treatments-xml/data/59/7A/87/597A878EFFB9FF95FF66F4A68257F578.xml
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<document id="7224DFF1A36A9D76B66B1878396A71B9" ID-CLB-Dataset="7470" ID-DOI="10.11646/zootaxa.4948.3.4" ID-GBIF-Dataset="6bfd846c-0304-4330-8941-95c406630230" ID-ISSN="1175-5326" ID-Zenodo-Dep="4629100" ID-ZooBank="8E0BF97A-D9E1-4F2E-BB63-82A6ED47E5DC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616485698777" checkinUser="plazi" docAuthor="Xu, Deliang, Webb, Michael D., Shang, Suqin &amp; Zhang, Yalin" docDate="2021" docId="597A878EFFB9FF95FF66F4A68257F578" docLanguage="en" docName="zootaxa.4948.3.4.pdf" docOrigin="Zootaxa 4948 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hecaloidella bimaculata Zhang &amp; Webb 2021, sp. nov." docType="treatment" docVersion="8" lastPageNumber="386" masterDocId="A543FFF6FFBCFF90FFF1F728870FF30F" masterDocTitle="Review of the Pacific leafhopper genus Hecaloidella (Hemiptera: Cicadellidae Deltocephalinae: Selenocephalini) with description of sixteen new species" masterLastPageNumber="403" masterPageNumber="381" pageNumber="386" updateTime="1698934487255" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="CE0C79888D66499E564B21CAB9271500">Review of the Pacific leafhopper genus Hecaloidella (Hemiptera: Cicadellidae Deltocephalinae: Selenocephalini) with description of sixteen new species</mods:title>
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<mods:namePart id="FFBFD62C9D8CEC65EC43ECF5A9964345">Xu, Deliang</mods:namePart>
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<mods:affiliation id="AAAC0BB4C7EC6E2D1BE58B7F7338C7BA">Key Laboratory of Plant Protection Resources and Pest Management, Ministry of Education, Entomological Museum, College of Plant Protection, Northwest A &amp; F University, Yangling, Shaanxi 712100, China &amp; 18710305867 @ 163. com; https: // orcid. org / 0000 - 0003 - 0106 - 5570</mods:affiliation>
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<mods:namePart id="4FB35E29A816676339E80AA7D6C85D23">Webb, Michael D.</mods:namePart>
<mods:nameIdentifier id="96C0B551CE7960C8648C2213A67B97CC" type="ORCID">0000-0002-1312-6142</mods:nameIdentifier>
<mods:affiliation id="E02139A12743A6E72FC9A40C26D5332E">Department of Life Sciences, Natural History Museum, Cromwell Road, London, ME 15 6 QX, UK &amp; mdwebb 04 @ gmail. com; https: // orcid. org / 0000 - 0002 - 1312 - 6142</mods:affiliation>
<mods:nameIdentifier id="0A52B54DB62A4A9056CF34DD45109A3C" type="email">mdwebb04@gmail.com</mods:nameIdentifier>
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<mods:namePart id="26DBF2C92D7370067D3B24DD47D2AEE7">Shang, Suqin</mods:namePart>
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<mods:affiliation id="1CB4578E766590CF8C3D0C678BEDE23C">Biological Engineering Laboratory of Crop Diseases and Pests of Gansu Province, College of Plant Protection, Gansu Agricultural University, Lanzhou, Gansu 730070, China &amp; shangsq @ gsau. edu. cn; https: // orcid. org / 0000 - 0001 - 9376 - 0735</mods:affiliation>
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<mods:namePart id="F93D90B360584E900474E18041F23A54">Zhang, Yalin</mods:namePart>
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<mods:affiliation id="7A1BB57648BD63948BAAE8D2CCF14690">Key Laboratory of Plant Protection Resources and Pest Management, Ministry of Education, Entomological Museum, College of Plant Protection, Northwest A &amp; F University, Yangling, Shaanxi 712100, China &amp; yalinzh @ nwsuaf. edu. cn; https: // orcid. org / 0000 - 0002 - 1204 - 9181</mods:affiliation>
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<subSubSection id="99C96513FFB9FF95FF66F4A6857DF0A8" box="[151,626,909,936]" pageId="5" pageNumber="386" type="nomenclature">
<paragraph id="D16C3698FFB9FF95FF66F4A6857DF0A8" blockId="5.[151,626,909,971]" box="[151,626,909,936]" pageId="5" pageNumber="386">
<heading id="8A2481F4FFB9FF95FF66F4A6857DF0A8" bold="true" box="[151,626,909,936]" fontSize="11" level="1" pageId="5" pageNumber="386" reason="1">
<emphasis id="E3A7EA8AFFB9FF95FF66F4A6857DF0A8" bold="true" box="[151,626,909,936]" pageId="5" pageNumber="386">
<taxonomicName id="16D34D1BFFB9FF95FF66F4A68503F0A8" ID-CoL="8KMZL" authority="Zhang &amp; Webb" authorityName="Zhang &amp; Webb" authorityYear="2021" box="[151,524,909,936]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="386" phylum="Arthropoda" rank="species" species="bimaculata" status="sp. nov.">
<emphasis id="E3A7EA8AFFB9FF95FF66F4A687A1F0A7" bold="true" box="[151,174,910,936]" italics="true" pageId="5" pageNumber="386">H</emphasis>
.
<emphasis id="E3A7EA8AFFB9FF95FF4CF4A5864BF0A7" bold="true" box="[189,324,909,936]" italics="true" pageId="5" pageNumber="386">bimaculata</emphasis>
Zhang &amp; Webb
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<taxonomicNameLabel id="F89457F1FFB9FF95FDE5F4A5857DF0A8" box="[532,626,909,935]" pageId="5" pageNumber="386" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="99C96513FFB9FF95FF66F4998498F658" pageId="5" pageNumber="386" type="description">
<paragraph id="D16C3698FFB9FF95FF66F4998645F0C4" blockId="5.[151,626,909,971]" box="[151,330,945,971]" pageId="5" pageNumber="386">
<figureCitation id="49E82A1DFFB9FF95FF66F49987F5F0C4" box="[151,250,945,971]" captionStart="FIGURE 1" captionStartId="6.[151,250,1870,1894]" captionTargetBox="[223,1380,183,1845]" captionTargetId="figure-19@6.[205,1382,181,1846]" captionTargetPageId="6" captionText="FIGURE 1. Habitus images of Hecaloidella species; A, H. bimaculata (holotype); B, H. caudospina (holotype); C, H. nitida (holotype); D, H. nudapenis (holotype); EG, H. pacifica (paratypes and immature); H, H. borneoensis (holotype); I, H. brevipenis (holotype); J, H. brevispina (holotype); K, H. longipenis (holotype); L, H. quadrispina (holotype); M, H. spinosa (holotype); N, H. brevis (holotype); O, H. brunnifacialis (topotype); P, H. knighti (holotype); Q, H. quadripicta (paratype); R, H. sandakanensis (holotype); S, H. singaporensis (paratype); T, H. brunifasciata (holotype)." figureDoi="http://doi.org/10.5281/zenodo.4629102" httpUri="https://zenodo.org/record/4629102/files/figure.png" pageId="5" pageNumber="386">Figs. 1A</figureCitation>
,
<figureCitation id="49E82A1DFFB9FF95FEF7F4998645F0C4" box="[262,330,945,971]" captionStart="FIGURE 2" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[191,1403,188,1905]" captionTargetId="figure-17@7.[179,1408,181,1920]" captionTargetPageId="7" captionText="FIGURE 2. H. bimaculata; AB, habitus, dorsal and lateral view; C, face, D, head and thorax, anterior view; E, male genital capsule, lateral view; F, style; G, valve, subgenital plates, styles and connective, dorsal view; H, aedeagus, lateral view; I aedeagus, caudal view; J, anal tube, lateral view; K, connective, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4629104" httpUri="https://zenodo.org/record/4629104/files/figure.png" pageId="5" pageNumber="386">2AK</figureCitation>
</paragraph>
<paragraph id="D16C3698FFB9FF95FF66F4D1833EF71B" blockId="5.[151,1437,1017,1655]" box="[151,1073,1017,1044]" pageId="5" pageNumber="386">
<emphasis id="E3A7EA8AFFB9FF95FF66F4D18626F71C" bold="true" box="[151,297,1017,1043]" pageId="5" pageNumber="386">Description.</emphasis>
Body length (including tegmina), male:
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; female:
<quantity id="162B9B7DFFB9FF95FC2BF4D18322F71B" box="[986,1069,1017,1044]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="5" pageNumber="386" unit="mm" value="5.3">5.3mm</quantity>
.
</paragraph>
<paragraph id="D16C3698FFB9FF95FF36F3358492F770" blockId="5.[151,1437,1017,1655]" pageId="5" pageNumber="386">Stramineous to sordid yellow. Vertex with a large brown spot anteriorly each side of midline. Face with a medial dark brown inverted Y-shaped mark; antennal pedicel dark brown. Forewings brownish hyaline; veins yellowish with brown marking restricted to a small spot at apex of claval veins.</paragraph>
<paragraph id="D16C3698FFB9FF95FF36F3A284D2F7AB" blockId="5.[151,1437,1017,1655]" box="[199,989,1161,1188]" pageId="5" pageNumber="386">Vertex relatively long, nearly twice as long medially than next to eyes.</paragraph>
<paragraph id="D16C3698FFB9FF95FF36F3858276F63C" blockId="5.[151,1437,1017,1655]" pageId="5" pageNumber="386">Male genitalia with subgenital plate very broadly curved basally with very short digitate apex, with a few short fine setae dorsolaterally. Style apical process rostrate. Connective Y-shaped with short stem and relatively short sharply upturned caudal extension.Aedeagus relatively large, disassociated from connective, with stout ventral process; shaft moderately long, laterally compressed and tapered to acute apex without processes, gonopore apical.</paragraph>
<paragraph id="D16C3698FFB9FF95FF36F2158498F658" blockId="5.[151,1437,1017,1655]" box="[199,919,1341,1367]" pageId="5" pageNumber="386">Female pregenital sternite with posterior margin almost straight.</paragraph>
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<emphasis id="E3A7EA8AFFB9FF95FF36F24986BCF674" bold="true" box="[199,435,1377,1403]" pageId="5" pageNumber="386">Material examined.</emphasis>
<materialsCitation id="61BB3CC5FFB9FF95FE4CF24984C2F690" ID-GBIF-Occurrence="3068500303" collectingDate="1992-10" collectionCode="BMNH" collectorName="J. H. Martin" country="Brunei" elevation="300" location="Kuala Belalong" pageId="5" pageNumber="386" specimenCount="1" specimenCount-male="1" stateProvince="Temburong" typeStatus="holotype">
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:
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,
<collectingCountry id="A9C47608FFB9FF95FDADF24985A4F674" box="[604,683,1377,1403]" name="Brunei" pageId="5" pageNumber="386">Brunei</collectingCountry>
,
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, ridge NE of
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, approx.
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alt.,
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<collectingDate id="B529E9B0FFB9FF95FF3BF2AD8615F690" box="[202,282,1413,1439]" pageId="5" pageNumber="386" value="1992-10">x.1992</collectingDate>
</date>
,
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, m.
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v.
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</taxonomicName>
trap, NHMUK 010591776 (
<collectionCode id="B7C2AE5DFFB9FF95FC9AF2AD84CAF690" box="[875,965,1413,1439]" pageId="5" pageNumber="386">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="61BB3CC5FFB9FF95FC28F2AE87E1F6E8" ID-GBIF-Occurrence="3068500330" collectingDate="2006-10-16" collectingDateMax="2006-10-22" collectingDateMin="2006-10-16" collectionCode="INHS" collectorName="J. R. Cryan &amp; J. M. Urban" country="Malaysia" elevation="40" latitude="4.0424724" location="Mulu National Park" longLatPrecision="1" longitude="114.81297" pageId="5" pageNumber="386" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="0E68883AFFB9FF95FC28F2AE8345F6AF" box="[985,1098,1414,1440]" pageId="5" pageNumber="386" type="paratype">Paratypes</typeStatus>
:
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,
<collectingCountry id="A9C47608FFB9FF95FB7BF2AD83FCF690" box="[1162,1267,1413,1439]" name="Malaysia" pageId="5" pageNumber="386">Malaysia</collectingCountry>
, Sarawak, Gu- nung
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,
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<elevation id="5AFED1ABFFB9FF95FE31F2818515F6CB" box="[448,538,1449,1476]" metricMagnitude="1" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="5" pageNumber="386" unit="m" value="40.0" valueMax="50.0" valueMin="30.0">30-50m</elevation>
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, elevation,
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,
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,
<date id="A56D1058FFB9FF95FC12F2818374F6CC" box="[995,1147,1449,1475]" pageId="5" pageNumber="386" value="2006-10-16" valueMax="2006-10-22" valueMin="2006-10-16">
<collectingDate id="B529E9B0FFB9FF95FC12F2818374F6CC" box="[995,1147,1449,1475]" pageId="5" pageNumber="386" value="2006-10-16" valueMax="2006-10-22" valueMin="2006-10-16">16-22.x.2006</collectingDate>
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,
<collectorName id="7C26534EFFB9FF95FB77F28283F4F6CC" box="[1158,1275,1449,1476]" pageId="5" pageNumber="386">J.R. Cryan</collectorName>
&amp;
<collectorName id="7C26534EFFB9FF95FAEFF2828293F6CC" box="[1310,1436,1449,1476]" pageId="5" pageNumber="386">J.M. Urban</collectorName>
(
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</materialsCitation>
;
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<specimenCount id="C7D5FD11FFB9FF95FF0BF2E58612F6E8" box="[250,285,1485,1511]" pageId="5" pageNumber="386" type="female">1♀</specimenCount>
,
<collectingCountry id="A9C47608FFB9FF95FEDAF2E5869BF6E8" box="[299,404,1485,1511]" name="Malaysia" pageId="5" pageNumber="386">Malaysia</collectingCountry>
, Sarawak,
<location id="D40C6043FFB9FF95FDFFF2E5845EF6E8" LSID="urn:lsid:plazi:treatment:597A878EFFB9FF95FF66F4A68257F578:D40C6043FFB9FF95FDFFF2E5845EF6E8" box="[526,849,1485,1511]" country="Malaysia" county="Mulu" municipality="Camp" name="Gunung Mulu National Park" pageId="5" pageNumber="386">Gunung Mulu National Park</location>
,
<location id="D40C6043FFB9FF95FCADF2E58487F6E8" LSID="urn:lsid:plazi:treatment:597A878EFFB9FF95FF66F4A68257F578:D40C6043FFB9FF95FCADF2E58487F6E8" box="[860,904,1485,1511]" country="Malaysia" county="Mulu" municipality="Camp" name="Site" pageId="5" pageNumber="386">Site</location>
10,
<location id="D40C6043FFB9FF95FC48F2E58310F6E8" LSID="urn:lsid:plazi:treatment:597A878EFFB9FF95FF66F4A68257F578:D40C6043FFB9FF95FC48F2E58310F6E8" box="[953,1055,1485,1511]" country="Malaysia" county="Mulu" municipality="Camp" name="February" pageId="5" pageNumber="386">February</location>
,
<collectingMunicipality id="3108ACE2FFB9FF95FBDBF2E58362F6E8" box="[1066,1133,1485,1511]" pageId="5" pageNumber="386">Camp</collectingMunicipality>
1
<collectingCounty id="380D4E14FFB9FF95FB78F2E583C7F6E8" box="[1161,1224,1485,1511]" pageId="5" pageNumber="386">Mulu</collectingCounty>
,
<quantity id="162B9B7DFFB9FF95FB25F2E58218F6E8" box="[1236,1303,1485,1511]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="5" pageNumber="386" unit="m" value="160.0">
<elevation id="5AFED1ABFFB9FF95FB25F2E58218F6E8" box="[1236,1303,1485,1511]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="5" pageNumber="386" unit="m" value="160.0">160m</elevation>
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, mixed dip- terocarp forest,
<collectorName id="7C26534EFFB9FF95FEB7F2D98664F504" box="[326,363,1521,1547]" pageId="5" pageNumber="386">Ac.</collectorName>
-understorey,
<collectorName id="7C26534EFFB9FF95FDF6F2DA85A3F504" box="[519,684,1521,1548]" pageId="5" pageNumber="386">J.D. Holloway</collectorName>
,
<collectorName id="7C26534EFFB9FF95FD47F2D983B7F504" box="[694,1208,1521,1547]" pageId="5" pageNumber="386">Royal Geographical Society Mulu Expedition</collectorName>
(
<collectionCode id="B7C2AE5DFFB9FF95FB37F2D9822FF504" box="[1222,1312,1521,1547]" pageId="5" pageNumber="386">BMNH</collectionCode>
)
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.
</paragraph>
</subSubSection>
<subSubSection id="99C96513FFB9FF95FF36F13D8363F53F" box="[199,1132,1557,1584]" pageId="5" pageNumber="386" type="etymology">
<paragraph id="D16C3698FFB9FF95FF36F13D8363F53F" blockId="5.[151,1437,1017,1655]" box="[199,1132,1557,1584]" pageId="5" pageNumber="386">
<emphasis id="E3A7EA8AFFB9FF95FF36F13D8641F520" bold="true" box="[199,334,1557,1583]" pageId="5" pageNumber="386">Etymology.</emphasis>
This species is named after the pair of large brown spots on the vertex.
</paragraph>
</subSubSection>
<subSubSection id="99C96513FFB9FF95FF36F1118257F578" pageId="5" pageNumber="386" type="discussion">
<paragraph id="D16C3698FFB9FF95FF36F1118257F578" blockId="5.[151,1437,1017,1655]" pageId="5" pageNumber="386">
<emphasis id="E3A7EA8AFFB9FF95FF36F1118634F55C" bold="true" box="[199,315,1593,1619]" pageId="5" pageNumber="386">Remarks.</emphasis>
This species differs from other species of
<taxonomicName id="16D34D1BFFB9FF95FCEBF1128471F55C" authorityName="Osborn" authorityYear="1934" box="[794,894,1593,1620]" class="Insecta" family="Cicadellidae" genus="Hecaloidella" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="386" phylum="Arthropoda" rank="species" species="nitida">
<emphasis id="E3A7EA8AFFB9FF95FCEBF1128421F55C" box="[794,814,1594,1619]" italics="true" pageId="5" pageNumber="386">H</emphasis>
.
<emphasis id="E3A7EA8AFFB9FF95FCCCF1118471F55C" box="[829,894,1593,1619]" italics="true" pageId="5" pageNumber="386">nitida</emphasis>
</taxonomicName>
group by its large head spots, longer vertex and in the male genitalia by its larger aedeagus and shorter caudal extension of the connective strongly upturned.
</paragraph>
</subSubSection>
</treatment>
</document>