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<mods:title id="931B193199C3D2CB1BCE952DCB38B45D">Taxonomic revision of the Neotropical genus Heteragrion Selys, 1862 (Zygoptera: Heteragrionidae): male morphology, new species and illustrated key</mods:title>
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<mods:affiliation id="59AE9F740D33ED917B11E80C31D1A57E">Grupo de Entomología Universidad de Antioquia (GEUA), Universidad de Antioquia, Medellin 50010, Colombia corneliobota @ gmail. com; https: // orcid. org / 0000 - 0002 - 6555 - 7913</mods:affiliation>
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<mods:affiliation id="F9381F48F6EEB4595469082E727F2599">Division of Invertebrate Zoology, American Museum of Natural History, USA jware @ amnh. org; https: // orcid. org / 0000 - 0002 - 4066 - 7681</mods:affiliation>
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<taxonomicName id="3CED87AFFFDD91366AD7F902A9F3A7E4" ID-CoL="3L74B" authority="Williamson, 1919" authorityName=": Williamson" authorityYear="1919" box="[151,644,1750,1776]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="ictericum">
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<emphasis id="C999203EFFDD91366AD7F902AAD2A7E4" bold="true" box="[151,421,1750,1776]" italics="true" pageId="53" pageNumber="54">Heteragrion ictericum</emphasis>
<bibRefCitation id="9F7C81DDFFDD91366BECF902A9F3A7E4" author="Williamson, E. B." box="[428,644,1750,1776]" pageId="53" pageNumber="54" pagination="1 - 87" refId="ref67331" refString="Williamson, E. B. (1919) Results of the University of Michigan - Williamson Expedition to Colombia, 1916 - 1917. IV. Notes on species of the genus Heteragrion Selys with description of new species (Odonata). Occasional Papers Museum of Zoology, University of Michigan, 68, 1 - 87." type="journal article" year="1919">Williamson, 1919</bibRefCitation>
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(
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)
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<paragraph id="FB52FC2CFFDD91366AD7F894A9C4A643" blockId="53.[151,691,1855,1880]" box="[151,691,1855,1880]" pageId="53" pageNumber="54">[Key locator: Key to Group A species, couplet 5 (4)]</paragraph>
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<taxonomicName id="3CED87AFFFDD91366AD7F852A9C1A68B" ID-CoL="3L74B" authority=": Williamson 1919: 17 - 20" authorityName=": Williamson" authorityPageNumber="17 - 20" authorityYear="1919" box="[151,694,1925,1952]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="ictericum">
<emphasis id="C999203EFFDD91366AD7F852AAE4A68B" box="[151,403,1926,1951]" italics="true" pageId="53" pageNumber="54">Heteragrion ictericum</emphasis>
:
<bibRefCitation id="9F7C81DDFFDD91366BE0F851A9C1A68B" author="Williamson, E. B." box="[416,694,1925,1951]" pageId="53" pageNumber="54" pagination="1 - 87" refId="ref67331" refString="Williamson, E. B. (1919) Results of the University of Michigan - Williamson Expedition to Colombia, 1916 - 1917. IV. Notes on species of the genus Heteragrion Selys with description of new species (Odonata). Occasional Papers Museum of Zoology, University of Michigan, 68, 1 - 87." type="journal article" year="1919">Williamson 1919: 1720</bibRefCitation>
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(key to species, ♁ in couplet c1,
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in couplet c2; description of ♁ and
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, illustrations of wings, ♁ and
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head, genital valves in ventral view, head profile, appendages in mediodorsal and lateral view, ♁ and
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prothorax; figs. 8, 1415, 54, 65, 7475, 134135);—
<treatmentCitation id="7A4CDA3DFFDD91366E7DF819AE3CA6F3" author="De Marmels, J." box="[1085,1355,1997,2024]" page="229" pageId="53" pageNumber="54" year="1987">
<bibRefCitation id="9F7C81DDFFDD91366E7DF819AE3CA6F3" author="De Marmels, J." box="[1085,1355,1997,2024]" pageId="53" pageNumber="54" pagination="225 - 238" refId="ref61368" refString="De Marmels, J. (1987) On the type specimens of some neotropical Megapodagrionidae, with a description of Heteragrion pemon spec. nov. and Oxystigma caerulans spec. nov. from Venezuela (Zygoptera). Odonatologica, 16 (3), 225 - 238." type="journal article" year="1987">De Marmels 1987: 229</bibRefCitation>
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(states that
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<emphasis id="C999203EFFDE91356A89FF43AA21A1A4" box="[201,342,151,176]" italics="true" pageId="54" pageNumber="55">H. ictericum</emphasis>
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differs from
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<emphasis id="C999203EFFDE91356BADFF43A905A1A5" box="[493,626,151,177]" italics="true" pageId="54" pageNumber="55">H. silvarum</emphasis>
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in color pattern of thorax and in denticulation of
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valves; comparison with
<taxonomicName id="3CED87AFFFDE91356A94FF6FAA33A1C0" ID-CoL="3L74S" authorityName=": De Marmels" authorityYear="1987" box="[212,324,187,212]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="pemon">
<emphasis id="C999203EFFDE91356A94FF6FAA33A1C0" box="[212,324,187,212]" italics="true" pageId="54" pageNumber="55">H. pemon</emphasis>
</taxonomicName>
);—
<treatmentCitation id="7A4CDA3DFFDE91356B2FFF6FA908A1C1" author="De Marmels, J." box="[367,639,186,213]" page="335" pageId="54" pageNumber="55" year="1990">
<bibRefCitation id="9F7C81DDFFDE91356B2FFF6FA908A1C1" author="De Marmels, J." box="[367,639,186,213]" pageId="54" pageNumber="55" pagination="333 - 345" refId="ref61468" refString="De Marmels, J. (1990) An updated checklist of the Odonata of Venezuela. Odonatologica, 19 (4), 333 - 345." type="journal article" year="1990">De Marmels 1990: 335</bibRefCitation>
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(species recorded in
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);—
<treatmentCitation id="7A4CDA3DFFDE91356E59FF6FAE44A1C1" author="Garrison, R. W. &amp; von Ellenrieder, N. &amp; O'Brien, M. F." box="[1049,1331,186,213]" page="15" pageId="54" pageNumber="55" year="2003">
<bibRefCitation id="9F7C81DDFFDE91356E59FF6FAE44A1C1" author="Garrison, R. W. &amp; von Ellenrieder, N. &amp; O'Brien, M. F." box="[1049,1331,186,213]" pageId="54" pageNumber="55" pagination="1 - 73" refId="ref62149" refString="Garrison, R. W., von Ellenrieder, N. &amp; O'Brien, M. F. (2003) An annotated list of the name - bearing types of species - group names in Odonata preserved in the University of Michigan Museum of Zoology. Occasional papers of the Museum of Zoology University of Michigan, 736, 1 - 73." type="journal article" year="2003">
Garrison
<emphasis id="C999203EFFDE91356EC7FF6FAFB4A1C1" box="[1159,1219,187,213]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
2003: 15
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(data on
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);—
<treatmentCitation id="7A4CDA3DFFDE91356B64FF0BA977A1ED" author="Lencioni, F. A. A." box="[292,512,222,249]" page="132" pageId="54" pageNumber="55" year="2005">
<bibRefCitation id="9F7C81DDFFDE91356B64FF0BA977A1ED" author="Lencioni, F. A. A." box="[292,512,222,249]" pageId="54" pageNumber="55" refId="ref63138" refString="Lencioni, F. A. A. (2005) Damselflies of Brazil. An illustrated identification guide. 1 - Non - Coenagrionidae families. All Print Editora, Sao Paulo, iv + 324 pp." type="book" year="2005">Lencioni 2005: 132</bibRefCitation>
</treatmentCitation>
(data on description and distribution, illustration from
<bibRefCitation id="9F7C81DDFFDE91356E21FF0BAE53A1ED" author="Williamson, E. B." box="[1121,1316,223,249]" pageId="54" pageNumber="55" pagination="1 - 87" refId="ref67331" refString="Williamson, E. B. (1919) Results of the University of Michigan - Williamson Expedition to Colombia, 1916 - 1917. IV. Notes on species of the genus Heteragrion Selys with description of new species (Odonata). Occasional Papers Museum of Zoology, University of Michigan, 68, 1 - 87." type="journal article" year="1919">Williamson 1919</bibRefCitation>
; figs. 81a e);—
<treatmentCitation id="7A4CDA3DFFDE91356A8DFED7AAAAA009" author="Garrison, R. W. &amp; von Ellenrieder, N. &amp; Louton, J. A." box="[205,477,259,285]" page="88" pageId="54" pageNumber="55" year="2010">
<bibRefCitation id="9F7C81DDFFDE91356A8DFED7AAAAA009" author="Garrison, R. W. &amp; von Ellenrieder, N. &amp; Louton, J. A." box="[205,477,259,285]" pageId="54" pageNumber="55" refId="ref62214" refString="Garrison, R. W., von Ellenrieder, N. &amp; Louton, J. A. (2010) Damselfly genera of the New World. An illustrated and Annotated Key to the Zygoptera. The Johns Hopkins University Press, Baltimore, Maryland, xiv + 490 pp." type="book" year="2010">
Garrison
<emphasis id="C999203EFFDE91356B7AFED7AA04A009" box="[314,371,259,285]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
2010: 88
</bibRefCitation>
</treatmentCitation>
(list of
<taxonomicName id="3CED87AFFFDE91356877FED7A9B6A008" ID-CoL="4W75" authorityName="Selys" authorityYear="1862" box="[567,705,259,284]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="C999203EFFDE91356877FED7A9B6A008" box="[567,705,259,284]" italics="true" pageId="54" pageNumber="55">Heteragrion</emphasis>
</taxonomicName>
species);—
<treatmentCitation id="7A4CDA3DFFDE91356906FED7AF16A009" author="von Ellenrieder, N." box="[838,1121,258,285]" page="59" pageId="54" pageNumber="55" year="2012">
<bibRefCitation id="9F7C81DDFFDE91356906FED7AF16A009" author="von Ellenrieder, N." box="[838,1121,258,285]" pageId="54" pageNumber="55" refId="ref67036" refString="https: // doi. org / 10.2305 / IUCN. UK. 2009 - 2. RLTS. T 159091 A 5309976. en von Ellenrieder, N. (2012) Odonata (dragonflies and damselflies) of the Kwamalasamutu region, Suriname. In: Conservation International (Ed.), A rapid biological assessment of the Kwamalasamutu region, southwestern Suriname. Conservation International, Arlington, Virginia, pp. 56 - 78. https: // doi. org / 10.1896 / 054.063.0105" type="book" year="2012">von Ellenrieder 2012: 59</bibRefCitation>
</treatmentCitation>
, 67, 73 (list of species from
<collectingCountry id="83FABCBCFFDE91356AD7FEF3AA71A055" box="[151,262,295,321]" name="Suriname" pageId="54" pageNumber="55">Suriname</collectingCountry>
; Plate III a);—
<treatmentCitation id="7A4CDA3DFFDE91356BEEFEF3A9F5A055" author="Lencioni, F. A. A." box="[430,642,295,321]" httpUri="http://treatment.plazi.org/id/9F43878D7D16FFEE65FDFDAFFD38AA65" page="10" pageId="54" pageNumber="55" year="2013">
<bibRefCitation id="9F7C81DDFFDE91356BEEFEF3A9F5A055" author="Lencioni, F. A. A." box="[430,642,295,321]" pageId="54" pageNumber="55" pagination="1 - 80" refId="ref63177" refString="Lencioni, F. A. A. (2013) Diagnoses and discussion of the group 1 and 2 Brazilian species of Heteragrion, with descriptions of four new species (Odonata: Megapodagrionidae). Zootaxa, 3685 (1), 1 - 80. https: // doi. org / 10.11646 / zootaxa. 3685.1.1" type="journal article" year="2013">Lencioni 2013: 10</bibRefCitation>
</treatmentCitation>
(information on types, distribution, diagnosis; figs. 17, 33ac, 46df,
<quantity id="3C1551C9FFDE91356AD7FE9FABB5A071" box="[151,194,331,357]" metricMagnitude="-2" metricUnit="kg" metricValue="4.8" pageId="54" pageNumber="55" unit="g" value="48.0">48g</quantity>
h, 57, 72);—
<treatmentCitation id="7A4CDA3DFFDE91356B22FE9FA9C0A071" author="von Ellenrieder, N. &amp; Castro, B. W. &amp; Svensson, E." box="[354,695,331,357]" page="3" pageId="54" pageNumber="55" year="2017">
<bibRefCitation id="9F7C81DDFFDE91356B22FE9FA9C0A071" author="von Ellenrieder, N. &amp; Castro, B. W. &amp; Svensson, E." box="[354,695,331,357]" pageId="54" pageNumber="55" pagination="2104" refId="ref67270" refString="von Ellenrieder, N., Castro, B. W. &amp; Svensson, E. (2017) Checklist of the dragonflies and damselflies from Guyana (Insecta: Odonata), with new records from the country. Check List, 13 (2), 2104. https: // doi. org / 10.15560 / 13.2.2104" type="journal article" year="2017">
von Ellenrieder
<emphasis id="C999203EFFDE9135685CFE9FA920A071" box="[540,599,331,357]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
2017: 3
</bibRefCitation>
</treatmentCitation>
(list of species from
<collectingCountry id="83FABCBCFFDE913569F0FE9FAF7EA071" box="[944,1033,331,357]" name="Guyana" pageId="54" pageNumber="55">Guyana</collectingCountry>
);—Garcia-Jr.
<emphasis id="C999203EFFDE91356EECFE9FAF91A071" box="[1196,1254,331,357]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
2021: 4 (list of species from
<collectingRegion id="392932CEFFDE91356B6BFEBBAACEA09D" box="[299,441,367,393]" country="Brazil" name="Amapa" pageId="54" pageNumber="55">Amapá state</collectingRegion>
,
<collectingCountry id="83FABCBCFFDE91356B84FEBBA97DA09D" box="[452,522,367,393]" name="Brazil" pageId="54" pageNumber="55">Brazil</collectingCountry>
);—
<treatmentCitation id="7A4CDA3DFFDE91356873FEBBA848A09D" author="Rochas, P. &amp; Minot, M. &amp; Meziere, N. &amp; Renoult, J. &amp; Uriot, Q." box="[563,831,367,393]" page="200" pageId="54" pageNumber="55" year="2022">
<bibRefCitation id="9F7C81DDFFDE91356873FEBBA848A09D" author="Rochas, P. &amp; Minot, M. &amp; Meziere, N. &amp; Renoult, J. &amp; Uriot, Q." box="[563,831,367,393]" pageId="54" pageNumber="55" pagination="175 - 224" refId="ref64861" refString="Rochas, P., Minot, M., Meziere, N., Renoult, J. &amp; Uriot, Q. (2022) Check-list of Odonata from French Guiana with notes on their distribution, ecology, and new state records. Odonatologica, 51 (3 / 4), 175 - 224. https: // doi. org / 10.60024 / odon. v 51 i 3 - 4. a 1" type="journal article" year="2022">
Rochas
<emphasis id="C999203EFFDE913568CFFEBBA9B0A09D" box="[655,711,367,393]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
2022: 200
</bibRefCitation>
</treatmentCitation>
(species recorded from French Guiana).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B3F7AFA7FFDE91356A87FE46AA65A329" pageId="54" pageNumber="55" type="materials_examined">
<paragraph id="FB52FC2CFFDE91356A87FE46AFBEA0E1" blockId="54.[151,1437,151,717]" pageId="54" pageNumber="55">
<emphasis id="C999203EFFDE91356A87FE46AAC2A0B8" bold="true" box="[199,437,402,428]" pageId="54" pageNumber="55">Material examined.</emphasis>
<materialsCitation id="4B85F671FFDE91356B80FE40AB84A0C5" ID-GBIF-Occurrence="4428728329" collectingDate="1957-02" collectionCode="ABMM" collectorName="A. B. M. Machado" country="Brazil" location="Represa" municipality="Serra do Navio" pageId="54" pageNumber="55" specimenCount="1" stateProvince="Amapa">
1♁
<collectingCountry id="83FABCBCFFDE91356BAFFE47A920A0B9" box="[495,599,403,429]" name="Brazil" pageId="54" pageNumber="55">BRAZIL</collectingCountry>
,
<collectingRegion id="392932CEFFDE91356825FE47A980A0B9" box="[613,759,403,429]" country="Brazil" name="Amapa" pageId="54" pageNumber="55">Amapá state</collectingRegion>
,
<collectingMunicipality id="1B366656FFDE91356947FE47A8CBA0B9" box="[775,956,403,429]" pageId="54" pageNumber="55">Serra do Navio</collectingMunicipality>
,
<location id="FE32AAF7FFDE9135698CFE47AF5EA0B9" LSID="urn:lsid:plazi:treatment:73444D3AFFDD91356AD7F902A8EDA3D9:FE32AAF7FFDE9135698CFE47AF5EA0B9" box="[972,1065,403,429]" country="Brazil" municipality="Serra do Navio" name="Represa" pageId="54" pageNumber="55" stateProvince="Amapa">Represa</location>
,
<date id="8F53DAECFFDE91356E79FE46AFFEA0B9" box="[1081,1161,402,429]" pageId="54" pageNumber="55" value="1957-02">
<collectingDate id="9F172304FFDE91356E79FE46AFFEA0B9" box="[1081,1161,402,429]" pageId="54" pageNumber="55" value="1957-02">ii.1957</collectingDate>
</date>
,
<collectorName id="561899FAFFDE91356ED8FE47AE17A0B9" box="[1176,1376,403,429]" pageId="54" pageNumber="55">A.B.M. Machado</collectorName>
leg.,
<collectionCode id="9DFC64E9FFDE91356AD7FE63AB84A0C5" box="[151,243,439,465]" pageId="54" pageNumber="55">ABMM</collectionCode>
</materialsCitation>
;
<materialsCitation id="4B85F671FFDE91356ABEFE6CAF37A0C5" ID-GBIF-Occurrence="4428728334" box="[254,1088,439,467]" collectingDate="2017-11-28" collectionCode="LABECO" collectorName="T. B. Miguel" country="Brazil" location="Resex Cajari" municipality="Laranjal do Jari" pageId="54" pageNumber="55" specimenCount="1" stateProvince="Amapa state">
1♁
<collectingMunicipality id="1B366656FFDE91356B68FE63AAADA0C5" box="[296,474,439,465]" pageId="54" pageNumber="55">Laranjal do Jari</collectingMunicipality>
,
<location id="FE32AAF7FFDE91356BA5FE63A900A0C5" LSID="urn:lsid:plazi:treatment:73444D3AFFDD91356AD7F902A8EDA3D9:FE32AAF7FFDE91356BA5FE63A900A0C5" box="[485,631,439,465]" country="Brazil" municipality="Laranjal do Jari" name="Resex Cajari" pageId="54" pageNumber="55" stateProvince="Amapa state">Resex Cajari</location>
,
<date id="8F53DAECFFDE913568C2FE63A98BA0C5" box="[642,764,439,465]" pageId="54" pageNumber="55" value="2017-11-28">
<collectingDate id="9F172304FFDE913568C2FE63A98BA0C5" box="[642,764,439,465]" pageId="54" pageNumber="55" value="2017-11-28">28.xi.2017</collectingDate>
</date>
,
<collectorName id="561899FAFFDE91356947FE63A8F9A0C5" box="[775,910,439,465]" pageId="54" pageNumber="55">T.B. Miguel</collectorName>
leg.,
<collectionCode id="9DFC64E9FFDE9135698CFE63AF37A0C5" box="[972,1088,439,465]" pageId="54" pageNumber="55">LABECO</collectionCode>
</materialsCitation>
;
<materialsCitation id="4B85F671FFDE91356E0AFE6CAB84A0E1" ID-GBIF-Occurrence="4428728446" collectingDate="1943-08-21" collectionCode="ABMM" country="Brazil" pageId="54" pageNumber="55" specimenCount="1" stateProvince="Amapa state">
1♁
<date id="8F53DAECFFDE91356E34FE63AF89A0C5" box="[1140,1278,439,465]" pageId="54" pageNumber="55" value="1943-08-21">
<collectingDate id="9F172304FFDE91356E34FE63AF89A0C5" box="[1140,1278,439,465]" pageId="54" pageNumber="55" value="1943-08-21">21.viii.1943</collectingDate>
</date>
, “Wilh”? leg.,
<collectionCode id="9DFC64E9FFDE91356AD7FE0FAB84A0E1" box="[151,243,475,501]" pageId="54" pageNumber="55">ABMM</collectionCode>
</materialsCitation>
;
<materialsCitation id="4B85F671FFDE91356ABFFE08AFB2A0E1" ID-GBIF-Occurrence="4428728327" box="[255,1221,475,503]" collectingDate="1922-06-26" collectorName="J. H. Williamson &amp; J. W. Strohm" country="Brazil" location="Manaus" municipality="Amazonas" pageId="54" pageNumber="55" specimenCount="1" stateProvince="Amapa state">
1♁
<collectingMunicipality id="1B366656FFDE91356B67FE0FAAEAA0E1" box="[295,413,475,501]" pageId="54" pageNumber="55">Amazonas</collectingMunicipality>
state,
<location id="FE32AAF7FFDE91356BA6FE0FA936A0E1" LSID="urn:lsid:plazi:treatment:73444D3AFFDD91356AD7F902A8EDA3D9:FE32AAF7FFDE91356BA6FE0FA936A0E1" box="[486,577,475,501]" country="Brazil" municipality="Amazonas" name="Manaus" pageId="54" pageNumber="55" stateProvince="Amapa state">Manaus</location>
,
<date id="8F53DAECFFDE9135680CFE0FA9B0A0E1" box="[588,711,475,501]" pageId="54" pageNumber="55" value="1922-06-26">
<collectingDate id="9F172304FFDE9135680CFE0FA9B0A0E1" box="[588,711,475,501]" pageId="54" pageNumber="55" value="1922-06-26">26.vi.1922</collectingDate>
</date>
,
<collectorName id="561899FAFFDE91356892FE0FA8F8A0E1" box="[722,911,475,501]" pageId="54" pageNumber="55">J. H. Williamson</collectorName>
&amp;
<collectorName id="561899FAFFDE913569F2FE0FAF4BA0E1" box="[946,1084,475,501]" pageId="54" pageNumber="55">J.W. Strohm</collectorName>
leg., FAAL
</materialsCitation>
.
</paragraph>
<paragraph id="FB52FC2CFFDE91356A87FE2BAA65A329" blockId="54.[151,1437,151,717]" pageId="54" pageNumber="55">
<emphasis id="C999203EFFDE91356A87FE2BAAB5A30C" bold="true" box="[199,450,510,537]" pageId="54" pageNumber="55">Known distribution.</emphasis>
<collectingCountry id="83FABCBCFFDE91356B91FE2BA962A30D" box="[465,533,511,537]" name="Brazil" pageId="54" pageNumber="55">Brazil</collectingCountry>
(states of Amazonas and
<collectingRegion id="392932CEFFDE91356919FE2BA8D9A30D" box="[857,942,511,537]" country="Brazil" name="Amapa" pageId="54" pageNumber="55">Amapá</collectingRegion>
),
<collectingCountry id="83FABCBCFFDE91356989FE2BAF54A30D" box="[969,1059,511,537]" name="Guyana" pageId="54" pageNumber="55">Guyana</collectingCountry>
, French Guiana,
<collectingCountry id="83FABCBCFFDE91356EBAFE2BAE12A30D" box="[1274,1381,511,537]" name="Suriname" pageId="54" pageNumber="55">Suriname</collectingCountry>
and
<collectingCountry id="83FABCBCFFDE91356AD7FDF7AA79A329" box="[151,270,547,573]" name="Venezuela" pageId="54" pageNumber="55">Venezuela</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="B3F7AFA7FFDE91356A87FD92A8EDA3D9" pageId="54" pageNumber="55" type="diagnosis">
<paragraph id="FB52FC2CFFDE91356A87FD92A8EDA3D9" blockId="54.[151,1437,151,717]" pageId="54" pageNumber="55">
<emphasis id="C999203EFFDE91356A87FD92AAA8A375" bold="true" box="[199,479,582,609]" pageId="54" pageNumber="55">Diagnosis and remarks.</emphasis>
This species possesses a tubercle or apophysis on the ventral BP portion, a feature that is shared only with
<taxonomicName id="3CED87AFFFDE91356B39FDBFA96BA391" box="[377,540,619,645]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="triangulare">
<emphasis id="C999203EFFDE91356B39FDBFA96BA391" box="[377,540,619,645]" italics="true" pageId="54" pageNumber="55">H. triangulare</emphasis>
</taxonomicName>
and
<taxonomicName id="3CED87AFFFDE91356815FDBFA9B3A390" authorityName=": De Marmels" authorityYear="1987" box="[597,708,619,644]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="pemon">
<emphasis id="C999203EFFDE91356815FDBFA9B3A390" box="[597,708,619,644]" italics="true" pageId="54" pageNumber="55">H. pemon</emphasis>
</taxonomicName>
. The following character combination distinguishes this species from the remainder of Group A species: BP the shortest portion of the cercus (
<figureCitation id="63D6E0A9FFDE91356E4AFD5BAFE5A3BD" box="[1034,1170,655,681]" captionStart="FIGURE 38" captionStartId="53.[151,250,1608,1632]" captionTargetBox="[244,1343,785,1572]" captionTargetId="figure-184@53.[231,1355,772,1584]" captionTargetPageId="53" captionText="FIGURE 38. Heteragrion ictericum: (a) anterior view of head; (b) lateral view of pterothorax; (c) mediodorsal view of cercus; (d) lateral view of cercus. BP: basal portion; ML: medial lobe." figureDoi="http://doi.org/10.5281/zenodo.10008636" httpUri="https://zenodo.org/record/10008636/files/figure.png" pageId="54" pageNumber="55">Figs. 38cd</figureCitation>
); MP the longest, ML a broad plate, bearing small teeth, roughly squared apex (
<figureCitation id="63D6E0A9FFDE91356946FD67A8F9A3D9" box="[774,910,691,717]" captionStart="FIGURE 38" captionStartId="53.[151,250,1608,1632]" captionTargetBox="[244,1343,785,1572]" captionTargetId="figure-184@53.[231,1355,772,1584]" captionTargetPageId="53" captionText="FIGURE 38. Heteragrion ictericum: (a) anterior view of head; (b) lateral view of pterothorax; (c) mediodorsal view of cercus; (d) lateral view of cercus. BP: basal portion; ML: medial lobe." figureDoi="http://doi.org/10.5281/zenodo.10008636" httpUri="https://zenodo.org/record/10008636/files/figure.png" pageId="54" pageNumber="55">Figs. 38cd</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>