282 lines
38 KiB
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282 lines
38 KiB
XML
<document id="405A42380E026413F290F4F327B3823C" ID-CLB-Dataset="21249" ID-DOI="10.5281/zenodo.3985221" ID-GBIF-Dataset="a2d1f7b5-8049-43dd-8166-d60a2216ca70" ID-ISSN="0253-116X" ID-Zenodo-Dep="3985221" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1597410763446" checkinUser="valdenar" docAuthor="Assing, Volker" docDate="2018" docId="50607F34FFECFF86EFA1FD2317BBFA41" docLanguage="en" docName="Linzer.biol.Beitr.50.1.0025-0040.pdf.imf" docOrigin="Linzer biologische Beiträge 50 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Othius burmensis Assing 2018, nov.sp." docType="treatment" docVersion="8" lastPageNumber="38" masterDocId="AC59074CFFE0FF8BEF13FF9B1611FFAC" masterDocTitle="A revision of Othiini. XX. Two new species of Othius, the first record of the genus from Myanmar, and additional records (Coleoptera: Staphylinidae: Staphylininae)" masterLastPageNumber="40" masterPageNumber="25" pageNumber="37" updateTime="1698844491833" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="532BBB14AED20F076121E30DAB0F25FB">
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<mods:title id="102AC201DED42448C56608B64017B67C">A revision of Othiini. XX. Two new species of Othius, the first record of the genus from Myanmar, and additional records (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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<mods:name id="3673BCDF528A6B283ED11132DE9A1E65" type="personal">
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<mods:namePart id="64D3ADFAAB6453EC7C7D28388CDC3087">Assing, Volker</mods:namePart>
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<mods:typeOfResource id="974A7AF4D51A4BC7F05AA785D2A57BF7">text</mods:typeOfResource>
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<mods:title id="7619F52DFCBFF1A41D5E55CD791AD98C">Linzer biologische Beiträge</mods:title>
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<mods:part id="E65A7E4D6D51452F7A77059147DD432B">
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<mods:date id="60108135090C00F105D6BDCF6358B9FB">2018</mods:date>
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<mods:number id="8E0DAF39A371343BFF130D4FC1527FF5">2018-07-27</mods:number>
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<mods:number id="96F6E3A5DE107B7C41175ED82499169C">50</mods:number>
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<mods:start id="AF5ACDCB98AC21E7FE9342852C5CF13E">25</mods:start>
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<mods:identifier id="A262DE305CCD4FE656BDE7DD81EBEE01" type="DOI">10.5281/zenodo.3985221</mods:identifier>
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<treatment id="50607F34FFECFF86EFA1FD2317BBFA41" ID-DOI="http://doi.org/10.5281/zenodo.4330295" ID-GBIF-Taxon="167284516" ID-Zenodo-Dep="4330295" LSID="urn:lsid:plazi:treatment:50607F34FFECFF86EFA1FD2317BBFA41" httpUri="http://treatment.plazi.org/id/50607F34FFECFF86EFA1FD2317BBFA41" lastPageId="13" lastPageNumber="38" pageId="12" pageNumber="37">
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<subSubSection id="90D39DA9FFECFF87EFA1FD231498FD63" box="[178,649,696,719]" pageId="12" pageNumber="37" type="nomenclature">
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<paragraph id="D876CE22FFECFF87EFA1FD231498FD63" blockId="12.[178,649,696,719]" box="[178,649,696,719]" pageId="12" pageNumber="37">
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<heading id="833E794EFFECFF87EFA1FD231498FD63" box="[178,649,696,719]" fontSize="8" level="2" pageId="12" pageNumber="37" reason="2">
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<emphasis id="EABD1230FFECFF87EFA1FD2317AFFD62" bold="true" box="[178,446,696,718]" pageId="12" pageNumber="37">
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<taxonomicName id="1FC9B5A1FFECFF87EFA1FD23177AFD62" ID-CoL="75CNY" authority="Assing, 2018" authorityName="Assing" authorityYear="2018" box="[178,363,696,718]" class="Insecta" family="Staphylinidae" genus="Othius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="burmensis" status="sp. nov.">
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<emphasis id="EABD1230FFECFF87EFA1FD23177AFD62" bold="true" box="[178,363,696,718]" italics="true" pageId="12" pageNumber="37">Othius burmensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="F18EAF4BFFECFF87EE62FD2217AFFD62" box="[369,446,697,718]" pageId="12" pageNumber="37" rank="species">nov.sp.</taxonomicNameLabel>
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</emphasis>
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(
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<figureCitation id="40F2D2A7FFECFF87EED8FD23142DFD62" box="[459,572,696,718]" captionStart="Figs 15-23" captionStartId="9.[178,218,1503,1525]" captionTargetBox="[191,1085,212,1496]" captionTargetId="figure@9.[178,1085,212,1497]" captionTargetPageId="9" captionText="Figs 15-23: Othius burmensis: (15) head and pronotum; (16) head; (17) median portion of head; (18) sutural portion of right elytron; (19) male sternite IX; (20) aedeagus in ventral view (internal sac fully extruded); (21) lateral internal structures of aedeagus; (22-23) basal internal structure. Scale bars: 15-16: 1.0 mm; 19-20: 0.5 mm; 17-18, 21: 0.2 mm; 22-23: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3985233" httpUri="https://zenodo.org/record/3985233/files/figure.png" pageId="12" pageNumber="37">Figs 15-23</figureCitation>
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,
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<figureCitation id="40F2D2A7FFECFF87ED55FD221492FD63" box="[582,643,697,719]" captionStart="Figs 24-29" captionStartId="10.[178,218,1314,1336]" captionTargetBox="[178,1085,212,1305]" captionTargetId="figure@10.[178,1085,212,1309]" captionTargetPageId="10" captionText="Figs 24-29: Othius arater (24-26) and O. burmensis (27-29): (24-26) internal structures of aedeagus in ventral view; (27-28) aedeagus in lateral and in ventral view (without extruded internal sac); (29) apical median internal structure of the aedeagus. Scale bars: 27-28: 0.5 mm; 24-26, 29: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3985235" httpUri="https://zenodo.org/record/3985235/files/figure.png" pageId="12" pageNumber="37">27-29</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="90D39DA9FFECFF87EFA1FD7817BEFC91" pageId="12" pageNumber="37" type="materials_examined">
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<paragraph id="D876CE22FFECFF87EFA1FD7817BEFC91" blockId="12.[178,1086,739,829]" pageId="12" pageNumber="37">
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<materialsCitation id="68A1C47FFFECFF87EFA1FD7817BEFC91" ID-GBIF-Occurrence="2846537311" collectingDate="2010-10-06" collectionCode="T, TF, V, NHMW" collectorName="Michael Langer, S & S. Loffler" country="Myanmar" elevation="2" location="Provinz" longitude="25.731222" municipality="Burma" pageId="12" pageNumber="37" specimenCount="2" specimenCount-male="2" stateProvince="Kachin State" typeStatus="holotype">
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<collectionCode id="BED856E7FFECFF87EFA1FD7816D1FD54" box="[178,192,739,760]" name="Tavera, Department of Geology and Geophysics" pageId="12" pageNumber="37">T</collectionCode>
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y p e m a t e r i a l:
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<typeStatus id="07727080FFECFF87EEB4FD7817EAFD55" box="[423,507,739,761]" pageId="12" pageNumber="37" type="holotype">Holotype</typeStatus>
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<specimenCount id="CECF05ABFFECFF87ED12FD781404FD54" box="[513,533,739,760]" pageId="12" pageNumber="37" type="male">♂</specimenCount>
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: "
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<collectingCountry id="A0DE8EB2FFECFF87ED35FD7814B0FD54" box="[550,673,739,760]" name="Myanmar" pageId="12" pageNumber="37">MYANMAR</collectingCountry>
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(
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<collectingMunicipality id="38125458FFECFF87EDBEFD7814FEFD55" box="[685,751,739,761]" pageId="12" pageNumber="37">Burma</collectingMunicipality>
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),
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<location id="DD1698F9FFECFF87EDECFD781554FD55" LSID="urn:lsid:plazi:treatment:50607F34FFECFF86EFA1FD2317BBFA41:DD1698F9FFECFF87EDECFD781554FD55" box="[767,837,739,761]" country="Myanmar" longitude="25.731222" municipality="Burma" name="Provinz" pageId="12" pageNumber="37" stateProvince="Kachin State">Provinz</location>
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<collectingRegion id="1A0D00C0FFECFF87EC5FFD7815D2FD54" box="[844,963,739,761]" country="Myanmar" name="Kachin" pageId="12" pageNumber="37">Kachin State</collectingRegion>
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, ca.
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<locationDeviation id="C9C4D700FFECFF87ECFEFD781735FCA3" location="Pangwa" pageId="12" pageNumber="37">
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<quantity id="1F3163C7FFECFF87ECFEFD781236FD55" box="[1005,1063,739,761]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="37" unit="km" value="30.0">30 km</quantity>
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N von Pangwa
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</locationDeviation>
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, 29.-30.IX. +
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<date id="AC77E8E2FFECFF87EEB0FD6217E8FCA3" box="[419,505,761,783]" pageId="12" pageNumber="37" value="2010-10-06">
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<collectingDate id="BC33110AFFECFF87EEB0FD6217E8FCA3" box="[419,505,761,783]" pageId="12" pageNumber="37" value="2010-10-06">6.X.2010</collectingDate>
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</date>
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,
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<geoCoordinate id="BDFDA8E5FFECFF87ED10FD611503FCA2" box="[515,786,761,783]" pageId="12" pageNumber="37">N 25°43'52.4'' E 098°24'06.0''</geoCoordinate>
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(H =
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<quantity id="1F3163C7FFECFF87EC5BFD621584FCA3" box="[840,917,761,783]" metricMagnitude="0" metricUnit="m" metricValue="2.255" pageId="12" pageNumber="37" unit="m" value="2.255">
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<elevation id="53E42911FFECFF87EC5BFD621584FCA3" box="[840,917,761,783]" metricMagnitude="0" metricUnit="m" metricValue="2.255" pageId="12" pageNumber="37" unit="m" value="2.255">2.255 m</elevation>
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</quantity>
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;
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<collectionCode id="BED856E7FFECFF87EC8DFD6115A9FCA3" box="[926,952,762,783]" country="Thailand" name="Department of Mineral Resources" pageId="12" pageNumber="37">TF</collectionCode>
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- under wood, stones), leg.
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<collectorName id="753CABF4FFECFF87EE34FC8B17C2FC89" box="[295,467,784,806]" pageId="12" pageNumber="37">Michael Langer, S</collectorName>
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. Naumann &
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<collectorName id="753CABF4FFECFF87ED47FC8B14BCFC8A" box="[596,685,784,806]" pageId="12" pageNumber="37">S. Löffler</collectorName>
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/
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<typeStatus id="07727080FFECFF87EDD3FC8B150FFC8A" box="[704,798,784,806]" pageId="12" pageNumber="37" type="holotype">Holotypus</typeStatus>
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<specimenCount id="CECF05ABFFECFF87EC36FC8B1527FC89" box="[805,822,784,805]" pageId="12" pageNumber="37" type="male">♂</specimenCount>
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<taxonomicName id="1FC9B5A1FFECFF87EC2EFC8B15CCFC89" authority="Assing, 2018" authorityName="Assing" authorityYear="2018" box="[829,989,784,805]" class="Insecta" family="Staphylinidae" genus="Othius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="burmensis" status="sp. nov.">
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<emphasis id="EABD1230FFECFF87EC2EFC8B15CCFC89" box="[829,989,784,805]" italics="true" pageId="12" pageNumber="37">Othius burmensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="F18EAF4BFFECFF87ECF7FC8B1207FC89" box="[996,1046,784,805]" pageId="12" pageNumber="37" rank="species">sp. n.</taxonomicNameLabel>
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det.
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<collectionCode id="BED856E7FFECFF87EFA1FCBC16D4FC90" box="[178,197,807,828]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="37" type="Museum">V</collectionCode>
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.
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<bibRefCitation id="BC58B3D3FFECFF87EFDDFCBC1750FC90" author="ASSING V" box="[206,321,806,829]" pageId="12" pageNumber="37" pagination="1235 - 1250" refId="ref9084" refString="ASSING V. (2015): A revision of Othiini. XIX. Two new species of Othius, the first record of the genus from Vietnam, and additional records (Coleoptera: Staphylinidae: Staphylininae). - Linzer Biologische Beitrage 47 (2): 1235 - 1250." type="journal article" year="2015">Assing 2015</bibRefCitation>
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" (
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<collectionCode id="BED856E7FFECFF87EE46FCBC17B4FC91" box="[341,421,807,829]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="12" pageNumber="37" type="Museum">NHMW</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="90D39DA9FFECFF87EFA1FCD014F7FCD1" pageId="12" pageNumber="37" type="etymology">
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<paragraph id="D876CE22FFECFF87EFA1FCD014F7FCD1" blockId="12.[178,1085,843,894]" pageId="12" pageNumber="37">
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E t y m o l o g y: The specific epithet is an adjective derived from
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<collectingCountry id="A0DE8EB2FFECFF87EC6CFCD015D6FCCC" box="[895,967,843,864]" name="Myanmar" pageId="12" pageNumber="37">Burma</collectingCountry>
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, where this species is currently the sole representative of the genus.
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</paragraph>
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</subSubSection>
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<subSubSection id="90D39DA9FFECFF86EFA1FC161751FAC5" lastPageId="13" lastPageNumber="38" pageId="12" pageNumber="37" type="description">
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<paragraph id="D876CE22FFECFF87EFA1FC161477FC71" blockId="12.[178,1085,909,990]" pageId="12" pageNumber="37">D e s c r i p t i o n: Measurements (mm) and ratios: HL: 1.59; HW: 1.53; PW: 1.65; PL: 1.95; EL: 1.37; TiL: 1.40; TaL: 1.16; TL: 11.4; HL/HW: 1.04; HW/PW: 0.93; PL/PW: 1.19; EL/PL: 0.70; TaL/TiL: 0.83.</paragraph>
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<paragraph id="D876CE22FFECFF87EFA1FC761200FB92" blockId="12.[178,1085,1005,1086]" pageId="12" pageNumber="37">Moderately large species (see measurements). Coloration: head, pronotum, and abdomen black; elytra blackish-brown with a dark-reddish hue; legs dark-brown with reddish tarsi; antennae dark-reddish, with antennomeres I, III, and the apical portion of II blackish.</paragraph>
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<paragraph id="D876CE22FFECFF87EFA1FBD61731FA9C" blockId="12.[178,1085,1101,1328]" pageId="12" pageNumber="37">
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Head (
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<figureCitation id="40F2D2A7FFECFF87EFE5FBD61777FBCF" box="[246,358,1101,1123]" captionStart="Figs 15-23" captionStartId="9.[178,218,1503,1525]" captionTargetBox="[191,1085,212,1496]" captionTargetId="figure@9.[178,1085,212,1497]" captionTargetPageId="9" captionText="Figs 15-23: Othius burmensis: (15) head and pronotum; (16) head; (17) median portion of head; (18) sutural portion of right elytron; (19) male sternite IX; (20) aedeagus in ventral view (internal sac fully extruded); (21) lateral internal structures of aedeagus; (22-23) basal internal structure. Scale bars: 15-16: 1.0 mm; 19-20: 0.5 mm; 17-18, 21: 0.2 mm; 22-23: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3985233" httpUri="https://zenodo.org/record/3985233/files/figure.png" pageId="12" pageNumber="37">Figs 15-16</figureCitation>
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) weakly oblong, weakly widened posteriorly, and somewhat narrower than pronotum (see ratios HL/HW and HW/PW); anterior pair of frontal punctures distinct, with additional smaller punctures; posterior pair of frontal punctures absent; median dorsal portion impunctate; lateral and posterior dorsal portions with rather sparse and coarse punctures of somewhat variable size; integument with distinct and fine transverse microsculpture (
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<figureCitation id="40F2D2A7FFECFF87EED5FB7B1400FB5A" box="[454,529,1248,1270]" captionStart="Figs 15-23" captionStartId="9.[178,218,1503,1525]" captionTargetBox="[191,1085,212,1496]" captionTargetId="figure@9.[178,1085,212,1497]" captionTargetPageId="9" captionText="Figs 15-23: Othius burmensis: (15) head and pronotum; (16) head; (17) median portion of head; (18) sutural portion of right elytron; (19) male sternite IX; (20) aedeagus in ventral view (internal sac fully extruded); (21) lateral internal structures of aedeagus; (22-23) basal internal structure. Scale bars: 15-16: 1.0 mm; 19-20: 0.5 mm; 17-18, 21: 0.2 mm; 22-23: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3985233" httpUri="https://zenodo.org/record/3985233/files/figure.png" pageId="12" pageNumber="37">Fig. 17</figureCitation>
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). Eyes of moderate size, approximately half as long as postocular region in dorsal view. Antenna
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<quantity id="1F3163C7FFECFF87ED90FB6614C7FABE" box="[643,726,1277,1298]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="12" pageNumber="37" unit="mm" value="2.9">2.9 mm</quantity>
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long; antennomeres V-X weakly transverse.
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</paragraph>
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<paragraph id="D876CE22FFECFF87EFA1FADB1731FA66" blockId="12.[178,1085,1344,1482]" pageId="12" pageNumber="37">
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Pronotum (
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<figureCitation id="40F2D2A7FFECFF87EE3BFADB1764FAFA" box="[296,373,1344,1366]" captionStart="Figs 15-23" captionStartId="9.[178,218,1503,1525]" captionTargetBox="[191,1085,212,1496]" captionTargetId="figure@9.[178,1085,212,1497]" captionTargetPageId="9" captionText="Figs 15-23: Othius burmensis: (15) head and pronotum; (16) head; (17) median portion of head; (18) sutural portion of right elytron; (19) male sternite IX; (20) aedeagus in ventral view (internal sac fully extruded); (21) lateral internal structures of aedeagus; (22-23) basal internal structure. Scale bars: 15-16: 1.0 mm; 19-20: 0.5 mm; 17-18, 21: 0.2 mm; 22-23: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3985233" httpUri="https://zenodo.org/record/3985233/files/figure.png" pageId="12" pageNumber="37">Fig. 15</figureCitation>
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) moderately oblong (see ratio PL/PW) and with subparallel lateral margins in dorsal view; discal punctation pattern similar to that of most other East Palaearctic congeners; posterior discal puncture (i. e., puncture near posterior angle) separated from posterior angle by a distance less than its diameter; microsculpture finely transverse.
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</paragraph>
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<paragraph id="D876CE22FFECFF87EFA1FA4115EBF986" blockId="12.[178,1085,1498,1579]" pageId="12" pageNumber="37">
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Elytra (
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<figureCitation id="40F2D2A7FFECFF87EFECFA41175BFA5C" box="[255,330,1498,1520]" captionStart="Figs 15-23" captionStartId="9.[178,218,1503,1525]" captionTargetBox="[191,1085,212,1496]" captionTargetId="figure@9.[178,1085,212,1497]" captionTargetPageId="9" captionText="Figs 15-23: Othius burmensis: (15) head and pronotum; (16) head; (17) median portion of head; (18) sutural portion of right elytron; (19) male sternite IX; (20) aedeagus in ventral view (internal sac fully extruded); (21) lateral internal structures of aedeagus; (22-23) basal internal structure. Scale bars: 15-16: 1.0 mm; 19-20: 0.5 mm; 17-18, 21: 0.2 mm; 22-23: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3985233" httpUri="https://zenodo.org/record/3985233/files/figure.png" pageId="12" pageNumber="37">Fig. 18</figureCitation>
|
|
) shorter than pronotum (see ratio EL/PL); macropunctation rather sparse and shallow; interstices on average several times as broad as diameter of punctures, with very dense micropunctation rendering the surface nearly matt. Hind wings present.
|
|
</paragraph>
|
|
<paragraph id="D876CE22FFEDFF86EFA1FF401221FEA2" blockId="13.[178,1085,219,270]" pageId="13" pageNumber="38">Abdomen slightly narrower than elytra; punctation fine and moderately dense; interstices with fine transverse microsculpture; posterior margin of tergite VII with palisade fringe.</paragraph>
|
|
<paragraph id="D876CE22FFEDFF86EFA1FE85147FFD90" blockId="13.[178,1085,285,572]" pageId="13" pageNumber="38">
|
|
<specimenCount id="CECF05ABFFEDFF86EFA1FE8516D9FE9F" box="[178,200,286,307]" pageId="13" pageNumber="38" type="male">♂</specimenCount>
|
|
: protarsomeres I-IV strongly dilated; sternites V-VII unmodified; sternite VIII posteriorly triangularly depressed, posterior margin weakly concave; posterior processes of hemitergites IX short, apically without tooth-like processes, not reaching posterior margin of tergite X; sternite IX (
|
|
<figureCitation id="40F2D2A7FFEDFF86ED12FEEE145DFE20" box="[513,588,373,396]" captionStart="Figs 15-23" captionStartId="9.[178,218,1503,1525]" captionTargetBox="[191,1085,212,1496]" captionTargetId="figure@9.[178,1085,212,1497]" captionTargetPageId="9" captionText="Figs 15-23: Othius burmensis: (15) head and pronotum; (16) head; (17) median portion of head; (18) sutural portion of right elytron; (19) male sternite IX; (20) aedeagus in ventral view (internal sac fully extruded); (21) lateral internal structures of aedeagus; (22-23) basal internal structure. Scale bars: 15-16: 1.0 mm; 19-20: 0.5 mm; 17-18, 21: 0.2 mm; 22-23: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3985233" httpUri="https://zenodo.org/record/3985233/files/figure.png" pageId="13" pageNumber="38">Fig. 19</figureCitation>
|
|
) broad, anteriorly deeply bifid, posterior margin strongly concave, postero-lateral angles without spine-like processes; aedeagus (
|
|
<figureCitation id="40F2D2A7FFEDFF86ECFAFE081228FE04" box="[1001,1081,403,425]" captionStart="Figs 15-23" captionStartId="9.[178,218,1503,1525]" captionTargetBox="[191,1085,212,1496]" captionTargetId="figure@9.[178,1085,212,1497]" captionTargetPageId="9" captionText="Figs 15-23: Othius burmensis: (15) head and pronotum; (16) head; (17) median portion of head; (18) sutural portion of right elytron; (19) male sternite IX; (20) aedeagus in ventral view (internal sac fully extruded); (21) lateral internal structures of aedeagus; (22-23) basal internal structure. Scale bars: 15-16: 1.0 mm; 19-20: 0.5 mm; 17-18, 21: 0.2 mm; 22-23: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3985233" httpUri="https://zenodo.org/record/3985233/files/figure.png" pageId="13" pageNumber="38">Figs 20</figureCitation>
|
|
,
|
|
<figureCitation id="40F2D2A7FFEDFF86EFA1FE2B16FFFE6A" box="[178,238,432,454]" captionStart="Figs 24-29" captionStartId="10.[178,218,1314,1336]" captionTargetBox="[178,1085,212,1305]" captionTargetId="figure@10.[178,1085,212,1309]" captionTargetPageId="10" captionText="Figs 24-29: Othius arater (24-26) and O. burmensis (27-29): (24-26) internal structures of aedeagus in ventral view; (27-28) aedeagus in lateral and in ventral view (without extruded internal sac); (29) apical median internal structure of the aedeagus. Scale bars: 27-28: 0.5 mm; 24-26, 29: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3985235" httpUri="https://zenodo.org/record/3985235/files/figure.png" pageId="13" pageNumber="38">27-28</figureCitation>
|
|
)
|
|
<quantity id="1F3163C7FFEDFF86EFEDFE2B175CFE69" box="[254,333,432,453]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="13" pageNumber="38" unit="mm" value="1.4">1.4 mm</quantity>
|
|
long; parameres apically not distinctly dilated, each with 4 apical setae; internal sac (extruded in
|
|
<typeStatus id="07727080FFEDFF86EEAFFE561407FE4F" box="[444,534,461,483]" pageId="13" pageNumber="38" type="holotype">holotype</typeStatus>
|
|
) with six sclerotized internal structures: a short and apically weakly curved median basal structure (
|
|
<figureCitation id="40F2D2A7FFEDFF86EDB7FE701508FDAD" box="[676,793,491,513]" captionStart="Figs 15-23" captionStartId="9.[178,218,1503,1525]" captionTargetBox="[191,1085,212,1496]" captionTargetId="figure@9.[178,1085,212,1497]" captionTargetPageId="9" captionText="Figs 15-23: Othius burmensis: (15) head and pronotum; (16) head; (17) median portion of head; (18) sutural portion of right elytron; (19) male sternite IX; (20) aedeagus in ventral view (internal sac fully extruded); (21) lateral internal structures of aedeagus; (22-23) basal internal structure. Scale bars: 15-16: 1.0 mm; 19-20: 0.5 mm; 17-18, 21: 0.2 mm; 22-23: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3985233" httpUri="https://zenodo.org/record/3985233/files/figure.png" pageId="13" pageNumber="38">Figs 22-23</figureCitation>
|
|
), a distinctly longer median apical structure with a hook-shaped apex (pointing ventrad) (
|
|
<figureCitation id="40F2D2A7FFEDFF86EC3CFD93156CFDB2" box="[815,893,520,542]" captionStart="Figs 24-29" captionStartId="10.[178,218,1314,1336]" captionTargetBox="[178,1085,212,1305]" captionTargetId="figure@10.[178,1085,212,1309]" captionTargetPageId="10" captionText="Figs 24-29: Othius arater (24-26) and O. burmensis (27-29): (24-26) internal structures of aedeagus in ventral view; (27-28) aedeagus in lateral and in ventral view (without extruded internal sac); (29) apical median internal structure of the aedeagus. Scale bars: 27-28: 0.5 mm; 24-26, 29: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3985235" httpUri="https://zenodo.org/record/3985235/files/figure.png" pageId="13" pageNumber="38">Fig. 29</figureCitation>
|
|
), and two pairs of long and slender lateral structures (
|
|
<figureCitation id="40F2D2A7FFEDFF86ED05FDBE1470FD90" box="[534,609,549,572]" captionStart="Figs 15-23" captionStartId="9.[178,218,1503,1525]" captionTargetBox="[191,1085,212,1496]" captionTargetId="figure@9.[178,1085,212,1497]" captionTargetPageId="9" captionText="Figs 15-23: Othius burmensis: (15) head and pronotum; (16) head; (17) median portion of head; (18) sutural portion of right elytron; (19) male sternite IX; (20) aedeagus in ventral view (internal sac fully extruded); (21) lateral internal structures of aedeagus; (22-23) basal internal structure. Scale bars: 15-16: 1.0 mm; 19-20: 0.5 mm; 17-18, 21: 0.2 mm; 22-23: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3985233" httpUri="https://zenodo.org/record/3985233/files/figure.png" pageId="13" pageNumber="38">Fig. 21</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="D876CE22FFEDFF86EFA1FDC81231FCB5" blockId="13.[178,1085,595,793]" pageId="13" pageNumber="38">
|
|
C o m p a r a t i v e n o t e s: The similarly modified sculpture of the elytra (dense micropunctation) and the shared absence of a pair of posterior frontal punctures on the head suggests that
|
|
<taxonomicName id="1FC9B5A1FFEDFF86EE62FD1517E9FD0E" authorityName="Assing" authorityYear="2018" box="[369,504,653,675]" class="Insecta" family="Staphylinidae" genus="Othius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="38" phylum="Arthropoda" rank="species" species="burmensis">
|
|
<emphasis id="EABD1230FFEDFF86EE62FD1517E9FD0E" box="[369,504,653,675]" italics="true" pageId="13" pageNumber="38">O. burmensis</emphasis>
|
|
</taxonomicName>
|
|
is closely related to
|
|
<taxonomicName id="1FC9B5A1FFEDFF86EDD9FD151575FD0F" authorityName="Cameron" authorityYear="1939" box="[714,868,654,675]" class="Insecta" family="Staphylinidae" genus="Othius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="38" phylum="Arthropoda" rank="species" species="opacipennis">
|
|
<emphasis id="EABD1230FFEDFF86EDD9FD151575FD0F" box="[714,868,654,675]" italics="true" pageId="13" pageNumber="38">O. opacipennis</emphasis>
|
|
</taxonomicName>
|
|
CAMERON, 1939 and allied species (
|
|
<taxonomicName id="1FC9B5A1FFEDFF86EE5DFD3017FBFD6C" authorityName="Assing" authorityYear="2003" box="[334,490,683,704]" class="Insecta" family="Staphylinidae" genus="Othius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="38" phylum="Arthropoda" rank="species" species="sericipennis">
|
|
<emphasis id="EABD1230FFEDFF86EE5DFD3017FBFD6C" box="[334,490,683,704]" italics="true" pageId="13" pageNumber="38">O. sericipennis</emphasis>
|
|
</taxonomicName>
|
|
ASSING, 2003;
|
|
<emphasis id="EABD1230FFEDFF86ED81FD30151EFD6C" box="[658,783,683,704]" italics="true" pageId="13" pageNumber="38">O. spoliatus</emphasis>
|
|
<bibRefCitation id="BC58B3D3FFEDFF86EC0BFD3015BBFD6D" author="ASSING V" box="[792,938,683,705]" pageId="13" pageNumber="38" pagination="245 - 263" refId="ref8956" refString="ASSING V. (2008): A revision of Othiini. XVI. Four new species of Othius from the Himalaya and China, and additional records (Coleoptera: Staphylinidae, Staphylininae). - Koleopterologische Rundschau 78: 245 - 263." type="journal article" year="2008">ASSING, 2008</bibRefCitation>
|
|
), all of which are distributed in southwestern
|
|
<collectingCountry id="A0DE8EB2FFEDFF86ED10FD53142EFD72" box="[515,575,712,734]" name="China" pageId="13" pageNumber="38">China</collectingCountry>
|
|
(
|
|
<collectingRegion id="1A0D00C0FFEDFF86ED42FD5314B4FD71" box="[593,677,712,733]" country="China" name="Yunnan" pageId="13" pageNumber="38">Yunnan</collectingRegion>
|
|
,
|
|
<collectingRegion id="1A0D00C0FFEDFF86EDA7FD53151BFD72" box="[692,778,712,734]" country="China" name="Sichuan" pageId="13" pageNumber="38">Sichuan</collectingRegion>
|
|
).
|
|
<taxonomicName id="1FC9B5A1FFEDFF86EC0CFD5315C4FD71" authorityName="Assing" authorityYear="2018" box="[799,981,712,733]" class="Insecta" family="Staphylinidae" genus="Othius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="38" phylum="Arthropoda" rank="species" species="burmensis">
|
|
<emphasis id="EABD1230FFEDFF86EC0CFD5315C4FD71" box="[799,981,712,733]" italics="true" pageId="13" pageNumber="38">Othius burmensis</emphasis>
|
|
</taxonomicName>
|
|
is distinguished from all of them by a less derived morphology of the aedeagus (less elongate and with the full set of internal structures). In addition, it differs from them as follows:
|
|
</paragraph>
|
|
<paragraph id="D876CE22FFEDFF86EFA1FCAB15CBFC17" blockId="13.[178,1085,816,955]" pageId="13" pageNumber="38">
|
|
from
|
|
<taxonomicName id="1FC9B5A1FFEDFF86EFE4FCAB178CFCEA" authorityName="Cameron" authorityYear="1939" box="[247,413,816,838]" class="Insecta" family="Staphylinidae" genus="Othius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="38" phylum="Arthropoda" rank="species" species="opacipennis">
|
|
<emphasis id="EABD1230FFEDFF86EFE4FCAB178CFCEA" box="[247,413,816,838]" italics="true" pageId="13" pageNumber="38">O. opacipennis</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<collectingRegion id="1A0D00C0FFEDFF86EEA4FCAB141AFCE9" box="[439,523,816,837]" country="China" name="Yunnan" pageId="13" pageNumber="38">Yunnan</collectingRegion>
|
|
,
|
|
<collectingRegion id="1A0D00C0FFEDFF86ED30FCAB1467FCEA" box="[547,630,816,838]" country="China" name="Sichuan" pageId="13" pageNumber="38">Sichuan</collectingRegion>
|
|
) by distinctly bicoloured antennae (
|
|
<taxonomicName id="1FC9B5A1FFEDFF86EB37FCAB173DFCCF" authorityName="Cameron" authorityYear="1939" class="Insecta" family="Staphylinidae" genus="Othius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="38" phylum="Arthropoda" rank="species" species="opacipennis">
|
|
<emphasis id="EABD1230FFEDFF86EB37FCAB173DFCCF" italics="true" pageId="13" pageNumber="38">O. opacipennis</emphasis>
|
|
</taxonomicName>
|
|
: antennae blackish-brown to black), a posteriorly dilated and less oblong head, subparallel lateral margins of the pronotum (
|
|
<taxonomicName id="1FC9B5A1FFEDFF86EDDCFCF0157AFC2C" authorityName="Cameron" authorityYear="1939" box="[719,875,875,896]" class="Insecta" family="Staphylinidae" genus="Othius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="38" phylum="Arthropoda" rank="species" species="opacipennis">
|
|
<emphasis id="EABD1230FFEDFF86EDDCFCF0157AFC2C" box="[719,875,875,896]" italics="true" pageId="13" pageNumber="38">O. opacipennis</emphasis>
|
|
</taxonomicName>
|
|
: pronotum tapering posteriorly), and the shape of the male sternite IX (
|
|
<taxonomicName id="1FC9B5A1FFEDFF86EDCFFC131568FC32" authorityName="Cameron" authorityYear="1939" box="[732,889,904,926]" class="Insecta" family="Staphylinidae" genus="Othius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="38" phylum="Arthropoda" rank="species" species="opacipennis">
|
|
<emphasis id="EABD1230FFEDFF86EDCFFC131568FC32" box="[732,889,904,926]" italics="true" pageId="13" pageNumber="38">O. opacipennis</emphasis>
|
|
</taxonomicName>
|
|
: sternite IX much longer and more slender, anteriorly less deeply bifid, and posteriorly bisinuate);
|
|
</paragraph>
|
|
<paragraph id="D876CE22FFEDFF86EFA1FC4815FAFBEC" blockId="13.[178,1085,979,1089]" pageId="13" pageNumber="38">
|
|
from
|
|
<taxonomicName id="1FC9B5A1FFEDFF86EFF8FC481794FC44" authorityName="Assing" authorityYear="2003" box="[235,389,979,1000]" class="Insecta" family="Staphylinidae" genus="Othius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="38" phylum="Arthropoda" rank="species" species="sericipennis">
|
|
<emphasis id="EABD1230FFEDFF86EFF8FC481794FC44" box="[235,389,979,1000]" italics="true" pageId="13" pageNumber="38">O. sericipennis</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<collectingRegion id="1A0D00C0FFEDFF86EE87FC4817F9FC44" box="[404,488,979,1000]" country="China" name="Yunnan" pageId="13" pageNumber="38">Yunnan</collectingRegion>
|
|
,
|
|
<collectingRegion id="1A0D00C0FFEDFF86EEE0FC481457FC45" box="[499,582,979,1001]" country="China" name="Sichuan" pageId="13" pageNumber="38">Sichuan</collectingRegion>
|
|
) by a posteriorly dilated, much broader, and less oblong head, subparallel lateral margins of the pronotum (
|
|
<taxonomicName id="1FC9B5A1FFEDFF86EC3AFC6B15D6FBAA" authorityName="Assing" authorityYear="2003" box="[809,967,1008,1030]" class="Insecta" family="Staphylinidae" genus="Othius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="38" phylum="Arthropoda" rank="species" species="sericipennis">
|
|
<emphasis id="EABD1230FFEDFF86EC3AFC6B15D6FBAA" box="[809,967,1008,1030]" italics="true" pageId="13" pageNumber="38">O. sericipennis</emphasis>
|
|
</taxonomicName>
|
|
: pronotum tapering posteriorly), and the shape of the male sternite IX (
|
|
<taxonomicName id="1FC9B5A1FFEDFF86EC36FB9515AEFB8F" authorityName="Assing" authorityYear="2003" box="[805,959,1038,1059]" class="Insecta" family="Staphylinidae" genus="Othius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="38" phylum="Arthropoda" rank="species" species="sericipennis">
|
|
<emphasis id="EABD1230FFEDFF86EC36FB9515AEFB8F" box="[805,959,1038,1059]" italics="true" pageId="13" pageNumber="38">O. sericipennis</emphasis>
|
|
</taxonomicName>
|
|
: sternite IX much longer, more slender, anteriorly not bifid, and posteriorly weakly concave);
|
|
</paragraph>
|
|
<paragraph id="D876CE22FFEDFF86EFA1FBC31707FAB2" blockId="13.[178,1085,1112,1310]" pageId="13" pageNumber="38">
|
|
from
|
|
<emphasis id="EABD1230FFEDFF86EFFDFBC3177CFBC1" box="[238,365,1112,1133]" italics="true" pageId="13" pageNumber="38">O. spoliatus</emphasis>
|
|
(
|
|
<collectingRegion id="1A0D00C0FFEDFF86EE6DFBC317C0FBC2" box="[382,465,1112,1134]" country="China" name="Yunnan" pageId="13" pageNumber="38">Yunnan</collectingRegion>
|
|
) by distinctly bicoloured antennae (
|
|
<emphasis id="EABD1230FFEDFF86EC47FBC315C3FBC1" box="[852,978,1112,1133]" italics="true" pageId="13" pageNumber="38">O. spoliatus</emphasis>
|
|
: antennomeres IV-X dark-brown), less transverse preapical antennomeres, a less broad and posteriorly less strongly dilated head, much longer elytra, the presence of hind wings (reduced in
|
|
<emphasis id="EABD1230FFEDFF86EE21FB2B17A1FB69" box="[306,432,1200,1221]" italics="true" pageId="13" pageNumber="38">O. spoliatus</emphasis>
|
|
), the presence of a palisade fringe at the posterior margin of tergite VII (absent in
|
|
<emphasis id="EABD1230FFEDFF86EE87FB551401FB4E" box="[404,528,1229,1251]" italics="true" pageId="13" pageNumber="38">O. spoliatus</emphasis>
|
|
), and the shape of the male sternite IX (
|
|
<emphasis id="EABD1230FFEDFF86ECA9FB551227FB4E" box="[954,1078,1229,1251]" italics="true" pageId="13" pageNumber="38">O. spoliatus</emphasis>
|
|
: sternite IX much longer, more slender, and posteriorly nearly truncate with median excision).
|
|
</paragraph>
|
|
<paragraph id="D876CE22FFEDFF86EFA1FAAD1751FAC5" blockId="13.[178,1085,1333,1385]" pageId="13" pageNumber="38">
|
|
For illustrations of
|
|
<taxonomicName id="1FC9B5A1FFEDFF86EE64FAAD1400FAE7" authorityName="Cameron" authorityYear="1939" box="[375,529,1334,1355]" class="Insecta" family="Staphylinidae" genus="Othius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="38" phylum="Arthropoda" rank="species" species="opacipennis">
|
|
<emphasis id="EABD1230FFEDFF86EE64FAAD1400FAE7" box="[375,529,1334,1355]" italics="true" pageId="13" pageNumber="38">O. opacipennis</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="1FC9B5A1FFEDFF86ED0CFAAD14A8FAE7" authorityName="Assing" authorityYear="2003" box="[543,697,1334,1355]" class="Insecta" family="Staphylinidae" genus="Othius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="38" phylum="Arthropoda" rank="species" species="sericipennis">
|
|
<emphasis id="EABD1230FFEDFF86ED0CFAAD14A8FAE7" box="[543,697,1334,1355]" italics="true" pageId="13" pageNumber="38">O. sericipennis</emphasis>
|
|
</taxonomicName>
|
|
, and
|
|
<emphasis id="EABD1230FFEDFF86EDE1FAAD157CFAE6" box="[754,877,1333,1355]" italics="true" pageId="13" pageNumber="38">O. spoliatus</emphasis>
|
|
see
|
|
<bibRefCitation id="BC58B3D3FFEDFF86EC8FFAAD1229FAE7" author="ASSING V" box="[924,1080,1333,1355]" pageId="13" pageNumber="38" pagination="661 - 691" refId="ref8725" refString="ASSING V. (1999 b): A revision of Othius STEPHENS (Coleoptera, Staphylinidae). VIII. Further records, new species, and a new synonym. - Linzer Biologische B eitrage 31 (2): 661 - 691." type="journal article" year="1999">ASSING (1999b</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="BC58B3D3FFEDFF86EFA1FAC816E5FAC5" author="ASSING V" box="[178,244,1363,1385]" pageId="13" pageNumber="38" pagination="69 - 88" refId="ref8863" refString="ASSING V. (2003 b): A revision of Othiini. XIII. Horizontal and vertical distribution of Othius, new species, and additional records (Coleoptera: Staphylinidae: Staphylininae). - Entomological Problems 33 (1 - 2): 69 - 88." type="journal article" year="2003">2003b</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="BC58B3D3FFEDFF86EFEDFAC81724FAC5" author="ASSING V" box="[254,309,1363,1385]" pageId="13" pageNumber="38" pagination="245 - 263" refId="ref8956" refString="ASSING V. (2008): A revision of Othiini. XVI. Four new species of Othius from the Himalaya and China, and additional records (Coleoptera: Staphylinidae, Staphylininae). - Koleopterologische Rundschau 78: 245 - 263." type="journal article" year="2008">2008</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="90D39DA9FFEDFF86EFA1FA1B17BBFA41" pageId="13" pageNumber="38" type="distribution">
|
|
<paragraph id="D876CE22FFEDFF86EFA1FA1B17BBFA41" blockId="13.[178,1085,1408,1518]" pageId="13" pageNumber="38">
|
|
D i s t r i b u t i o n a n d n a t u r a l h i s t o r y:
|
|
<taxonomicName id="1FC9B5A1FFEDFF86EC14FA1B15AEFA39" authorityName="Assing" authorityYear="2018" box="[775,959,1408,1429]" class="Insecta" family="Staphylinidae" genus="Othius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="38" phylum="Arthropoda" rank="species" species="burmensis">
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<emphasis id="EABD1230FFEDFF86EC14FA1B15AEFA39" box="[775,959,1408,1429]" italics="true" pageId="13" pageNumber="38">Othius burmensis</emphasis>
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</taxonomicName>
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is the first representative of the genus to be recorded from
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<collectingCountry id="A0DE8EB2FFEDFF86ED89FA051511FA1F" box="[666,768,1438,1459]" name="Myanmar" pageId="13" pageNumber="38">Myanmar</collectingCountry>
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. The
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<typeStatus id="07727080FFEDFF86EC2BFA051575FA1F" box="[824,868,1438,1459]" pageId="13" pageNumber="38">type</typeStatus>
|
|
locality is situated in the northeast of the country, not far from the border with the Chinese province Yunnan, at an altitude of
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|
<quantity id="1F3163C7FFEDFF86EE44FA4317B7FA41" box="[343,422,1496,1518]" metricMagnitude="3" metricUnit="m" metricValue="2.255" pageId="13" pageNumber="38" unit="m" value="2255.0">2255 m</quantity>
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|
.
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</paragraph>
|
|
</subSubSection>
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|
</treatment>
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|
</document> |