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<document ID-DOI="10.7554/eLife.09560" ID-GBIF-Dataset="9f78d18a-b7d7-4b2e-94b3-9a970823b6a9" ID-PMC="PMC4559886" ID-PubMed="26354291" checkinTime="1441993702984" checkinUser="donat" docAuthor="Lee R Berger, John Hawks, Darryl J de Ruiter, Steven E Churchill, Peter Schmid, Lucas K Delezene, Tracy L Kivell, Heather M Garvin, Scott A Williams, Jeremy M DeSilva, Matthew M Skinner, Charles M Musiba, Noel Cameron, Trenton W Holliday, William Harcourt-Smith, Rebecca R Ackermann, Markus Bastir, Barry Bogin, Debra Bolter, Juliet Brophy, Zachary D Cofran, Kimberly A Congdon, Andrew S Deane, Mana Dembo, Michelle Drapeau, Marina C Elliott, , Elen M Feuerriegel, Daniel Garcia-Martinez, David J Green, Alia Gurtov, Joel D Irish, Ashley Kruger, Myra F Laird, Damiano Marchi, Marc R Meyer, Shahed Nalla, Enquye W Negash, Caley M Orr, Davorka Radovcic, Lauren Schroeder, Jill E Scott, Zachary Throckmorton, Matthew W Tocheri, Caroline VanSickle, Christopher S Walker, Pianpian Wei &amp; Bernhard Zipfel" docDate="2015" docId="822A19638D30FFC1FE35FABCFA93F914" docLanguage="en" docName="e09560.full.pdf.gs.imf" docOrigin="eLife e 09560 4" docTitle="Homo habilis" docType="treatment" docVersion="13" lastPageNumber="26" masterDocId="7E13611B8D29FFD8FFF4FF93FFACFFBD" masterDocTitle="Homo naledi, a new species of the genus Homo from the Dinaledi Chamber, South Africa" masterLastPageNumber="35" masterPageNumber="1" pageNumber="26" updateTime="1668128208704" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Homo naledi, a new species of the genus Homo from the Dinaledi Chamber, South Africa</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lee R Berger</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>John Hawks</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Darryl J de Ruiter</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Steven E Churchill</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Peter Schmid</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lucas K Delezene</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tracy L Kivell</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Heather M Garvin</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Scott A Williams</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jeremy M DeSilva</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Matthew M Skinner</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Charles M Musiba</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Noel Cameron</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Trenton W Holliday</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>William Harcourt-Smith</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rebecca R Ackermann</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Markus Bastir</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Barry Bogin</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Debra Bolter</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Juliet Brophy</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zachary D Cofran</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kimberly A Congdon</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Andrew S Deane</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mana Dembo</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Michelle Drapeau</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Marina C Elliott</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>, Elen M Feuerriegel</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Daniel Garcia-Martinez</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>David J Green</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Alia Gurtov</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Joel D Irish</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ashley Kruger</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Myra F Laird</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Damiano Marchi</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Marc R Meyer</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shahed Nalla</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Enquye W Negash</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Caley M Orr</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Davorka Radovcic</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lauren Schroeder</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jill E Scott</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zachary Throckmorton</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Matthew W Tocheri</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Caroline VanSickle</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Christopher S Walker</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pianpian Wei</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bernhard Zipfel</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>eLife e 09560</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="volume">
<mods:number>4</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>35</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.7554/eLife.09560</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6106401" ID-GBIF-Taxon="114920999" ID-Zenodo-Dep="6106401" LSID="urn:lsid:plazi:treatment:822A19638D30FFC1FE35FABCFA93F914" httpUri="http://treatment.plazi.org/id/822A19638D30FFC1FE35FABCFA93F914" lastPageNumber="26" pageId="25" pageNumber="26">
<subSubSection box="[449,638,1326,1355]" pageId="25" pageNumber="26" type="nomenclature">
<paragraph box="[449,638,1326,1355]" pageId="25" pageNumber="26">
<heading box="[449,638,1326,1355]" fontSize="12" level="3" pageId="25" pageNumber="26" reason="1">
<taxonomicName box="[449,638,1326,1355]" class="Mammalia" family="Hominidae" genus="Homo" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Primates" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="habilis">
<emphasis box="[449,638,1326,1355]" italics="true" pageId="25" pageNumber="26">Homo habilis</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="25" pageNumber="26" type="materials_examined">
<paragraph pageId="25" pageNumber="26">
Samples from Olduvai Gorge, East Lake Turkana, the Omo Shungura sequence, Hadar, and Sterkfontein were included in this study. Original Olduvai Gorge and East Lake Turkana fossils were examined first-hand, while for the Omo and Hadar materials we relied on our original observations on casts and originals and published reports (
<emphasis bold="true" box="[925,1195,1460,1481]" italics="true" pageId="25" pageNumber="26">Boaz and Howell, 1977</emphasis>
;
<emphasis bold="true" box="[1212,1360,1460,1481]" italics="true" pageId="25" pageNumber="26">Tobias, 1991</emphasis>
;
<emphasis bold="true" italics="true" pageId="25" pageNumber="26">Kimbel et al., 1997</emphasis>
). We include the following fossils in the hypodigm of
<taxonomicName box="[1074,1172,1491,1513]" class="Mammalia" family="Hominidae" genus="Homo" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Primates" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="habilis">
<emphasis box="[1074,1092,1492,1513]" italics="true" pageId="25" pageNumber="26">H</emphasis>
.
<emphasis box="[1104,1172,1491,1513]" italics="true" pageId="25" pageNumber="26">habilis</emphasis>
</taxonomicName>
: A.L. 666-1, KNM-ER 1478, KNM- ER 1501, KNM-ER 1502, KNM-ER 1805, KNM-ER 1813, KNM-ER 3735, OH 4, OH 6, OH 7, OH 8, OH 13, OH 15, OH 16, OH 21, OH 24, OH 27, OH 31, OH 35, OH 37, OH 39, OH 42, OH 44, OH 45, OH 62, OMO-L894-1, and Stw 53. We recognize that some authors (including some of the authors of this paper) prefer to classify OH 62, Stw 53 and A.L. 666-1 outside of
<taxonomicName box="[1238,1343,1619,1641]" class="Mammalia" family="Hominidae" genus="Homo" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Primates" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="habilis">
<emphasis box="[1238,1343,1619,1641]" italics="true" pageId="25" pageNumber="26">H. habilis</emphasis>
</taxonomicName>
, (e.g., as
<taxonomicName class="Mammalia" family="Hominidae" genus="Homo" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Primates" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="gautengensis">
<emphasis italics="true" pageId="25" pageNumber="26">Homo gautengensis</emphasis>
</taxonomicName>
which we do not recognize as valid), or even outside the genus
<taxonomicName box="[1277,1343,1652,1673]" class="Mammalia" family="Hominidae" genus="Homo" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Primates" pageId="25" pageNumber="26" phylum="Chordata" rank="genus">
<emphasis box="[1277,1343,1652,1673]" italics="true" pageId="25" pageNumber="26">Homo</emphasis>
</taxonomicName>
; these specimens expand the morphological and temporal variability encompassed within
<emphasis box="[1235,1343,1683,1705]" italics="true" pageId="25" pageNumber="26">
<taxonomicName box="[1235,1339,1683,1705]" class="Mammalia" family="Hominidae" genus="Homo" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Primates" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="habilis">H. habilis</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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