp; Bernhard Zipfel" docDate="2015" docId="822A19638D33FFC2FE35FAD8FA87F93A" docLanguage="en" docName="e09560.full.pdf.gs.imf" docOrigin="eLife e 09560 4" docTitle="Homo" docType="treatment" docVersion="13" lastPageNumber="27" masterDocId="7E13611B8D29FFD8FFF4FF93FFACFFBD" masterDocTitle="Homo naledi, a new species of the genus Homo from the Dinaledi Chamber, South Africa" masterLastPageNumber="35" masterPageNumber="1" pageNumber="27" updateTime="1668128208704" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Homo naledi, a new species of the genus Homo from the Dinaledi Chamber, South Africa</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lee R Berger</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>John Hawks</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Darryl J de Ruiter</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Steven E Churchill</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Peter Schmid</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lucas K Delezene</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tracy L Kivell</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Heather M Garvin</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Scott A Williams</mods:namePart>
</mods:name>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jeremy M DeSilva</mods:namePart>
</mods:name>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Matthew M Skinner</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Charles M Musiba</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Noel Cameron</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Trenton W Holliday</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>William Harcourt-Smith</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rebecca R Ackermann</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Markus Bastir</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Barry Bogin</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Debra Bolter</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Juliet Brophy</mods:namePart>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zachary D Cofran</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kimberly A Congdon</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Andrew S Deane</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mana Dembo</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Michelle Drapeau</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Marina C Elliott</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>, Elen M Feuerriegel</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Daniel Garcia-Martinez</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>David J Green</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Alia Gurtov</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Joel D Irish</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ashley Kruger</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Myra F Laird</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Damiano Marchi</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Marc R Meyer</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shahed Nalla</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Enquye W Negash</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Caley M Orr</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Davorka Radovcic</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lauren Schroeder</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jill E Scott</mods:namePart>
</mods:name>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zachary Throckmorton</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Matthew W Tocheri</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Caroline VanSickle</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Christopher S Walker</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pianpian Wei</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bernhard Zipfel</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>eLife e 09560</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="volume">
<mods:number>4</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>35</mods:end>
</mods:extent>
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</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.7554/eLife.09560</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6106411" ID-GBIF-Taxon="114920996" ID-Zenodo-Dep="6106411" LSID="urn:lsid:plazi:treatment:822A19638D33FFC2FE35FAD8FA87F93A" httpUri="http://treatment.plazi.org/id/822A19638D33FFC2FE35FAD8FA87F93A" lastPageNumber="27" pageId="26" pageNumber="27">
<subSubSection box="[449,822,1355,1384]" pageId="26" pageNumber="27" type="nomenclature">
<paragraph box="[449,822,1355,1384]" pageId="26" pageNumber="27">
<heading box="[449,822,1355,1384]" fontSize="12" level="3" pageId="26" pageNumber="27" reason="1">
Middle Pleistocene
<taxonomicName box="[733,822,1356,1384]" class="Mammalia" family="Hominidae" genus="Homo" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Primates" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">
<emphasis box="[733,822,1356,1384]" italics="true" pageId="26" pageNumber="27">Homo</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="26" pageNumber="27" type="discussion">
<paragraph pageId="26" pageNumber="27">
Specimens from the latest Lower Pleistocene and MP of Europe and Africa that cannot be attributed to
<taxonomicName box="[481,592,1425,1446]" class="Mammalia" family="Hominidae" genus="Homo" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Primates" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="erectus">
<emphasis box="[481,592,1425,1446]" italics="true" pageId="26" pageNumber="27">H. erectus</emphasis>
</taxonomicName>
were included in our comparisons. These include fossils that have been attributed to
<taxonomicName box="[449,650,1456,1478]" class="Mammalia" family="Hominidae" genus="Homo" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Primates" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="heidelbergensis">
<emphasis box="[449,650,1456,1478]" italics="true" pageId="26" pageNumber="27">H. heidelbergensis</emphasis>
</taxonomicName>
,
<taxonomicName box="[664,828,1456,1478]" class="Mammalia" family="Hominidae" genus="Homo" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Primates" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="rhodesiensis">
<emphasis box="[664,828,1456,1478]" italics="true" pageId="26" pageNumber="27">H. rhodesiensis</emphasis>
</taxonomicName>
, ‘archaic
<taxonomicName box="[931,1042,1457,1478]" class="Mammalia" family="Hominidae" genus="Homo" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Primates" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="sapiens">
<emphasis box="[931,1042,1457,1478]" italics="true" pageId="26" pageNumber="27">H. sapiens</emphasis>
</taxonomicName>
’, or ‘evolved
<taxonomicName box="[1191,1301,1457,1478]" class="Mammalia" family="Hominidae" genus="Homo" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Primates" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="erectus">
<emphasis box="[1191,1301,1457,1478]" italics="true" pageId="26" pageNumber="27">H. erectus</emphasis>
</taxonomicName>
’ by a variety of other authors. Specimens attributed to MP
<taxonomicName box="[861,927,1489,1510]" class="Mammalia" family="Hominidae" genus="Homo" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Primates" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">
<emphasis box="[861,927,1489,1510]" italics="true" pageId="26" pageNumber="27">Homo</emphasis>
</taxonomicName>
include materials from Eliye Springs, Arago, Atapuerca Sima de los Huesos, Bodo, Broken Hill, Cave of Hearths, Ceprano, Dali, Elandsfontein, Jinniushan, Kapthurin, Mauer, Narmada, Ndutu, Petralona, Reilingen-Schwetzingen, Solo, Steinheim, Swanscombe. This grouping includes the following specimens: KNM-ES 11693, Arago 2, Arago 13, Arago 21, Atapuerca 1, Atapuerca 2, Atapuerca 4, Atapuerca 5, Atapuerca 6, Cave of Hearths, SAM-PQ- EH1, Kabwe, Mauer, Ndutu, Sale´, Petralona, Reilingen-Schwetzingen, Steinheim.
</paragraph>
</subSubSection>
</treatment>
</document>
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