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<document id="F1EF1DA4E6DAE7AEAB07C826BEB2ABEE" ID-CLB-Dataset="273743" ID-DOI="10.11646/zootaxa.5356.1.1" ID-GBIF-Dataset="38e48899-447b-40cc-b9bd-e84bb0ac221e" ID-ISSN="1175-5334" ID-Zenodo-Dep="10008529" ID-ZooBank="F3CE1E00-45BB-44C8-8911-1A355BFD223C" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1697461253574" checkinUser="plazi" docAuthor="Vilela, Diogo Silva, Lencioni, Frederico A. A., Bota-Sierra, Cornelio A., Ware, Jessica L. &amp; Bispo, Pitágoras C." docDate="2023" docId="73444D3AFFF4911E6AD7FB44AE4FA6F3" docLanguage="en" docName="zootaxa.5356.1.1.pdf" docOrigin="Zootaxa 5356 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5356.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Heteragrion bickorum Daigle 2005" docType="treatment" docVersion="9" lastPageNumber="30" masterDocId="8F7D3542FFE891036A40FFD4AB77A114" masterDocTitle="Taxonomic revision of the Neotropical genus Heteragrion Selys, 1862 (Zygoptera: Heteragrionidae): male morphology, new species and illustrated key" masterLastPageNumber="96" masterPageNumber="1" pageNumber="29" updateTime="1699697007752" updateUser="plazi">
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<mods:title id="246B878D9ECF80E99BE47E383A4229B2">Taxonomic revision of the Neotropical genus Heteragrion Selys, 1862 (Zygoptera: Heteragrionidae): male morphology, new species and illustrated key</mods:title>
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<mods:namePart id="A4A92D62EB86FF515AFAD5CA15E8219D">Vilela, Diogo Silva</mods:namePart>
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<mods:affiliation id="1DA9FC1ACBC795D47502848A88E04EC0">Laboratório de Biologia Aquática, Departamento de Ciências Biológicas, Faculdade de Ciências e Letras de Assis, Universidade Estadual Paulista, Assis, SP, Brazil deeogoo @ gmail. com; https: // orcid. org / 0000 - 0001 - 6510 - 7018</mods:affiliation>
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<mods:namePart id="CCE05C464599A5593CEEC0B1075698AE">Lencioni, Frederico A. A.</mods:namePart>
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<mods:affiliation id="33104E14D8FB6D0399F17E1D0B37E688">Rua Anibal, 216, Jardim Coleginho, CEP 12310 - 780, Jacareí, SP, Brazil lencioni. odonata @ gmail. com; https: // orcid. org / 0000 - 0002 - 3912 - 9181</mods:affiliation>
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<mods:namePart id="81ED59FF90D7424234D672BF3C23C3D0">Bota-Sierra, Cornelio A.</mods:namePart>
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<mods:affiliation id="934763117232E942DF1E84BDC44DEB29">Grupo de Entomología Universidad de Antioquia (GEUA), Universidad de Antioquia, Medellin 50010, Colombia corneliobota @ gmail. com; https: // orcid. org / 0000 - 0002 - 6555 - 7913</mods:affiliation>
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<mods:namePart id="62AC6C60A66CC474FC9A078102ACD8DB">Ware, Jessica L.</mods:namePart>
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<mods:affiliation id="E37C1C7C8CB29F7F95F3908D7D9DBB94">Division of Invertebrate Zoology, American Museum of Natural History, USA jware @ amnh. org; https: // orcid. org / 0000 - 0002 - 4066 - 7681</mods:affiliation>
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<mods:namePart id="C10C54B5996C628FC0F7325E676B8559">Bispo, Pitágoras C.</mods:namePart>
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<mods:affiliation id="C1BE723185E6E6FE1EF5361330E1BC4A">Laboratório de Biologia Aquática, Departamento de Ciências Biológicas, Faculdade de Ciências e Letras de Assis, Universidade Estadual Paulista, Assis, SP, Brazil deeogoo @ gmail. com; https: // orcid. org / 0000 - 0001 - 6510 - 7018 &amp; Laboratório de Biologia Aquática, Departamento de Ciências Biológicas, Faculdade de Ciências e Letras de Assis, Universidade Estadual Paulista, Assis, SP, Brazil deeogoo @ gmail. com; https: // orcid. org / 0000 - 0001 - 6510 - 7018 &amp; pitagoras. bispo @ unesp. br; https: // orcid. org / 0000 - 0002 - 7356 - 8882 * Corresponding author</mods:affiliation>
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<treatment id="73444D3AFFF4911E6AD7FB44AE4FA6F3" ID-GBIF-Taxon="214231486" LSID="urn:lsid:plazi:treatment:73444D3AFFF4911E6AD7FB44AE4FA6F3" httpUri="http://treatment.plazi.org/id/73444D3AFFF4911E6AD7FB44AE4FA6F3" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
<subSubSection id="B3F7AFA7FFF4911F6AD7FB44A930A5BE" box="[151,583,1168,1195]" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="FB52FC2CFFF4911F6AD7FB44A930A5BE" blockId="28.[151,583,1168,1230]" box="[151,583,1168,1195]" pageId="28" pageNumber="29">
<heading id="A01A4B40FFF4911F6AD7FB44A930A5BE" bold="true" box="[151,583,1168,1195]" fontSize="11" level="2" pageId="28" pageNumber="29" reason="4">
<taxonomicName id="3CED87AFFFF4911F6AD7FB44A930A5BE" ID-CoL="3L73V" authority="Daigle, 2005" authorityName="Daigle" authorityYear="2005" box="[151,583,1168,1195]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="bickorum">
<emphasis id="C999203EFFF4911F6AD7FB44A930A5BE" bold="true" box="[151,583,1168,1195]" pageId="28" pageNumber="29">
<emphasis id="C999203EFFF4911F6AD7FB44AAD0A5BF" bold="true" box="[151,423,1168,1195]" italics="true" pageId="28" pageNumber="29">Heteragrion bickorum</emphasis>
<bibRefCitation id="9F7C81DDFFF4911F6BEEFB44A930A5BE" author="Daigle, J. J." box="[430,583,1168,1194]" pageId="28" pageNumber="29" pagination="165 - 168" refId="ref61242" refString="Daigle, J. J. (2005) Heteragrion bickorum spec. nov. from Ecuador (Zygoptera: Megapodagrionidae). Odonatologica, 34 (2), 165 - 168." type="journal article" year="2005">Daigle, 2005</bibRefCitation>
</emphasis>
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<paragraph id="FB52FC2CFFF4911F6AD7FB60AA5BA5DA" blockId="28.[151,583,1168,1230]" box="[151,300,1204,1230]" pageId="28" pageNumber="29">
(
<figureCitation id="63D6E0A9FFF4911F6ADFFB60AA53A5DA" box="[159,292,1204,1230]" captionStart="FIGURE 16" captionStartId="29.[151,250,1567,1591]" captionTargetBox="[242,1342,194,1530]" captionTargetId="figure-17@29.[231,1356,181,1543]" captionTargetPageId="29" captionText="FIGURE 16. Heteragrion bickorum: (a–b) anterior view of head; (c) lateral view of pterothorax; (d) mediodorsal view of cercus; (e) close up of MP-AP in medial view, showing the elevated crest and the ridge above ML; (f) medial view of cercus; (g) dorsal view of cercus." figureDoi="http://doi.org/10.5281/zenodo.10008579" httpUri="https://zenodo.org/record/10008579/files/figure.png" pageId="28" pageNumber="29">Figs. 16a–g</figureCitation>
)
</paragraph>
<paragraph id="FB52FC2CFFF4911F6AD7FB2DA9EAA405" blockId="28.[151,669,1273,1297]" box="[151,669,1273,1297]" pageId="28" pageNumber="29">[Key locator: Key to Group B species, couplet 17’]</paragraph>
</subSubSection>
<subSubSection id="B3F7AFA7FFF4911F6AD7FA94AA59A725" pageId="28" pageNumber="29" type="reference_group">
<paragraph id="FB52FC2CFFF4911F6AD7FA94AA59A725" blockId="28.[151,1437,1343,1945]" pageId="28" pageNumber="29">
<treatmentCitationGroup id="DBFDDB02FFF4911F6AD7FA94AA59A725" pageId="28" pageNumber="29">
<treatmentCitation id="7A4CDA3DFFF4911F6AD7FA94A9D6A44E" author="Daigle, J. J." box="[151,673,1343,1370]" page="168" pageId="28" pageNumber="29" year="2005">
<taxonomicName id="3CED87AFFFF4911F6AD7FA94A9D6A44E" ID-CoL="3L73V" authority="Daigle 2005: 165 - 168" authorityName="Daigle" authorityPageNumber="165 - 168" authorityYear="2005" box="[151,673,1343,1370]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="bickorum">
<emphasis id="C999203EFFF4911F6AD7FA94AAE3A44D" box="[151,404,1343,1369]" italics="true" pageId="28" pageNumber="29">Heteragrion bickorum</emphasis>
:
<bibRefCitation id="9F7C81DDFFF4911F6BE4FAEBA9D6A44E" author="Daigle, J. J." box="[420,673,1343,1370]" pageId="28" pageNumber="29" pagination="165 - 168" refId="ref61242" refString="Daigle, J. J. (2005) Heteragrion bickorum spec. nov. from Ecuador (Zygoptera: Megapodagrionidae). Odonatologica, 34 (2), 165 - 168." type="journal article" year="2005">Daigle 2005: 165–168</bibRefCitation>
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(description of ♁ and
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from
<collectingCountry id="83FABCBCFFF4911F6E42FAEBAF14A44D" box="[1026,1123,1343,1369]" name="Ecuador" pageId="28" pageNumber="29">Ecuador</collectingCountry>
; comparison with
<taxonomicName id="3CED87AFFFF4911F6F74FA94AEEBA44D" ID-CoL="3L73T" authorityName="De Marmels" authorityYear="1989" box="[1332,1436,1343,1369]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="bariai">
<emphasis id="C999203EFFF4911F6F74FA94AEEBA44D" box="[1332,1436,1343,1369]" italics="true" pageId="28" pageNumber="29">H. bariai</emphasis>
</taxonomicName>
and
<taxonomicName id="3CED87AFFFF4911F6A86FAB0AA6FA469" ID-CoL="3L74D" authorityName=": Calvert" authorityYear="1909" box="[198,280,1380,1405]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="inca">
<emphasis id="C999203EFFF4911F6A86FAB0AA6FA469" box="[198,280,1380,1405]" italics="true" pageId="28" pageNumber="29">H. inca</emphasis>
</taxonomicName>
; figs. 1–5);—
<treatmentCitation id="7A4CDA3DFFF4911F6BEEFAB7A9C8A469" author="Garrison, R. W. &amp; von Ellenrieder, N. &amp; Louton, J. A." box="[430,703,1379,1406]" page="88" pageId="28" pageNumber="29" year="2010">
<bibRefCitation id="9F7C81DDFFF4911F6BEEFAB7A9C8A469" author="Garrison, R. W. &amp; von Ellenrieder, N. &amp; Louton, J. A." box="[430,703,1379,1406]" pageId="28" pageNumber="29" refId="ref62214" refString="Garrison, R. W., von Ellenrieder, N. &amp; Louton, J. A. (2010) Damselfly genera of the New World. An illustrated and Annotated Key to the Zygoptera. The Johns Hopkins University Press, Baltimore, Maryland, xiv + 490 pp." type="book" year="2010">
Garrison
<emphasis id="C999203EFFF4911F685EFAB0A921A469" box="[542,598,1379,1405]" italics="true" pageId="28" pageNumber="29">et al.</emphasis>
2010: 88
</bibRefCitation>
</treatmentCitation>
(list of
<taxonomicName id="3CED87AFFFF4911F6956FAB0A8D7A469" ID-CoL="4W75" authorityName="Selys" authorityYear="1862" box="[790,928,1380,1405]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="C999203EFFF4911F6956FAB0A8D7A469" box="[790,928,1380,1405]" italics="true" pageId="28" pageNumber="29">Heteragrion</emphasis>
</taxonomicName>
species);—
<treatmentCitation id="7A4CDA3DFFF4911F6E64FAB7ABC4A4B5" author="von Ellenrieder, N. &amp; Garrison, R. W." page="40" pageId="28" pageNumber="29" year="2011">
<bibRefCitation id="9F7C81DDFFF4911F6E64FAB7ABC4A4B5" author="von Ellenrieder, N. &amp; Garrison, R. W." pageId="28" pageNumber="29" pagination="40 - 46" refId="ref67235" refString="von Ellenrieder, N. &amp; Garrison, R. W. (2011) Odonata from Tiputini, Orellana Province, Ecuador. Agrion, 15 (2), 40 - 46." type="journal article" year="2011">von Ellenrieder &amp; Garrison 2011: 40</bibRefCitation>
</treatmentCitation>
(list of species for Tiputini,
<collectingCountry id="83FABCBCFFF4911F6BB3FA53A921A4B5" box="[499,598,1415,1441]" name="Ecuador" pageId="28" pageNumber="29">Ecuador</collectingCountry>
);
<treatmentCitation id="7A4CDA3DFFF4911F6828FA53A8D6A4B5" author="Bota-Sierra, C. A. &amp; Corso, A. &amp; Janni, O. &amp; Sandoval-H. J. &amp; Vigano, M." box="[616,929,1415,1441]" page="120" pageId="28" pageNumber="29" year="2018">
<bibRefCitation id="9F7C81DDFFF4911F6828FA53A8D6A4B5" author="Bota-Sierra, C. A. &amp; Corso, A. &amp; Janni, O. &amp; Sandoval-H. J. &amp; Vigano, M." box="[616,929,1415,1441]" pageId="28" pageNumber="29" pagination="115 - 127" refId="ref60351" refString="Bota-Sierra, C. A., Corso, A., Janni, O., Sandoval-H. J. &amp; Vigano, M. (2018) Seventeen new dragonfly records from Colombia and the confirmation of the synonymy of Philogenia monotis and P. tinalandia (Insecta: Odonata). International Journal of Odonatology, 21 (2), 115 - 127. https: // doi. org / 10.1080 / 13887890.2018.1462262" type="journal article" year="2018">
Bota-Sierra
<emphasis id="C999203EFFF4911F68B1FA5CA85EA4B5" box="[753,809,1415,1441]" italics="true" pageId="28" pageNumber="29">et al.</emphasis>
2018: 120
</bibRefCitation>
</treatmentCitation>
(new record for
<collectingCountry id="83FABCBCFFF4911F6E1EFA53AFB9A4B5" box="[1118,1230,1415,1441]" name="Colombia" pageId="28" pageNumber="29">Colombia</collectingCountry>
);—
<treatmentCitation id="7A4CDA3DFFF4911F6EB7FA53AA3BA4D1" ID-DOI="http://doi.org/10.5281/zenodo.6527908" ID-Zenodo-Dep="6527908" author="Stand-Perez, M. A. &amp; Bota-Sierra, C. A. &amp; Perez-Gutierrez, L. A." httpUri="http://treatment.plazi.org/id/03B6085D6916FFCBFCC8F94565B3F81B" page="92" pageId="28" pageNumber="29" year="2019">
<bibRefCitation id="9F7C81DDFFF4911F6EB7FA53AA3BA4D1" author="Stand-Perez, M. A. &amp; Bota-Sierra, C. A. &amp; Perez-Gutierrez, L. A." pageId="28" pageNumber="29" pagination="90 - 112" refId="ref65416" refString="Stand-Perez, M. A., Bota-Sierra, C. A. &amp; Perez-Gutierrez, L. A. (2019) Heteragrion demarmelsi sp. nov., with taxonomic notes on Colombian Heteragrion species (Odonata: Heteragrionidae). Zootaxa, 4623 (1), 90 - 112. https: // doi. org / 10.11646 / zootaxa. 4623.1.6" type="journal article" year="2019">
Stand-Pérez
<emphasis id="C999203EFFF4911F6FC8FA5CABC3A4D1" italics="true" pageId="28" pageNumber="29">et al.</emphasis>
2019: 92–93
</bibRefCitation>
</treatmentCitation>
, 97–98 (key to species, ♁ in couplet 7,
<specimenCount id="EDEB37A5FFF4911F694FFA78A853A4D2" box="[783,804,1452,1478]" pageId="28" pageNumber="29" type="female">♀</specimenCount>
in couplet 7; remarks and comparison with
<taxonomicName id="3CED87AFFFF4911F6F59FA78AE1BA4D1" ID-CoL="3L74D" authorityName=": Calvert" authorityYear="1909" box="[1305,1388,1452,1477]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="inca">
<emphasis id="C999203EFFF4911F6F59FA78AE1BA4D1" box="[1305,1388,1452,1477]" italics="true" pageId="28" pageNumber="29">H. inca</emphasis>
</taxonomicName>
and
<taxonomicName id="3CED87AFFFF4911F6AD7FA04AA4AA4FD" authorityName="Stand-Perez, Bota-Sierra &amp; Perez-Gutierrez" authorityYear="2019" box="[151,317,1487,1513]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="demarmelsi">
<emphasis id="C999203EFFF4911F6AD7FA04AA4AA4FD" box="[151,317,1487,1513]" italics="true" pageId="28" pageNumber="29">H. demarmelsi</emphasis>
</taxonomicName>
; figs.
<quantity id="3C1551C9FFF4911F6BC4FA1BAAD7A4FD" box="[388,416,1487,1513]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="28" pageNumber="29" unit="g" value="4.0">4g</quantity>
–h, 8a–c, 13a–c, 17);—
<treatmentCitation id="7A4CDA3DFFF4911F68E1FA1BA891A4FD" author="Stand-Perez, M. A. &amp; Montes-Fontalvo, J. &amp; Perez-Gutierrez, L. A." box="[673,998,1487,1513]" page="381" pageId="28" pageNumber="29" year="2020">
<bibRefCitation id="9F7C81DDFFF4911F68E1FA1BA891A4FD" author="Stand-Perez, M. A. &amp; Montes-Fontalvo, J. &amp; Perez-Gutierrez, L. A." box="[673,998,1487,1513]" pageId="28" pageNumber="29" pagination="378 - 388" refId="ref65482" refString="Stand-Perez, M. A., Montes-Fontalvo, J. &amp; Perez-Gutierrez, L. A. (2020) Sixteen new records of Odonata for Colombia from the Araracuara Region (Perilestidae, Calopterygidae, Heteragrionidae, Megapodagrionidae, Polythoridae, Coenagrionidae, Aeshnidae, Gomphidae, Libellulidae). Notulae odonatologicae, 9 (6), 378 - 388. https: // doi. org / 10.60024 / zenodo. 5702965" type="journal article" year="2020">
Stand-Pérez
<emphasis id="C999203EFFF4911F6976FA04A819A4FD" box="[822,878,1487,1513]" italics="true" pageId="28" pageNumber="29">et al.</emphasis>
2020: 381
</bibRefCitation>
</treatmentCitation>
(list of species recorded in Serranía de Chiribiquete National Natural Park,
<collectingCountry id="83FABCBCFFF4911F6877FA27A9D0A719" box="[567,679,1523,1549]" name="Colombia" pageId="28" pageNumber="29">Colombia</collectingCountry>
);—Miranda-Filho
<emphasis id="C999203EFFF4911F69C0FA20A8CDA719" box="[896,954,1523,1549]" italics="true" pageId="28" pageNumber="29">et al.</emphasis>
2022: 9 (list of species occurring in
<collectingRegion id="392932CEFFF4911F6F26FA27ABBAA726" country="Brazil" name="Acre" pageId="28" pageNumber="29">Acre state</collectingRegion>
,
<collectingCountry id="83FABCBCFFF4911F6A99F9C3AA55A725" box="[217,290,1559,1585]" name="Brazil" pageId="28" pageNumber="29">Brazil</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B3F7AFA7FFF4911F6A87F9EFAFABA68D" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="FB52FC2CFFF4911F6A87F9EFAA6CA661" blockId="28.[151,1437,1343,1945]" pageId="28" pageNumber="29">
<emphasis id="C999203EFFF4911F6A87F9EFAAD8A741" bold="true" box="[199,431,1595,1621]" pageId="28" pageNumber="29">Material examined.</emphasis>
<materialsCitation id="4B85F671FFF4911F6BF4F9E8AABAA76D" ID-GBIF-Occurrence="4428728410" collectingDate="2012-12-07" collectionCode="ECOEVO" collectorName="A. C. Rivera" country="Ecuador" elevation="240" latitude="-0.6321" location="Tiputini Biodiversity Station" longLatPrecision="7" longitude="-76.1451" pageId="28" pageNumber="29" specimenCount="1">
3♁♁
<collectingCountry id="83FABCBCFFF4911F6BB1F9EFA90CA741" box="[497,635,1595,1621]" name="Ecuador" pageId="28" pageNumber="29">ECUADOR</collectingCountry>
,
<location id="FE32AAF7FFF4911F68C4F9EFA8B3A741" LSID="urn:lsid:plazi:treatment:73444D3AFFF4911E6AD7FB44AE4FA6F3:FE32AAF7FFF4911F68C4F9EFA8B3A741" box="[644,964,1595,1621]" country="Ecuador" latitude="-0.6321" longLatPrecision="7" longitude="-76.1451" name="Tiputini Biodiversity Station" pageId="28" pageNumber="29">Tiputini Biodiversity Station</location>
,
<date id="8F53DAECFFF4911F698EF9EFAF35A741" box="[974,1090,1595,1621]" pageId="28" pageNumber="29" value="2012-12-07">
<collectingDate id="9F172304FFF4911F698EF9EFAF35A741" box="[974,1090,1595,1621]" pageId="28" pageNumber="29" value="2012-12-07">7.xii.2012</collectingDate>
</date>
, (
<geoCoordinate id="9ED99AEBFFF4911F6E13F9EFAFDAA741" box="[1107,1197,1595,1621]" degrees="0.6321" direction="south" orientation="latitude" pageId="28" pageNumber="29" precision="5" value="-0.6321">-0.6321</geoCoordinate>
,
<geoCoordinate id="9ED99AEBFFF4911F6EF7F9EFAE68A742" box="[1207,1311,1595,1622]" degrees="76.1451" direction="west" orientation="longitude" pageId="28" pageNumber="29" precision="5" value="-76.1451">-76.1451</geoCoordinate>
,
<elevation id="70C01B1FFFF4911F6F68F9EFAEF8A741" box="[1320,1423,1595,1621]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="28" pageNumber="29" unit="m" value="240.0">
<quantity id="3C1551C9FFF4911F6F68F9EFAE1FA741" box="[1320,1384,1595,1621]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="28" pageNumber="29" unit="m" value="240.0">240m</quantity>
asl
</elevation>
),
<collectorName id="561899FAFFF4911F6AD7F98BAA6AA76D" box="[151,285,1631,1657]" pageId="28" pageNumber="29">A.C. Rivera</collectorName>
leg.,
<collectionCode id="9DFC64E9FFF4911F6B18F98BAABAA76D" box="[344,461,1631,1657]" pageId="28" pageNumber="29">ECOEVO</collectionCode>
</materialsCitation>
; 6♁♁
<materialsCitation id="4B85F671FFF4911F6855F9B4AA34A789" ID-GBIF-Occurrence="4428728443" collectingDate="2014-11-11" collectionCode="ECOEVO" collectorName="I. Sanmartin-Villar" country="Ecuador" elevation="437" latitude="-1.0769" location="Jatun Sacha" longLatPrecision="7" longitude="-77.6086" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Napo">
<collectingRegion id="392932CEFFF4911F6855F9B4A923A76E" box="[533,596,1632,1658]" country="Ecuador" name="Napo" pageId="28" pageNumber="29">Napo</collectingRegion>
,
<location id="FE32AAF7FFF4911F681EF98BA992A76D" LSID="urn:lsid:plazi:treatment:73444D3AFFF4911E6AD7FB44AE4FA6F3:FE32AAF7FFF4911F681EF98BA992A76D" box="[606,741,1631,1657]" country="Ecuador" latitude="-1.0769" longLatPrecision="7" longitude="-77.6086" name="Jatun Sacha" pageId="28" pageNumber="29" stateProvince="Napo">Jatún Sacha</location>
,
<date id="8F53DAECFFF4911F68AFF98BA81FA76D" box="[751,872,1631,1657]" pageId="28" pageNumber="29" value="2014-11-11">
<collectingDate id="9F172304FFF4911F68AFF98BA81FA76D" box="[751,872,1631,1657]" pageId="28" pageNumber="29" value="2014-11-11">11.xi.2014</collectingDate>
</date>
, (
<geoCoordinate id="9ED99AEBFFF4911F693AF98BA8A3A76D" box="[890,980,1631,1657]" degrees="1.0769" direction="south" orientation="latitude" pageId="28" pageNumber="29" precision="5" value="-1.0769">-1.0769</geoCoordinate>
,
<geoCoordinate id="9ED99AEBFFF4911F699EF98BAF31A76D" box="[990,1094,1631,1657]" degrees="77.6086" direction="west" orientation="longitude" pageId="28" pageNumber="29" precision="5" value="-77.6086">-77.6086</geoCoordinate>
,
<elevation id="70C01B1FFFF4911F6E0FF98BAFC0A76D" box="[1103,1207,1631,1657]" metricMagnitude="2" metricUnit="m" metricValue="4.37" pageId="28" pageNumber="29" unit="m" value="437.0">
<quantity id="3C1551C9FFF4911F6E0FF98BAFF8A76D" box="[1103,1167,1631,1657]" metricMagnitude="2" metricUnit="m" metricValue="4.37" pageId="28" pageNumber="29" unit="m" value="437.0">437m</quantity>
asl
</elevation>
),
<collectorName id="561899FAFFF4911F6E8AF9B4AEEBA76D" box="[1226,1436,1631,1658]" pageId="28" pageNumber="29">I. Sanmartín-Villar</collectorName>
leg.,
<collectionCode id="9DFC64E9FFF4911F6A8EF957AA34A789" box="[206,323,1667,1693]" pageId="28" pageNumber="29">ECOEVO</collectionCode>
</materialsCitation>
; 1♁
<materialsCitation id="4B85F671FFF4911F6B39F957AA61A7D5" ID-GBIF-Occurrence="4428728408" collectingDate="2022-07-18" collectionCode="INPA" collectorName="G. Desiderio" country="Brazil" elevation="220" latitude="-7.4589" location="Igarape do Veu" longLatPrecision="7" longitude="-73.6327" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Acre">
<collectingCountry id="83FABCBCFFF4911F6B39F957AA96A789" box="[377,481,1667,1693]" name="Brazil" pageId="28" pageNumber="29">BRAZIL</collectingCountry>
,
<collectingRegion id="392932CEFFF4911F6BAAF957A92AA78A" box="[490,605,1667,1694]" country="Brazil" name="Acre" pageId="28" pageNumber="29">Acre state</collectingRegion>
,
<location id="FE32AAF7FFF4911F6829F957A860A789" LSID="urn:lsid:plazi:treatment:73444D3AFFF4911E6AD7FB44AE4FA6F3:FE32AAF7FFF4911F6829F957A860A789" box="[617,791,1667,1693]" country="Brazil" latitude="-7.4589" longLatPrecision="7" longitude="-73.6327" name="Igarape do Veu" pageId="28" pageNumber="29" stateProvince="Acre">Igarapé do Véu</location>
,
<date id="8F53DAECFFF4911F6963F957A8D3A789" box="[803,932,1667,1693]" pageId="28" pageNumber="29" value="2022-07-18">
<collectingDate id="9F172304FFF4911F6963F957A8D3A789" box="[803,932,1667,1693]" pageId="28" pageNumber="29" value="2022-07-18">18.vii.2022</collectingDate>
</date>
, (
<geoCoordinate id="9ED99AEBFFF4911F69F8F957AF64A78A" box="[952,1043,1667,1694]" degrees="7.4589" direction="south" orientation="latitude" pageId="28" pageNumber="29" precision="5" value="-7.4589">-7.4589</geoCoordinate>
,
<geoCoordinate id="9ED99AEBFFF4911F6E5EF957AFF2A789" box="[1054,1157,1667,1693]" degrees="73.6327" direction="west" orientation="longitude" pageId="28" pageNumber="29" precision="5" value="-73.6327">-73.6327</geoCoordinate>
,
<elevation id="70C01B1FFFF4911F6ED0F957AF8EA789" box="[1168,1273,1667,1693]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="28" pageNumber="29" unit="m" value="220.0">
<quantity id="3C1551C9FFF4911F6ED0F957AFA7A789" box="[1168,1232,1667,1693]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="28" pageNumber="29" unit="m" value="220.0">220m</quantity>
asl
</elevation>
),
<collectorName id="561899FAFFF4911F6F4DF957AEEBA789" box="[1293,1436,1667,1693]" pageId="28" pageNumber="29">G. Desidério</collectorName>
leg.,
<collectionCode id="9DFC64E9FFF4911F6A93F973AA61A7D5" box="[211,278,1703,1729]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="28" pageNumber="29">INPA</collectionCode>
</materialsCitation>
;
<materialsCitation id="4B85F671FFF4911F6B65F97CAF00A7D7" ID-GBIF-Occurrence="4428728482" box="[293,1143,1703,1731]" collectingDate="2022-07-19" collectingDateMax="2022-07-25" collectingDateMin="2022-07-19" collectionCode="INPA" collectorName="G. Desiderio" country="Brazil" elevation="220" latitude="-7.4589" location="Igarape do Veu" longLatPrecision="7" longitude="-73.6327" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Acre">
2♁♁ same data, but
<date id="8F53DAECFFF4911F685FF973A9D6A7D5" box="[543,673,1703,1729]" pageId="28" pageNumber="29" value="2022-07-19">
<collectingDate id="9F172304FFF4911F685FF973A9D6A7D5" box="[543,673,1703,1729]" pageId="28" pageNumber="29" value="2022-07-19">19.vii.2022</collectingDate>
</date>
; 4♁♁ same data, but
<date id="8F53DAECFFF4911F69ECF973AF58A7D6" box="[940,1071,1703,1730]" pageId="28" pageNumber="29" value="2022-07-25">
<collectingDate id="9F172304FFF4911F69ECF973AF58A7D6" box="[940,1071,1703,1730]" pageId="28" pageNumber="29" value="2022-07-25">25.vii.2022</collectingDate>
</date>
; 5♁♁
</materialsCitation>
<materialsCitation id="4B85F671FFF4911F6EC3F973AEEFA7F1" ID-GBIF-Occurrence="4428728501" collectionCode="UARC" collectorName="L. Perez &amp; Montes &amp; J. Villamil" country="Colombia" elevation="650" location="Quebrada Sangoyaco" municipality="Mocoa Municipality" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Putumayo">
<collectingCountry id="83FABCBCFFF4911F6EC3F973AE6CA7D5" box="[1155,1307,1703,1729]" name="Colombia" pageId="28" pageNumber="29">COLOMBIA</collectingCountry>
,
<collectingRegion id="392932CEFFF4911F6F6BF97CAA69A7F2" country="Colombia" name="Putumayo" pageId="28" pageNumber="29">Putumayo Department</collectingRegion>
,
<collectingMunicipality id="1B366656FFF4911F6B69F918A978A7F1" box="[297,527,1739,1766]" pageId="28" pageNumber="29">Mocoa Municipality</collectingMunicipality>
,
<location id="FE32AAF7FFF4911F685AF91FA87CA7F1" LSID="urn:lsid:plazi:treatment:73444D3AFFF4911E6AD7FB44AE4FA6F3:FE32AAF7FFF4911F685AF91FA87CA7F1" box="[538,779,1739,1765]" country="Colombia" municipality="Mocoa Municipality" name="Quebrada Sangoyaco" pageId="28" pageNumber="29" stateProvince="Putumayo">Quebrada Sangoyaco</location>
, (
<elevation id="70C01B1FFFF4911F695DF91FA8F0A7F1" box="[797,903,1739,1766]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="28" pageNumber="29" unit="m" value="650.0">
<quantity id="3C1551C9FFF4911F695DF91FA828A7F2" box="[797,863,1739,1766]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="28" pageNumber="29" unit="m" value="650.0">650m</quantity>
asl
</elevation>
),
<collectorName id="561899FAFFF4911F69DBF918A88CA7F1" box="[923,1019,1739,1766]" pageId="28" pageNumber="29">L. Pérez</collectorName>
, J,
<collectorName id="561899FAFFF4911F6E5EF918AF05A7F2" box="[1054,1138,1740,1766]" pageId="28" pageNumber="29">Montes</collectorName>
&amp;
<collectorName id="561899FAFFF4911F6ED5F918AE7EA7F1" box="[1173,1289,1739,1766]" pageId="28" pageNumber="29">J. Villamil</collectorName>
leg.,
<collectionCode id="9DFC64E9FFF4911F6F07F91FAEEFA7F1" box="[1351,1432,1739,1765]" pageId="28" pageNumber="29">UARC</collectionCode>
</materialsCitation>
; 1♁
<materialsCitation id="4B85F671FFF4911F6A81F924A9A6A639" ID-GBIF-Occurrence="4428728457" collectingDate="2017-01-17" collectionCode="CEUA" collectorName="M. Vigano &amp; A. Corso &amp; O. Janni" country="Colombia" elevation="850" latitude="0.657" location="Isla Escondida Natural Reserve" longLatPrecision="55" longitude="-77.0722" municipality="Orito Municipality" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Putumayo">
<collectingRegion id="392932CEFFF4911F6A81F924AAB6A61E" box="[193,449,1776,1802]" country="Colombia" name="Putumayo" pageId="28" pageNumber="29">Putumayo Department</collectingRegion>
,
<collectingMunicipality id="1B366656FFF4911F6B8DF93BA9D6A61D" box="[461,673,1775,1801]" pageId="28" pageNumber="29">Orito Municipality</collectingMunicipality>
,
<location id="FE32AAF7FFF4911F68ECF93BAF67A61E" LSID="urn:lsid:plazi:treatment:73444D3AFFF4911E6AD7FB44AE4FA6F3:FE32AAF7FFF4911F68ECF93BAF67A61E" box="[684,1040,1775,1802]" country="Colombia" latitude="0.657" longLatPrecision="55" longitude="-77.0722" municipality="Orito Municipality" name="Isla Escondida Natural Reserve" pageId="28" pageNumber="29" stateProvince="Putumayo">Isla Escondida Natural Reserve</location>
,
<date id="8F53DAECFFF4911F6E5BF93BAFFFA61D" box="[1051,1160,1775,1801]" pageId="28" pageNumber="29" value="2017-01-17">
<collectingDate id="9F172304FFF4911F6E5BF93BAFFFA61D" box="[1051,1160,1775,1801]" pageId="28" pageNumber="29" value="2017-01-17">17.i.2017</collectingDate>
</date>
, (
<geoCoordinate id="9ED99AEBFFF4911F6EDBF93BAFA9A61E" box="[1179,1246,1775,1802]" degrees="0.657" direction="north" orientation="latitude" pageId="28" pageNumber="29" precision="55" value="0.657">0.657</geoCoordinate>
,
<geoCoordinate id="9ED99AEBFFF4911F6EAAF93BAE26A61D" box="[1258,1361,1775,1801]" degrees="77.0722" direction="west" orientation="longitude" pageId="28" pageNumber="29" precision="5" value="-77.0722">-77.0722</geoCoordinate>
,
<elevation id="70C01B1FFFF4911F6F1CF93BABCEA639" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="28" pageNumber="29" unit="m" value="850.0">
<quantity id="3C1551C9FFF4911F6F1CF93BAEEBA61E" box="[1372,1436,1775,1802]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="28" pageNumber="29" unit="m" value="850.0">850m</quantity>
asl
</elevation>
),
<collectorName id="561899FAFFF4911F6A8FF8C0AA3CA639" box="[207,331,1811,1838]" pageId="28" pageNumber="29">M. Vigano</collectorName>
,
<collectorName id="561899FAFFF4911F6B15F8C7AACBA639" box="[341,444,1811,1837]" pageId="28" pageNumber="29">A. Corso</collectorName>
&amp;
<collectorName id="561899FAFFF4911F6BA2F8C7A937A639" box="[482,576,1811,1837]" pageId="28" pageNumber="29">O. Janni</collectorName>
leg.,
<collectionCode id="9DFC64E9FFF4911F68C1F8C7A9A6A639" box="[641,721,1811,1837]" pageId="28" pageNumber="29">CEUA</collectionCode>
</materialsCitation>
;
<materialsCitation id="4B85F671FFF4911F689DF8C0A805A645" ID-GBIF-Occurrence="4428728375" collectingDate="2017-01-18" collectionCode="CEUA" collectorName="M. Vigano &amp; A. Corso &amp; O. Janni" country="Colombia" elevation="890" latitude="0.6613" location="Nuevo Mundo island" longLatPrecision="7" longitude="-77.0767" pageId="28" pageNumber="29" specimenCount="1">
5♁♁ road to
<location id="FE32AAF7FFF4911F6936F8C0AF1FA639" LSID="urn:lsid:plazi:treatment:73444D3AFFF4911E6AD7FB44AE4FA6F3:FE32AAF7FFF4911F6936F8C0AF1FA639" box="[886,1128,1811,1838]" country="Colombia" latitude="0.6613" longLatPrecision="7" longitude="-77.0767" name="Nuevo Mundo island" pageId="28" pageNumber="29">Nuevo Mundo island</location>
,
<date id="8F53DAECFFF4911F6E34F8C7AE44A639" box="[1140,1331,1811,1838]" pageId="28" pageNumber="29" value="2017-01-18">
<collectingDate id="9F172304FFF4911F6E34F8C7AE44A639" box="[1140,1331,1811,1838]" pageId="28" pageNumber="29" value="2017-01-18">18 January 2017</collectingDate>
</date>
, (
<geoCoordinate id="9ED99AEBFFF4911F6F07F8C7AEEFA639" box="[1351,1432,1811,1837]" degrees="0.6613" direction="north" orientation="latitude" pageId="28" pageNumber="29" precision="5" value="0.6613">0.6613</geoCoordinate>
,
<geoCoordinate id="9ED99AEBFFF4911F6AD7F8E3AB88A645" box="[151,255,1847,1873]" degrees="77.0767" direction="west" orientation="longitude" pageId="28" pageNumber="29" precision="5" value="-77.0767">-77.0767</geoCoordinate>
,
<elevation id="70C01B1FFFF4911F6B48F8E3AA07A645" box="[264,368,1847,1873]" metricMagnitude="2" metricUnit="m" metricValue="8.9" pageId="28" pageNumber="29" unit="m" value="890.0">
<quantity id="3C1551C9FFF4911F6B48F8E3AA3FA645" box="[264,328,1847,1873]" metricMagnitude="2" metricUnit="m" metricValue="8.9" pageId="28" pageNumber="29" unit="m" value="890.0">890m</quantity>
asl
</elevation>
),
<collectorName id="561899FAFFF4911F6BC3F8ECAA8BA645" box="[387,508,1847,1874]" pageId="28" pageNumber="29">M. Vigano</collectorName>
,
<collectorName id="561899FAFFF4911F6844F8E3A91FA645" box="[516,616,1847,1873]" pageId="28" pageNumber="29">A. Corso</collectorName>
&amp;
<collectorName id="561899FAFFF4911F68CAF8E3A991A645" box="[650,742,1847,1873]" pageId="28" pageNumber="29">O. Janni</collectorName>
leg.,
<collectionCode id="9DFC64E9FFF4911F6962F8E3A805A645" box="[802,882,1847,1873]" pageId="28" pageNumber="29">CEUA</collectionCode>
</materialsCitation>
; 1♁
<collectingRegion id="392932CEFFF4911F69E4F8E3AFFAA646" box="[932,1165,1847,1874]" country="Colombia" name="Caqueta" pageId="28" pageNumber="29">Caquetá Department</collectingRegion>
, Florencia Municipality, ANDES–E.
</paragraph>
<paragraph id="FB52FC2CFFF4911F6A87F8ABAFABA68D" blockId="28.[151,1437,1343,1945]" box="[199,1244,1919,1945]" pageId="28" pageNumber="29">
<emphasis id="C999203EFFF4911F6A87F8ABAACDA68D" bold="true" box="[199,442,1919,1945]" pageId="28" pageNumber="29">Known distribution.</emphasis>
<collectingCountry id="83FABCBCFFF4911F6B81F8ABA972A68D" box="[449,517,1919,1945]" name="Brazil" pageId="28" pageNumber="29">Brazil</collectingCountry>
(
<collectingRegion id="392932CEFFF4911F6854F8ABA9FDA68D" box="[532,650,1919,1945]" country="Brazil" name="Acre" pageId="28" pageNumber="29">Acre state</collectingRegion>
),
<collectingCountry id="83FABCBCFFF4911F68DEF8ABA87BA68D" box="[670,780,1919,1945]" name="Colombia" pageId="28" pageNumber="29">Colombia</collectingCountry>
(Eastern Andean foothills), and
<collectingCountry id="83FABCBCFFF4911F6E38F8ABAFA0A68D" box="[1144,1239,1919,1945]" name="Ecuador" pageId="28" pageNumber="29">Ecuador</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="AF92ACA4FFF5911E6AD7F9CBAAEDA76A" ID-DOI="http://doi.org/10.5281/zenodo.10008579" ID-Zenodo-Dep="10008579" httpUri="https://zenodo.org/record/10008579/files/figure.png" pageId="29" pageNumber="30" startId="29.[151,250,1567,1591]" targetBox="[242,1342,194,1530]" targetPageId="29" targetType="figure">
<paragraph id="FB52FC2CFFF5911E6AD7F9CBAAEDA76A" blockId="29.[151,1436,1567,1663]" pageId="29" pageNumber="30">
<emphasis id="C999203EFFF5911E6AD7F9CBAA54A723" bold="true" box="[151,291,1567,1591]" pageId="29" pageNumber="30">FIGURE 16.</emphasis>
<taxonomicName id="3CED87AFFFF5911E6B6CF9F4A964A723" authorityName=": Daigle" authorityYear="2005" box="[300,531,1567,1591]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="bickorum">
<emphasis id="C999203EFFF5911E6B6CF9F4A964A723" box="[300,531,1567,1591]" italics="true" pageId="29" pageNumber="30">Heteragrion bickorum</emphasis>
</taxonomicName>
: (a–b) anterior view of head; (c) lateral view of pterothorax; (d) mediodorsal view of cercus; (e) close up of MP-AP in medial view, showing the elevated crest and the ridge above ML; (f) medial view of cercus; (g) dorsal view of cercus.
</paragraph>
</caption>
<subSubSection id="B3F7AFA7FFF5911E6A87F905AE4FA6F3" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="FB52FC2CFFF5911E6A87F905AE4FA6F3" blockId="29.[151,1437,1745,2024]" pageId="29" pageNumber="30">
<emphasis id="C999203EFFF5911E6A87F905AA96A7FF" bold="true" box="[199,481,1745,1771]" pageId="29" pageNumber="30">Diagnosis and remarks.</emphasis>
Some degree of morphological variation was observed in the material from
<collectingCountry id="83FABCBCFFF5911E6F79F905AEEFA7FF" box="[1337,1432,1745,1771]" name="Ecuador" pageId="29" pageNumber="30">Ecuador</collectingCountry>
,
<collectingCountry id="83FABCBCFFF5911E6AD7F921AA7FA61B" box="[151,264,1781,1807]" name="Colombia" pageId="29" pageNumber="30">Colombia</collectingCountry>
, and
<collectingCountry id="83FABCBCFFF5911E6B02F921AAFDA61B" box="[322,394,1781,1807]" name="Brazil" pageId="29" pageNumber="30">Brazil</collectingCountry>
. Whereas specimens from
<collectingCountry id="83FABCBCFFF5911E68F4F921A866A61B" box="[692,785,1781,1807]" name="Ecuador" pageId="29" pageNumber="30">Ecuador</collectingCountry>
and
<collectingCountry id="83FABCBCFFF5911E6906F921A8C3A61B" box="[838,948,1781,1807]" name="Colombia" pageId="29" pageNumber="30">Colombia</collectingCountry>
presented a certain degree of BP convexity, the Brazilian specimens have this structure nearly straight or poorly convex. The remaining structures of the cercus (
<emphasis id="C999203EFFF5911E6AE1F8EAABB4A643" box="[161,195,1854,1879]" italics="true" pageId="29" pageNumber="30">i.e.</emphasis>
, the crest, ML with acute tip) and overall coloration match the description of
<taxonomicName id="3CED87AFFFF5911E6E63F8EAAFD9A643" authorityName=": Daigle" authorityYear="2005" box="[1059,1198,1853,1879]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="bickorum">
<emphasis id="C999203EFFF5911E6E63F8EAAFD9A643" box="[1059,1198,1853,1879]" italics="true" pageId="29" pageNumber="30">H. bickorum</emphasis>
</taxonomicName>
; therefore, they were considered conspecific. This species, which shares morphological similarities with
<taxonomicName id="3CED87AFFFF5911E6E70F8B6AFE2A66F" authorityName="De Marmels" authorityYear="1989" box="[1072,1173,1889,1915]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="bariai">
<emphasis id="C999203EFFF5911E6E70F8B6AFE2A66F" box="[1072,1173,1889,1915]" italics="true" pageId="29" pageNumber="30">H. bariai</emphasis>
</taxonomicName>
,
<taxonomicName id="3CED87AFFFF5911E6EE2F8B6AE31A66F" authorityName="Stand-Perez, Bota-Sierra &amp; Perez-Gutierrez" authorityYear="2019" box="[1186,1350,1889,1915]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="demarmelsi">
<emphasis id="C999203EFFF5911E6EE2F8B6AE31A66F" box="[1186,1350,1889,1915]" italics="true" pageId="29" pageNumber="30">H. demarmelsi</emphasis>
</taxonomicName>
, and
<taxonomicName id="3CED87AFFFF5911E6FC1F8B6ABB0A68B" authorityName=": Calvert" authorityYear="1909" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="inca">
<emphasis id="C999203EFFF5911E6FC1F8B6ABB0A68B" italics="true" pageId="29" pageNumber="30">H. inca</emphasis>
</taxonomicName>
, can be distinguished from these and other Group B species based on the following combination of characters: BP nearly straight or only slightly convex; AP longer than MP (
<figureCitation id="63D6E0A9FFF5911E6913F87DA8C4A6D7" box="[851,947,1961,1987]" captionStart="FIGURE 16" captionStartId="29.[151,250,1567,1591]" captionTargetBox="[242,1342,194,1530]" captionTargetId="figure-17@29.[231,1356,181,1543]" captionTargetPageId="29" captionText="FIGURE 16. Heteragrion bickorum: (a–b) anterior view of head; (c) lateral view of pterothorax; (d) mediodorsal view of cercus; (e) close up of MP-AP in medial view, showing the elevated crest and the ridge above ML; (f) medial view of cercus; (g) dorsal view of cercus." figureDoi="http://doi.org/10.5281/zenodo.10008579" httpUri="https://zenodo.org/record/10008579/files/figure.png" pageId="29" pageNumber="30">
Fig.
<quantity id="3C1551C9FFF5911E69C6F87DA8C4A6D7" box="[902,947,1961,1987]" metricMagnitude="-2" metricUnit="kg" metricValue="1.6" pageId="29" pageNumber="30" unit="g" value="16.0">16g</quantity>
</figureCitation>
); MP bearing an elevated crest on its medial margin (
<figureCitation id="63D6E0A9FFF5911E6AB5F819AA0AA6F3" box="[245,381,1997,2023]" captionStart="FIGURE 16" captionStartId="29.[151,250,1567,1591]" captionTargetBox="[242,1342,194,1530]" captionTargetId="figure-17@29.[231,1356,181,1543]" captionTargetPageId="29" captionText="FIGURE 16. Heteragrion bickorum: (a–b) anterior view of head; (c) lateral view of pterothorax; (d) mediodorsal view of cercus; (e) close up of MP-AP in medial view, showing the elevated crest and the ridge above ML; (f) medial view of cercus; (g) dorsal view of cercus." figureDoi="http://doi.org/10.5281/zenodo.10008579" httpUri="https://zenodo.org/record/10008579/files/figure.png" pageId="29" pageNumber="30">Figs. 16d–e</figureCitation>
); ML small, with acute apex (
<figureCitation id="63D6E0A9FFF5911E688BF819A85CA6F3" box="[715,811,1997,2023]" captionStart="FIGURE 16" captionStartId="29.[151,250,1567,1591]" captionTargetBox="[242,1342,194,1530]" captionTargetId="figure-17@29.[231,1356,181,1543]" captionTargetPageId="29" captionText="FIGURE 16. Heteragrion bickorum: (a–b) anterior view of head; (c) lateral view of pterothorax; (d) mediodorsal view of cercus; (e) close up of MP-AP in medial view, showing the elevated crest and the ridge above ML; (f) medial view of cercus; (g) dorsal view of cercus." figureDoi="http://doi.org/10.5281/zenodo.10008579" httpUri="https://zenodo.org/record/10008579/files/figure.png" pageId="29" pageNumber="30">Fig. 16e</figureCitation>
); AP with a longitudinal depression (
<figureCitation id="63D6E0A9FFF5911E6E89F819AE5CA6F3" box="[1225,1323,1997,2023]" captionStart="FIGURE 16" captionStartId="29.[151,250,1567,1591]" captionTargetBox="[242,1342,194,1530]" captionTargetId="figure-17@29.[231,1356,181,1543]" captionTargetPageId="29" captionText="FIGURE 16. Heteragrion bickorum: (a–b) anterior view of head; (c) lateral view of pterothorax; (d) mediodorsal view of cercus; (e) close up of MP-AP in medial view, showing the elevated crest and the ridge above ML; (f) medial view of cercus; (g) dorsal view of cercus." figureDoi="http://doi.org/10.5281/zenodo.10008579" httpUri="https://zenodo.org/record/10008579/files/figure.png" pageId="29" pageNumber="30">Fig. 16d</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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