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<document ID-DOI="10.5252/cryptogamie-bryologie2020v41a18" ID-ISSN="1776-0992" ID-Zenodo-Dep="7822194" checkinTime="1681310790419" checkinUser="felipe" docAuthor="Bruggeman-Nannenga, Maria A. &amp; Müller, Frank" docDate="2020" docId="752687AF8070E90CFEB0164CFA0F806A" docLanguage="en" docName="CryptogamieBryologie.41.18.229-238.pdf" docOrigin="Cryptogamie, Bryologie 20 (18)" docSource="http://dx.doi.org/10.11646/zootaxa.4878.3.2" docStyle="DocumentStyle:3F4A000F5B047ABA0EAB2943AA9373AE:CryptogamieBryologie.2019-.journal_article" docStyleId="3F4A000F5B047ABA0EAB2943AA9373AE" docStyleName="CryptogamieBryologie.2019-.journal_article" docTitle="Fissidens pseudoanomalus Bruggeman-Nannenga &amp; Müller 2020, sp. nov." docType="treatment" docVersion="2" lastPageNumber="233" masterDocId="891FFFD78076E90AFF8A1518FFA78606" masterDocTitle="A contribution to the Fissidens Hedw. (Musci, Fissidentaceae) of Myanmar, including F. strictidens sp. nov. and F. pseudoanomalus sp. nov." masterLastPageNumber="238" masterPageNumber="229" pageNumber="233" updateTime="1681311199742" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A contribution to the Fissidens Hedw. (Musci, Fissidentaceae) of Myanmar, including F. strictidens sp. nov. and F. pseudoanomalus sp. nov.</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bruggeman-Nannenga, Maria A.</mods:namePart>
<mods:affiliation>Griffensteijnseplein 23, NL- 3703 BE Zeist (The Netherlands)</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Müller, Frank</mods:namePart>
<mods:affiliation>TU Dresden, Institut für Botanik, D- 01062 Dresden (Germany) frank. mueller @ tu-dresden. de (corresponding author)</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Cryptogamie, Bryologie</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-11-10</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>20</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>18</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>229</mods:start>
<mods:end>238</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.4878.3.2</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5252/cryptogamie-bryologie2020v41a18</mods:identifier>
<mods:identifier type="ISSN">1776-0992</mods:identifier>
<mods:identifier type="Zenodo-Dep">7822194</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7822214" ID-Zenodo-Dep="7822214" LSID="urn:lsid:plazi:treatment:752687AF8070E90CFEB0164CFA0F806A" httpUri="http://treatment.plazi.org/id/752687AF8070E90CFEB0164CFA0F806A" lastPageNumber="233" pageId="6" pageNumber="233">
<subSubSection pageId="6" pageNumber="233" type="nomenclature">
<paragraph blockId="6.[247,660,852,911]" pageId="6" pageNumber="233">
<heading centered="true" fontSize="11" level="2" pageId="6" pageNumber="233" reason="4">
<taxonomicName authority="Brugg." authorityName="Bruggeman-Nannenga &amp; Müller" authorityYear="2020" class="Bryopsida" family="Fissidentaceae" genus="Fissidens" kingdom="Plantae" order="Fissidentales" pageId="6" pageNumber="233" phylum="Bryophyta" rank="species" species="pseudoanomalus">
<emphasis bold="true" box="[314,593,852,879]" italics="true" pageId="6" pageNumber="233">Fissidens pseudoanomalus</emphasis>
Brugg.
</taxonomicName>
-Nann. &amp; Frank Müll.,
<taxonomicNameLabel box="[578,660,885,911]" pageId="6" pageNumber="233" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[401,506,917,944]" pageId="6" pageNumber="233" type="description">
<paragraph blockId="6.[401,506,917,944]" box="[401,506,917,944]" pageId="6" pageNumber="233">
<heading box="[401,506,917,944]" centered="true" fontSize="11" level="3" pageId="6" pageNumber="233" reason="3">
(
<figureCitation box="[408,497,917,944]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="7.[132,143,1962,1980]" captionStartId-1="8.[132,143,1962,1980]" captionStartId-2="10.[132,143,1985,2003]" captionTargetBox-0="[151,1434,216,1934]" captionTargetBox-1="[189,1306,220,1917]" captionTargetBox-2="[383,1214,214,1954]" captionTargetId-0="figure-26@7.[701,1254,788,1362]" captionTargetId-1="figure-19@8.[189,1399,215,1923]" captionTargetId-2="figure-19@10.[383,1214,214,1964]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="10" captionText-0="FIG. 3. — Fissidens pseudoanomalus sp. nov.: A, leaf near insertion (left) and mid leaf (right); B, leaf apex; C, trans section of leaf above the vaginant lamina; D, margins of mid leaf. A, B, C, D (right and left) from holotype G. Miehe et al. 13-076-075-F, D (middle) from G. Miehe et al.13-093-202-B. Scale bars: A, 200 µm; B, 100 µm; C, D, 50 µm." captionText-1="FIG. 4. — Fissidens pseudoanomalus sp. nov.: A, perichaetial branches; B, stem with perichaetial branches; C, part of trans section of leaf above vaginant laminae showing the bistratose border zone; D, leaves. A, C (two top cross sections), D from holotype G. Miehe et al. 13-076-075-F; B, C (lowest cross section) from G. Miehe et al. 13-093-202-B. Scale bars: A, 100 µm; B, D, 1 mm; C, 50 µm." captionText-2="FIG. 5. — Fissidens pseudoanomalus sp. nov.: A, sporophyte; B, upper part of peristome teeth; C, median part of peristome teeth; D, exterior side of basal part of peristome teeth. A-D from holotype Miehe et al. 13-076-075-F. Scale bars: A, 1 mm; B-D, 50 µm." figureDoi-0="http://doi.org/10.5281/zenodo.7822204" figureDoi-1="http://doi.org/10.5281/zenodo.7822208" figureDoi-2="http://doi.org/10.5281/zenodo.7822212" httpUri-0="https://zenodo.org/record/7822204/files/figure.png" httpUri-1="https://zenodo.org/record/7822208/files/figure.png" httpUri-2="https://zenodo.org/record/7822212/files/figure.png" pageId="6" pageNumber="233">Figs 3-5</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="233" type="diagnosis">
<paragraph blockId="6.[132,777,974,1051]" pageId="6" pageNumber="233">
DIAGNOSIS. — The species closely resembles
<taxonomicName authority="Mont." authorityName="Mont." class="Bryopsida" family="Fissidentaceae" genus="Fissidens" kingdom="Plantae" order="Fissidentales" pageId="6" pageNumber="233" phylum="Bryophyta" rank="species" species="anomalus">
<emphasis box="[594,777,974,998]" italics="true" pageId="6" pageNumber="233">Fissidens anomalus</emphasis>
Mont.
</taxonomicName>
, but differs in the structure of the leaf border zone which is in part bistratose and contains elongated cells.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="233" type="materials_examined">
<paragraph blockId="6.[130,775,1074,1151]" pageId="6" pageNumber="233">
<materialsCitation collectingDate="2013-10-30" collectionCode="G, F, DR, L" country="Myanmar" elevation="2632" latitude="27.616608" location="Hponyin Razi" longLatPrecision="1" longitude="96.98285" municipality="Magnolia - Rhododendron forest" pageId="6" pageNumber="233" specimenCount="1" typeStatus="holotype">
<typeStatus box="[132,240,1075,1098]" pageId="6" pageNumber="233">HOLOTYPE</typeStatus>
. —
<collectingCountry box="[288,386,1075,1099]" name="Myanmar" pageId="6" pageNumber="233">Myanmar</collectingCountry>
,
<location LSID="urn:lsid:plazi:treatment:752687AF8070E90CFEB0164CFA0F806A:F85060628070E90CFE04112AFDB8824C" box="[398,543,1074,1098]" country="Myanmar" latitude="27.616608" longLatPrecision="1" longitude="96.98285" municipality="Magnolia - Rhododendron forest" name="Hponyin Razi" pageId="6" pageNumber="233">Hponyin Razi</location>
,
<emphasis italics="true" pageId="6" pageNumber="233">
<taxonomicName authorityName="Lendenfeld" authorityYear="1884" box="[557,666,1074,1098]" class="Magnoliopsida" family="Fagaceae" genus="Lithocarpus" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="233" phylum="Tracheophyta" rank="genus">Lithocarpus</taxonomicName>
-
<collectingMunicipality pageId="6" pageNumber="233">Magnolia - Rhododendron</collectingMunicipality>
</emphasis>
forest,
<quantity box="[327,402,1101,1125]" metricMagnitude="3" metricUnit="m" metricValue="2.632" pageId="6" pageNumber="233" unit="m" value="2632.0">
<elevation box="[327,402,1101,1125]" metricMagnitude="3" metricUnit="m" metricValue="2.632" pageId="6" pageNumber="233" unit="m" value="2632.0">2632 m</elevation>
</quantity>
,
<geoCoordinate box="[409,534,1101,1125]" degrees="27.616607" direction="north" orientation="latitude" pageId="6" pageNumber="233" precision="1" value="27.616608">27.616607N</geoCoordinate>
,
<geoCoordinate box="[541,661,1101,1125]" degrees="96.982848" direction="east" orientation="longitude" pageId="6" pageNumber="233" precision="1" value="96.98285">96.982848E</geoCoordinate>
,
<date box="[668,771,1101,1125]" pageId="6" pageNumber="233" value="2013-10-30">
<collectingDate box="[668,771,1101,1125]" pageId="6" pageNumber="233" value="2013-10-30">30.X.2013</collectingDate>
</date>
,
<emphasis box="[130,417,1127,1151]" italics="true" pageId="6" pageNumber="233">
<collectionCode box="[130,151,1128,1151]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="6" pageNumber="233" type="Herbarium">G</collectionCode>
. Miehe et al. 13-076-075-
<collectionCode box="[400,417,1127,1151]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="6" pageNumber="233" type="Herbarium">F</collectionCode>
</emphasis>
(holo-,
<collectionCode box="[498,536,1128,1151]" country="Germany" lsid="urn:lsid:biocol.org:col:15173" name="Technische Universität Dresden" pageId="6" pageNumber="233" type="Herbarium">DR</collectionCode>
; iso-,
<collectionCode box="[595,612,1127,1151]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="6" pageNumber="233" type="Herbarium">L</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph blockId="6.[132,777,1175,1358]" pageId="6" pageNumber="233">
FURTHER SPECIMENS EXAMINED (
<typeStatus box="[454,546,1175,1199]" pageId="6" pageNumber="233">paratypes</typeStatus>
). —
<collectingCountry box="[598,695,1175,1199]" name="Myanmar" pageId="6" pageNumber="233">Myanmar</collectingCountry>
,
<collectingRegion country="Myanmar" name="Kachin" pageId="6" pageNumber="233">Kachin State</collectingRegion>
, Hponkan Razi, evergreen broadleaved forest,
<quantity box="[685,770,1201,1225]" metricMagnitude="3" metricUnit="m" metricValue="1.3599999999999999" pageId="6" pageNumber="233" unit="m" value="1360.0">1360 m</quantity>
,
<geoCoordinate box="[132,250,1228,1252]" degrees="27.58421" direction="north" orientation="latitude" pageId="6" pageNumber="233" precision="1" value="27.58421">27.58421N</geoCoordinate>
,
<geoCoordinate box="[259,371,1228,1252]" degrees="97.10447" direction="east" orientation="longitude" pageId="6" pageNumber="233" precision="1" value="97.10447">97.10447E</geoCoordinate>
,
<date box="[381,488,1228,1252]" pageId="6" pageNumber="233" value="2014-10-24">24.X.2014</date>
,
<emphasis italics="true" pageId="6" pageNumber="233">
<collectionCode box="[497,509,1228,1251]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="6" pageNumber="233" type="Herbarium">J</collectionCode>
. Kluge &amp;
<collectionCode box="[601,616,1228,1251]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="6" pageNumber="233" type="Herbarium">P</collectionCode>
.
<collectionCode box="[619,637,1228,1251]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="6" pageNumber="233" type="Herbarium">K</collectionCode>
. Kine 14-050- 12e-A
</emphasis>
(
<collectionCode box="[203,242,1254,1278]" country="Germany" lsid="urn:lsid:biocol.org:col:15173" name="Technische Universität Dresden" pageId="6" pageNumber="233" type="Herbarium">DR</collectionCode>
, private herbarium Bruggeman-Nannenga);
<collectingCountry box="[674,771,1255,1279]" name="Myanmar" pageId="6" pageNumber="233">Myanmar</collectingCountry>
,
<collectingRegion box="[132,259,1281,1305]" country="Myanmar" name="Kachin" pageId="6" pageNumber="233">Kachin State</collectingRegion>
, Hponkan Razi,
<emphasis box="[428,601,1281,1305]" italics="true" pageId="6" pageNumber="233">
<taxonomicName box="[428,503,1281,1304]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="233" phylum="Tracheophyta" rank="genus">Quercus</taxonomicName>
-Magnolia
</emphasis>
-
<taxonomicName box="[608,708,1281,1305]" class="Magnoliopsida" family="Araliaceae" kingdom="Plantae" order="Apiales" pageId="6" pageNumber="233" phylum="Tracheophyta" rank="family">Araliaceae</taxonomicName>
forest,
<quantity box="[133,213,1307,1331]" metricMagnitude="3" metricUnit="m" metricValue="1.833" pageId="6" pageNumber="233" unit="m" value="1833.0">1833 m</quantity>
,
<geoCoordinate box="[225,355,1307,1331]" degrees="27.603756" direction="north" orientation="latitude" pageId="6" pageNumber="233" precision="1" value="27.603756">27.603756N</geoCoordinate>
,
<geoCoordinate box="[366,491,1307,1331]" degrees="96.988924" direction="east" orientation="longitude" pageId="6" pageNumber="233" precision="1" value="96.98892">96.988924E</geoCoordinate>
,
<date box="[503,619,1307,1331]" pageId="6" pageNumber="233" value="2013-11-16">16.XI.2013</date>
,
<emphasis italics="true" pageId="6" pageNumber="233">
<collectionCode box="[631,652,1308,1331]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="6" pageNumber="233" type="Herbarium">G</collectionCode>
. Miehe et al. 13-093-202-
<collectionCode box="[257,273,1334,1358]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="6" pageNumber="233" type="Herbarium">B</collectionCode>
</emphasis>
(
<collectionCode box="[286,325,1334,1358]" country="Germany" lsid="urn:lsid:biocol.org:col:15173" name="Technische Universität Dresden" pageId="6" pageNumber="233" type="Herbarium">DR</collectionCode>
, private herbarium Bruggeman-Nannenga).
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="233" type="description">
<paragraph blockId="6.[132,277,1396,1422]" box="[132,277,1396,1422]" pageId="6" pageNumber="233">
<heading box="[132,277,1396,1422]" fontSize="9" level="3" pageId="6" pageNumber="233" reason="5">DESCRIPTION</heading>
</paragraph>
<paragraph blockId="6.[131,775,1427,1517]" box="[132,190,1427,1453]" pageId="6" pageNumber="233">
<heading box="[132,190,1427,1453]" fontSize="11" level="3" pageId="6" pageNumber="233" reason="9">
<emphasis box="[132,190,1427,1453]" italics="true" pageId="6" pageNumber="233">Stems</emphasis>
</heading>
</paragraph>
<paragraph blockId="6.[131,775,1427,1517]" pageId="6" pageNumber="233">
Up to
<quantity box="[200,278,1459,1486]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="233" unit="mm" value="35.0">35 mm</quantity>
long and up to
<quantity box="[446,510,1459,1485]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="233" unit="mm" value="8.0">8 mm</quantity>
wide, pinnately foliated, sparsely to heavily branched, central strand well developed.
</paragraph>
<paragraph blockId="6.[132,220,1554,1580]" box="[132,220,1554,1580]" pageId="6" pageNumber="233">
<heading box="[132,220,1554,1580]" fontSize="11" level="3" pageId="6" pageNumber="233" reason="9">
<emphasis box="[132,220,1554,1580]" italics="true" pageId="6" pageNumber="233">Rhizoids</emphasis>
</heading>
</paragraph>
<paragraph blockId="6.[132,607,1586,1612]" box="[132,607,1586,1612]" pageId="6" pageNumber="233">Dark red-brown, heavily branched, smooth.</paragraph>
<paragraph blockId="6.[132,298,1650,1676]" box="[132,298,1650,1676]" pageId="6" pageNumber="233">
<heading box="[132,298,1650,1676]" fontSize="11" level="3" pageId="6" pageNumber="233" reason="9">
<emphasis box="[132,298,1650,1676]" italics="true" pageId="6" pageNumber="233">Axillary nodules</emphasis>
</heading>
</paragraph>
<paragraph blockId="6.[132,335,1682,1709]" box="[132,335,1682,1709]" pageId="6" pageNumber="233">Not differentiated.</paragraph>
<paragraph blockId="6.[132,198,1747,1772]" box="[132,198,1747,1772]" pageId="6" pageNumber="233">
<heading box="[132,198,1747,1772]" fontSize="11" level="3" pageId="6" pageNumber="233" reason="9">
<emphasis box="[132,198,1747,1772]" italics="true" pageId="6" pageNumber="233">Leaves</emphasis>
</heading>
</paragraph>
<paragraph blockId="6.[130,776,1778,2027]" pageId="6" pageNumber="233">
Close to distant, up to 30 pairs, crispate when dry, lanceolate, often falcate, with acute-acuminate apex, 2.4-5.6 ×
<quantity metricMagnitude="-4" metricUnit="m" metricValue="6.75" metricValueMax="9.5" metricValueMin="4.0" pageId="6" pageNumber="233" unit="mm" value="0.675" valueMax="0.95" valueMin="0.4">0.4- 0.95 mm</quantity>
, 5-6 times as long as wide, margin coarsely serrate on dorsal and apical lamina, on the vaginant laminae serrulate with a few distant larger teeth; apical and dorsal lamina with pale, thickened, 3-5 cells wide, uni- to bistratose border of isodiametric and partly elongated cells, margin of vaginant lamina unistratose, hardly differentiated; vaginant laminae
</paragraph>
<paragraph blockId="6.[811,1455,214,306]" pageId="6" pageNumber="233">½-ª/5 the leaf length, rounded at insertion, unistratose, slightly unequal; dorsal lamina rounded-truncate below, reaching the insertion, not decurrent; dorsal and apical lamina unistratose.</paragraph>
<paragraph blockId="6.[811,868,343,369]" box="[811,868,343,369]" pageId="6" pageNumber="233">
<heading box="[811,868,343,369]" fontSize="11" level="3" pageId="6" pageNumber="233" reason="9">
<emphasis box="[811,868,343,369]" italics="true" pageId="6" pageNumber="233">
<collectingRegion box="[811,868,343,369]" country="New Zealand" name="West Coast" pageId="6" pageNumber="233">Costa</collectingRegion>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="6.[813,1325,375,401]" box="[813,1325,375,401]" pageId="6" pageNumber="233">Long excurrent, in cross-section taxifolius-type.</paragraph>
<paragraph blockId="6.[813,896,439,464]" box="[813,896,439,464]" pageId="6" pageNumber="233">
<heading box="[813,896,439,464]" fontSize="11" level="3" pageId="6" pageNumber="233" reason="9">
<emphasis box="[813,896,439,464]" italics="true" pageId="6" pageNumber="233">Lumina</emphasis>
</heading>
</paragraph>
<paragraph blockId="6.[810,1456,470,625]" pageId="6" pageNumber="233">Of mid dorsal laminal cells (6.5-)9.5-10.5(-13.0) × (5-)6.5- 7.5(-9) µm, walls smooth, convex, thin, 1-1.5 µm wide; lumina of mid vaginant laminal cells (8.5-)10-10.7(-12.1) × (5-)6- 6.9(-8.2) µm; walls 2-3 µm wide, smooth; marginal cells of vaginant lamina with more incrassate, 3-4(-5) µm wide walls.</paragraph>
<paragraph blockId="6.[811,903,662,688]" box="[811,903,662,688]" pageId="6" pageNumber="233">
<heading box="[811,903,662,688]" fontSize="11" level="3" pageId="6" pageNumber="233" reason="9">
<emphasis box="[811,903,662,688]" italics="true" pageId="6" pageNumber="233">Gemmae</emphasis>
</heading>
</paragraph>
<paragraph blockId="6.[813,966,693,720]" box="[813,966,693,720]" pageId="6" pageNumber="233">Not observed.</paragraph>
<paragraph blockId="6.[813,935,757,783]" box="[813,935,757,783]" pageId="6" pageNumber="233">
<heading box="[813,935,757,783]" fontSize="11" level="3" pageId="6" pageNumber="233" reason="9">
<emphasis box="[813,935,757,783]" italics="true" pageId="6" pageNumber="233">Fertile parts</emphasis>
</heading>
</paragraph>
<paragraph blockId="6.[811,1457,789,1071]" pageId="6" pageNumber="233">
Perigonia not observed. Perichaetia terminal on axillary, short,
<quantity box="[811,944,821,848]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.5" metricValueMin="1.6" pageId="6" pageNumber="233" unit="mm" value="2.05" valueMax="2.5" valueMin="1.6">1.6-2.5 mm</quantity>
long branches, archegonia 350-450 µm long, perichaetial leaves
<quantity box="[1014,1145,853,880]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" metricValueMax="2.1" metricValueMin="1.5" pageId="6" pageNumber="233" unit="mm" value="1.8" valueMax="2.1" valueMin="1.5">1.5-2.1 mm</quantity>
long; calyptra not observed. Sporophyte, seta
<quantity box="[1003,1070,885,911]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="233" unit="mm" value="3.0">3 mm</quantity>
long, smooth, 1 per perichaetium; capsule erect, 1.5 × 1.0 mm, with ± 94 columns of oblong exothecial cells with thick vertical walls, with stomata; peristome taxifolius-type, teeth deeply divided, ± 500 µm long, tooth-base 42-51 µm wide; operculum not observed; spores subglobose, 18.5-22(-25) × 16-20 µm, papillose.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="233" type="discussion">
<paragraph blockId="6.[813,942,1109,1135]" box="[813,942,1109,1135]" pageId="6" pageNumber="233">
<heading box="[813,942,1109,1135]" fontSize="9" level="3" pageId="6" pageNumber="233" reason="5">DISCUSSION</heading>
</paragraph>
<paragraph blockId="6.[810,1457,1140,1644]" pageId="6" pageNumber="233">
The new species is easily recognized by its large size, the elimbate leaves with a pale, partly bistratose border-zone with elongated cells, coarsely serrate margins and lateral sporophytes with short setae and the reduced taxifolius-type peristome. It is closely resembles
<taxonomicName authority="Mont." authorityName="Mont." box="[925,1129,1267,1294]" class="Bryopsida" family="Fissidentaceae" genus="Fissidens" kingdom="Plantae" order="Fissidentales" pageId="6" pageNumber="233" phylum="Bryophyta" rank="species" species="anomalus">
<emphasis box="[925,1051,1267,1293]" italics="true" pageId="6" pageNumber="233">F. anomalus</emphasis>
Mont.
</taxonomicName>
that has a similar sporophyte and a similar peristome. The two differ in the structure of the border zone. That of the new species is in part bistratose and contains elongated cells next to isodiametric cells. In
<taxonomicName box="[815,938,1395,1421]" class="Bryopsida" family="Fissidentaceae" genus="Fissidens" kingdom="Plantae" order="Fissidentales" pageId="6" pageNumber="233" phylum="Bryophyta" rank="species" species="anomalus">
<emphasis box="[815,938,1395,1421]" italics="true" pageId="6" pageNumber="233">F. anomalus</emphasis>
</taxonomicName>
the border zone is unistratose and composed of isodiametric cells. One of the
<typeStatus box="[1130,1233,1428,1454]" pageId="6" pageNumber="233">paratypes</typeStatus>
,
<emphasis italics="true" pageId="6" pageNumber="233">Miehe et al. 13-093- 202-B</emphasis>
, differs from the other two in having bistratose areas in the dorsal lamina (
<figureCitation box="[1003,1082,1491,1517]" captionStart="FIG" captionStartId="8.[132,143,1962,1980]" captionTargetBox="[189,1306,220,1917]" captionTargetId="figure-19@8.[189,1399,215,1923]" captionTargetPageId="8" captionText="FIG. 4. — Fissidens pseudoanomalus sp. nov.: A, perichaetial branches; B, stem with perichaetial branches; C, part of trans section of leaf above vaginant laminae showing the bistratose border zone; D, leaves. A, C (two top cross sections), D from holotype G. Miehe et al. 13-076-075-F; B, C (lowest cross section) from G. Miehe et al. 13-093-202-B. Scale bars: A, 100 µm; B, D, 1 mm; C, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.7822208" httpUri="https://zenodo.org/record/7822208/files/figure.png" pageId="6" pageNumber="233">Fig. 4C</figureCitation>
lowest cross section). In this respect it resembles
<taxonomicName authority="P. Beauv." authorityName="P. Beauv." box="[943,1131,1523,1549]" class="Bryopsida" family="Fissidentaceae" genus="Fissidens" kingdom="Plantae" order="Fissidentales" pageId="6" pageNumber="233" phylum="Bryophyta" rank="species" species="dubius">
<emphasis box="[943,1036,1523,1549]" italics="true" pageId="6" pageNumber="233">F. dubius</emphasis>
P.Beauv.
</taxonomicName>
and it will key out as that species (e.g.,
<bibRefCitation author="LI Z. - H. &amp; IWATSUKI Z." box="[918,1130,1554,1581]" pageId="6" pageNumber="233" refId="ref4809" refString="LI Z. - H. &amp; IWATSUKI Z. 2001. - Fissidentaceae, in LI X. - J. &amp; CROSBY M. R. (eds), Moss Flora of China. Volume 2. Beijing &amp; New York, Science Press &amp; St. Louis, Missouri Botanical Garden, 283 p." type="book" year="2001">Li &amp; Iwatsuki 2001</bibRefCitation>
). It can be distinguished from
<taxonomicName box="[813,905,1586,1612]" class="Bryopsida" family="Fissidentaceae" genus="Fissidens" kingdom="Plantae" order="Fissidentales" pageId="6" pageNumber="233" phylum="Bryophyta" rank="species" species="dubius">
<emphasis box="[813,905,1586,1612]" italics="true" pageId="6" pageNumber="233">F. dubius</emphasis>
</taxonomicName>
by the structure of the border zone. That of
<taxonomicName box="[1362,1454,1586,1612]" class="Bryopsida" family="Fissidentaceae" genus="Fissidens" kingdom="Plantae" order="Fissidentales" pageId="6" pageNumber="233" phylum="Bryophyta" rank="species" species="dubius">
<emphasis box="[1362,1454,1586,1612]" italics="true" pageId="6" pageNumber="233">F. dubius</emphasis>
</taxonomicName>
is unistratose and is composed entirely of isodiametric cells.
</paragraph>
</subSubSection>
</treatment>
</document>
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