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<document id="44B4F0C1E626AFAE2C5567839EC862DE" ID-CLB-Dataset="273743" ID-DOI="10.11646/zootaxa.5356.1.1" ID-GBIF-Dataset="38e48899-447b-40cc-b9bd-e84bb0ac221e" ID-ISSN="1175-5334" ID-Zenodo-Dep="10008529" ID-ZooBank="F3CE1E00-45BB-44C8-8911-1A355BFD223C" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1697461253574" checkinUser="plazi" docAuthor="Vilela, Diogo Silva, Lencioni, Frederico A. A., Bota-Sierra, Cornelio A., Ware, Jessica L. &amp; Bispo, Pitágoras C." docDate="2023" docId="73444D3AFFFD91156AD7F8ACA98AA329" docLanguage="en" docName="zootaxa.5356.1.1.pdf" docOrigin="Zootaxa 5356 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5356.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Heteragrion archon : De Marmels 2008" docType="treatment" docVersion="9" lastPageNumber="23" masterDocId="8F7D3542FFE891036A40FFD4AB77A114" masterDocTitle="Taxonomic revision of the Neotropical genus Heteragrion Selys, 1862 (Zygoptera: Heteragrionidae): male morphology, new species and illustrated key" masterLastPageNumber="96" masterPageNumber="1" pageNumber="22" updateTime="1699697007752" updateUser="plazi">
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<mods:title id="F5FE137B087A5D5C2FDA64083CB93E8F">Taxonomic revision of the Neotropical genus Heteragrion Selys, 1862 (Zygoptera: Heteragrionidae): male morphology, new species and illustrated key</mods:title>
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<mods:namePart id="2536E1E353C58C6E206229D646CB8A6F">Vilela, Diogo Silva</mods:namePart>
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<mods:namePart id="9CB2914C97149E49EEFEC3F6A756F4BB">Lencioni, Frederico A. A.</mods:namePart>
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<mods:affiliation id="C1AA1C5EDEC9A8EF177D40F47103487D">Rua Anibal, 216, Jardim Coleginho, CEP 12310 - 780, Jacareí, SP, Brazil lencioni. odonata @ gmail. com; https: // orcid. org / 0000 - 0002 - 3912 - 9181</mods:affiliation>
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<mods:namePart id="0072E9346DFB09FB8140B136EDBD7CB6">Bota-Sierra, Cornelio A.</mods:namePart>
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<mods:affiliation id="434DDCB6C002AE662745208780768563">Grupo de Entomología Universidad de Antioquia (GEUA), Universidad de Antioquia, Medellin 50010, Colombia corneliobota @ gmail. com; https: // orcid. org / 0000 - 0002 - 6555 - 7913</mods:affiliation>
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<mods:namePart id="2850B31DCEA1E5C526CE60BFC22345F8">Ware, Jessica L.</mods:namePart>
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<mods:affiliation id="842D43A47DFC0181C0816D16842D0DFB">Division of Invertebrate Zoology, American Museum of Natural History, USA jware @ amnh. org; https: // orcid. org / 0000 - 0002 - 4066 - 7681</mods:affiliation>
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<mods:affiliation id="4686BDC89FFEE40D122CF8A4704328C8">Laboratório de Biologia Aquática, Departamento de Ciências Biológicas, Faculdade de Ciências e Letras de Assis, Universidade Estadual Paulista, Assis, SP, Brazil deeogoo @ gmail. com; https: // orcid. org / 0000 - 0001 - 6510 - 7018 &amp; Laboratório de Biologia Aquática, Departamento de Ciências Biológicas, Faculdade de Ciências e Letras de Assis, Universidade Estadual Paulista, Assis, SP, Brazil deeogoo @ gmail. com; https: // orcid. org / 0000 - 0001 - 6510 - 7018 &amp; pitagoras. bispo @ unesp. br; https: // orcid. org / 0000 - 0002 - 7356 - 8882 * Corresponding author</mods:affiliation>
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<treatment id="73444D3AFFFD91156AD7F8ACA98AA329" ID-DOI="http://doi.org/10.5281/zenodo.10008282" ID-GBIF-Taxon="214231528" ID-Zenodo-Dep="10008282" LSID="urn:lsid:plazi:treatment:73444D3AFFFD91156AD7F8ACA98AA329" httpUri="http://treatment.plazi.org/id/73444D3AFFFD91156AD7F8ACA98AA329" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<subSubSection id="B3F7AFA7FFFD91166AD7F8ACA903A686" box="[151,628,1911,1938]" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="FB52FC2CFFFD91166AD7F8ACA903A686" blockId="21.[151,628,1911,1974]" box="[151,628,1911,1938]" pageId="21" pageNumber="22">
<heading id="A01A4B40FFFD91166AD7F8ACA903A686" bold="true" box="[151,628,1911,1938]" fontSize="11" level="2" pageId="21" pageNumber="22" reason="4">
<taxonomicName id="3CED87AFFFFD91166AD7F8ACA903A686" ID-CoL="3L73P" authority="De Marmels, 2008" authorityName=": De Marmels" authorityYear="2008" box="[151,628,1911,1938]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="archon">
<emphasis id="C999203EFFFD91166AD7F8ACA903A686" bold="true" box="[151,628,1911,1938]" pageId="21" pageNumber="22">
<emphasis id="C999203EFFFD91166AD7F8ACAAFEA686" bold="true" box="[151,393,1911,1938]" italics="true" pageId="21" pageNumber="22">Heteragrion archon</emphasis>
<bibRefCitation id="9F7C81DDFFFD91166BD1F8ACA903A686" author="De Marmels, J." box="[401,628,1912,1938]" pageId="21" pageNumber="22" pagination="151 - 155" refId="ref61591" refString="De Marmels, J. (2008) Heteragrion archon spec. nov. from the coastal Cordillera of Venezuela (Zygoptera: Megapodagrionidae). Odonatologica, 37 (2), 151 - 155." type="journal article" year="2008">De Marmels, 2008</bibRefCitation>
</emphasis>
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</paragraph>
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<subSubSection id="B3F7AFA7FFFD91156AD7F848A8B4A009" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="description">
<paragraph id="FB52FC2CFFFD91166AD7F848AA5CA6A2" blockId="21.[151,628,1911,1974]" box="[151,299,1948,1974]" pageId="21" pageNumber="22">
(
<figureCitation id="63D6E0A9FFFD91166ADFF848AA53A6A2" box="[159,292,1948,1974]" captionStart="FIGURE 11" captionStartId="22.[151,250,1612,1636]" captionTargetBox="[243,1347,610,1575]" captionTargetId="figure-177@22.[230,1358,597,1588]" captionTargetPageId="22" captionText="FIGURE 11. Heteragrion archon: (a) anterior view of head; (b) lateral view of pterothorax; (c) lateral view of anal appendages; (d) mediodorsal view of anal appendages; (ab) adapted from the original drawings by De Marmels (2008). Figures cd provided by Rosser Garrison and Natalia von Ellenrieder." figureDoi="http://doi.org/10.5281/zenodo.10008565" httpUri="https://zenodo.org/record/10008565/files/figure.png" pageId="21" pageNumber="22">Figs. 11ad</figureCitation>
)
</paragraph>
<paragraph id="FB52FC2CFFFD91156AD7F835A8B4A009" blockId="21.[151,657,2016,2041]" lastBlockId="22.[151,1437,150,573]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
[Key locator: Key to Group B species, couplet 4]
<taxonomicName id="3CED87AFFFFE91156AD7FF43A9BAA1A5" authority=": De Marmels 2008: 151 - 155" authorityName=": De Marmels" authorityPageNumber="151 - 155" authorityYear="2008" box="[151,717,150,177]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="archon">
<emphasis id="C999203EFFFE91156AD7FF43AA0EA1A5" box="[151,377,151,177]" italics="true" pageId="22" pageNumber="23">Heteragrion archon</emphasis>
:
<bibRefCitation id="9F7C81DDFFFE91156BCBFF43A9BAA1A5" author="De Marmels, J." box="[395,717,150,177]" pageId="22" pageNumber="23" pagination="151 - 155" refId="ref61591" refString="De Marmels, J. (2008) Heteragrion archon spec. nov. from the coastal Cordillera of Venezuela (Zygoptera: Megapodagrionidae). Odonatologica, 37 (2), 151 - 155." type="journal article" year="2008">De Marmels 2008: 151155</bibRefCitation>
</taxonomicName>
(description of ♁
<typeStatus id="2456428EFFFE911569E5FF43AF70A1A5" box="[933,1031,151,177]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
from
<collectingCountry id="83FABCBCFFFE91156E0FFF43AFB1A1A5" box="[1103,1222,151,177]" name="Venezuela" pageId="22" pageNumber="23">Venezuela</collectingCountry>
; comparisons with
<taxonomicName id="3CED87AFFFFE91156AD7FF6FAA41A1C1" authorityName=": Hartung" authorityYear="2002" box="[151,310,187,213]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="palmichale">
<emphasis id="C999203EFFFE91156AD7FF6FAA41A1C1" box="[151,310,187,213]" italics="true" pageId="22" pageNumber="23">H. palmichale</emphasis>
</taxonomicName>
);—
<bibRefCitation id="9F7C81DDFFFE91156B22FF6FA9FDA1C1" author="von Ellenrieder, N." box="[354,650,187,213]" pageId="22" pageNumber="23" pagination="58 - 71" refId="ref66897" refString="https: // doi. org / 10.2305 / IUCN. UK. 2021 - 1. RLTS. T 111006521 A 111006523. en von Ellenrieder, N. (2009 a) Databasing dragonflies: State of knowledge in the Neotropical region. Agrion, 13 (2), 58 - 71." type="journal article" year="2009">von Ellenrieder 2009a: 63</bibRefCitation>
(listed among Neotropical species described between 20042009);—
<bibRefCitation id="9F7C81DDFFFE91156AD7FF0BAAD4A1ED" author="Garrison, R. W. &amp; von Ellenrieder, N. &amp; Louton, J. A." box="[151,419,223,249]" pageId="22" pageNumber="23" refId="ref62214" refString="Garrison, R. W., von Ellenrieder, N. &amp; Louton, J. A. (2010) Damselfly genera of the New World. An illustrated and Annotated Key to the Zygoptera. The Johns Hopkins University Press, Baltimore, Maryland, xiv + 490 pp." type="book" year="2010">
Garrison
<emphasis id="C999203EFFFE91156B40FF0BAA46A1ED" box="[256,305,223,249]" italics="true" pageId="22" pageNumber="23">et al</emphasis>
. 2010: 88
</bibRefCitation>
, 90 (list of
<taxonomicName id="3CED87AFFFFE91156862FF0BA87FA1ED" box="[546,776,223,249]" pageId="22" pageNumber="23">
<emphasis id="C999203EFFFE91156862FF0BA9DBA1EC" box="[546,684,223,248]" italics="true" pageId="22" pageNumber="23">Heteragrion</emphasis>
species
</taxonomicName>
; figs. 383384);—
<bibRefCitation id="9F7C81DDFFFE91156996FF0BAEE0A1ED" author="Vivas-Santeliz, J. &amp; De Marmels, J." box="[982,1431,222,249]" pageId="22" pageNumber="23" pagination="35 - 54" refId="ref66738" refString="Vivas-Santeliz, J. &amp; De Marmels, J. (2017) Current knowledge of Odonata in Venezuela: Diversity and distribution of endemic species. Odonatologica, 46 (1), 35 - 54." type="journal article" year="2017">Vivas-Santeliz &amp; De Marmels 2017: 45</bibRefCitation>
, 50 (listed among endemic Venezuelan species being assessed by IUCN).
</paragraph>
</subSubSection>
<subSubSection id="B3F7AFA7FFFE91156A87FEF2A862A09D" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="FB52FC2CFFFE91156A87FEF2AB99A071" blockId="22.[151,1437,150,573]" pageId="22" pageNumber="23">
<emphasis id="C999203EFFFE91156A87FEF2AAC3A054" bold="true" box="[199,436,294,320]" pageId="22" pageNumber="23">Material examined.</emphasis>
Known only from type material; examined through original description and
<typeStatus id="2456428EFFFE91156F7AFEF3AEEBA055" box="[1338,1436,295,321]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
images.
</paragraph>
<paragraph id="FB52FC2CFFFE91156A87FEBBA862A09D" blockId="22.[151,1437,150,573]" box="[199,789,366,393]" pageId="22" pageNumber="23">
<emphasis id="C999203EFFFE91156A87FEBBAACDA09C" bold="true" box="[199,442,366,393]" pageId="22" pageNumber="23">Known distribution.</emphasis>
<collectingCountry id="83FABCBCFFFE91156B80FEBBA944A09D" box="[448,563,367,393]" name="Venezuela" pageId="22" pageNumber="23">Venezuela</collectingCountry>
(coastal cordillera).
</paragraph>
</subSubSection>
<subSubSection id="B3F7AFA7FFFE91156A87FE46A98AA329" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="FB52FC2CFFFE91156A87FE46A98AA329" blockId="22.[151,1437,150,573]" pageId="22" pageNumber="23">
<emphasis id="C999203EFFFE91156A87FE46AA90A0B9" bold="true" box="[199,487,402,429]" pageId="22" pageNumber="23">Diagnosis and remarks.</emphasis>
See the morphological comments on this species in the section of
<taxonomicName id="3CED87AFFFFE91156EB4FE47AEE2A0B9" authorityName=": Hartung" authorityYear="2002" box="[1268,1429,403,429]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="palmichale">
<emphasis id="C999203EFFFE91156EB4FE47AEE2A0B9" box="[1268,1429,403,429]" italics="true" pageId="22" pageNumber="23">H. palmichale</emphasis>
</taxonomicName>
.
<taxonomicName id="3CED87AFFFFE91156AD7FE63AA0EA0C5" authorityName=": De Marmels" authorityYear="2008" box="[151,377,439,465]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="archon">
<emphasis id="C999203EFFFE91156AD7FE63AA0EA0C5" box="[151,377,439,465]" italics="true" pageId="22" pageNumber="23">Heteragrion archon</emphasis>
</taxonomicName>
was assessed as CR by the IUCN Red List, as “recent visits to its
<typeStatus id="2456428EFFFE91156EC5FE63AFC2A0C5" box="[1157,1205,439,465]" pageId="22" pageNumber="23">type</typeStatus>
locality and similar habitats in the surrounding region have failed to find this species, which is threatened by tourism and fires” (
<bibRefCitation id="9F7C81DDFFFE91156F11FE0FAA08A30D" author="Vivas-Santeliz, J. J. &amp; De Marmels, J. &amp; Bota-Sierra, C. A." pageId="22" pageNumber="23" pagination="111006505" refId="ref66776" refString="Vivas-Santeliz, J. J., De Marmels, J., Sandoval-H, J. &amp; Bota-Sierra, C. A. (2021 a) Heteragrion archon. The IUCN Red List of Threatened Species, 2021, e. T 111006503 A 111006505. https: // doi. org / 10.2305 / IUCN. UK. 2021 - 1. RLTS. T 111006503 A 111006505. en" type="journal article" year="2021">
Vivas-Santeliz
<emphasis id="C999203EFFFE91156AB8FE2BAA47A30D" box="[248,304,511,537]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
2021a
</bibRefCitation>
). We hope that additional surveys are conducted within the
<typeStatus id="2456428EFFFE91156E56FE2BAF31A30D" box="[1046,1094,511,537]" pageId="22" pageNumber="23">type</typeStatus>
locality and the adjacent areas to assess the present conservation status of the species.
</paragraph>
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