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<mods:title id="8FBCE2F519E32A6302FF9A7D7C3BD38C">Annotated checklist of the snapping shrimps of the genus Alpheus Fabricius, 1798 (Crustacea: Decapoda: Alpheidae) from the Persian Gulf and Gulf of Oman, Iran</mods:title>
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<mods:namePart id="B6B68E38156CE2DBE4178CEEDFE55085">Sari, Alireza</mods:namePart>
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<treatment id="730587B4FF883C1EFF06FE07FEC8FA99" ID-DOI="http://doi.org/10.5281/zenodo.5255778" ID-GBIF-Taxon="154530494" ID-Zenodo-Dep="5255778" LSID="urn:lsid:plazi:treatment:730587B4FF883C1EFF06FE07FEC8FA99" httpUri="http://treatment.plazi.org/id/730587B4FF883C1EFF06FE07FEC8FA99" lastPageNumber="484" pageId="5" pageNumber="484">
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<paragraph id="FB1336A2FF883C1EFF06FE07FDA1FD71" blockId="5.[151,564,511,571]" box="[151,564,511,538]" pageId="5" pageNumber="484">
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<taxonomicName id="3CAC4D21FF883C1EFF06FE07FDA1FD71" ID-CoL="85LH9" authority="De Man, 1888" authorityName="De Man" authorityYear="1888" box="[151,564,511,538]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="484" phylum="Arthropoda" rank="species" species="edamensis">
<emphasis id="C9D8EAB0FF883C1EFF06FE07FDA1FD71" bold="true" box="[151,564,511,538]" pageId="5" pageNumber="484">
<emphasis id="C9D8EAB0FF883C1EFF06FE07FEEBFD72" bold="true" box="[151,382,511,537]" italics="true" pageId="5" pageNumber="484">Alpheus edamensis</emphasis>
De Man, 1888
</emphasis>
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<subSubSection id="B3B66529FF883C1EFF06FDDAFE8FFD50" box="[151,282,546,571]" pageId="5" pageNumber="484" type="description">
<paragraph id="FB1336A2FF883C1EFF06FDDAFE8FFD50" blockId="5.[151,564,511,571]" box="[151,282,546,571]" pageId="5" pageNumber="484">
(
<figureCitation id="63972A27FF883C1EFF0EFDDAFF74FD50" box="[159,225,546,571]" captionStart="FIGURE 2" captionStartId="14.[151,250,1922,1944]" captionTargetBox="[151,1436,210,1873]" captionTargetId="figure@14.[151,1436,210,1873]" captionTargetPageId="14" captionText="FIGURE 2. Alpheus deuteropus Hilgendorf, 1879, A. Male (CL. 8.6 mm); B. Female (CL. 8.3 mm), Abu-Musa Island. Alpheus djeddensis Coutière, 1897, C. Male (CL. 10 mm); D. Male pale morph (CL. 9.8 mm), Tis, Chabahar Bay. Alpheus edamensis De Man, 1888, E. Large male (CL. 20 mm); F. Medium male (CL. 8.9 mm), Tis, Chabahar Bay, Gulf of Oman." httpUri="https://zenodo.org/record/2618479/files/figure.png" pageId="5" pageNumber="484">Fig. 2</figureCitation>
E–F)
</paragraph>
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<treatmentCitationGroup id="DBBC118CFF883C1EFF06FD92FD58FDEB" box="[151,717,618,640]" pageId="5" pageNumber="484">
<taxonomicName id="3CAC4D21FF883C1EFF06FD92FD5CFDEB" ID-CoL="85LH9" authority="De Man, 1888 a: 518" authorityName="De Man" authorityPageNumber="518" authorityYear="1888" box="[151,713,618,640]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="484" phylum="Arthropoda" rank="variety" species="hippothoe" variety="edamensis">
<emphasis id="C9D8EAB0FF883C1EFF06FD92FEC1FDEB" box="[151,340,618,640]" italics="true" pageId="5" pageNumber="484">Alpheus hippothoë</emphasis>
var.
<emphasis id="C9D8EAB0FF883C1EFE17FD92FE65FDEB" box="[390,496,618,640]" italics="true" pageId="5" pageNumber="484">edamensis</emphasis>
<treatmentCitation id="7A0D10B3FF883C1EFE64FD93FD5CFDEB" author="De Man, J. G." box="[501,713,618,640]" page="518" pageId="5" pageNumber="484" year="1888">
<bibRefCitation id="9F3D4B53FF883C1EFE64FD93FD03FDEB" author="De Man, J. G." box="[501,662,618,640]" pageId="5" pageNumber="484" pagination="289 - 600" refId="ref14091" refString="De Man, J. G. (1888 a) Bericht uber die von Herrn Dr. J. Brock im indischen Archipel gesammelten Decapoden und Stomatopoden. Archiv fur Naturgeschichte, 53, 289 - 600. https: // doi. org / 10.5962 / bhl. part. 4747" type="journal article" year="1888">De Man, 1888a</bibRefCitation>
: 518
</treatmentCitation>
</taxonomicName>
.
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</paragraph>
<paragraph id="FB1336A2FF883C1EFF06FD71FDCDFDF4" blockId="5.[151,1436,618,732]" box="[151,600,649,671]" pageId="5" pageNumber="484">
<treatmentCitationGroup id="DBBC118CFF883C1EFF06FD71FDCDFDF4" box="[151,600,649,671]" pageId="5" pageNumber="484">
<taxonomicName id="3CAC4D21FF883C1EFF06FD71FDC1FDF4" ID-CoL="85LDB" authority="Ortmann, 1890: 474" authorityName="Ortmann" authorityPageNumber="474" authorityYear="1890" box="[151,596,649,671]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="484" phylum="Arthropoda" rank="species" species="acanthomerus">
<emphasis id="C9D8EAB0FF883C1EFF06FD71FEEBFDF4" box="[151,382,649,671]" italics="true" pageId="5" pageNumber="484">Alpheus acanthomerus</emphasis>
<treatmentCitation id="7A0D10B3FF883C1EFE15FD71FDC1FDF4" author="Ortmann, A." box="[388,596,649,671]" page="474" pageId="5" pageNumber="484" year="1890">
<bibRefCitation id="9F3D4B53FF883C1EFE15FD71FDB7FDF4" author="Ortmann, A." box="[388,546,649,671]" pageId="5" pageNumber="484" pagination="437 - 542" refId="ref15863" refString="Ortmann, A. (1890) Die Decapoden-Krebse des Strassburger Museums, mit besonderer Berucksichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. I. Theil. Die Unterordnung Natantia (Boas) (Abtheilungen: Penaeidae und Eucyphidea Caridae der Autoren). Zoologische Jahrbucher. Abtheilung fur Systematik, Geographie und Biologie der Thiere, 5, 437 - 542." type="journal article" year="1890">Ortmann, 1890</bibRefCitation>
: 474
</treatmentCitation>
</taxonomicName>
.
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</paragraph>
<paragraph id="FB1336A2FF883C1EFF06FD50FAD1FDB7" blockId="5.[151,1436,618,732]" pageId="5" pageNumber="484">
<treatmentCitationGroup id="DBBC118CFF883C1EFF06FD50FAD1FDB7" pageId="5" pageNumber="484">
<taxonomicName id="3CAC4D21FF883C1EFF06FD50FECFFDD5" ID-CoL="85LH9" authorityName="De Man" authorityYear="1888" box="[151,346,680,702]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="484" phylum="Arthropoda" rank="species" species="edamensis">
<emphasis id="C9D8EAB0FF883C1EFF06FD50FECFFDD5" box="[151,346,680,702]" italics="true" pageId="5" pageNumber="484">Alpheus edamensis</emphasis>
</taxonomicName>
—
<treatmentCitation id="7A0D10B3FF883C1EFEE5FD51FDD6FDD5" author="De Man, J. G." box="[372,579,680,702]" page="437" pageId="5" pageNumber="484" year="1911">
<bibRefCitation id="9F3D4B53FF883C1EFEE5FD51FD98FDD5" author="De Man, J. G." box="[372,525,680,702]" pageId="5" pageNumber="484" pagination="133 - 465" refId="ref14475" refString="De Man, J. G. (1911) The Decapoda of the Siboga Expedition. Part II. Family Alpheidae. Siboga Expeditie, 39 a (2), 133 - 465. https: // doi. org / 10.5962 / bhl. title. 10512" type="journal article" year="1911">De Man, 1911</bibRefCitation>
: 437
</treatmentCitation>
;
<treatmentCitation id="7A0D10B3FF883C1EFDDEFD51FC19FDD5" author="Banner, A. H. &amp; Banner, D. M." box="[591,908,680,702]" page="157" pageId="5" pageNumber="484" year="1966">
<bibRefCitation id="9F3D4B53FF883C1EFDDEFD51FCC2FDD5" author="Banner, A. H. &amp; Banner, D. M." box="[591,855,680,702]" pageId="5" pageNumber="484" pagination="1 - 168" refId="ref12330" refString="Banner, A. H. &amp; Banner, D. M. (1966 a) The alpheid shrimp of Thailand. The Siam Society Monograph Series, 3, 1 - 168." type="journal article" year="1966">Banner &amp; Banner, 1966a</bibRefCitation>
: 157
</treatmentCitation>
;
<treatmentCitation id="7A0D10B3FF883C1EFC08FD50FBCCFDD5" author="Johnson, D. S." box="[921,1113,680,702]" page="37" pageId="5" pageNumber="484" year="1979">
<bibRefCitation id="9F3D4B53FF883C1EFC08FD50FBA4FDD5" author="Johnson, D. S." box="[921,1073,680,702]" pageId="5" pageNumber="484" pagination="1 - 54" refId="ref15178" refString="Johnson, D. S. (1979) Prawns of the Malacca Straits and Singapore waters. Journal of the Marine Biological Association of India, 18, 1 - 54. [for 1976]" type="journal article" year="1979">Johnson, 1979</bibRefCitation>
: 37
</treatmentCitation>
;
<treatmentCitation id="7A0D10B3FF883C1EFBF6FD51FA0DFDD5" author="Banner, D. M. &amp; Banner, A. H." box="[1127,1432,680,702]" page="222" pageId="5" pageNumber="484" year="1978">
<bibRefCitation id="9F3D4B53FF883C1EFBF6FD51FAF7FDD5" author="Banner, D. M. &amp; Banner, A. H." box="[1127,1378,680,702]" pageId="5" pageNumber="484" pagination="215 - 257" refId="ref12817" refString="Banner, D. M. &amp; Banner, A. H. (1978) Annotated checklist of alpheid and ogyridid shrimp from the Philippine Archipelago and the South China Sea. Micronesica, 14 (2), 215 - 257." type="journal article" year="1978">Banner &amp; Banner, 1978</bibRefCitation>
: 222
</treatmentCitation>
;
<treatmentCitation id="7A0D10B3FF883C1EFF56FD3FFE7BFDB7" author="Banner, D. M. &amp; Banner, A. H." box="[199,494,710,732]" page="188" pageId="5" pageNumber="484" year="1982">
<bibRefCitation id="9F3D4B53FF883C1EFF56FD3FFE2EFDB7" author="Banner, D. M. &amp; Banner, A. H." box="[199,443,710,732]" pageId="5" pageNumber="484" pagination="1 - 357" refId="ref12861" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia. Part III: The remaining alpheids, principally the genus Alpheus and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner, 1982</bibRefCitation>
: 188
</treatmentCitation>
;
<treatmentCitation id="7A0D10B3FF883C1EFE69FD3FFCBAFDB7" author="Banner, D. M. &amp; Banner, A. H." box="[504,815,710,732]" page="1985" pageId="5" pageNumber="484" year="1985">
<bibRefCitation id="9F3D4B53FF883C1EFE69FD3FFD79FDB7" author="Banner, D. M. &amp; Banner, A. H." box="[504,748,710,732]" pageId="5" pageNumber="484" pagination="1 - 79" refId="ref12925" refString="Banner, D. M. &amp; Banner, A. H. (1985) The alpheid shrimp of Indonesia, based upon J. G. De Man's &quot; The Decapoda of the Siboga Expedition, Part II. Family Alpheidae. &quot; (1911). Marine Research in Indonesia, 25, 1 - 79." type="journal article" year="1985">Banner &amp; Banner, 1985</bibRefCitation>
:
<treatmentCitation id="7A0D10B3FF883C1EFD67FD3EFCBAFDB7" author="Banner, D. M. &amp; Banner, A. H." box="[758,815,710,732]" page="15" pageId="5" pageNumber="484" year="1985">
<bibRefCitation id="9F3D4B53FF883C1EFD67FD3EFCBAFDB7" author="Banner, D. M. &amp; Banner, A. H." box="[758,815,710,732]" pageId="5" pageNumber="484" pagination="1 - 79" refId="ref12925" refString="Banner, D. M. &amp; Banner, A. H. (1985) The alpheid shrimp of Indonesia, based upon J. G. De Man's &quot; The Decapoda of the Siboga Expedition, Part II. Family Alpheidae. &quot; (1911). Marine Research in Indonesia, 25, 1 - 79." type="journal article" year="1985">1985</bibRefCitation>
</treatmentCitation>
</treatmentCitation>
:15;
<treatmentCitation id="7A0D10B3FF883C1EFCC6FD3EFC6BFDB7" author="Chace, F. A. Jr." box="[855,1022,710,732]" page="24" pageId="5" pageNumber="484" year="1988">
<bibRefCitation id="9F3D4B53FF883C1EFCC6FD3EFC4CFDB7" author="Chace, F. A. Jr." box="[855,985,710,732]" pageId="5" pageNumber="484" pagination="1 - 99" refId="ref13206" refString="Chace, F. A. Jr. (1988) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 5: family Alpheidae. Smithsonian Contributions to Zoology, 466, 1 - 99." type="journal article" year="1988">Chace, 1988</bibRefCitation>
: 24
</treatmentCitation>
;
<treatmentCitation id="7A0D10B3FF883C1EFB98FD3EFAD5FDB7" author="Anker, A. &amp; De Grave, S." box="[1033,1344,710,732]" page="351" pageId="5" pageNumber="484" year="2016">
<bibRefCitation id="9F3D4B53FF883C1EFB98FD3EFA98FDB7" author="Anker, A. &amp; De Grave, S." box="[1033,1293,710,732]" pageId="5" pageNumber="484" pagination="343 - 454" refId="ref11825" refString="Anker, A. &amp; De Grave, S. (2016) An updated and annotated checklist of marine and brackish caridean shrimps of Singapore (Crustacea, Decapoda). Raffles Bulletin of Zoology, 34, 343 - 454." type="journal article" year="2016">Anker &amp; De Grave, 2016</bibRefCitation>
: 351
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="B3B66529FF883C1EFF06FCF1FDCEFB29" pageId="5" pageNumber="484" type="materials_examined">
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<emphasis id="C9D8EAB0FF883C1EFF06FCF1FE2CFC4A" bold="true" box="[151,441,777,802]" pageId="5" pageNumber="484">
Material examined.
<emphasis id="C9D8EAB0FF883C1EFE05FCF1FE2CFC4A" bold="true" box="[404,441,777,801]" italics="true" pageId="5" pageNumber="484">PG</emphasis>
</emphasis>
:
<materialsCitation id="4BC43CFFFF883C1EFE5EFCF2FE31FC2D" ID-GBIF-Occurrence="2147525587" collectingDate="2013-10-14" collectionCode="ZUTC" collectorName="Coll. R. Naderloo" location="Bandar-Dayyer" pageId="5" pageNumber="484" specimenCode="ZUTC 6714" specimenCount="5" specimenCount-female="3" specimenCount-male="2">
<specimenCount id="EDAAFD2BFF883C1EFE5EFCF2FDBBFC49" box="[463,558,777,802]" pageId="5" pageNumber="484" type="male">2 males</specimenCount>
,
<specimenCount id="EDAAFD2BFF883C1EFDD1FCF2FD26FC49" box="[576,691,777,802]" pageId="5" pageNumber="484" type="female">3 females</specimenCount>
(
<specimenCode id="AB0A9ED9FF883C1EFD59FCF2FCC9FC49" box="[712,860,777,802]" collectionCode="ZUTC" pageId="5" pageNumber="484">ZUTC 6714</specimenCode>
),
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, sandy/boulder shores,
<collectorName id="56595374FF883C1EFAAFFCF1FE97FC2D" pageId="5" pageNumber="484">Coll. R. Naderloo</collectorName>
,
<collectingDate id="9F56E98AFF883C1EFE9FFCD6FE31FC2D" box="[270,420,814,838]" pageId="5" pageNumber="484" value="2013-10-14">
<date id="8F121062FF883C1EFE9FFCD6FE31FC2D" box="[270,420,814,838]" pageId="5" pageNumber="484" value="2013-10-14">14 Oct. 2013</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="4BC43CFFFF883C1EFE21FCD6FEA5FC01" ID-GBIF-Occurrence="2147525826" collectingDate="2016-05-14" collectionCode="ZUTC" collectorName="Coll. A. Dehghani" country="Iran" location="Hormuz" pageId="5" pageNumber="484" specimenCode="ZUTC 6716" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="EDAAFD2BFF883C1EFE21FCD6FE6AFC2D" box="[432,511,813,838]" pageId="5" pageNumber="484" type="male">1 male</specimenCount>
,
<specimenCount id="EDAAFD2BFF883C1EFD9DFCD6FDF8FC2D" box="[524,621,813,838]" pageId="5" pageNumber="484" type="female">1 female</specimenCount>
(
<specimenCode id="AB0A9ED9FF883C1EFDECFCD6FC99FC2D" box="[637,780,813,838]" collectionCode="ZUTC" pageId="5" pageNumber="484">ZUTC 6716</specimenCode>
),
<collectingCountry id="83BB7632FF883C1EFCB1FCD6FCE9FC2E" box="[800,892,814,837]" name="Iran" pageId="5" pageNumber="484">Hormuz</collectingCountry>
I., in crevices of dead coral,
<collectorName id="56595374FF883C1EFB57FCD5FA0DFC2D" box="[1222,1432,813,838]" pageId="5" pageNumber="484">Coll. A. Dehghani</collectorName>
,
<collectingDate id="9F56E98AFF883C1EFF06FCAAFEA5FC01" box="[151,304,849,874]" pageId="5" pageNumber="484" value="2016-05-14">
<date id="8F121062FF883C1EFF06FCAAFEA5FC01" box="[151,304,849,874]" pageId="5" pageNumber="484" value="2016-05-14">14 May 2016</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="4BC43CFFFF883C1EFEAAFCAAFE51FCE5" ID-GBIF-Occurrence="2147525580" collectingDate="2012-11-28" collectionCode="ZUTC" collectorName="Coll. A. &amp; Sari" country="Iran" location="Bandar-Genaveh" pageId="5" pageNumber="484" specimenCode="ZUTC 6771" specimenCount="4" specimenCount-female="3" specimenCount-male="1">
<specimenCount id="EDAAFD2BFF883C1EFEAAFCAAFE1CFC01" box="[315,393,849,874]" pageId="5" pageNumber="484" type="male">1 male</specimenCount>
,
<specimenCount id="EDAAFD2BFF883C1EFE04FCAAFD94FC01" box="[405,513,849,874]" pageId="5" pageNumber="484" type="female">3 females</specimenCount>
(
<specimenCode id="AB0A9ED9FF883C1EFD81FCAAFD0BFC01" box="[528,670,849,874]" collectionCode="ZUTC" pageId="5" pageNumber="484">ZUTC 6771</specimenCode>
),
<location id="FE736079FF883C1EFD23FCA9FCE7FC01" LSID="urn:lsid:plazi:treatment:730587B4FF883C1EFF06FE07FEC8FA99:FE736079FF883C1EFD23FCA9FCE7FC01" box="[690,882,849,874]" country="Iran" name="Bandar-Genaveh" pageId="5" pageNumber="484">Bandar-Genaveh</location>
, under rocks and from burrows,
<collectorName id="56595374FF883C1EFB4EFCA9FAF9FC01" box="[1247,1388,849,874]" pageId="5" pageNumber="484">Coll. A. Sari</collectorName>
and H. Salehi,
<collectingDate id="9F56E98AFF883C1EFE8AFC8EFE51FCE5" box="[283,452,886,910]" pageId="5" pageNumber="484" value="2012-11-28">
<date id="8F121062FF883C1EFE8AFC8EFE51FCE5" box="[283,452,886,910]" pageId="5" pageNumber="484" value="2012-11-28">28 Nov. 2012</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="4BC43CFFFF883C1EFE46FC8EFE31FCD9" ID-GBIF-Occurrence="2147525919" collectingDate="2016-05-14" collectionCode="SMF" collectorName="Coll. A. Dehghani" country="Iran" location="Hormuz" pageId="5" pageNumber="484" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="EDAAFD2BFF883C1EFE46FC8EFDB9FCE5" box="[471,556,885,910]" pageId="5" pageNumber="484" type="male">1 male</specimenCount>
,
<specimenCount id="EDAAFD2BFF883C1EFDAFFC8EFD32FCE5" box="[574,679,885,910]" pageId="5" pageNumber="484" type="female">1 female</specimenCount>
(
<collectionCode id="9DBDAE67FF883C1EFD2CFC8EFD62FCE5" box="[701,759,886,910]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="484">SMF</collectionCode>
51378),
<collectingCountry id="83BB7632FF883C1EFCF8FC8EFC50FCE6" box="[873,965,886,909]" name="Iran" pageId="5" pageNumber="484">Hormuz</collectingCountry>
I., in crevices of dead coral,
<collectorName id="56595374FF883C1EFAAAFC8DFE9CFCD9" pageId="5" pageNumber="484">Coll. A. Dehghani</collectorName>
,
<collectingDate id="9F56E98AFF883C1EFE9CFC61FE31FCD9" box="[269,420,921,946]" pageId="5" pageNumber="484" value="2016-05-14">
<date id="8F121062FF883C1EFE9CFC61FE31FCD9" box="[269,420,921,946]" pageId="5" pageNumber="484" value="2016-05-14">14 May 2016</date>
</collectingDate>
</materialsCitation>
.
</paragraph>
<paragraph id="FB1336A2FF883C1EFF56FC45FDCEFB29" blockId="5.[151,1436,777,1522]" pageId="5" pageNumber="484">
<emphasis id="C9D8EAB0FF883C1EFF56FC45FF6DFCBE" box="[199,248,957,981]" italics="true" pageId="5" pageNumber="484">
<emphasis id="C9D8EAB0FF883C1EFF56FC45FF7AFCBE" bold="true" box="[199,239,957,981]" italics="true" pageId="5" pageNumber="484">GO</emphasis>
:
</emphasis>
<materialsCitation id="4BC43CFFFF883C1EFE94FC46FDDDFC91" ID-GBIF-Occurrence="2147525830" collectingDate="2016-10-02" collectionCode="ZUTC" collectorName="Coll. A. Dehghani &amp; H. Ashrafi" location="Chabahar Bay" pageId="5" pageNumber="484" specimenCode="ZUTC 6715" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<specimenCount id="EDAAFD2BFF883C1EFE94FC46FEF1FCBD" box="[261,356,957,982]" pageId="5" pageNumber="484" type="male">2 males</specimenCount>
,
<specimenCount id="EDAAFD2BFF883C1EFEE4FC46FE48FCBD" box="[373,477,957,982]" pageId="5" pageNumber="484" type="female">1 female</specimenCount>
(
<specimenCode id="AB0A9ED9FF883C1EFE60FC46FD10FCBD" box="[497,645,957,982]" collectionCode="ZUTC" pageId="5" pageNumber="484">ZUTC 6715</specimenCode>
),
<location id="FE736079FF883C1EFD0FFC45FCD1FCBD" LSID="urn:lsid:plazi:treatment:730587B4FF883C1EFF06FE07FEC8FA99:FE736079FF883C1EFD0FFC45FCD1FCBD" box="[670,836,957,982]" name="Chabahar Bay" pageId="5" pageNumber="484">Chabahar Bay</location>
, Tis, under large rocks and coral rubble,
<collectorName id="56595374FF883C1EFAACFC45FE91FC91" pageId="5" pageNumber="484">Coll. A. Dehghani</collectorName>
and
<collectorName id="56595374FF883C1EFEAAFC1AFE20FC91" box="[315,437,993,1018]" pageId="5" pageNumber="484">H. Ashrafi</collectorName>
,
<collectingDate id="9F56E98AFF883C1EFE51FC1AFDDDFC91" box="[448,584,993,1018]" pageId="5" pageNumber="484" value="2016-10-02">
<date id="8F121062FF883C1EFE51FC1AFDDDFC91" box="[448,584,993,1018]" pageId="5" pageNumber="484" value="2016-10-02">2 Oct. 2016</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="4BC43CFFFF883C1EFDC2FC1AFDC0FB75" ID-GBIF-Occurrence="2147525913" collectingDate="2016-05-02" collectionCode="ZUTC" collectorName="Coll. A. Dehghani &amp; H. Ashrafi" location="Abd Village" pageId="5" pageNumber="484" specimenCode="ZUTC 6772" specimenCount="1" specimenCount-male="1">
<specimenCount id="EDAAFD2BFF883C1EFDC2FC1AFD37FC91" box="[595,674,993,1018]" pageId="5" pageNumber="484" type="male">1 male</specimenCount>
, 2 ovig. females (
<specimenCode id="AB0A9ED9FF883C1EFCF8FC1AFC62FC91" box="[873,1015,993,1018]" collectionCode="ZUTC" pageId="5" pageNumber="484">ZUTC 6772</specimenCode>
),
<location id="FE736079FF883C1EFB9AFC19FB02FC91" LSID="urn:lsid:plazi:treatment:730587B4FF883C1EFF06FE07FEC8FA99:FE736079FF883C1EFB9AFC19FB02FC91" box="[1035,1175,993,1018]" name="Abd Village" pageId="5" pageNumber="484">Abd Village</location>
, rocky/cobble,
<collectorName id="56595374FF883C1EFAD3FC19FE91FB75" pageId="5" pageNumber="484">Coll. A. Dehghani</collectorName>
&amp;
<collectorName id="56595374FF883C1EFEA3FBFEFE25FB75" box="[306,432,1029,1054]" pageId="5" pageNumber="484">H. Ashrafi</collectorName>
,
<collectingDate id="9F56E98AFF883C1EFE51FBFEFDC0FB75" box="[448,597,1029,1054]" pageId="5" pageNumber="484" value="2016-05-02">
<date id="8F121062FF883C1EFE51FBFEFDC0FB75" box="[448,597,1029,1054]" pageId="5" pageNumber="484" value="2016-05-02">2 May 2016</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="4BC43CFFFF883C1EFDF7FBFEFDCDFB29" ID-GBIF-Occurrence="2147525910" collectingDate="2017-05-25" collectionCode="ZUTC" collectorName="Coll. A. Dehghani &amp; H. Ashrafi" location="Djod Village" pageId="5" pageNumber="484" specimenCode="ZUTC 6773" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="EDAAFD2BFF883C1EFDF7FBFEFD2CFB75" box="[614,697,1029,1054]" pageId="5" pageNumber="484" type="male">1 male</specimenCount>
,
<specimenCount id="EDAAFD2BFF883C1EFD5BFBFEFCA5FB75" box="[714,816,1029,1054]" pageId="5" pageNumber="484" type="female">1 female</specimenCount>
(
<specimenCode id="AB0A9ED9FF883C1EFCD5FBFEFC42FB75" box="[836,983,1029,1054]" collectionCode="ZUTC" pageId="5" pageNumber="484">ZUTC 6773</specimenCode>
),
<location id="FE736079FF883C1EFC61FBFDFB1DFB75" LSID="urn:lsid:plazi:treatment:730587B4FF883C1EFF06FE07FEC8FA99:FE736079FF883C1EFC61FBFDFB1DFB75" box="[1008,1160,1029,1054]" name="Djod Village" pageId="5" pageNumber="484">Djod Village</location>
, rocky/cobble,
<collectorName id="56595374FF883C1EFAACFBFDFE91FB29" pageId="5" pageNumber="484">Coll. A. Dehghani</collectorName>
and
<collectorName id="56595374FF883C1EFEABFBD2FE26FB29" box="[314,435,1065,1090]" pageId="5" pageNumber="484">H. Ashrafi</collectorName>
,
<collectingDate id="9F56E98AFF883C1EFE2FFBD1FDCDFB29" box="[446,600,1065,1090]" pageId="5" pageNumber="484" value="2017-05-25">
<date id="8F121062FF883C1EFE2FFBD1FDCDFB29" box="[446,600,1065,1090]" pageId="5" pageNumber="484" value="2017-05-25">25 May 2017</date>
</collectingDate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B3B66529FF883C1EFF56FBB5FA92FBC5" pageId="5" pageNumber="484" type="distribution">
<paragraph id="FB1336A2FF883C1EFF56FBB5FCE1FBE1" blockId="5.[151,1436,777,1522]" pageId="5" pageNumber="484">
<emphasis id="C9D8EAB0FF883C1EFF56FBB5FEF5FB0D" bold="true" box="[199,352,1101,1126]" pageId="5" pageNumber="484">Distribution.</emphasis>
Indo-West Pacific:
<collectingCountry id="83BB7632FF883C1EFDD1FBB5FD5CFB0D" box="[576,713,1101,1126]" name="Madagascar" pageId="5" pageNumber="484">Madagascar</collectingCountry>
,
<collectingCountry id="83BB7632FF883C1EFD45FBB5FCD1FB0D" box="[724,836,1101,1126]" name="Mauritius" pageId="5" pageNumber="484">Mauritius</collectingCountry>
,
<collectingCountry id="83BB7632FF883C1EFCDEFBB5FC24FB0D" box="[847,945,1101,1126]" name="Réunion" pageId="5" pageNumber="484">Réunion</collectingCountry>
,
<collectingCountry id="83BB7632FF883C1EFC2CFBB5FBADFB0D" box="[957,1080,1101,1126]" name="Seychelles" pageId="5" pageNumber="484">Seychelles</collectingCountry>
, Red Sea, Persian Gulf, Gulf of
<collectingCountry id="83BB7632FF883C1EFF06FB8AFF4BFBE1" box="[151,222,1138,1162]" name="Oman" pageId="5" pageNumber="484">Oman</collectingCountry>
,
<collectingCountry id="83BB7632FF883C1EFF78FB89FEC7FBE1" box="[233,338,1137,1162]" name="Malaysia" pageId="5" pageNumber="484">Malaysia</collectingCountry>
,
<collectingCountry id="83BB7632FF883C1EFECFFB89FE51FBE1" box="[350,452,1137,1162]" name="Thailand" pageId="5" pageNumber="484">Thailand</collectingCountry>
,
<collectingCountry id="83BB7632FF883C1EFE5EFB89FDAAFBE1" box="[463,575,1137,1162]" name="Indonesia" pageId="5" pageNumber="484">Indonesia</collectingCountry>
,
<collectingCountry id="83BB7632FF883C1EFDDBFB89FD5EFBE1" box="[586,715,1137,1162]" name="Philippines" pageId="5" pageNumber="484">Philippines</collectingCountry>
, and
<collectingCountry id="83BB7632FF883C1EFC97FB89FCE5FBE1" box="[774,880,1137,1162]" name="Australia" pageId="5" pageNumber="484">Australia</collectingCountry>
.
</paragraph>
<paragraph id="FB1336A2FF883C1EFF56FB6DFA92FBC5" blockId="5.[151,1436,777,1522]" box="[199,1287,1173,1198]" pageId="5" pageNumber="484">
<emphasis id="C9D8EAB0FF883C1EFF56FB6DFE53FBC5" bold="true" box="[199,454,1173,1198]" pageId="5" pageNumber="484">Regional records. PG</emphasis>
:
<collectingCountry id="83BB7632FF883C1EFE44FB6EFD97FBC6" box="[469,514,1174,1197]" name="Iran" pageId="5" pageNumber="484">Iran</collectingCountry>
(
<bibRefCitation id="9F3D4B53FF883C1EFD81FB6DFC43FBC5" author="Naderloo, R. &amp; Turkay, M." box="[528,982,1173,1198]" pageId="5" pageNumber="484" pagination="1 - 67" refId="ref15603" refString="Naderloo, R. &amp; Turkay, M. (2012) Decapod crustaceans of the littoral and shallow sublittoral Iranian coast of the Persian Gulf: faunistics, biodiversity and zoogeography. Zootaxa, 3374 (1), 1 - 67." type="journal article" year="2012">Naderloo &amp; Türkay 2012; present study</bibRefCitation>
);
<emphasis id="C9D8EAB0FF883C1EFC78FB6DFB80FBC5" bold="true" box="[1001,1045,1173,1198]" pageId="5" pageNumber="484">GO</emphasis>
:
<collectingCountry id="83BB7632FF883C1EFBB5FB6EFBC4FBC6" box="[1060,1105,1174,1197]" name="Iran" pageId="5" pageNumber="484">Iran</collectingCountry>
(present study).
</paragraph>
</subSubSection>
<subSubSection id="B3B66529FF883C1EFF56FB41FF46FB9D" pageId="5" pageNumber="484" type="description">
<paragraph id="FB1336A2FF883C1EFF56FB41FF46FB9D" blockId="5.[151,1436,777,1522]" pageId="5" pageNumber="484">
<emphasis id="C9D8EAB0FF883C1EFF56FB41FEBFFBB9" bold="true" box="[199,298,1209,1234]" pageId="5" pageNumber="484">Habitat.</emphasis>
Sandy with rubble and rocky-sand flats, under rocks and large pieces of coral rubble; intertidal to
<quantity id="3C549B47FF883C1EFAE3FB41FA09FBBA" box="[1394,1436,1209,1234]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="484" unit="m" value="5.0">5 m</quantity>
deep.
</paragraph>
</subSubSection>
<subSubSection id="B3B66529FF883C1EFF56FAF9FEC8FA99" pageId="5" pageNumber="484" type="discussion">
<paragraph id="FB1336A2FF883C1EFF56FAF9FEC8FA99" blockId="5.[151,1436,777,1522]" pageId="5" pageNumber="484">
<emphasis id="C9D8EAB0FF883C1EFF56FAF9FEAEFA71" bold="true" box="[199,315,1281,1306]" pageId="5" pageNumber="484">Remarks.</emphasis>
This species is one of largest
<taxonomicName id="3CAC4D21FF883C1EFD0FFAF9FD6DFA71" authorityName="Fabricius" authorityYear="1798" box="[670,760,1281,1306]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="C9D8EAB0FF883C1EFD0FFAF9FD6DFA71" box="[670,760,1281,1306]" italics="true" pageId="5" pageNumber="484">Alpheus</emphasis>
</taxonomicName>
in
<emphasis id="C9D8EAB0FF883C1EFCB3FAF9FCDCFA71" bold="true" box="[802,841,1281,1306]" pageId="5" pageNumber="484">PG</emphasis>
and
<emphasis id="C9D8EAB0FF883C1EFC15FAF9FC25FA71" bold="true" box="[900,944,1281,1306]" pageId="5" pageNumber="484">GO</emphasis>
(TL:
<quantity id="3C549B47FF883C1EFC66FAFAFBD6FA71" box="[1015,1091,1282,1306]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="484" unit="cm" value="10.0">10 cm</quantity>
). Small-sized specimens were free-living (
<figureCitation id="63972A27FF883C1EFEB1FADDFEECFA55" box="[288,377,1317,1342]" captionStart="FIGURE 2" captionStartId="14.[151,250,1922,1944]" captionTargetBox="[151,1436,210,1873]" captionTargetId="figure@14.[151,1436,210,1873]" captionTargetPageId="14" captionText="FIGURE 2. Alpheus deuteropus Hilgendorf, 1879, A. Male (CL. 8.6 mm); B. Female (CL. 8.3 mm), Abu-Musa Island. Alpheus djeddensis Coutière, 1897, C. Male (CL. 10 mm); D. Male pale morph (CL. 9.8 mm), Tis, Chabahar Bay. Alpheus edamensis De Man, 1888, E. Large male (CL. 20 mm); F. Medium male (CL. 8.9 mm), Tis, Chabahar Bay, Gulf of Oman." httpUri="https://zenodo.org/record/2618479/files/figure.png" pageId="5" pageNumber="484">Fig. 2F</figureCitation>
), but larger specimens (
<figureCitation id="63972A27FF883C1EFD02FADDFD79FA55" box="[659,748,1317,1342]" captionStart="FIGURE 2" captionStartId="14.[151,250,1922,1944]" captionTargetBox="[151,1436,210,1873]" captionTargetId="figure@14.[151,1436,210,1873]" captionTargetPageId="14" captionText="FIGURE 2. Alpheus deuteropus Hilgendorf, 1879, A. Male (CL. 8.6 mm); B. Female (CL. 8.3 mm), Abu-Musa Island. Alpheus djeddensis Coutière, 1897, C. Male (CL. 10 mm); D. Male pale morph (CL. 9.8 mm), Tis, Chabahar Bay. Alpheus edamensis De Man, 1888, E. Large male (CL. 20 mm); F. Medium male (CL. 8.9 mm), Tis, Chabahar Bay, Gulf of Oman." httpUri="https://zenodo.org/record/2618479/files/figure.png" pageId="5" pageNumber="484">Fig. 2E</figureCitation>
) were found associated with the ophiuroid
<taxonomicName id="3CAC4D21FF883C1EFB7DFADDFE1DFA09" authority="Clark, 1938" authorityName="Clark" authorityYear="1938" class="Ophiuroidea" family="Ophiotrichidae" genus="Macrophiothrix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="5" pageNumber="484" phylum="Echinodermata" rank="species" species="elongata">
<emphasis id="C9D8EAB0FF883C1EFB7DFADDFF6CFA09" italics="true" pageId="5" pageNumber="484">Macrophiothrix elongata</emphasis>
<bibRefCitation id="9F3D4B53FF883C1EFE91FAB1FE1DFA09" author="Clark, H. L." box="[256,392,1353,1378]" pageId="5" pageNumber="484" pagination="1 - 597" refId="ref13152" refString="Clark, H. L. (1938) Echinoderms from Australia, an account of collections made in 1929 and 1932. Memoirs of the Museum of Comparative Zoology at Harvard College. 55, 1 - 597. https: // doi. org / 10.5962 / bhl. title. 49349" type="journal article" year="1938">Clark, 1938</bibRefCitation>
</taxonomicName>
. Furthermore,
<emphasis id="C9D8EAB0FF883C1EFDBEFAB1FDC3FA09" bold="true" box="[559,598,1353,1378]" pageId="5" pageNumber="484">PG</emphasis>
specimens differ from Singaporean specimens in color pattern (see
<bibRefCitation id="9F3D4B53FF883C1EFAC7FAB1FEFBFAED" author="Anker, A. &amp; De Grave, S." pageId="5" pageNumber="484" pagination="343 - 454" refId="ref11825" refString="Anker, A. &amp; De Grave, S. (2016) An updated and annotated checklist of marine and brackish caridean shrimps of Singapore (Crustacea, Decapoda). Raffles Bulletin of Zoology, 34, 343 - 454." type="journal article" year="2016">Anker &amp; De Grave, 2016</bibRefCitation>
). To clarify the species identity, a molecular analysis and thorough comparison of morphology and color patterns were applied on the present study material. Accordingly, these specimens were precisely categorized into “symbiont
<emphasis id="C9D8EAB0FF883C1EFE59FA4EFE42FAA6" box="[456,471,1462,1485]" italics="true" pageId="5" pageNumber="484">”</emphasis>
and
<emphasis id="C9D8EAB0FF883C1EFD80FA4EFDB5FAA6" box="[529,544,1462,1485]" italics="true" pageId="5" pageNumber="484">“</emphasis>
non-symbiont
<emphasis id="C9D8EAB0FF883C1EFD2DFA4EFCFDFAA5" box="[700,872,1461,1486]" italics="true" pageId="5" pageNumber="484">
”
<taxonomicName id="3CAC4D21FF883C1EFD42FA4EFCFDFAA5" authorityName="De Man" authorityYear="1888" box="[723,872,1461,1486]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="484" phylum="Arthropoda" rank="species" species="edamensis">A. edamensis</taxonomicName>
</emphasis>
(unpublished molecular data by authors based on COI sequencing).
</paragraph>
</subSubSection>
</treatment>
</document>
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