304 lines
35 KiB
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304 lines
35 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.907.36563" ID-GBIF-Dataset="69422977-aa4d-4fe3-a450-8ef54d6fa905" ID-PMC="PMC7002455" ID-Pensoft-Pub="1313-2970-907-1" ID-Pensoft-UUID="9920267E73CF5E00B644DED1F101D965" ID-PubMed="32063727" ID-ZooBank="DBF339E5EBBC461994388359C769473F" ModsDocID="1313-2970-907-1" checkinTime="1580398733216" checkinUser="pensoft" docAuthor="Leger, Theo, Kehlmaier, Christian, Vairappan, Charles S. & Nuss, Matthias" docDate="2020" docId="6CD65D63F8AC5F2FBBC0A38A9DFD2A40" docLanguage="en" docName="ZooKeys 907: 1-99" docOrigin="ZooKeys 907" docSource="http://dx.doi.org/10.3897/zookeys.907.36563" docTitle="Hoploscopa albipuncta Leger & Nuss 2020, sp. nov." docType="treatment" docUuid="26FE78DF-5D7E-4034-A730-FA5C7177466E" docUuidSource="ZooBank" docVersion="4" id="9920267E73CF5E00B644DED1F101D965" lastPageNumber="1" masterDocId="9920267E73CF5E00B644DED1F101D965" masterDocTitle="Twenty-six new species of Hoploscopa (Lepidoptera, Crambidae) from South-East Asia revealed by morphology and DNA barcoding" masterLastPageNumber="99" masterPageNumber="1" pageNumber="1" updateTime="1668168367651" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Twenty-six new species of Hoploscopa (Lepidoptera, Crambidae) from South-East Asia revealed by morphology and DNA barcoding</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Leger, Theo</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Kehlmaier, Christian</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Vairappan, Charles S.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Nuss, Matthias</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>907</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>99</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.907.36563</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.907.36563</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-907-1</mods:identifier>
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<mods:identifier type="ZooBank">DBF339E5EBBC461994388359C769473F</mods:identifier>
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<treatment ID-GBIF-Taxon="161926055" LSID="urn:lsid:zoobank.org:act:26FE78DF-5D7E-4034-A730-FA5C7177466E" httpUri="http://treatment.plazi.org/id/6CD65D63F8AC5F2FBBC0A38A9DFD2A40" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<taxonomicName LSID="6CD65D63-F8AC-5F2F-BBC0-A38A9DFD2A40" authority="Léger & Nuss" authorityName="Leger & Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa albipuncta" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="albipuncta" status="sp. nov.">
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Hoploscopa albipuncta
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<normalizedToken originalValue="Léger">Leger</normalizedToken>
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& Nuss
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</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figures 4–18" captionStartId="F2" captionText="Figures 4 - 18. Forewing of Hoploscopa species. 4 Hoploscopa albipuncta sp. nov., holotype, ♂, Malaysia, Sabah, Kinabalu Park HQ, Timpohon Gate, 700 m from Liwagu Trail starting point, near Liwagu River, 6 ° 1 ' 40 " N, 116 ° 32 ' 59 " E, 1760 m, 18. vi. 2015 (T. Leger & R. Mally) (genitalia on slide TL 335 ♂) 5 Hoploscopa sepanggi sp. nov., holotype, ♂, Malaysia, Sabah, Mesilau Nature Resort, 6 ° 2 ' 43.71 " N, 116 ° 35 ' 48.03 ' E, 1925 m, at lamps, 30. v. 2015 (T. Leger & R. Mally) 6 Hoploscopa cynodonta sp. nov., holotype, ♂, Malaysia, Sabah, Kinabalu Park HQ, junction Kiau View- and Pandanus Trail, 6 ° 0 ' 32.84 " N, 116 ° 32 ' 14.94 " E, 1690 m, UV light, 07. vi. 2015 (T. Leger & R. Mally) (genitalia on slide TL 327 ♂) 7 Hoploscopa parvimacula sp. nov., paratype, ♀, Sabah, Mesilau Nature Resort, 6 ° 2 ' 43.71 " N, 116 ° 35 ' 48.03 " E, 1925 m, at lamps, 30. v. 2015 (T. Leger & R. Mally) (genitalia on slide TL 312 ♀. 8 Hoploscopa kinabaluensis sp. nov., paratype, ♀, Sabah, Kinabalu Park HQ, Timpohon Gate, 700 m from Liwagu trail starting point, near Liwagu River, 6 ° 1 ' 40 " N, 116 ° 32 ' 59 " E, 1760 m, UV light, 18. vi. 2015 (T. Leger & R. Mally) (genitalia on slide TL 326 ♀) 9 Hoploscopa luteomacula Nuss, paratype, ♀, Indonesia, Sumatra, Barat, N-Padangpanjang, Mt Singgalang, 2100 m, 10 - 11. ii. 1996 (A. Kallies) (genitalia on slide GU prep. Nuss 743) 10 Hoploscopa obliqua Rotschild, abdomen missing, Papua New Guinea, Madang Province, Wanang village, 05 ° 15 ' S, 145 ° 17 ' E, reared from Diplazium esculentum, 12. vi. 2007 11 Hoploscopa niveofascia sp. nov., holotype, ♂, Papua New Guinea, Morobe Prov., nr. Bulolo, Mt. Susu Nat. Res., 975 m, 27 - 28. viii. 1983, Araucaria For. (S. Miller) (genitalia on slide TL 442 ♂) 12 Hoploscopa gombongi sp. nov., paratype, ♂, Papua New Guinea, Yawan village, 06 ° 10 ' S, 146 ° 5 ' E, NG Binatang Res. Ctr. (B Gewa, J. Kua, S Sau, A. Kinibel) 13 Hoploscopa tonsepi sp. nov., holotype, ♂, Papua New Guinea, Yawan village, 06 ° 10 ' S, 146 ° 5 ' E, NG Binatang Res. Ctr. (J. Valeba, J. Auga, M. Dilu, F. Philip, R. Lilip) (genitalia on slide TL 655 ♂) 14 Hoploscopa marijoweissae sp. nov., ♂, Papua New Guinea: Morobe Province, Mount Kaindi, 2350 m, 11. xii. 1976 (G. F. Hevel & R. E. Dietz) (genitalia on slide TL 437 ♂) 15 Hoploscopa titika sp. nov., holotype, ♂, Sumatra-Holzweg, 25 km SSW-Pematangsiantar, Strasse nach Prapat, 13. ii. 1996 (A. Kallies) (genitalia on slide TL 505 ♂) 16 Hoploscopa pangrangoensis sp. nov., paratype, ♀, Indonesia, Java, Mt. Pangrange, SE Bogor, 6.30 S 107.10 E, 1625 m, primary forest, 16 - 20. ii. 1996 (Siniaev & Afonin) (genitalia on slide TL 627 ♀) 17 Hoploscopa isarogensis sp. nov., paratype, ♀, Philippines, South Luzon, Mt Isarog, 13 ° 40 ' N, 123 ° 20 ' E, 530 m, submontane forest, at light, 22. iii. 2000 (M. Nuss) (genitalia on slide TL 523 ♀) 18 Hoploscopa ypsilon sp. nov., paratype, ♀, Philippines, Luzon, Mountain Province, Chatol, 2100 m, 16 - 18. xi. 1997 (Mey, Ebert & Nuss)." figureDoi="10.3897/zookeys.907.36563.figures4-18" httpUri="https://binary.pensoft.net/fig/375757" pageId="0" pageNumber="1">Figs 4</figureCitation>
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<figureCitation captionStart="Figures 46–51" captionStartId="F5" captionText="Figures 46 - 51. Male genitalia features of Hoploscopa. For each figure left above: genitalia without phallus; left below: phallus; right above: gnathos drawing; right below: juxta drawing. 46 Hoploscopa albipuncta sp. nov., paratype, TL 336 ♂ (genitalia without phallus), TL 635 ♂ (phallus) 47 Hoploscopa matheae sp. nov., paratype, TL 730 ♂ 48 Hoploscopa sepanggi sp. nov., paratype, TL 641 ♂ 49 Hoploscopa cynodonta sp. nov., paratype, TL 531 ♂ 50 Hoploscopa parvimacula sp. nov., holotype, TL 510 ♂ 51 Hoploscopa kinabaluensis sp. nov., paratype, TL 342 ♂. Abbreviations: c (cornutus), de (ductus ejaculatorius), g (gnathos), jx (juxta), s (saccus), ta (tegumen arm), v (valva), vc (vinculum), uc (uncus)." figureDoi="10.3897/zookeys.907.36563.figures46-51" httpUri="https://binary.pensoft.net/fig/375760" pageId="0" pageNumber="1">, 46</figureCitation>
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<figureCitation captionStart="Figures 87–90" captionStartId="F12" captionText="Figures 87 - 90. Female genitalia features of Hoploscopa. 87 Hoploscopa albipuncta sp. nov., paratype, TL 609 ♀ 88 Hoploscopa matheae sp. nov., paratype, TL 599 ♀ 89 Hoploscopa sepanggi sp. nov., paratype, TL 547 ♀ 90 Hoploscopa cynodonta sp. nov., paratype, TL 625 ♀. Abbreviations: aa (anterior apophyses), at (antrum), cb (corpus bursae), cs (corpus sclerotization), db (ductus bursae), ds (ductus seminalis), pa (posterior apophyses), pp (papillae anales), segm. VIII (segment VIII), t (thorn)." figureDoi="10.3897/zookeys.907.36563.figures87-90" httpUri="https://binary.pensoft.net/fig/375767" pageId="0" pageNumber="1">, 87</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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: ♂, with labels: "Malaysia: Sabah, Kinabalu Park H[ead]Q[uarter], | Timpohon Gate, 700m from Liwagu | Trail starting point, near Liwagu River, | 6°1'40"N, 116°32'59"E, 1760m, UV light, | 18.vi.2015, leg. T.
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<normalizedToken originalValue="Léger">Leger</normalizedToken>
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& R. Mally"; "TL335 | ♂". Deposited in BORN.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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: 13 ♂, 1 ♀. Malaysia: 8 ♂ (1 with genitalia on slide TL336 ♂), same data as holotype; 3 ♂ (1 with genitalia on slide TL536 ♂), Sabah, Kinabalu National Park, Timpohon Gate, 300 m from Ligawu trail starting point,
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<geoCoordinate degrees="6" direction="north" minutes="1" orientation="latitude" precision="15" seconds="41" value="6.0280557">6°1'41"N</geoCoordinate>
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,
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<geoCoordinate degrees="116" direction="east" minutes="32" orientation="longitude" precision="15" seconds="54" value="116.54833">116°32'54"E</geoCoordinate>
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, 1820 m, UV light, 18.vi.2015, leg. T.
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<normalizedToken originalValue="Léger">Leger</normalizedToken>
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& R. Mally (MTD); 1 ♂, Sabah, Mesilau, Kopogon, 24.ii.2006, light trap, leg. W. & M. Mey; 1 ♂ (genitalia on slide TL635 ♂), Sabah, Mesilau River, 24-25.ii.2006, light trap, leg. W. & M. Mey; 1 ♀ (DNA voucher MTD7434, genitalia on slide TL609 ♀), Sabah, Mesilau, 2000 m, 14-17.xi.2006, light trap, leg. W. Mey & K. Ebert (MFNB).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Other specimens examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">1 ♂. Malaysia: 1 ♂ (DNA voucher ITBC09, genitalia on slide TL309 ♂), same data as holotype.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Leger & Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa albipuncta" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="albipuncta">
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<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa albipuncta</emphasis>
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</taxonomicName>
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sp. nov. displays a basal white well-rounded spot on the forewing. In male genitalia, the dorsal margin of the valva is protruded, the juxta displays two conspicuous tips and the phallus bears an anvil-shaped cornutus. In female genitalia, the antrum is membranous and forms a rounded pouch.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="similar species">
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<paragraph pageId="0" pageNumber="1">Similar species.</paragraph>
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<paragraph pageId="0" pageNumber="1">No similar species known.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
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Antennae dorsally with brown scales. Proboscis pale yellow. Maxillary palpi brown, basally light brown. Labial palpi brown, ventrally pale yellow.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Thorax</emphasis>
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(Fig.
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<figureCitation captionStart="Figures 4–18" captionStartId="F2" captionText="Figures 4 - 18. Forewing of Hoploscopa species. 4 Hoploscopa albipuncta sp. nov., holotype, ♂, Malaysia, Sabah, Kinabalu Park HQ, Timpohon Gate, 700 m from Liwagu Trail starting point, near Liwagu River, 6 ° 1 ' 40 " N, 116 ° 32 ' 59 " E, 1760 m, 18. vi. 2015 (T. Leger & R. Mally) (genitalia on slide TL 335 ♂) 5 Hoploscopa sepanggi sp. nov., holotype, ♂, Malaysia, Sabah, Mesilau Nature Resort, 6 ° 2 ' 43.71 " N, 116 ° 35 ' 48.03 ' E, 1925 m, at lamps, 30. v. 2015 (T. Leger & R. Mally) 6 Hoploscopa cynodonta sp. nov., holotype, ♂, Malaysia, Sabah, Kinabalu Park HQ, junction Kiau View- and Pandanus Trail, 6 ° 0 ' 32.84 " N, 116 ° 32 ' 14.94 " E, 1690 m, UV light, 07. vi. 2015 (T. Leger & R. Mally) (genitalia on slide TL 327 ♂) 7 Hoploscopa parvimacula sp. nov., paratype, ♀, Sabah, Mesilau Nature Resort, 6 ° 2 ' 43.71 " N, 116 ° 35 ' 48.03 " E, 1925 m, at lamps, 30. v. 2015 (T. Leger & R. Mally) (genitalia on slide TL 312 ♀. 8 Hoploscopa kinabaluensis sp. nov., paratype, ♀, Sabah, Kinabalu Park HQ, Timpohon Gate, 700 m from Liwagu trail starting point, near Liwagu River, 6 ° 1 ' 40 " N, 116 ° 32 ' 59 " E, 1760 m, UV light, 18. vi. 2015 (T. Leger & R. Mally) (genitalia on slide TL 326 ♀) 9 Hoploscopa luteomacula Nuss, paratype, ♀, Indonesia, Sumatra, Barat, N-Padangpanjang, Mt Singgalang, 2100 m, 10 - 11. ii. 1996 (A. Kallies) (genitalia on slide GU prep. Nuss 743) 10 Hoploscopa obliqua Rotschild, abdomen missing, Papua New Guinea, Madang Province, Wanang village, 05 ° 15 ' S, 145 ° 17 ' E, reared from Diplazium esculentum, 12. vi. 2007 11 Hoploscopa niveofascia sp. nov., holotype, ♂, Papua New Guinea, Morobe Prov., nr. Bulolo, Mt. Susu Nat. Res., 975 m, 27 - 28. viii. 1983, Araucaria For. (S. Miller) (genitalia on slide TL 442 ♂) 12 Hoploscopa gombongi sp. nov., paratype, ♂, Papua New Guinea, Yawan village, 06 ° 10 ' S, 146 ° 5 ' E, NG Binatang Res. Ctr. (B Gewa, J. Kua, S Sau, A. Kinibel) 13 Hoploscopa tonsepi sp. nov., holotype, ♂, Papua New Guinea, Yawan village, 06 ° 10 ' S, 146 ° 5 ' E, NG Binatang Res. Ctr. (J. Valeba, J. Auga, M. Dilu, F. Philip, R. Lilip) (genitalia on slide TL 655 ♂) 14 Hoploscopa marijoweissae sp. nov., ♂, Papua New Guinea: Morobe Province, Mount Kaindi, 2350 m, 11. xii. 1976 (G. F. Hevel & R. E. Dietz) (genitalia on slide TL 437 ♂) 15 Hoploscopa titika sp. nov., holotype, ♂, Sumatra-Holzweg, 25 km SSW-Pematangsiantar, Strasse nach Prapat, 13. ii. 1996 (A. Kallies) (genitalia on slide TL 505 ♂) 16 Hoploscopa pangrangoensis sp. nov., paratype, ♀, Indonesia, Java, Mt. Pangrange, SE Bogor, 6.30 S 107.10 E, 1625 m, primary forest, 16 - 20. ii. 1996 (Siniaev & Afonin) (genitalia on slide TL 627 ♀) 17 Hoploscopa isarogensis sp. nov., paratype, ♀, Philippines, South Luzon, Mt Isarog, 13 ° 40 ' N, 123 ° 20 ' E, 530 m, submontane forest, at light, 22. iii. 2000 (M. Nuss) (genitalia on slide TL 523 ♀) 18 Hoploscopa ypsilon sp. nov., paratype, ♀, Philippines, Luzon, Mountain Province, Chatol, 2100 m, 16 - 18. xi. 1997 (Mey, Ebert & Nuss)." figureDoi="10.3897/zookeys.907.36563.figures4-18" httpUri="https://binary.pensoft.net/fig/375757" pageId="0" pageNumber="1">4</figureCitation>
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). Collar white. Forewing length: 9-10 mm (♂ & ♀); forewing ground colour brown; rounded snow white basal cubital spot; median discoidal stigma rhomboid to ellipsoid, dark red, edged with pale yellow; postmedian spot trapezoid, dark red, proximally pale yellow; subterminal line snow white, parallel to termen, slightly angled at M1, broader at costa; fringes brown, with white dots. Hindwing dirty pale yellow. Forelegs bronze. Midlegs brown, with tibia and tarsi segments distally white. Hindlegs brown, with tibia and tarsi segments distally white.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Abdomen.</emphasis>
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Male sternum A8 posterior margin straight.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Male genitalia</emphasis>
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(
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<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 5) (Fig.
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<figureCitation captionStart="Figures 46–51" captionStartId="F5" captionText="Figures 46 - 51. Male genitalia features of Hoploscopa. For each figure left above: genitalia without phallus; left below: phallus; right above: gnathos drawing; right below: juxta drawing. 46 Hoploscopa albipuncta sp. nov., paratype, TL 336 ♂ (genitalia without phallus), TL 635 ♂ (phallus) 47 Hoploscopa matheae sp. nov., paratype, TL 730 ♂ 48 Hoploscopa sepanggi sp. nov., paratype, TL 641 ♂ 49 Hoploscopa cynodonta sp. nov., paratype, TL 531 ♂ 50 Hoploscopa parvimacula sp. nov., holotype, TL 510 ♂ 51 Hoploscopa kinabaluensis sp. nov., paratype, TL 342 ♂. Abbreviations: c (cornutus), de (ductus ejaculatorius), g (gnathos), jx (juxta), s (saccus), ta (tegumen arm), v (valva), vc (vinculum), uc (uncus)." figureDoi="10.3897/zookeys.907.36563.figures46-51" httpUri="https://binary.pensoft.net/fig/375760" pageId="0" pageNumber="1">46</figureCitation>
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). Uncus long and slender, gently narrowing on basal half, apex tongue-shaped. Gnathos projection broad, triangular, with rounded apex. Valva ventral margin straight, dorsal margin strongly protruded dorsad, apex rounded. Juxta with base rounded, medially narrowed, apex split into two conspicuous tips. Saccus small, pointing dorsad. Phallus with sclerotised, anvil-shaped cornutus.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Female genitalia</emphasis>
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(
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<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 1) (Fig.
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<figureCitation captionStart="Figures 87–90" captionStartId="F12" captionText="Figures 87 - 90. Female genitalia features of Hoploscopa. 87 Hoploscopa albipuncta sp. nov., paratype, TL 609 ♀ 88 Hoploscopa matheae sp. nov., paratype, TL 599 ♀ 89 Hoploscopa sepanggi sp. nov., paratype, TL 547 ♀ 90 Hoploscopa cynodonta sp. nov., paratype, TL 625 ♀. Abbreviations: aa (anterior apophyses), at (antrum), cb (corpus bursae), cs (corpus sclerotization), db (ductus bursae), ds (ductus seminalis), pa (posterior apophyses), pp (papillae anales), segm. VIII (segment VIII), t (thorn)." figureDoi="10.3897/zookeys.907.36563.figures87-90" httpUri="https://binary.pensoft.net/fig/375767" pageId="0" pageNumber="1">87</figureCitation>
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). Anterior apophyses with dorsal bump at posterior 1/3. Antrum forming a membranous rounded pouch. Ductus bursae of medium length, slender, straight. Corpus bursae small, globular, reticulate, with barely marked sclerotisation at thorn base. Thorn curved, with small dents pointing toward thorn apex on its inner side, glabrous on the outer side.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">Known from the slopes of the Mount Kinabalu (4,095 m) on Borneo, at altitudes between 1,700 m and 2,000 m.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="dna barcoding">
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<paragraph pageId="0" pageNumber="1">DNA barcoding.</paragraph>
|
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<paragraph pageId="0" pageNumber="1">Specimen MTD7430 from Luzon shows an K2P-distance of 4.5-4.9% with the two specimens from Borneo.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="etymology">
|
||
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
The species name
|
||
<taxonomicName authorityName="Leger & Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="albipuncta" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="albipuncta">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">albipuncta</emphasis>
|
||
</taxonomicName>
|
||
refers to the Latin
|
||
<emphasis italics="true" pageId="0" pageNumber="1">albus</emphasis>
|
||
, white, and
|
||
<emphasis italics="true" pageId="0" pageNumber="1">punctus</emphasis>
|
||
, forming a point.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="remarks">
|
||
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">Two female specimens from the Philippines with similar habitus and genitalia but a COI barcode divergence of 4.5-4.9% are recovered as different MOTU in the species delimitation analyses. Unfortunately, no male was available for this study, thus we refrained from describing a new species based on these two specimens.</paragraph>
|
||
<caption doi="10.3897/zookeys.907.36563.figures4-18" httpUri="https://binary.pensoft.net/fig/375757" pageId="0" pageNumber="1" start="Figures 4–18" startId="F2">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figures 4-18.</emphasis>
|
||
Forewing of
|
||
<taxonomicName authorityName="Meyrick" authorityYear="1886" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
<emphasis bold="true" pageId="0" pageNumber="1">4</emphasis>
|
||
<taxonomicName authorityName="Leger & Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa albipuncta" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="albipuncta">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa albipuncta</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype, ♂, Malaysia, Sabah, Kinabalu Park HQ, Timpohon Gate, 700 m from Liwagu Trail starting point, near Liwagu River,
|
||
<geoCoordinate degrees="6" direction="north" minutes="1" orientation="latitude" precision="15" seconds="40" value="6.027778">6°1'40"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="116" direction="east" minutes="32" orientation="longitude" precision="15" seconds="59" value="116.54972">116°32'59"E</geoCoordinate>
|
||
, 1760 m, 18.vi.2015 (T.
|
||
<normalizedToken originalValue="Léger">Leger</normalizedToken>
|
||
& R. Mally) (genitalia on slide TL335 ♂)
|
||
<emphasis bold="true" pageId="0" pageNumber="1">5</emphasis>
|
||
<taxonomicName authorityName="Leger & Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa sepanggi" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sepanggi">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa sepanggi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype, ♂, Malaysia, Sabah, Mesilau Nature Resort,
|
||
<geoCoordinate degrees="6" direction="north" minutes="2" orientation="latitude" precision="1" seconds="43.71" value="6.045475">6°2'43.71"N</geoCoordinate>
|
||
, 116°35'48.03'E, 1925 m, at lamps, 30.v.2015 (T.
|
||
<normalizedToken originalValue="Léger">Leger</normalizedToken>
|
||
& R. Mally)
|
||
<emphasis bold="true" pageId="0" pageNumber="1">6</emphasis>
|
||
<taxonomicName authorityName="Leger & Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa cynodonta" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="cynodonta">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa cynodonta</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype, ♂, Malaysia, Sabah, Kinabalu Park HQ, junction Kiau View- and Pandanus Trail,
|
||
<geoCoordinate degrees="6" direction="north" minutes="0" orientation="latitude" precision="1" seconds="32.84" value="6.0091224">6°0'32.84"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="116" direction="east" minutes="32" orientation="longitude" precision="1" seconds="14.94" value="116.53748">116°32'14.94"E</geoCoordinate>
|
||
, 1690m, UV light, 07.vi.2015 (T.
|
||
<normalizedToken originalValue="Léger">Leger</normalizedToken>
|
||
& R. Mally) (genitalia on slide TL327 ♂)
|
||
<emphasis bold="true" pageId="0" pageNumber="1">7</emphasis>
|
||
<taxonomicName authorityName="Leger & Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa parvimacula" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="parvimacula">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa parvimacula</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., paratype, ♀, Sabah, Mesilau Nature Resort,
|
||
<geoCoordinate degrees="6" direction="north" minutes="2" orientation="latitude" precision="1" seconds="43.71" value="6.045475">6°2'43.71"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="116" direction="east" minutes="35" orientation="longitude" precision="1" seconds="48.03" value="116.59668">116°35'48.03"E</geoCoordinate>
|
||
, 1925 m, at lamps, 30.v.2015 (T.
|
||
<normalizedToken originalValue="Léger">Leger</normalizedToken>
|
||
& R. Mally) (genitalia on slide TL312 ♀.
|
||
<emphasis bold="true" pageId="0" pageNumber="1">8</emphasis>
|
||
<taxonomicName authorityName="Leger & Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa kinabaluensis" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kinabaluensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa kinabaluensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., paratype, ♀, Sabah, Kinabalu Park HQ, Timpohon Gate, 700 m from Liwagu trail starting point, near Liwagu River,
|
||
<geoCoordinate degrees="6" direction="north" minutes="1" orientation="latitude" precision="15" seconds="40" value="6.027778">6°1'40"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="116" direction="east" minutes="32" orientation="longitude" precision="15" seconds="59" value="116.54972">116°32'59"E</geoCoordinate>
|
||
, 1760 m, UV light, 18.vi.2015 (T.
|
||
<normalizedToken originalValue="Léger">Leger</normalizedToken>
|
||
& R. Mally) (genitalia on slide TL326 ♀)
|
||
<emphasis bold="true" pageId="0" pageNumber="1">9</emphasis>
|
||
<taxonomicName authorityName="Nuss" authorityYear="1998" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa luteomacula" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="luteomacula">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa luteomacula</emphasis>
|
||
</taxonomicName>
|
||
Nuss, paratype, ♀, Indonesia, Sumatra, Barat, N-Padangpanjang, Mt Singgalang, 2100 m, 10-11.ii.1996 (A. Kallies) (genitalia on slide GU prep. Nuss 743)
|
||
<emphasis bold="true" pageId="0" pageNumber="1">10</emphasis>
|
||
<taxonomicName baseAuthorityName="Rothschild" baseAuthorityYear="1915" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa obliqua" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="obliqua">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa obliqua</emphasis>
|
||
</taxonomicName>
|
||
Rotschild, abdomen missing, Papua New Guinea, Madang Province, Wanang village,
|
||
<geoCoordinate degrees="05" direction="south" minutes="15" orientation="latitude" precision="925" value="-5.25">05°15'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="145" direction="east" minutes="17" orientation="longitude" precision="925" value="145.28334">145°17'E</geoCoordinate>
|
||
, reared from
|
||
<taxonomicName class="Polypodiopsida" family="Athyriaceae" genus="Diplazium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Diplazium esculentum" order="Polypodiales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="esculentum">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Diplazium esculentum</emphasis>
|
||
</taxonomicName>
|
||
, 12.vi.2007
|
||
<emphasis bold="true" pageId="0" pageNumber="1">11</emphasis>
|
||
<taxonomicName authorityName="Leger & Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa niveofascia" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="niveofascia">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa niveofascia</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype, ♂, Papua New Guinea, Morobe Prov., nr. Bulolo, Mt. Susu Nat. Res., 975 m, 27-28.viii. 1983,
|
||
<taxonomicName class="Pinopsida" family="Araucariaceae" genus="Araucaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Araucaria" order="Pinales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Araucaria</emphasis>
|
||
</taxonomicName>
|
||
For. (S. Miller) (genitalia on slide TL442 ♂)
|
||
<emphasis bold="true" pageId="0" pageNumber="1">12</emphasis>
|
||
<taxonomicName authorityName="Leger & Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa gombongi" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="gombongi">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa gombongi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., paratype, ♂, Papua New Guinea, Yawan village,
|
||
<geoCoordinate degrees="06" direction="south" minutes="10" orientation="latitude" precision="925" value="-6.1666665">06°10'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="146" direction="east" minutes="5" orientation="longitude" precision="925" value="146.08333">146°5'E</geoCoordinate>
|
||
, NG Binatang Res. Ctr. (B Gewa, J. Kua, S Sau, A. Kinibel)
|
||
<emphasis bold="true" pageId="0" pageNumber="1">13</emphasis>
|
||
<taxonomicName authorityName="Leger & Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa tonsepi" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tonsepi">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa tonsepi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype, ♂, Papua New Guinea, Yawan village,
|
||
<geoCoordinate degrees="06" direction="south" minutes="10" orientation="latitude" precision="925" value="-6.1666665">06°10'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="146" direction="east" minutes="5" orientation="longitude" precision="925" value="146.08333">146°5'E</geoCoordinate>
|
||
, NG Binatang Res. Ctr. (J. Valeba, J. Auga, M. Dilu, F. Philip, R. Lilip) (genitalia on slide TL655 ♂)
|
||
<emphasis bold="true" pageId="0" pageNumber="1">14</emphasis>
|
||
<taxonomicName authorityName="Leger & Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa marijoweissae" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="marijoweissae">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa marijoweissae</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., ♂, Papua New Guinea: Morobe Province, Mount Kaindi, 2350 m, 11.xii.1976 (G. F. Hevel & R. E. Dietz) (genitalia on slide TL437 ♂)
|
||
<emphasis bold="true" pageId="0" pageNumber="1">15</emphasis>
|
||
<taxonomicName authorityName="Leger & Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa titika" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="titika">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa titika</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype, ♂, Sumatra-Holzweg, 25 km SSW-Pematangsiantar,
|
||
<normalizedToken originalValue="Straße">Strasse</normalizedToken>
|
||
nach Prapat, 13.ii.1996 (A. Kallies) (genitalia on slide TL505 ♂)
|
||
<emphasis bold="true" pageId="0" pageNumber="1">16</emphasis>
|
||
<taxonomicName authorityName="Leger & Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa pangrangoensis" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pangrangoensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa pangrangoensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., paratype, ♀, Indonesia, Java, Mt. Pangrange, SE Bogor, 6.30S 107.10E, 1625 m, primary forest, 16-20.ii.1996 (Siniaev & Afonin) (genitalia on slide TL627 ♀)
|
||
<emphasis bold="true" pageId="0" pageNumber="1">17</emphasis>
|
||
<taxonomicName authorityName="Leger & Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa isarogensis" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="isarogensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa isarogensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., paratype, ♀, Philippines, South Luzon, Mt Isarog,
|
||
<geoCoordinate degrees="13" direction="north" minutes="40" orientation="latitude" precision="925" value="13.666667">13°40'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="123" direction="east" minutes="20" orientation="longitude" precision="925" value="123.333336">123°20'E</geoCoordinate>
|
||
, 530 m, submontane forest, at light, 22.iii.2000 (M. Nuss) (genitalia on slide TL523 ♀)
|
||
<emphasis bold="true" pageId="0" pageNumber="1">18</emphasis>
|
||
<taxonomicName authorityName="Leger & Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa ypsilon" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ypsilon">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa ypsilon</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., paratype, ♀, Philippines, Luzon, Mountain Province, Chatol, 2100 m, 16-18.xi.1997 (Mey, Ebert & Nuss).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |