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<document id="63430C30DCEC1A71EB13668BF4F984C8" ID-CLB-Dataset="273743" ID-DOI="10.11646/zootaxa.5356.1.1" ID-GBIF-Dataset="38e48899-447b-40cc-b9bd-e84bb0ac221e" ID-ISSN="1175-5334" ID-Zenodo-Dep="10008529" ID-ZooBank="F3CE1E00-45BB-44C8-8911-1A355BFD223C" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1697461253574" checkinUser="plazi" docAuthor="Vilela, Diogo Silva, Lencioni, Frederico A. A., Bota-Sierra, Cornelio A., Ware, Jessica L. &amp; Bispo, Pitágoras C." docDate="2023" docId="73444D3AFFC0912B6AD7FF43AFE5A7DC" docLanguage="en" docName="zootaxa.5356.1.1.pdf" docOrigin="Zootaxa 5356 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5356.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Heteragrion cooki : Daigle &amp; Tennessen 2000" docType="treatment" docVersion="8" lastPageNumber="41" masterDocId="8F7D3542FFE891036A40FFD4AB77A114" masterDocTitle="Taxonomic revision of the Neotropical genus Heteragrion Selys, 1862 (Zygoptera: Heteragrionidae): male morphology, new species and illustrated key" masterLastPageNumber="96" masterPageNumber="1" pageNumber="41" updateTime="1699697007752" updateUser="plazi">
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<mods:title id="2F03C55962772FCD5992640455509894">Taxonomic revision of the Neotropical genus Heteragrion Selys, 1862 (Zygoptera: Heteragrionidae): male morphology, new species and illustrated key</mods:title>
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<mods:namePart id="1B377972B42272C463CC2194F74CAF0D">Vilela, Diogo Silva</mods:namePart>
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<mods:namePart id="7BDE965AAA081B2307C6F1662F89F2B6">Lencioni, Frederico A. A.</mods:namePart>
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<mods:affiliation id="20FAF082F3FBD8D1152F735E75911051">Rua Anibal, 216, Jardim Coleginho, CEP 12310 - 780, Jacareí, SP, Brazil lencioni. odonata @ gmail. com; https: // orcid. org / 0000 - 0002 - 3912 - 9181</mods:affiliation>
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<mods:affiliation id="3562BAD0014AD1CCB10A7740E694DC7B">Division of Invertebrate Zoology, American Museum of Natural History, USA jware @ amnh. org; https: // orcid. org / 0000 - 0002 - 4066 - 7681</mods:affiliation>
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<mods:affiliation id="B97F66B5BC91953AF3D7A386CBB0EF27">Laboratório de Biologia Aquática, Departamento de Ciências Biológicas, Faculdade de Ciências e Letras de Assis, Universidade Estadual Paulista, Assis, SP, Brazil deeogoo @ gmail. com; https: // orcid. org / 0000 - 0001 - 6510 - 7018 &amp; Laboratório de Biologia Aquática, Departamento de Ciências Biológicas, Faculdade de Ciências e Letras de Assis, Universidade Estadual Paulista, Assis, SP, Brazil deeogoo @ gmail. com; https: // orcid. org / 0000 - 0001 - 6510 - 7018 &amp; pitagoras. bispo @ unesp. br; https: // orcid. org / 0000 - 0002 - 7356 - 8882 * Corresponding author</mods:affiliation>
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<treatment id="73444D3AFFC0912B6AD7FF43AFE5A7DC" ID-GBIF-Taxon="214231529" LSID="urn:lsid:plazi:treatment:73444D3AFFC0912B6AD7FF43AFE5A7DC" httpUri="http://treatment.plazi.org/id/73444D3AFFC0912B6AD7FF43AFE5A7DC" lastPageNumber="41" pageId="40" pageNumber="41">
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<heading id="A01A4B40FFC0912B6AD7FF43A9CCA1A5" bold="true" box="[151,699,151,178]" fontSize="11" level="2" pageId="40" pageNumber="41" reason="4">
<taxonomicName id="3CED87AFFFC0912B6AD7FF43A9CCA1A5" ID-CoL="3L744" authority="Daigle &amp; Tennessen, 2000" authorityName=": Daigle &amp; Tennessen" authorityYear="2000" box="[151,699,151,178]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="cooki">
<emphasis id="C999203EFFC0912B6AD7FF43A9CCA1A5" bold="true" box="[151,699,151,178]" pageId="40" pageNumber="41">
<emphasis id="C999203EFFC0912B6AD7FF43AA03A1A6" bold="true" box="[151,372,151,178]" italics="true" pageId="40" pageNumber="41">Heteragrion cooki</emphasis>
<bibRefCitation id="9F7C81DDFFC0912B6B3BFF43A9CCA1A5" author="Daigle, J. J. &amp; Tennessen, K. J." box="[379,699,151,177]" pageId="40" pageNumber="41" pagination="255 - 259" refId="ref61201" refString="Daigle, J. J. &amp; Tennessen, K. J. (2000) Heteragrion cooki spec. nov. from Ecuador (Zygoptera: Megapodagrionidae). Odonatologica, 29 (3), 255 - 259." type="journal article" year="2000">Daigle &amp; Tennessen, 2000</bibRefCitation>
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(
<figureCitation id="63D6E0A9FFC0912B6ADFFF6FAA52A1C1" box="[159,293,187,213]" captionStart="FIGURE 26" captionStartId="40.[151,250,1293,1317]" captionTargetBox="[246,1342,519,1257]" captionTargetId="figure-109@40.[233,1354,506,1270]" captionTargetPageId="40" captionText="FIGURE 26. Heteragrion cooki: (a) anterior view of head; (b) lateral view of pterothorax; (c–d) mediodorsal view of cercus. ML: medial lobe." figureDoi="http://doi.org/10.5281/zenodo.10008605" httpUri="https://zenodo.org/record/10008605/files/figure.png" pageId="40" pageNumber="41">Figs. 26a–d</figureCitation>
)
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<paragraph id="FB52FC2CFFC0912B6AD7FF2DA9F8A005" blockId="40.[151,655,249,273]" box="[151,655,249,273]" pageId="40" pageNumber="41">[Key locator: Key to Group A species, couplet 8’]</paragraph>
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<treatmentCitation id="7A4CDA3DFFC0912B6AD7FEEDA867A047" author="Daigle, J. J. &amp; Tennessen, K. J." box="[151,784,312,339]" page="259" pageId="40" pageNumber="41" year="2000">
<taxonomicName id="3CED87AFFFC0912B6AD7FEEDA867A047" ID-CoL="3L744" authority=": Daigle &amp; Tennessen 2000: 255 - 259" authorityName=": Daigle &amp; Tennessen" authorityPageNumber="255 - 259" authorityYear="2000" box="[151,784,312,339]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="cooki">
<emphasis id="C999203EFFC0912B6AD7FEEDAA10A046" box="[151,359,312,338]" italics="true" pageId="40" pageNumber="41">Heteragrion cooki</emphasis>
:
<bibRefCitation id="9F7C81DDFFC0912B6B37FEECA867A047" author="Daigle, J. J. &amp; Tennessen, K. J." box="[375,784,312,339]" pageId="40" pageNumber="41" pagination="255 - 259" refId="ref61201" refString="Daigle, J. J. &amp; Tennessen, K. J. (2000) Heteragrion cooki spec. nov. from Ecuador (Zygoptera: Megapodagrionidae). Odonatologica, 29 (3), 255 - 259." type="journal article" year="2000">Daigle &amp; Tennessen 2000: 255–259</bibRefCitation>
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(description of ♁ and
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; diagnosis and comparison with
<taxonomicName id="3CED87AFFFC0912B6AD7FE89AA62A062" ID-CoL="3L73L" authorityName=": Williamson" authorityYear="1919" box="[151,277,348,374]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="alienum">
<emphasis id="C999203EFFC0912B6AD7FE89AA62A062" box="[151,277,348,374]" italics="true" pageId="40" pageNumber="41">H. alienum</emphasis>
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and
<taxonomicName id="3CED87AFFFC0912B6B0FFE89A975A062" ID-CoL="3L73R" authorityName=": Selys" authorityYear="1862" box="[335,514,349,374]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="aurantiacum">
<emphasis id="C999203EFFC0912B6B0FFE89A975A062" box="[335,514,349,374]" italics="true" pageId="40" pageNumber="41">H. aurantiacum</emphasis>
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; figs. 1–10);—
<treatmentCitation id="7A4CDA3DFFC0912B68EDFE89A8CEA063" author="De Marmels, J." box="[685,953,348,375]" page="447" pageId="40" pageNumber="41" year="2004">
<bibRefCitation id="9F7C81DDFFC0912B68EDFE89A8CEA063" author="De Marmels, J." box="[685,953,348,375]" pageId="40" pageNumber="41" pagination="439 - 458" refId="ref61531" refString="De Marmels, J. (2004) Heteragrion makiritare sp. nov., with descriptions of hitherto unknown females and larvae of other species from Venezuela (Odonata: Megapodagrionidae, Lestidae). International Journal of Odonatology, 7 (3), 439 - 458. https: // doi. org / 10.1080 / 13887890.2004.9748230" type="journal article" year="2004">De Marmels 2004: 447</bibRefCitation>
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(comparison with
<taxonomicName id="3CED87AFFFC0912B6ED0FE89AE5EA062" ID-CoL="3L74H" authorityName=": De Marmels" authorityYear="2004" box="[1168,1321,348,374]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="makiritare">
<emphasis id="C999203EFFC0912B6ED0FE89AE5EA062" box="[1168,1321,348,374]" italics="true" pageId="40" pageNumber="41">H. makiritare</emphasis>
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and other
<taxonomicName id="3CED87AFFFC0912B6AD7FE55AA56A08E" ID-CoL="4W75" authorityName="Selys" authorityYear="1862" box="[151,289,385,410]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="C999203EFFC0912B6AD7FE55AA56A08E" box="[151,289,385,410]" italics="true" pageId="40" pageNumber="41">Heteragrion</emphasis>
</taxonomicName>
species; figs. 18–19);—
<treatmentCitation id="7A4CDA3DFFC0912B6870FE55A8C2A08F" author="Tennessen, K. J. &amp; Johnson, J. T." box="[560,949,384,411]" page="9" pageId="40" pageNumber="41" year="2008">
<bibRefCitation id="9F7C81DDFFC0912B6870FE55A8C2A08F" author="Tennessen, K. J. &amp; Johnson, J. T." box="[560,949,384,411]" pageId="40" pageNumber="41" pagination="9 - 11" refId="ref65761" refString="Tennessen, K. J. &amp; Johnson, J. T. (2008) Chasing dragonflies in southern Ecuador. ARGIA, The News Journal of the DSA, 20 (2), 9 - 11." type="journal article" year="2008">Tennessen &amp; Johnson 2008: 9–11</bibRefCitation>
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(list of species from southern
<collectingCountry id="83FABCBCFFC0912B6F52FE54AE04A08E" box="[1298,1395,384,410]" name="Ecuador" pageId="40" pageNumber="41">Ecuador</collectingCountry>
);—
<treatmentCitation id="7A4CDA3DFFC0912B6AD7FE70AAC2A0AB" author="Garrison, R. W. &amp; von Ellenrieder, N. &amp; Louton, J. A." box="[151,437,420,447]" page="88" pageId="40" pageNumber="41" year="2010">
<bibRefCitation id="9F7C81DDFFC0912B6AD7FE70AAC2A0AB" author="Garrison, R. W. &amp; von Ellenrieder, N. &amp; Louton, J. A." box="[151,437,420,447]" pageId="40" pageNumber="41" refId="ref62214" refString="Garrison, R. W., von Ellenrieder, N. &amp; Louton, J. A. (2010) Damselfly genera of the New World. An illustrated and Annotated Key to the Zygoptera. The Johns Hopkins University Press, Baltimore, Maryland, xiv + 490 pp." type="book" year="2010">
Garrison
<emphasis id="C999203EFFC0912B6B45FE71AA36A0AA" box="[261,321,420,446]" italics="true" pageId="40" pageNumber="41">et al.</emphasis>
2010: 88
</bibRefCitation>
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, 91 (list of
<taxonomicName id="3CED87AFFFC0912B6807FE71A9A6A0AA" ID-CoL="4W75" authorityName="Selys" authorityYear="1862" box="[583,721,421,446]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="C999203EFFC0912B6807FE71A9A6A0AA" box="[583,721,421,446]" italics="true" pageId="40" pageNumber="41">Heteragrion</emphasis>
</taxonomicName>
species; figs. 387–388);—
<treatmentCitation id="7A4CDA3DFFC0912B6E47FE70AEEBA0AB" author="Rivas-Torres, A. &amp; Sanmartin-Villar, I. &amp; Gabela-Flores, M. V. &amp; Cordero-Rivera, A." box="[1031,1436,420,447]" page="123" pageId="40" pageNumber="41" year="2017">
<bibRefCitation id="9F7C81DDFFC0912B6E47FE70AEEBA0AB" author="Rivas-Torres, A. &amp; Sanmartin-Villar, I. &amp; Gabela-Flores, M. V. &amp; Cordero-Rivera, A." box="[1031,1436,420,447]" pageId="40" pageNumber="41" pagination="123 - 135" refId="ref64796" refString="Rivas-Torres, A., Sanmartin-Villar, I., Gabela-Flores, M. V. &amp; Cordero-Rivera, A. (2017) Demographics and behaviour of Heteragrion cooki, a forest damselfly endemic to Ecuador (Odonata). International Journal of Odonatology, 20 (2), 123 - 135. https: // doi. org / 10.1080 / 13887890.2017.1336495" type="journal article" year="2017">
Rivas-Torres
<emphasis id="C999203EFFC0912B6EE7FE71AF93A0AA" box="[1191,1252,420,446]" italics="true" pageId="40" pageNumber="41">et al.</emphasis>
2017: 123–135
</bibRefCitation>
</treatmentCitation>
(demographics and behavioral ecology);—
<treatmentCitation id="7A4CDA3DFFC0912B6832FE1CA89BA0F6" author="Medina-Espinoza, E. F." box="[626,1004,456,483]" page="236" pageId="40" pageNumber="41" year="2022">
<bibRefCitation id="9F7C81DDFFC0912B6832FE1CA89BA0F6" author="Medina-Espinoza, E. F." box="[626,1004,456,483]" pageId="40" pageNumber="41" pagination="236 - 240" refId="ref63663" refString="Medina-Espinoza, E. F. (2022) First record of Heteragrion cooki from Peru (Odonata: Heteragrionidae). Notulae odonatologicae, 9 (6), 236 - 240. https: // doi. org / 10.60024 / zenodo. 4268583" type="journal article" year="2022">Medina-Espinoza 2022: 236–240</bibRefCitation>
</treatmentCitation>
(first record to
<collectingCountry id="83FABCBCFFC0912B6EDCFE1DAFA3A0F7" box="[1180,1236,457,483]" name="Peru" pageId="40" pageNumber="41">Peru</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="AF92ACA4FFC0912B6AD7FAD9AA3EA45E" ID-DOI="http://doi.org/10.5281/zenodo.10008605" ID-Zenodo-Dep="10008605" httpUri="https://zenodo.org/record/10008605/files/figure.png" pageId="40" pageNumber="41" startId="40.[151,250,1293,1317]" targetBox="[246,1342,519,1257]" targetPageId="40" targetType="figure">
<paragraph id="FB52FC2CFFC0912B6AD7FAD9AA3EA45E" blockId="40.[151,1437,1293,1354]" pageId="40" pageNumber="41">
<emphasis id="C999203EFFC0912B6AD7FAD9AA56A432" bold="true" box="[151,289,1293,1318]" pageId="40" pageNumber="41">FIGURE 26.</emphasis>
<taxonomicName id="3CED87AFFFC0912B6B68FADAAA94A432" authorityName=": Daigle &amp; Tennessen" authorityYear="2000" box="[296,483,1294,1318]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="cooki">
<emphasis id="C999203EFFC0912B6B68FADAAA94A432" box="[296,483,1294,1318]" italics="true" pageId="40" pageNumber="41">Heteragrion cooki</emphasis>
</taxonomicName>
: (a) anterior view of head; (b) lateral view of pterothorax; (c–d) mediodorsal view of cercus. ML: medial lobe.
</paragraph>
</caption>
<subSubSection id="B3F7AFA7FFC0912B6A87FA5AA9E7A72D" pageId="40" pageNumber="41" type="materials_examined">
<paragraph id="FB52FC2CFFC0912B6A87FA5AAAE1A701" blockId="40.[151,1436,1422,1737]" pageId="40" pageNumber="41">
<emphasis id="C999203EFFC0912B6A87FA5AAAD8A4BC" bold="true" box="[199,431,1422,1448]" pageId="40" pageNumber="41">Material examined.</emphasis>
<materialsCitation id="4B85F671FFC0912B6BF5FA44AAFEA4D9" ID-GBIF-Occurrence="4428728429" collectingDate="1941-07-04" collectionCode="ABMM" collectorName="D. B. Laddey" country="Ecuador" elevation="300" latitude="-3.6333" location="Piedras" longLatPrecision="7" longitude="-79.9166" pageId="40" pageNumber="41" specimenCount="1" stateProvince="El Oro">
1♁
<collectingCountry id="83FABCBCFFC0912B6B9EFA5BA91FA4BD" box="[478,616,1423,1449]" name="Ecuador" pageId="40" pageNumber="41">ECUADOR</collectingCountry>
,
<collectingRegion id="392932CEFFC0912B6832FA5BA85FA4BD" box="[626,808,1423,1449]" country="Ecuador" name="El Oro" pageId="40" pageNumber="41">El Oro Province</collectingRegion>
,
<location id="FE32AAF7FFC0912B6972FA5BA8F0A4BD" LSID="urn:lsid:plazi:treatment:73444D3AFFC0912B6AD7FF43AFE5A7DC:FE32AAF7FFC0912B6972FA5BA8F0A4BD" box="[818,903,1423,1449]" country="Ecuador" latitude="-3.6333" longLatPrecision="7" longitude="-79.9166" name="Piedras" pageId="40" pageNumber="41" stateProvince="El Oro">Piedras</location>
,
<date id="8F53DAECFFC0912B69D1FA5BAF72A4BD" box="[913,1029,1423,1449]" pageId="40" pageNumber="41" value="1941-07-04">
<collectingDate id="9F172304FFC0912B69D1FA5BAF72A4BD" box="[913,1029,1423,1449]" pageId="40" pageNumber="41" value="1941-07-04">4.vii.1941</collectingDate>
</date>
, (
<geoCoordinate id="9ED99AEBFFC0912B6E57FA5AAF05A4BC" box="[1047,1138,1422,1448]" degrees="3.6333" direction="south" orientation="latitude" pageId="40" pageNumber="41" precision="5" value="-3.6333">-3.6333</geoCoordinate>
,
<geoCoordinate id="9ED99AEBFFC0912B6E3CFA5AAF94A4BC" box="[1148,1251,1422,1448]" degrees="79.9166" direction="west" orientation="longitude" pageId="40" pageNumber="41" precision="5" value="-79.9166">-79.9166</geoCoordinate>
,
<elevation id="70C01B1FFFC0912B6EADFA5BAE23A4BD" box="[1261,1364,1423,1449]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="40" pageNumber="41" unit="m" value="300.0">
<quantity id="3C1551C9FFC0912B6EADFA5BAE5BA4BD" box="[1261,1324,1423,1449]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="40" pageNumber="41" unit="m" value="300.0">300m</quantity>
asl
</elevation>
),
<collectorName id="561899FAFFC0912B6F27FA5BAB9CA4D9" pageId="40" pageNumber="41">D.B. Laddey</collectorName>
leg.,
<collectionCode id="9DFC64E9FFC0912B6B6DFA67AAFEA4D9" box="[301,393,1459,1485]" pageId="40" pageNumber="41">ABMM</collectionCode>
</materialsCitation>
;
<materialsCitation id="4B85F671FFC0912B6BD7FA60AA7BA4E5" ID-GBIF-Occurrence="4428728326" collectingDate="2012-11-30" collectionCode="ECOEVO" collectorName="A. C. Rivera" country="Ecuador" elevation="65" latitude="-0.0834" location="Bosque Seco Lalo Loor" longLatPrecision="7" longitude="-80.1485" pageId="40" pageNumber="41" specimenCount="1" stateProvince="El Oro Province">
4♁♁
<location id="FE32AAF7FFC0912B6B99FA67A99CA4D9" LSID="urn:lsid:plazi:treatment:73444D3AFFC0912B6AD7FF43AFE5A7DC:FE32AAF7FFC0912B6B99FA67A99CA4D9" box="[473,747,1459,1485]" country="Ecuador" latitude="-0.0834" longLatPrecision="7" longitude="-80.1485" name="Bosque Seco Lalo Loor" pageId="40" pageNumber="41" stateProvince="El Oro Province">Bosque Seco Lalo Loor</location>
,
<date id="8F53DAECFFC0912B68B9FA67A804A4D9" box="[761,883,1459,1485]" pageId="40" pageNumber="41" value="2012-11-30">
<collectingDate id="9F172304FFC0912B68B9FA67A804A4D9" box="[761,883,1459,1485]" pageId="40" pageNumber="41" value="2012-11-30">30.xi.2012</collectingDate>
</date>
, (
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,
<geoCoordinate id="9ED99AEBFFC0912B69B2FA66AF2EA4D9" box="[1010,1113,1458,1485]" degrees="80.1485" direction="west" orientation="longitude" pageId="40" pageNumber="41" precision="5" value="-80.1485">-80.1485</geoCoordinate>
,
<elevation id="70C01B1FFFC0912B6E26FA66AFB3A4D9" box="[1126,1220,1458,1485]" metricMagnitude="1" metricUnit="m" metricValue="6.5" pageId="40" pageNumber="41" unit="m" value="65.0">
<quantity id="3C1551C9FFC0912B6E26FA66AFEEA4D9" box="[1126,1177,1458,1485]" metricMagnitude="1" metricUnit="m" metricValue="6.5" pageId="40" pageNumber="41" unit="m" value="65.0">65m</quantity>
asl
</elevation>
),
<collectorName id="561899FAFFC0912B6E99FA67AE14A4D9" box="[1241,1379,1459,1485]" pageId="40" pageNumber="41">A.C. Rivera</collectorName>
leg.,
<collectionCode id="9DFC64E9FFC0912B6AD7FA03AA7BA4E5" box="[151,268,1495,1521]" pageId="40" pageNumber="41">ECOEVO</collectionCode>
</materialsCitation>
; 6♁♁
<materialsCitation id="4B85F671FFC0912B6B17FA03AAE5A701" ID-GBIF-Occurrence="4428728421" collectingDate="2016-06-18" collectionCode="ECOEVO" collectorName="I. Sanmartin-Villar &amp; A. Rivas-Torres" country="Ecuador" elevation="532" latitude="-3.6529" location="Canton Pinas" longLatPrecision="7" longitude="-79.7652" pageId="40" pageNumber="41" specimenCount="1" stateProvince="El Oro">
<collectingRegion id="392932CEFFC0912B6B17FA03AAD1A4E5" box="[343,422,1495,1521]" country="Ecuador" name="El Oro" pageId="40" pageNumber="41">El Oro</collectingRegion>
,
<location id="FE32AAF7FFC0912B6BF2FA03A93EA4E5" LSID="urn:lsid:plazi:treatment:73444D3AFFC0912B6AD7FF43AFE5A7DC:FE32AAF7FFC0912B6BF2FA03A93EA4E5" box="[434,585,1495,1521]" country="Ecuador" latitude="-3.6529" longLatPrecision="7" longitude="-79.7652" name="Canton Pinas" pageId="40" pageNumber="41" stateProvince="El Oro">Cantón Piñas</location>
,
<date id="8F53DAECFFC0912B6815FA02A9B8A4E4" box="[597,719,1494,1520]" pageId="40" pageNumber="41" value="2016-06-18">
<collectingDate id="9F172304FFC0912B6815FA02A9B8A4E4" box="[597,719,1494,1520]" pageId="40" pageNumber="41" value="2016-06-18">18.vi.2016</collectingDate>
</date>
, (
<geoCoordinate id="9ED99AEBFFC0912B68A3FA02A849A4E5" box="[739,830,1494,1521]" degrees="3.6529" direction="south" orientation="latitude" pageId="40" pageNumber="41" precision="5" value="-3.6529">-3.6529</geoCoordinate>
,
<geoCoordinate id="9ED99AEBFFC0912B6909FA02A8C7A4E5" box="[841,944,1494,1521]" degrees="79.7652" direction="west" orientation="longitude" pageId="40" pageNumber="41" precision="5" value="-79.7652">-79.7652</geoCoordinate>
,
<elevation id="70C01B1FFFC0912B69FCFA02AF52A4E5" box="[956,1061,1494,1521]" metricMagnitude="2" metricUnit="m" metricValue="5.32" pageId="40" pageNumber="41" unit="m" value="532.0">
<quantity id="3C1551C9FFC0912B69FCFA02A88BA4E5" box="[956,1020,1494,1521]" metricMagnitude="2" metricUnit="m" metricValue="5.32" pageId="40" pageNumber="41" unit="m" value="532.0">532m</quantity>
asl
</elevation>
),
<collectorName id="561899FAFFC0912B6E7AFA03AE79A4E5" box="[1082,1294,1495,1521]" pageId="40" pageNumber="41">I. Sanmartín-Villar</collectorName>
&amp;
<collectorName id="561899FAFFC0912B6F71FA03ABA9A701" pageId="40" pageNumber="41">A. Rivas-Torres</collectorName>
leg.,
<collectionCode id="9DFC64E9FFC0912B6B5DFA2FAAE5A701" box="[285,402,1531,1557]" pageId="40" pageNumber="41">ECOEVO</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="FB52FC2CFFC0912B6A87F9CBA9E7A72D" blockId="40.[151,1436,1422,1737]" box="[199,656,1566,1593]" pageId="40" pageNumber="41">
<emphasis id="C999203EFFC0912B6A87F9CBAACDA72C" bold="true" box="[199,442,1566,1593]" pageId="40" pageNumber="41">Known distribution.</emphasis>
<collectingCountry id="83FABCBCFFC0912B6B81F9CBA969A72D" box="[449,542,1567,1593]" name="Ecuador" pageId="40" pageNumber="41">Ecuador</collectingCountry>
and
<collectingCountry id="83FABCBCFFC0912B6815F9CBA9FBA72D" box="[597,652,1567,1593]" name="Peru" pageId="40" pageNumber="41">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="B3F7AFA7FFC0912B6A87F996AFE5A7DC" pageId="40" pageNumber="41" type="diagnosis">
<paragraph id="FB52FC2CFFC0912B6A87F996AFE5A7DC" blockId="40.[151,1436,1422,1737]" pageId="40" pageNumber="41">
<emphasis id="C999203EFFC0912B6A87F996AA92A749" bold="true" box="[199,485,1602,1629]" pageId="40" pageNumber="41">Diagnosis and remarks.</emphasis>
This species shares with
<taxonomicName id="3CED87AFFFC0912B694DF997A8E4A749" authorityName=": Sjostedt" authorityYear="1918" box="[781,915,1603,1629]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="silvarum">
<emphasis id="C999203EFFC0912B694DF997A8E4A749" box="[781,915,1603,1629]" italics="true" pageId="40" pageNumber="41">H. silvarum</emphasis>
</taxonomicName>
and
<taxonomicName id="3CED87AFFFC0912B698DF997AFE2A748" baseAuthorityName="Williamson" baseAuthorityYear="1919" box="[973,1173,1603,1628]" class="Insecta" family="Megapodagrionidae" genus="Heteragrion" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="cinnamomeum">
<emphasis id="C999203EFFC0912B698DF997AFE2A748" box="[973,1173,1603,1628]" italics="true" pageId="40" pageNumber="41">H. cinnamomeum</emphasis>
</taxonomicName>
the cercus with a long BP, ending in subequal MP and AP, with a developed ML. The following character combination distinguishes this species from the remainder of Group A species: BP the longest cercus portion, nearly straight (
<figureCitation id="63D6E0A9FFC0912B6EFBF95FAE33A7B0" box="[1211,1348,1674,1701]" captionStart="FIGURE 26" captionStartId="40.[151,250,1293,1317]" captionTargetBox="[246,1342,519,1257]" captionTargetId="figure-109@40.[233,1354,506,1270]" captionTargetPageId="40" captionText="FIGURE 26. Heteragrion cooki: (a) anterior view of head; (b) lateral view of pterothorax; (c–d) mediodorsal view of cercus. ML: medial lobe." figureDoi="http://doi.org/10.5281/zenodo.10008605" httpUri="https://zenodo.org/record/10008605/files/figure.png" pageId="40" pageNumber="41">Figs. 26c–d</figureCitation>
); on the MP, ML broad, bearing an elevated ridge (
<figureCitation id="63D6E0A9FFC0912B6832F97BA9A4A7DC" box="[626,723,1710,1737]" captionStart="FIGURE 26" captionStartId="40.[151,250,1293,1317]" captionTargetBox="[246,1342,519,1257]" captionTargetId="figure-109@40.[233,1354,506,1270]" captionTargetPageId="40" captionText="FIGURE 26. Heteragrion cooki: (a) anterior view of head; (b) lateral view of pterothorax; (c–d) mediodorsal view of cercus. ML: medial lobe." figureDoi="http://doi.org/10.5281/zenodo.10008605" httpUri="https://zenodo.org/record/10008605/files/figure.png" pageId="40" pageNumber="41">Fig. 26c</figureCitation>
); ML apex with a small cleft (
<figureCitation id="63D6E0A9FFC0912B6E63F97BAFF1A7DC" box="[1059,1158,1710,1737]" captionStart="FIGURE 26" captionStartId="40.[151,250,1293,1317]" captionTargetBox="[246,1342,519,1257]" captionTargetId="figure-109@40.[233,1354,506,1270]" captionTargetPageId="40" captionText="FIGURE 26. Heteragrion cooki: (a) anterior view of head; (b) lateral view of pterothorax; (c–d) mediodorsal view of cercus. ML: medial lobe." figureDoi="http://doi.org/10.5281/zenodo.10008605" httpUri="https://zenodo.org/record/10008605/files/figure.png" pageId="40" pageNumber="41">Fig. 26d</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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