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<document id="60CFCE7CCE8F5BFB8A4B4EF203629536" ID-CLB-Dataset="7017" ID-DOI="10.1080/00222933.2012.737483" ID-GBIF-Dataset="f7104997-eb0e-4bc6-931c-61f6426d0b97" ID-ISSN="1464-5262" ID-Zenodo-Dep="4632021" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616524552656" checkinUser="carolina" docAuthor="Madinabeitia, Ione, Tang, Danny &amp; Nagasawa, Kazuya" docDate="2013" docId="813A87D1FF963A02564EFF134963FA6A" docLanguage="en" docName="JNATHIST.47.5-12.563-580.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 47 (5 - 12)" docSource="http://dx.doi.org/10.1080/00222933.2012.737483" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.6:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="6" docTitle="Colobomatus absens Madinabeitia &amp; Tang &amp; Nagasawa 2013, sp. nov." docType="treatment" docVersion="7" lastPageNumber="578" masterDocId="7D03FFA9FF863A135458FFBB480EFF92" masterDocTitle="Four new species of Colobomatus (Copepoda: Philichthyidae) parasitic in the lateral line system of marine finfishes captured off the Ryukyu Islands, Japan, with redescriptions of Colobomatus collettei Cressey, 1977 and Colobomatus pupa Izawa, 1974" masterLastPageNumber="580" masterPageNumber="563" pageNumber="577" updateTime="1698934689405" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="44C1EEBF678742E3FD84403D262BC6AD">Four new species of Colobomatus (Copepoda: Philichthyidae) parasitic in the lateral line system of marine finfishes captured off the Ryukyu Islands, Japan, with redescriptions of Colobomatus collettei Cressey, 1977 and Colobomatus pupa Izawa, 1974</mods:title>
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<mods:namePart id="4F1C0BA9349BD8DC22DFE2B1D3EB72B0">Madinabeitia, Ione</mods:namePart>
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<mods:namePart id="1AFC0ED604B0C0DA316B7A7264E6F909">Tang, Danny</mods:namePart>
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<mods:namePart id="A3FAF68AF19A86B19395ECB8ADA84E87">Nagasawa, Kazuya</mods:namePart>
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<mods:title id="A35E709A4C47D720E7155CE407030CAD">Journal of Natural History</mods:title>
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<mods:date id="DCFA7290AF779D7CCDDA5931891270AB">2013</mods:date>
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<mods:number id="5921931D2ABE0CAA2B0BB3400FEF83E9">2012-12-17</mods:number>
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<treatment id="813A87D1FF963A02564EFF134963FA6A" ID-DOI="http://doi.org/10.5281/zenodo.4631783" ID-GBIF-Taxon="181974082" ID-Zenodo-Dep="4631783" LSID="urn:lsid:plazi:treatment:813A87D1FF963A02564EFF134963FA6A" httpUri="http://treatment.plazi.org/id/813A87D1FF963A02564EFF134963FA6A" lastPageId="17" lastPageNumber="578" pageId="16" pageNumber="577">
<subSubSection id="4189654CFF963A03564EFF134B5FFF2D" box="[534,849,168,192]" pageId="16" pageNumber="577" type="nomenclature">
<paragraph id="092C36C7FF963A03564EFF134B5FFF2D" blockId="16.[534,849,168,223]" box="[534,849,168,192]" pageId="16" pageNumber="577">
<heading id="526481ABFF963A03564EFF134B5FFF2D" box="[534,849,168,192]" centered="true" fontSize="10" level="2" pageId="16" pageNumber="577" reason="2">
<taxonomicName id="CE934D44FF963A03564EFF134AF8FF52" authority="Madinabeitia &amp; Tang &amp; Nagasawa, 2013" authorityName="Madinabeitia &amp; Tang &amp; Nagasawa" authorityYear="2013" box="[534,758,168,192]" class="Maxillopoda" family="Philichthyidae" genus="Colobomatus" kingdom="Animalia" order="Poecilostomatoida" pageId="16" pageNumber="577" phylum="Arthropoda" rank="species" species="absens" status="sp. nov.">
<emphasis id="3BE7EAD5FF963A03564EFF134AF8FF52" bold="true" box="[534,758,168,192]" italics="true" pageId="16" pageNumber="577">Colobomatus absens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="20D457AEFF963A0356A5FF134B5FFF2D" box="[765,849,168,191]" pageId="16" pageNumber="577" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="4189654CFF963A03563AFF7C4B0BFF4D" box="[610,773,199,223]" pageId="16" pageNumber="577" type="description">
<paragraph id="092C36C7FF963A03563AFF7C4B0BFF4D" blockId="16.[534,849,168,223]" box="[610,773,199,223]" pageId="16" pageNumber="577">
(
<figureCitation id="91A82A42FF963A035631FF7C4AC6FF4D" box="[617,712,199,223]" captionStart="Figure 6" captionStartId="17.[143,210,1252,1274]" captionTargetBox="[143,1103,167,1219]" captionTargetId="figure-5@17.[143,1103,167,1219]" captionTargetPageId="17" captionText="Figure 6. Colobomatus absens sp. nov., adult female.(A) Habitus, ventral; (B) antennule, ventral; (C) buccal capsule and mouthparts, ventral; (D) leg 1, ventrolateral; (E) tip of thoracic process, ventral. Abbreviations: A, antenna; la, labium; mx, maxillule; Mx, maxilla; Mxp, maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4632033" httpUri="https://zenodo.org/record/4632033/files/figure.png" pageId="16" pageNumber="577">Figure 6</figureCitation>
AE)
</paragraph>
</subSubSection>
<subSubSection id="4189654CFF963A03548BFE9F4A52FE71" pageId="16" pageNumber="577" type="materials_examined">
<paragraph id="092C36C7FF963A03548BFE9F49ABFEAE" blockId="16.[211,1171,292,484]" box="[211,421,292,316]" pageId="16" pageNumber="577">
<heading id="526481ABFF963A03548BFE9F49ABFEAE" box="[211,421,292,316]" fontSize="10" level="3" pageId="16" pageNumber="577" reason="5">
<emphasis id="3BE7EAD5FF963A03548BFE9F49ABFEAE" box="[211,421,292,316]" italics="true" pageId="16" pageNumber="577">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="092C36C7FF963A03548BFEF04A52FE71" blockId="16.[211,1171,292,484]" pageId="16" pageNumber="577">
<typeStatus id="D6288865FF963A03548BFEF04930FEF1" box="[211,318,331,355]" pageId="16" pageNumber="577" type="holotype">Holotype</typeStatus>
female (NSMT-Cr 21863) and
<specimenCount id="1F95FD4EFF963A03569CFEF04BA8FEF1" box="[708,934,331,356]" pageId="16" pageNumber="577" type="female" typeStatus="paratype">2 paratype females</specimenCount>
(NSMT-Cr 21864), from head canals of two
<taxonomicName id="CE934D44FF963A0355A8FED74B5AFE11" authority="(Bleeker)" baseAuthorityName="Bleeker" baseAuthorityYear="1864" box="[496,852,363,388]" class="Actinopterygii" family="Caesionidae" genus="Pterocaesio" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="577" phylum="Chordata" rank="species" species="digramma">
<emphasis id="3BE7EAD5FF963A0355A8FED74AE9FE16" box="[496,743,364,388]" italics="true" pageId="16" pageNumber="577">Pterocaesio digramma</emphasis>
(Bleeker)
</taxonomicName>
(
<taxonomicName id="CE934D44FF963A03573CFED04BE0FE11" box="[868,1006,363,387]" class="Actinopterygii" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="577" phylum="Chordata" rank="order">Perciformes</taxonomicName>
:
<taxonomicName id="CE934D44FF963A0357A2FED04C86FE11" authorityName="Bonaparte" authorityYear="1831" box="[1018,1160,363,387]" class="Actinopterygii" family="Caesionidae" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="577" phylum="Chordata" rank="family">Caesionidae</taxonomicName>
), captured in the western North Pacific Ocean off Ishigaki-jima Island and subsequently purchased at Yaeyama Fisheries Cooperative (24
<superScript id="FEE69B8FFF963A03570AFE1C4B55FE2A" attach="none" box="[850,859,423,440]" fontSize="7" pageId="16" pageNumber="577"></superScript>
19
<superScript id="FEE69B8FFF963A03572FFE1C4B72FE2A" attach="left" box="[887,892,423,440]" fontSize="7" pageId="16" pageNumber="577"></superScript>
N, 124
<superScript id="FEE69B8FFF963A035796FE1C4BD9FE2A" attach="left" box="[974,983,423,440]" fontSize="7" pageId="16" pageNumber="577"></superScript>
0
<superScript id="FEE69B8FFF963A0357BEFE1C4BE5FE2A" attach="left" box="[998,1003,423,440]" fontSize="7" pageId="16" pageNumber="577"></superScript>
E) in Ishigaki, Ishigaki-jima Island,
<date id="7D2D1007FF963A03559BFE704A56FE71" box="[451,600,459,484]" pageId="16" pageNumber="577" value="2010-05-12">12 May 2010</date>
.
</paragraph>
</subSubSection>
<subSubSection id="4189654CFF963A03548BFD9D4C08FAF0" pageId="16" pageNumber="577" type="description">
<paragraph id="092C36C7FF963A03548BFD9D495CFDAF" blockId="16.[211,1172,550,1379]" box="[211,338,550,573]" pageId="16" pageNumber="577">
<heading id="526481ABFF963A03548BFD9D495CFDAF" box="[211,338,550,573]" fontSize="10" level="3" pageId="16" pageNumber="577" reason="5">
<emphasis id="3BE7EAD5FF963A03548BFD9D495CFDAF" box="[211,338,550,573]" italics="true" pageId="16" pageNumber="577">Description</emphasis>
</heading>
</paragraph>
<paragraph id="092C36C7FF963A03548BFDF64A62FB91" blockId="16.[211,1172,550,1379]" pageId="16" pageNumber="577">
<emphasis id="3BE7EAD5FF963A03548BFDF64965FDF7" box="[211,363,589,613]" italics="true" pageId="16" pageNumber="577">Adult female.</emphasis>
Body (
<figureCitation id="91A82A42FF963A0355E7FDF64A36FDF7" box="[447,568,589,613]" captionStart="Figure 6" captionStartId="17.[143,210,1252,1274]" captionTargetBox="[143,1103,167,1219]" captionTargetId="figure-5@17.[143,1103,167,1219]" captionTargetPageId="17" captionText="Figure 6. Colobomatus absens sp. nov., adult female.(A) Habitus, ventral; (B) antennule, ventral; (C) buccal capsule and mouthparts, ventral; (D) leg 1, ventrolateral; (E) tip of thoracic process, ventral. Abbreviations: A, antenna; la, labium; mx, maxillule; Mx, maxilla; Mxp, maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4632033" httpUri="https://zenodo.org/record/4632033/files/figure.png" pageId="16" pageNumber="577">Figure 6A</figureCitation>
) 1.89 (1.871.91) mm long (excluding cephalic processes and caudal rami) (
<emphasis id="3BE7EAD5FF963A0355AEFDD54A0DFD17" box="[502,515,622,645]" italics="true" pageId="16" pageNumber="577">n</emphasis>
= 3). Pre-oral area of cephalosome (
<figureCitation id="91A82A42FF963A0357EDFDD64C23FD17" box="[949,1069,621,645]" captionStart="Figure 6" captionStartId="17.[143,210,1252,1274]" captionTargetBox="[143,1103,167,1219]" captionTargetId="figure-5@17.[143,1103,167,1219]" captionTargetPageId="17" captionText="Figure 6. Colobomatus absens sp. nov., adult female.(A) Habitus, ventral; (B) antennule, ventral; (C) buccal capsule and mouthparts, ventral; (D) leg 1, ventrolateral; (E) tip of thoracic process, ventral. Abbreviations: A, antenna; la, labium; mx, maxillule; Mx, maxilla; Mxp, maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4632033" httpUri="https://zenodo.org/record/4632033/files/figure.png" pageId="16" pageNumber="577">Figure 6A</figureCitation>
) bearing one anterior, spinulose pair of anteriorly directed and pointed processes. Cephalosome round and constricted at posterior margin. First pedigerous somite cylindrical, forming short neck; first pair of legs on ventral surface near anterior constriction. Second to fourth pedigerous somites fused, elongated, widest at posterior end and bearing one pair of anterolateral processes; latter with spinulose ornamentation and tapering into pointed tips (
<figureCitation id="91A82A42FF963A0355C4FC974A1FFCD6" box="[412,529,812,836]" captionStart="Figure 6" captionStartId="17.[143,210,1252,1274]" captionTargetBox="[143,1103,167,1219]" captionTargetId="figure-5@17.[143,1103,167,1219]" captionTargetPageId="17" captionText="Figure 6. Colobomatus absens sp. nov., adult female.(A) Habitus, ventral; (B) antennule, ventral; (C) buccal capsule and mouthparts, ventral; (D) leg 1, ventrolateral; (E) tip of thoracic process, ventral. Abbreviations: A, antenna; la, labium; mx, maxillule; Mx, maxilla; Mxp, maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4632033" httpUri="https://zenodo.org/record/4632033/files/figure.png" pageId="16" pageNumber="577">Figure 6E</figureCitation>
). Fifth pedigerous somite short, narrower than preceding tagma. Genital somite (
<figureCitation id="91A82A42FF963A035583FCF74A41FCF6" box="[475,591,844,868]" captionStart="Figure 6" captionStartId="17.[143,210,1252,1274]" captionTargetBox="[143,1103,167,1219]" captionTargetId="figure-5@17.[143,1103,167,1219]" captionTargetPageId="17" captionText="Figure 6. Colobomatus absens sp. nov., adult female.(A) Habitus, ventral; (B) antennule, ventral; (C) buccal capsule and mouthparts, ventral; (D) leg 1, ventrolateral; (E) tip of thoracic process, ventral. Abbreviations: A, antenna; la, labium; mx, maxillule; Mx, maxilla; Mxp, maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4632033" httpUri="https://zenodo.org/record/4632033/files/figure.png" pageId="16" pageNumber="577">Figure 6A</figureCitation>
) bearing one ventrolateral pair of slender, spinulose and posterolaterally directed processes; latter as long as anterior pair of thoracic processes. Abdomen (
<figureCitation id="91A82A42FF963A0355FDFC304A12FC31" box="[421,540,907,931]" captionStart="Figure 6" captionStartId="17.[143,210,1252,1274]" captionTargetBox="[143,1103,167,1219]" captionTargetId="figure-5@17.[143,1103,167,1219]" captionTargetPageId="17" captionText="Figure 6. Colobomatus absens sp. nov., adult female.(A) Habitus, ventral; (B) antennule, ventral; (C) buccal capsule and mouthparts, ventral; (D) leg 1, ventrolateral; (E) tip of thoracic process, ventral. Abbreviations: A, antenna; la, labium; mx, maxillule; Mx, maxilla; Mxp, maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4632033" httpUri="https://zenodo.org/record/4632033/files/figure.png" pageId="16" pageNumber="577">Figure 6A</figureCitation>
) indistinctly three-segmented, with single dorsomedian process at posterior end projecting between caudal rami; process spinulose, pointed at tip and as long as caudal rami. Caudal ramus (
<figureCitation id="91A82A42FF963A0356ABFC704B67FC71" box="[755,873,971,995]" captionStart="Figure 6" captionStartId="17.[143,210,1252,1274]" captionTargetBox="[143,1103,167,1219]" captionTargetId="figure-5@17.[143,1103,167,1219]" captionTargetPageId="17" captionText="Figure 6. Colobomatus absens sp. nov., adult female.(A) Habitus, ventral; (B) antennule, ventral; (C) buccal capsule and mouthparts, ventral; (D) leg 1, ventrolateral; (E) tip of thoracic process, ventral. Abbreviations: A, antenna; la, labium; mx, maxillule; Mx, maxilla; Mxp, maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4632033" httpUri="https://zenodo.org/record/4632033/files/figure.png" pageId="16" pageNumber="577">Figure 6A</figureCitation>
) posterolaterally directed, spinulose, tapering into pointed tips.
</paragraph>
<paragraph id="092C36C7FF963A0354A6FBB04B81FA90" blockId="16.[211,1172,550,1379]" pageId="16" pageNumber="577">
Antennule (
<figureCitation id="91A82A42FF963A0355DDFBB049F4FBB1" box="[389,506,1035,1059]" captionStart="Figure 6" captionStartId="17.[143,210,1252,1274]" captionTargetBox="[143,1103,167,1219]" captionTargetId="figure-5@17.[143,1103,167,1219]" captionTargetPageId="17" captionText="Figure 6. Colobomatus absens sp. nov., adult female.(A) Habitus, ventral; (B) antennule, ventral; (C) buccal capsule and mouthparts, ventral; (D) leg 1, ventrolateral; (E) tip of thoracic process, ventral. Abbreviations: A, antenna; la, labium; mx, maxillule; Mx, maxilla; Mxp, maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4632033" httpUri="https://zenodo.org/record/4632033/files/figure.png" pageId="16" pageNumber="577">Figure 6B</figureCitation>
) arising near base of cephalic processes, apparently foursegmented with armature of 1, 8, 4 and 5 + 2 aesthetascs. Antenna (
<figureCitation id="91A82A42FF963A03504AFB904C85FBD1" box="[1042,1163,1067,1091]" captionStart="Figure 6" captionStartId="17.[143,210,1252,1274]" captionTargetBox="[143,1103,167,1219]" captionTargetId="figure-5@17.[143,1103,167,1219]" captionTargetPageId="17" captionText="Figure 6. Colobomatus absens sp. nov., adult female.(A) Habitus, ventral; (B) antennule, ventral; (C) buccal capsule and mouthparts, ventral; (D) leg 1, ventrolateral; (E) tip of thoracic process, ventral. Abbreviations: A, antenna; la, labium; mx, maxillule; Mx, maxilla; Mxp, maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4632033" httpUri="https://zenodo.org/record/4632033/files/figure.png" pageId="16" pageNumber="577">Figure 6C</figureCitation>
) modified, forming large, longitudinally divided anterior margin of buccal capsule; latter (
<figureCitation id="91A82A42FF963A03555DFBD04997FB11" box="[261,409,1131,1155]" captionStart="Figure 6" captionStartId="17.[143,210,1252,1274]" captionTargetBox="[143,1103,167,1219]" captionTargetId="figure-5@17.[143,1103,167,1219]" captionTargetPageId="17" captionText="Figure 6. Colobomatus absens sp. nov., adult female.(A) Habitus, ventral; (B) antennule, ventral; (C) buccal capsule and mouthparts, ventral; (D) leg 1, ventrolateral; (E) tip of thoracic process, ventral. Abbreviations: A, antenna; la, labium; mx, maxillule; Mx, maxilla; Mxp, maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4632033" httpUri="https://zenodo.org/record/4632033/files/figure.png" pageId="16" pageNumber="577">Figure 6A,C</figureCitation>
) tube-like, projecting ventrally. Labrum and mandibles not seen. Maxillule (
<figureCitation id="91A82A42FF963A03550AFB3149C4FB30" box="[338,458,1162,1186]" captionStart="Figure 6" captionStartId="17.[143,210,1252,1274]" captionTargetBox="[143,1103,167,1219]" captionTargetId="figure-5@17.[143,1103,167,1219]" captionTargetPageId="17" captionText="Figure 6. Colobomatus absens sp. nov., adult female.(A) Habitus, ventral; (B) antennule, ventral; (C) buccal capsule and mouthparts, ventral; (D) leg 1, ventrolateral; (E) tip of thoracic process, ventral. Abbreviations: A, antenna; la, labium; mx, maxillule; Mx, maxilla; Mxp, maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4632033" httpUri="https://zenodo.org/record/4632033/files/figure.png" pageId="16" pageNumber="577">Figure 6C</figureCitation>
) anteriorly located in buccal capsule and bearing one apical spine. Maxilla (
<figureCitation id="91A82A42FF963A0355D9FB1149FAFB50" box="[385,500,1194,1218]" captionStart="Figure 6" captionStartId="17.[143,210,1252,1274]" captionTargetBox="[143,1103,167,1219]" captionTargetId="figure-5@17.[143,1103,167,1219]" captionTargetPageId="17" captionText="Figure 6. Colobomatus absens sp. nov., adult female.(A) Habitus, ventral; (B) antennule, ventral; (C) buccal capsule and mouthparts, ventral; (D) leg 1, ventrolateral; (E) tip of thoracic process, ventral. Abbreviations: A, antenna; la, labium; mx, maxillule; Mx, maxilla; Mxp, maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4632033" httpUri="https://zenodo.org/record/4632033/files/figure.png" pageId="16" pageNumber="577">Figure 6C</figureCitation>
) large, apparently one-segmented and armed with claw apically. Maxilliped (
<figureCitation id="91A82A42FF963A0355FCFB714A12FB70" box="[420,540,1226,1250]" captionStart="Figure 6" captionStartId="17.[143,210,1252,1274]" captionTargetBox="[143,1103,167,1219]" captionTargetId="figure-5@17.[143,1103,167,1219]" captionTargetPageId="17" captionText="Figure 6. Colobomatus absens sp. nov., adult female.(A) Habitus, ventral; (B) antennule, ventral; (C) buccal capsule and mouthparts, ventral; (D) leg 1, ventrolateral; (E) tip of thoracic process, ventral. Abbreviations: A, antenna; la, labium; mx, maxillule; Mx, maxilla; Mxp, maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4632033" httpUri="https://zenodo.org/record/4632033/files/figure.png" pageId="16" pageNumber="577">Figure 6C</figureCitation>
) blunt and unarmed. Labium (
<figureCitation id="91A82A42FF963A0357C8FB714C06FB70" box="[912,1032,1226,1250]" captionStart="Figure 6" captionStartId="17.[143,210,1252,1274]" captionTargetBox="[143,1103,167,1219]" captionTargetId="figure-5@17.[143,1103,167,1219]" captionTargetPageId="17" captionText="Figure 6. Colobomatus absens sp. nov., adult female.(A) Habitus, ventral; (B) antennule, ventral; (C) buccal capsule and mouthparts, ventral; (D) leg 1, ventrolateral; (E) tip of thoracic process, ventral. Abbreviations: A, antenna; la, labium; mx, maxillule; Mx, maxilla; Mxp, maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4632033" httpUri="https://zenodo.org/record/4632033/files/figure.png" pageId="16" pageNumber="577">Figure 6C</figureCitation>
) undivided, anteriorly rounded. Posterior rim of buccal capsule undivided.
</paragraph>
<paragraph id="092C36C7FF963A0354A6FAB04C08FAF0" blockId="16.[211,1172,550,1379]" pageId="16" pageNumber="577">
Leg 1 (
<figureCitation id="91A82A42FF963A035516FAB149C8FAB0" box="[334,454,1290,1314]" captionStart="Figure 6" captionStartId="17.[143,210,1252,1274]" captionTargetBox="[143,1103,167,1219]" captionTargetId="figure-5@17.[143,1103,167,1219]" captionTargetPageId="17" captionText="Figure 6. Colobomatus absens sp. nov., adult female.(A) Habitus, ventral; (B) antennule, ventral; (C) buccal capsule and mouthparts, ventral; (D) leg 1, ventrolateral; (E) tip of thoracic process, ventral. Abbreviations: A, antenna; la, labium; mx, maxillule; Mx, maxilla; Mxp, maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4632033" httpUri="https://zenodo.org/record/4632033/files/figure.png" pageId="16" pageNumber="577">Figure 6D</figureCitation>
) uniramous; protopod completely fused to somite, represented by long, lateral surface seta; exopod indistinctly two-segmented, with unarmed basal segment and four elements on distal segment. Legs 2 and 3 not observed.
</paragraph>
</subSubSection>
<subSubSection id="4189654CFF963A03548BFA1F4B85F991" pageId="16" pageNumber="577" type="etymology">
<paragraph id="092C36C7FF963A03548BFA1F4945FA2E" blockId="16.[211,1171,1444,1539]" box="[211,331,1444,1468]" pageId="16" pageNumber="577">
<heading id="526481ABFF963A03548BFA1F4945FA2E" box="[211,331,1444,1468]" fontSize="10" level="3" pageId="16" pageNumber="577" reason="5">
<emphasis id="3BE7EAD5FF963A03548BFA1F4945FA2E" box="[211,331,1444,1468]" italics="true" pageId="16" pageNumber="577">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="092C36C7FF963A03548BFA704B85F991" blockId="16.[211,1171,1444,1539]" pageId="16" pageNumber="577">
The species epithet,
<taxonomicName id="CE934D44FF963A03559AFA774A04FA76" authorityName="Madinabeitia &amp; Tang &amp; Nagasawa" authorityYear="2013" box="[450,522,1484,1508]" class="Maxillopoda" family="Philichthyidae" genus="Colobomatus" kingdom="Animalia" order="Poecilostomatoida" pageId="16" pageNumber="577" phylum="Arthropoda" rank="species" species="absens">
<emphasis id="3BE7EAD5FF963A03559AFA774A04FA76" box="[450,522,1484,1508]" italics="true" pageId="16" pageNumber="577">absens</emphasis>
</taxonomicName>
, is taken from the Latin for “missing”. It refers to the absence of a posterior pair of thoracic processes in the female.
</paragraph>
</subSubSection>
<subSubSection id="4189654CFF963A02548BF9FE4963FA6A" lastPageId="17" lastPageNumber="578" pageId="16" pageNumber="577" type="discussion">
<paragraph id="092C36C7FF963A03548BF9FE493BF9CF" blockId="16.[211,1171,1605,1701]" box="[211,309,1605,1629]" pageId="16" pageNumber="577">
<heading id="526481ABFF963A03548BF9FE493BF9CF" box="[211,309,1605,1629]" fontSize="10" level="3" pageId="16" pageNumber="577" reason="5">
<emphasis id="3BE7EAD5FF963A03548BF9FE493BF9CF" box="[211,309,1605,1629]" italics="true" pageId="16" pageNumber="577">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="092C36C7FF963A02548BF9D64AA8FA2A" blockId="16.[211,1171,1605,1701]" lastBlockId="17.[143,1103,1376,1528]" lastPageId="17" lastPageNumber="578" pageId="16" pageNumber="577">
The presence of a posteriorly directed, median process at the posterior end of the abdomen of
<taxonomicName id="CE934D44FF963A035530F93549DBF937" authority="Madinabeitia &amp; Tang &amp; Nagasawa, 2013" authorityName="Madinabeitia &amp; Tang &amp; Nagasawa" authorityYear="2013" box="[360,469,1677,1701]" class="Maxillopoda" family="Philichthyidae" genus="Colobomatus" kingdom="Animalia" order="Poecilostomatoida" pageId="16" pageNumber="577" phylum="Arthropoda" rank="species" species="absens" status="sp. nov.">
<emphasis id="3BE7EAD5FF963A035530F93549DBF937" box="[360,469,1677,1701]" italics="true" pageId="16" pageNumber="577">C. absens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="20D457AEFF963A035587F9354A39F937" box="[479,567,1678,1701]" pageId="16" pageNumber="577" rank="species">sp. nov.</taxonomicNameLabel>
is a character shared with
<taxonomicName id="CE934D44FF963A035722F9354BF6F937" baseAuthorityName="Richiardi" baseAuthorityYear="1877" box="[890,1016,1677,1701]" class="Maxillopoda" family="Philichthyidae" genus="Colobomatus" kingdom="Animalia" order="Poecilostomatoida" pageId="16" pageNumber="577" phylum="Arthropoda" rank="species" species="haeckeli">
<emphasis id="3BE7EAD5FF963A035722F9354BF6F937" box="[890,1016,1677,1701]" italics="true" pageId="16" pageNumber="577">C. haeckeli</emphasis>
</taxonomicName>
,
<taxonomicName id="CE934D44FF963A035052F9354C85F937" authorityName="Cressey and Collette" authorityYear="1970" box="[1034,1163,1677,1701]" class="Maxillopoda" family="Philichthyidae" genus="Colobomatus" kingdom="Animalia" order="Poecilostomatoida" pageId="16" pageNumber="577" phylum="Arthropoda" rank="species" species="goodingi">
<emphasis id="3BE7EAD5FF963A035052F9354C85F937" box="[1034,1163,1677,1701]" italics="true" pageId="16" pageNumber="577">C. goodingi</emphasis>
</taxonomicName>
,
<taxonomicName id="CE934D44FF973A0254D7FADA48E5FAEB" authorityName="Izawa" authorityYear="1974" box="[143,235,1377,1401]" class="Maxillopoda" family="Philichthyidae" genus="Colobomatus" kingdom="Animalia" order="Poecilostomatoida" pageId="17" pageNumber="578" phylum="Arthropoda" rank="species" species="exilis">
<emphasis id="3BE7EAD5FF973A0254D7FADA48E5FAEB" box="[143,235,1377,1401]" italics="true" pageId="17" pageNumber="578">C. exilis</emphasis>
</taxonomicName>
,
<taxonomicName id="CE934D44FF973A0254A1FADA497DFAEB" authorityName="Cressey" authorityYear="1977" box="[249,371,1377,1401]" class="Maxillopoda" family="Philichthyidae" genus="Colobomatus" kingdom="Animalia" order="Poecilostomatoida" pageId="17" pageNumber="578" phylum="Arthropoda" rank="species" species="collettei">
<emphasis id="3BE7EAD5FF973A0254A1FADA497DFAEB" box="[249,371,1377,1401]" italics="true" pageId="17" pageNumber="578">C. collettei</emphasis>
</taxonomicName>
,
<taxonomicName id="CE934D44FF973A0255DAFADA4A05FAEA" authorityName="West" authorityYear="1992" box="[386,523,1377,1400]" class="Maxillopoda" family="Philichthyidae" genus="Colobomatus" kingdom="Animalia" order="Poecilostomatoida" pageId="17" pageNumber="578" phylum="Arthropoda" rank="species" species="creeveyae">
<emphasis id="3BE7EAD5FF973A0255DAFADA4A05FAEA" box="[386,523,1377,1400]" italics="true" pageId="17" pageNumber="578">C. creeveyae</emphasis>
</taxonomicName>
and
<taxonomicName id="CE934D44FF973A02561DFADA4AB6FAEA" authorityName="West" authorityYear="1992" box="[581,696,1377,1400]" class="Maxillopoda" family="Philichthyidae" genus="Colobomatus" kingdom="Animalia" order="Poecilostomatoida" pageId="17" pageNumber="578" phylum="Arthropoda" rank="species" species="ornatus">
<emphasis id="3BE7EAD5FF973A02561DFADA4AB6FAEA" box="[581,696,1377,1400]" italics="true" pageId="17" pageNumber="578">C. ornatus</emphasis>
</taxonomicName>
. However,
<taxonomicName id="CE934D44FF973A025760FADA4BACFAEB" authority="Madinabeitia &amp; Tang &amp; Nagasawa, 2013" authorityName="Madinabeitia &amp; Tang &amp; Nagasawa" authorityYear="2013" box="[824,930,1377,1401]" class="Maxillopoda" family="Philichthyidae" genus="Colobomatus" kingdom="Animalia" order="Poecilostomatoida" pageId="17" pageNumber="578" phylum="Arthropoda" rank="species" species="absens" status="sp. nov.">
<emphasis id="3BE7EAD5FF973A025760FADA4BACFAEB" box="[824,930,1377,1401]" italics="true" pageId="17" pageNumber="578">C. absens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="20D457AEFF973A0257F1FADA4BF1FAEA" box="[937,1023,1377,1400]" pageId="17" pageNumber="578" rank="species">sp. nov.</taxonomicNameLabel>
can be easily distinguished from these five species and all other congeners by having only one pair rather than two pairs of thoracic processes.
</paragraph>
<caption id="5DEC664FFF973A0254D7FB5F4C36FAA6" ID-DOI="http://doi.org/10.5281/zenodo.4632033" ID-Zenodo-Dep="4632033" httpUri="https://zenodo.org/record/4632033/files/figure.png" pageId="17" pageNumber="578" startId="17.[143,210,1252,1274]" targetBox="[143,1103,167,1219]" targetPageId="17">
<paragraph id="092C36C7FF973A0254D7FB5F4C36FAA6" blockId="17.[143,1104,1252,1332]" pageId="17" pageNumber="578">
Figure 6.
<taxonomicName id="CE934D44FF973A0254ADFB5F49B0FB6B" authority="Madinabeitia &amp; Tang &amp; Nagasawa, 2013" authorityName="Madinabeitia &amp; Tang &amp; Nagasawa" authorityYear="2013" box="[245,446,1252,1273]" class="Maxillopoda" family="Philichthyidae" genus="Colobomatus" kingdom="Animalia" order="Poecilostomatoida" pageId="17" pageNumber="578" phylum="Arthropoda" rank="species" species="absens" status="sp. nov.">
<emphasis id="3BE7EAD5FF973A0254ADFB5F49B0FB6B" box="[245,446,1252,1273]" italics="true" pageId="17" pageNumber="578">Colobomatus absens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="20D457AEFF973A02559AFB5F4A01FB6B" box="[450,527,1252,1273]" pageId="17" pageNumber="578" rank="species">sp. nov.</taxonomicNameLabel>
, adult female. (A) Habitus, ventral; (B) antennule, ventral; (C) buccal capsule and mouthparts, ventral; (D) leg 1, ventrolateral; (E) tip of thoracic process, ventral. Abbreviations: A, antenna; la, labium; mx, maxillule; Mx, maxilla; Mxp, maxilliped.
</paragraph>
</caption>
<paragraph id="092C36C7FF973A0254E2FA7B4963FA6A" blockId="17.[143,1103,1376,1528]" pageId="17" pageNumber="578">
This study represents the first report of a species of
<taxonomicName id="CE934D44FF973A025753FA7B4B93FA4A" authorityName="Hesse" authorityYear="1873" box="[779,925,1472,1496]" class="Maxillopoda" family="Philichthyidae" genus="Colobomatus" kingdom="Animalia" order="Poecilostomatoida" pageId="17" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="3BE7EAD5FF973A025753FA7B4B93FA4A" box="[779,925,1472,1496]" italics="true" pageId="17" pageNumber="578">Colobomatus</emphasis>
</taxonomicName>
parasitic in the family
<taxonomicName id="CE934D44FF973A025485FA5B4967FA6A" authorityName="Bonaparte" authorityYear="1831" box="[221,361,1504,1528]" class="Actinopterygii" family="Caesionidae" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="578" phylum="Chordata" rank="family">Caesionidae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>