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<document id="A5531DD631B0E352125153797AF72D6F" ID-DOI="10.5281/zenodo.196847" ID-GBIF-Dataset="5e5a206b-9cfe-4bde-bc1d-4de509a3a4e9" ID-ISSN="1175-5326" ID-Zenodo-Dep="196847" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460254280575" checkinUser="plazi" docAuthor="Pitts, James P., Wilson, Joseph S., Williams, Kevin A. &amp; Boehme, Nicole F." docDate="2010" docId="504687FBC749FFB5FF65F9AB3A46F925" docLanguage="en" docName="zt02553p034.pdf" docOrigin="Zootaxa 2553" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Sphaeropthalma fergusoni Pitts, NEW SPECIES" docType="treatment" docVersion="10" lastPageNumber="23" masterDocId="AC7FFF83C75DFFA3FFF2FFF4397FFF89" masterDocTitle="Nocturnal velvet ant males (Hymenoptera: Mutillidae) of Deep Canyon, California including four new species and a fifth new species from Owens Lake Valley, California" masterLastPageNumber="34" masterPageNumber="1" pageNumber="21" updateTime="1698243197414" updateUser="plazi">
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<mods:title id="694EED9C50188B5C2180EB21DA340AFA">Nocturnal velvet ant males (Hymenoptera: Mutillidae) of Deep Canyon, California including four new species and a fifth new species from Owens Lake Valley, California</mods:title>
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<mods:namePart id="EE9DA97C0E3E1B353A2DBB8739D5E91C">Pitts, James P.</mods:namePart>
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<mods:namePart id="4D7623F871D0A77996F81ADC38A2C3CF">Wilson, Joseph S.</mods:namePart>
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<mods:namePart id="D79EDA89CBA5168A42FCB99DE3D34C23">Williams, Kevin A.</mods:namePart>
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<mods:namePart id="AB47AB7B1285E7F4260D017FA2BAF7F0">Boehme, Nicole F.</mods:namePart>
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<mods:date id="7A9FB894525101DF024C18CACDFECED8">2010</mods:date>
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<mods:number id="F6AB7456A8201BF0BEA906D3E74EF01A">2553</mods:number>
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<treatment id="504687FBC749FFB5FF65F9AB3A46F925" ID-DOI="http://doi.org/10.5281/zenodo.5670792" ID-GBIF-Taxon="119392240" ID-Zenodo-Dep="5670792" LSID="urn:lsid:plazi:treatment:504687FBC749FFB5FF65F9AB3A46F925" httpUri="http://treatment.plazi.org/id/504687FBC749FFB5FF65F9AB3A46F925" lastPageId="22" lastPageNumber="23" pageId="20" pageNumber="21">
<subSubSection id="90F56566C749FFB7FF65F9AB3B8FF9F0" box="[151,752,1631,1657]" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="D85036EDC749FFB7FF65F9AB3B8FF9F0" blockId="20.[151,752,1631,1657]" box="[151,752,1631,1657]" pageId="20" pageNumber="21">
<heading id="83188181C749FFB7FF65F9AB3B8FF9F0" bold="true" box="[151,752,1631,1657]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
<emphasis id="EA9BEAFFC749FFB7FF65F9AB3B8FF9F0" bold="true" box="[151,752,1631,1657]" pageId="20" pageNumber="21">
<taxonomicName id="1FEF4D6EC749FFB7FF65F9AB3B63F9F0" authority="Pitts" authorityName="Pitts" box="[151,540,1631,1657]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="fergusoni" status="sp. nov.">
<emphasis id="EA9BEAFFC749FFB7FF65F9AB38A5F9F0" bold="true" box="[151,474,1631,1657]" italics="true" pageId="20" pageNumber="21">Sphaeropthalma fergusoni</emphasis>
Pitts
</taxonomicName>
,
<taxonomicNameLabel id="F1A85784C749FFB7FDDBF9AB3B8FF9F0" box="[553,752,1631,1657]" pageId="20" pageNumber="21" rank="species">NEW SPECIES</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="90F56566C749FFB7FF65F9533C4CF885" pageId="20" pageNumber="21" type="diagnosis">
<paragraph id="D85036EDC749FFB7FF65F9533C4CF885" blockId="20.[151,1437,1703,1991]" pageId="20" pageNumber="21">
<emphasis id="EA9BEAFFC749FFB7FF65F9533804F948" bold="true" box="[151,379,1703,1729]" pageId="20" pageNumber="21">Diagnosis of male.</emphasis>
This species is can be recognized by its quadrate head, the thickened apex of the clypeus (
<figureCitation id="40D42A68C749FFB7FF6DF9393989F96E" box="[159,246,1741,1767]" captionStart="FIGURES 38 43" captionStartId="16.[151,273,1981,2005]" captionTargetBox="[189,1401,194,1939]" captionTargetId="figure@16.[189,1402,194,1950]" captionTargetPageId="16" captionText="FIGURES 38 43. Sphaeropthalma chandleri, sp. nov.: 38. Head, anterior view; and 39. Habitus. Sphaeropthalma fergusoni, sp. nov.: 40. Pygidium, dorsal view; 41. Head anterior view; 42. Head, dorsal view; and 43. Habitus." httpUri="https://zenodo.org/record/196852/files/figure.png" pageId="20" pageNumber="21">Fig. 41</figureCitation>
), the mid coxa with a median denticle present on the inner margin and diagnostic genitalia with which it shares features only with
<taxonomicName id="1FEF4D6EC749FFB7FE28F9073B18F885" box="[474,615,1779,1804]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="arnalduri" status="sp. nov.">
<emphasis id="EA9BEAFFC749FFB7FE28F9073B18F885" box="[474,615,1779,1804]" italics="true" pageId="20" pageNumber="21">S. arnalduri</emphasis>
</taxonomicName>
,
<emphasis id="EA9BEAFFC749FFB7FD84F9063BABF885" bold="true" box="[630,724,1778,1804]" pageId="20" pageNumber="21">
<taxonomicNameLabel id="F1A85784C749FFB7FD84F9063BABF885" box="[630,724,1778,1804]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(Fig. 65), while lacking any mesosternal processes.
</paragraph>
</subSubSection>
<subSubSection id="90F56566C749FFB5FF37F8E338BFFC4B" lastPageId="22" lastPageNumber="23" pageId="20" pageNumber="21" type="description">
<paragraph id="D85036EDC749FFB7FF37F8E33AFDF84E" blockId="20.[151,1437,1703,1991]" pageId="20" pageNumber="21">
<emphasis id="EA9BEAFFC749FFB7FF37F8E338AFF8B8" bold="true" box="[197,464,1815,1841]" pageId="20" pageNumber="21">Description of male.</emphasis>
<emphasis id="EA9BEAFFC749FFB7FE2BF8EC3B1EF8B8" box="[473,609,1816,1841]" italics="true" pageId="20" pageNumber="21">Coloration</emphasis>
(
<figureCitation id="40D42A68C749FFB7FD80F8EC3A7FF8BB" box="[626,768,1816,1842]" captionStart="FIGURES 38 43" captionStartId="16.[151,273,1981,2005]" captionTargetBox="[189,1401,194,1939]" captionTargetId="figure@16.[189,1402,194,1950]" captionTargetPageId="16" captionText="FIGURES 38 43. Sphaeropthalma chandleri, sp. nov.: 38. Head, anterior view; and 39. Habitus. Sphaeropthalma fergusoni, sp. nov.: 40. Pygidium, dorsal view; 41. Head anterior view; 42. Head, dorsal view; and 43. Habitus." httpUri="https://zenodo.org/record/196852/files/figure.png" pageId="20" pageNumber="21">Figs 4143</figureCitation>
). Body testaceous; flagellum and legs stramineous to concolorous with body, sometime femur infuscated; metasoma varies from stramineous to testaceous. Body clothed with sparse, erect, brachyplumose, whitish setae. T1 with sparse plumose fringe at distal margin. T2 and S2 with sparse fringes of whitish plumose setae. T35 and S35 each with sparser, less conspicuous fringe of whitish plumose setae. Setae somewhat yellow tinged.
</paragraph>
<paragraph id="D85036EDC748FFB6FF65F95C3B4FF8D6" blockId="21.[151,1436,1704,1887]" pageId="21" pageNumber="22">
<emphasis id="EA9BEAFFC748FFB6FF65F95C3862F949" bold="true" box="[151,285,1704,1728]" pageId="21" pageNumber="22">Figs. 59-67:</emphasis>
<taxonomicName id="1FEF4D6EC748FFB6FED7F95C3B0FF936" box="[293,624,1704,1727]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="megagnathos">Sphaeropthalma megagnathos</taxonomicName>
: 59. Genitalia, ventral, dorsal, inner lateral view, and lateral view of penis valve (from
<bibRefCitation id="BC7E4B1CC748FFB6FED2F93C38EAF956" author="Pitts" box="[288,405,1736,1759]" pageId="21" pageNumber="34" refString="Pitts, J. P. (2006) Review of the Sphaeropthalma imperialis species-group (Hymenoptera: Mutillidae), with descriptions of females and taxonomic notes. Zootaxa, 1248, 1 - 20." type="journal article" year="2006">Pitts 2006</bibRefCitation>
).
<taxonomicName id="1FEF4D6EC748FFB6FE55F93C3BBDF956" box="[423,706,1736,1759]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="mendica">Sphaeropthalma mendica</taxonomicName>
: 60. Genitalia, dorsal, inner lateral view, and lateral view of penis valve (from Wilson &amp; Pitts 2009).
<taxonomicName id="1FEF4D6EC748FFB6FDE6F91C3A66F976" box="[532,793,1768,1791]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="pallida">Sphaeropthalma pallida</taxonomicName>
: 61. Genitalia, lateral view.
<taxonomicName id="1FEF4D6EC748FFB6FBBEF91C3C11F976" box="[1100,1390,1768,1791]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="sublobata">Sphaeropthalma sublobata</taxonomicName>
; 62. Genitalia, lateral view. Figs. 63-67, Genitalia, ventral view left, dorsal view right: 63.
<taxonomicName id="1FEF4D6EC748FFB6FBB3F8FC3C10F896" box="[1089,1391,1800,1823]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="tetracuspis">Sphaeropthalma tetracuspis</taxonomicName>
; 64.
<taxonomicName id="1FEF4D6EC748FFB6FF65F8DC38C5F8B6" box="[151,442,1832,1855]" class="Insecta" family="Formicidae" genus="Odontophotopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="hammetti" status="sp. nov.">Odontophotopsis hammetti</taxonomicName>
,
<taxonomicNameLabel id="F1A85784C748FFB6FE37F8DC3B65F8B6" box="[453,538,1832,1855]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
; 65.
<taxonomicName id="1FEF4D6EC748FFB6FDB9F8DC3A1DF8B6" box="[587,866,1832,1855]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="arnalduri" status="sp. nov.">Sphaeropthalma arnalduri</taxonomicName>
,
<taxonomicNameLabel id="F1A85784C748FFB6FC9EF8DC3ABEF8B6" box="[876,961,1832,1855]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
; 66.
<taxonomicName id="1FEF4D6EC748FFB6FC01F8DC3C73F8B6" box="[1011,1292,1832,1855]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="chandleri" status="sp. nov.">Sphaeropthalma chandleri</taxonomicName>
,
<taxonomicNameLabel id="F1A85784C748FFB6FAE5F8DC3C13F8B6" box="[1303,1388,1832,1855]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
; and 67.
<taxonomicName id="1FEF4D6EC748FFB6FF4DF8BC38ACF8D6" box="[191,467,1864,1887]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="mankelli" status="sp. nov.">Sphaeropthalma mankelli</taxonomicName>
,
<taxonomicNameLabel id="F1A85784C748FFB6FE2FF8BC3B4FF8D6" box="[477,560,1864,1887]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="D85036EDC748FFB5FF37F85A3821FECE" blockId="21.[151,1437,1966,2029]" lastBlockId="22.[151,1438,152,1708]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<emphasis id="EA9BEAFFC748FFB6FF37F85A3870F84E" box="[197,271,1966,1991]" italics="true" pageId="21" pageNumber="22">Head.</emphasis>
Head distinctly quadrate posteriorly. Mandible (
<figureCitation id="40D42A68C748FFB6FC96F85A3AC1F841" box="[868,958,1966,1992]" captionStart="FIGURES 38 43" captionStartId="16.[151,273,1981,2005]" captionTargetBox="[189,1401,194,1939]" captionTargetId="figure@16.[189,1402,194,1950]" captionTargetPageId="16" captionText="FIGURES 38 43. Sphaeropthalma chandleri, sp. nov.: 38. Head, anterior view; and 39. Habitus. Sphaeropthalma fergusoni, sp. nov.: 40. Pygidium, dorsal view; 41. Head anterior view; 42. Head, dorsal view; and 43. Habitus." httpUri="https://zenodo.org/record/196852/files/figure.png" pageId="21" pageNumber="22">Fig. 41</figureCitation>
) tridentate, with weak rounded ventral tooth, lacking excision apical to tooth; dorsal carina incomplete forming slight dorsal lamella towards termination; apex oblique; mandible appearing tapered beyond excision in strict frontal view. Clypeus depressed slightly below margin of mandible, median area concave; surface of clypeus polished, almost impunctate, with few erect setae; apex truncate and distinctly thickened (
<figureCitation id="40D42A68C74BFFB5FC15FF163D41FF75" box="[999,1086,226,252]" captionStart="FIGURES 38 43" captionStartId="16.[151,273,1981,2005]" captionTargetBox="[189,1401,194,1939]" captionTargetId="figure@16.[189,1402,194,1950]" captionTargetPageId="16" captionText="FIGURES 38 43. Sphaeropthalma chandleri, sp. nov.: 38. Head, anterior view; and 39. Habitus. Sphaeropthalma fergusoni, sp. nov.: 40. Pygidium, dorsal view; 41. Head anterior view; 42. Head, dorsal view; and 43. Habitus." httpUri="https://zenodo.org/record/196852/files/figure.png" pageId="22" pageNumber="23">Fig. 41</figureCitation>
). F1 approximately
<date id="AC51102DC74BFFB5FADAFF163C33FF75" box="[1320,1356,226,252]" pageId="22" pageNumber="23">1X</date>
length of F2. Ocelli moderate in size, ocellocular distance 1.11.2X greatest width of lateral ocellus (
<figureCitation id="40D42A68C74BFFB5FB02FEFC3C35FEAB" box="[1264,1354,264,290]" captionStart="FIGURES 38 43" captionStartId="16.[151,273,1981,2005]" captionTargetBox="[189,1401,194,1939]" captionTargetId="figure@16.[189,1402,194,1950]" captionTargetPageId="16" captionText="FIGURES 38 43. Sphaeropthalma chandleri, sp. nov.: 38. Head, anterior view; and 39. Habitus. Sphaeropthalma fergusoni, sp. nov.: 40. Pygidium, dorsal view; 41. Head anterior view; 42. Head, dorsal view; and 43. Habitus." httpUri="https://zenodo.org/record/196852/files/figure.png" pageId="22" pageNumber="23">Fig. 42</figureCitation>
). Head weakly punctate.
</paragraph>
<paragraph id="D85036EDC74BFFB5FF37FEA63A9DFD1E" blockId="22.[151,1438,152,1708]" pageId="22" pageNumber="23">
<taxonomicName id="1FEF4D6EC74BFFB5FF37FEA6383CFEE2" box="[197,323,338,363]" class="Crustacea" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="EA9BEAFFC74BFFB5FF37FEA6383CFEE2" box="[197,323,338,363]" italics="true" pageId="22" pageNumber="23">Mesosoma</emphasis>
</taxonomicName>
. Sides and dorsum of pronotum coarsely punctate, dorsum with moderate, shallow punctures, sides with somewhat larger, shallower punctures. Mesonotum with weak, separated, shallow punctures. Notaulus distinct, complete. Scutellum coarsely, confluently punctate. Axillae not projecting posteriorly. Dorsum and posterior face of propodeum conspicuously, shallow reticulate, reticulations extending on to sides of propodeum, either remaining reticulate laterally or becoming coarse, punctate-reticulate. Anterolateral area of mesopleuron mostly glabrous; remainder of mesopleuron with deeper, contiguous to confluent punctures; interstitial areas sometimes micropunctate. Metapleuron polished. Mesosternal processes absent. Metasternum bidentate. Mid femur not swollen. Mid coxa with distinct to indistinct denticle located medially on inner margin. Marginal cell on costa long, 1.75
<date id="AC51102DC74BFFB5FD02FD893A6BFD1E" box="[752,788,637,663]" pageId="22" pageNumber="23">2X</date>
length of stigma.
</paragraph>
<paragraph id="D85036EDC74BFFB5FF37FD5638C0FCFE" blockId="22.[151,1438,152,1708]" pageId="22" pageNumber="23">
<emphasis id="EA9BEAFFC74BFFB5FF37FD56383BFD32" box="[197,324,674,699]" italics="true" pageId="22" pageNumber="23">Metasoma</emphasis>
. First metasomal segment petiolate. Pygidium shortened, quadrate, polished, not strongly margined; S2 with sternal felt line 0.30.5X length of tergal felt line. Hypopygium quadrate. Posterior margin slightly dentate medially. Genitalia with paramere acicular, only extreme apex curving dorsally; cuspis elongate,&gt;0.9X free length of paramere, and cylindrical basally, dorsal face with central longitudinal swelling, apex spatulate, ventral face slightly concave, with basal pit, inner margin with shallow ventral notch anterior to spatulate area.
</paragraph>
<paragraph id="D85036EDC74BFFB5FF37FC7638D8FC15" blockId="22.[151,1438,152,1708]" box="[197,423,898,924]" pageId="22" pageNumber="23">
<emphasis id="EA9BEAFFC74BFFB5FF37FC76385AFC12" box="[197,293,898,923]" italics="true" pageId="22" pageNumber="23">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="D85036EDC74BFFB5FF37FC5C38BFFC4B" blockId="22.[151,1438,152,1708]" box="[197,448,936,962]" pageId="22" pageNumber="23">
<emphasis id="EA9BEAFFC74BFFB5FF37FC5C3861FC48" box="[197,286,936,961]" italics="true" pageId="22" pageNumber="23">Length.</emphasis>
11.5
<quantity id="1F179B08C74BFFB5FE9AFC5C38C3FC4B" box="[360,444,936,962]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="22" pageNumber="23" unit="mm" value="13.0">13 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="90F56566C74BFFB5FF37FC393892FB4E" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="D85036EDC74BFFB5FF37FC393892FB4E" blockId="22.[151,1438,152,1708]" pageId="22" pageNumber="23">
<emphasis id="EA9BEAFFC74BFFB5FF37FC3938C4FC6E" bold="true" box="[197,443,973,999]" pageId="22" pageNumber="23">Material examined.</emphasis>
<typeStatus id="0754884FC74BFFB5FE36FC393B48FC6E" box="[452,567,973,999]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
:
<emphasis id="EA9BEAFFC74BFFB5FDB0FC393BBBFC6E" bold="true" box="[578,708,973,999]" pageId="22" pageNumber="23">California</emphasis>
,
<emphasis id="EA9BEAFFC74BFFB5FD21FC393A04FC6F" box="[723,891,973,998]" italics="true" pageId="22" pageNumber="23">Riverside Co.,</emphasis>
Corn Springs,
<date id="AC51102DC74BFFB5FBC3FC393DBCFC6E" box="[1073,1219,973,999]" pageId="22" pageNumber="23" value="2004-06-24">24.Jun.2004</date>
, coll. K. Williams (
<collectionCode id="BEFEAE28C74BFFB5FF6DFC063987FB85" box="[159,248,1010,1036]" pageId="22" pageNumber="23">EMUS</collectionCode>
).
<typeStatus id="0754884FC74BFFB5FEFEFC0638FBFB85" box="[268,388,1010,1036]" pageId="22" pageNumber="23" type="paratype">Paratypes</typeStatus>
:
<emphasis id="EA9BEAFFC74BFFB5FE7DFC063B6EFB85" bold="true" box="[399,529,1010,1036]" pageId="22" pageNumber="23">California</emphasis>
,
<emphasis id="EA9BEAFFC74BFFB5FDD3FC063BB5FB82" box="[545,714,1010,1035]" italics="true" pageId="22" pageNumber="23">Riverside Co.,</emphasis>
Deep Canyon:
<specimenCount id="CEE9FD64C74BFFB5FC77FC063AA7FB85" box="[901,984,1010,1036]" pageId="22" pageNumber="23" type="male">1 male</specimenCount>
, 2.
<date id="AC51102DC74BFFB5FC0AFC063DEEFB85" box="[1016,1169,1010,1036]" pageId="22" pageNumber="23" value="1936-05-01">
May.1936,
<specimenCount id="CEE9FD64C74BFFB5FB70FC063DEEFB85" box="[1154,1169,1010,1036]" pageId="22" pageNumber="23" type="male">1</specimenCount>
</date>
male, 15.
<date id="AC51102DC74BFFB5FAF0FC063CE3FB85" box="[1282,1436,1010,1036]" pageId="22" pageNumber="23" value="1969-05-03">
May.1969,
<specimenCount id="CEE9FD64C74BFFB5FA7FFC063CE3FB85" box="[1421,1436,1010,1036]" pageId="22" pageNumber="23" type="male">3</specimenCount>
</date>
males, 16.
<date id="AC51102DC74BFFB5FEE1FBEC38CAFBBB" box="[275,437,1048,1074]" pageId="22" pageNumber="23" value="1963-05-01">
May.1963,
<specimenCount id="CEE9FD64C74BFFB5FE57FBEC38CAFBBB" box="[421,437,1048,1074]" pageId="22" pageNumber="23" type="male">1</specimenCount>
</date>
male, 19.
<date id="AC51102DC74BFFB5FDDFFBEC3BB0FBBB" box="[557,719,1048,1074]" pageId="22" pageNumber="23" value="1973-05-02">
May.1973,
<specimenCount id="CEE9FD64C74BFFB5FD4DFBEC3BB0FBBB" box="[703,719,1048,1074]" pageId="22" pageNumber="23" type="male">2</specimenCount>
</date>
males, 613.
<date id="AC51102DC74BFFB5FC87FBEC3D76FBBB" box="[885,1033,1048,1074]" pageId="22" pageNumber="23" value="1969-06-01">
Jun.1969,
<specimenCount id="CEE9FD64C74BFFB5FC0BFBEC3D76FBBB" box="[1017,1033,1048,1074]" pageId="22" pageNumber="23" type="male">1</specimenCount>
</date>
male, 13.
<date id="AC51102DC74BFFB5FB70FBEC3C67FBBB" box="[1154,1304,1048,1074]" pageId="22" pageNumber="23" value="1963-06-01">
Jun.1963,
<specimenCount id="CEE9FD64C74BFFB5FAFBFBEC3C67FBBB" box="[1289,1304,1048,1074]" pageId="22" pageNumber="23" type="male">1</specimenCount>
</date>
male, 20 24.
<date id="AC51102DC74BFFB5FF48FBC9383CFBDE" box="[186,323,1085,1111]" pageId="22" pageNumber="23" value="1969-06-02">
Jun.1969,
<specimenCount id="CEE9FD64C74BFFB5FEC6FBC9383CFBDE" box="[308,323,1085,1111]" pageId="22" pageNumber="23" type="male">2</specimenCount>
</date>
males, 2426.
<date id="AC51102DC74BFFB5FE1CFBC93B0AFBDE" box="[494,629,1085,1111]" pageId="22" pageNumber="23" value="1969-06-01">
Jun.1969,
<specimenCount id="CEE9FD64C74BFFB5FD94FBC93B0AFBDE" box="[614,629,1085,1111]" pageId="22" pageNumber="23" type="male">1</specimenCount>
</date>
male, 1.2.
<date id="AC51102DC74BFFB5FD1FFBC93A10FBDE" box="[749,879,1085,1111]" pageId="22" pageNumber="23" value="1969-07-01">
Jul.1969,
<specimenCount id="CEE9FD64C74BFFB5FC92FBC93A10FBDE" box="[864,879,1085,1111]" pageId="22" pageNumber="23" type="male">1</specimenCount>
</date>
male, 37.
<date id="AC51102DC74BFFB5FC1CFBC93D0EFBDE" box="[1006,1137,1085,1111]" pageId="22" pageNumber="23" value="1969-07-03">
Jul.1969,
<specimenCount id="CEE9FD64C74BFFB5FB91FBC93D0EFBDE" box="[1123,1137,1085,1111]" pageId="22" pageNumber="23" type="male">3</specimenCount>
</date>
males, 1012.
<date id="AC51102DC74BFFB5FAE8FBC93CE3FBDE" box="[1306,1436,1085,1111]" pageId="22" pageNumber="23" value="1969-07-02">
Jul.1969,
<specimenCount id="CEE9FD64C74BFFB5FA7FFBC93CE3FBDE" box="[1421,1436,1085,1111]" pageId="22" pageNumber="23" type="male">2</specimenCount>
</date>
males, 1921.
<date id="AC51102DC74BFFB5FEC8FB9638C0FBF5" box="[314,447,1122,1148]" pageId="22" pageNumber="23" value="1969-07-01">
Jul.1969,
<specimenCount id="CEE9FD64C74BFFB5FE42FB9638C0FBF5" box="[432,447,1122,1148]" pageId="22" pageNumber="23" type="male">1</specimenCount>
</date>
male, 2324.
<date id="AC51102DC74BFFB5FD90FB963B9BFBF5" box="[610,740,1122,1148]" pageId="22" pageNumber="23" value="1969-07-01">
Jul.1969,
<specimenCount id="CEE9FD64C74BFFB5FD27FB963B9BFBF5" box="[725,740,1122,1148]" pageId="22" pageNumber="23" type="male">1</specimenCount>
</date>
male, 2628.
<date id="AC51102DC74BFFB5FC75FB963D76FBF5" box="[903,1033,1122,1148]" pageId="22" pageNumber="23" value="1969-07-01">
Jul.1969,
<specimenCount id="CEE9FD64C74BFFB5FC08FB963D76FBF5" box="[1018,1033,1122,1148]" pageId="22" pageNumber="23" type="male">1</specimenCount>
</date>
male, 3031.
<date id="AC51102DC74BFFB5FB5BFB963C52FBF5" box="[1193,1325,1122,1148]" pageId="22" pageNumber="23" value="2007-07-01">
Jul.2007,
<specimenCount id="CEE9FD64C74BFFB5FAECFB963C52FBF5" box="[1310,1325,1122,1148]" pageId="22" pageNumber="23" type="male">1</specimenCount>
</date>
male, 7 9.
<date id="AC51102DC74BFFB5FF59FB7C3831FB2B" box="[171,334,1160,1186]" pageId="22" pageNumber="23" value="1969-08-01">
Aug.1969,
<specimenCount id="CEE9FD64C74BFFB5FECCFB7C3831FB2B" box="[318,334,1160,1186]" pageId="22" pageNumber="23" type="male">1</specimenCount>
</date>
male, 911.
<date id="AC51102DC74BFFB5FE1AFB7C3BF4FB2B" box="[488,651,1160,1186]" pageId="22" pageNumber="23" value="1969-08-02">
Aug.1969,
<specimenCount id="CEE9FD64C74BFFB5FD89FB7C3BF4FB2B" box="[635,651,1160,1186]" pageId="22" pageNumber="23" type="male">2</specimenCount>
</date>
male, 1113.
<date id="AC51102DC74BFFB5FCC7FB7C3AA7FB2B" box="[821,984,1160,1186]" pageId="22" pageNumber="23" value="1969-08-01">
Aug.1969,
<specimenCount id="CEE9FD64C74BFFB5FC3AFB7C3AA7FB2B" box="[968,984,1160,1186]" pageId="22" pageNumber="23" type="male">1</specimenCount>
</date>
male, 1617.
<date id="AC51102DC74BFFB5FB76FB7C3C5AFB2B" box="[1156,1317,1160,1186]" pageId="22" pageNumber="23" value="1969-08-01">
Aug.1969,
<specimenCount id="CEE9FD64C74BFFB5FAE7FB7C3C5AFB2B" box="[1301,1317,1160,1186]" pageId="22" pageNumber="23" type="male">1</specimenCount>
</date>
male,
<date id="AC51102DC74BFFB5FA8FFB7C3865FB4E" pageId="22" pageNumber="23" value="1969-09-05" valueMax="1969-09-09" valueMin="1969-09-05">5 9.Sep.1969</date>
(
<collectionCode id="BEFEAE28C74BFFB5FED8FB593800FB4E" box="[298,383,1197,1223]" httpUri="http://grbio.org/cool/m2jc-wmeq" name="University of California, Riverside" pageId="22" pageNumber="23">UCRC</collectionCode>
,
<collectionCode id="BEFEAE28C74BFFB5FE78FB59389FFB4E" box="[394,480,1197,1223]" pageId="22" pageNumber="23">EMUS</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="90F56566C74BFFB5FF37FB263ABDFB65" box="[197,962,1234,1260]" pageId="22" pageNumber="23" type="distribution">
<paragraph id="D85036EDC74BFFB5FF37FB263ABDFB65" blockId="22.[151,1438,152,1708]" box="[197,962,1234,1260]" pageId="22" pageNumber="23">
<emphasis id="EA9BEAFFC74BFFB5FF37FB26381AFB65" bold="true" box="[197,357,1234,1260]" pageId="22" pageNumber="23">Distribution.</emphasis>
Known only from Corn Springs and Deep Canyon.
</paragraph>
</subSubSection>
<subSubSection id="90F56566C74BFFB5FF37FB033A5DFABE" pageId="22" pageNumber="23" type="etymology">
<paragraph id="D85036EDC74BFFB5FF37FB033A5DFABE" blockId="22.[151,1438,152,1708]" pageId="22" pageNumber="23">
<emphasis id="EA9BEAFFC74BFFB5FF37FB03382DFA98" bold="true" box="[197,338,1271,1297]" pageId="22" pageNumber="23">Etymology.</emphasis>
Named after W.E. Ferguson who surveyed the mutillids of the Nevada Test Site in 1967 and published several critical papers on nocturnal mutillids.
</paragraph>
</subSubSection>
<subSubSection id="90F56566C74BFFB5FF37FAB63A46F925" pageId="22" pageNumber="23" type="discussion">
<paragraph id="D85036EDC74BFFB5FF37FAB63A3FFA45" blockId="22.[151,1438,152,1708]" pageId="22" pageNumber="23">
<emphasis id="EA9BEAFFC74BFFB5FF37FAB63841FAD5" bold="true" box="[197,318,1346,1372]" pageId="22" pageNumber="23">Remarks.</emphasis>
This species belongs in a new
<taxonomicName id="1FEF4D6EC74BFFB5FD58FAB73A4AFAD5" box="[682,821,1347,1372]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="fergusoni">
<emphasis id="EA9BEAFFC74BFFB5FD58FAB73A4AFAD5" box="[682,821,1347,1372]" italics="true" pageId="22" pageNumber="23">S. fergusoni</emphasis>
</taxonomicName>
species-group, which also includes
<taxonomicName id="1FEF4D6EC74BFFB5FB2EFAB73C15FAD5" box="[1244,1386,1347,1372]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="arnalduri" status="sp. nov.">
<emphasis id="EA9BEAFFC74BFFB5FB2EFAB73C15FAD5" box="[1244,1386,1347,1372]" italics="true" pageId="22" pageNumber="23">S. arnalduri</emphasis>
</taxonomicName>
,
<emphasis id="EA9BEAFFC74BFFB5FA8BFAB639B3FA08" bold="true" pageId="22" pageNumber="23">
<taxonomicNameLabel id="F1A85784C74BFFB5FA8BFAB639B3FA08" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, which is described later in this paper. The genitalia of
<taxonomicName id="1FEF4D6EC74BFFB5FC72FA9C3D6AFA08" box="[896,1045,1384,1409]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="arnalduri">
<emphasis id="EA9BEAFFC74BFFB5FC72FA9C3D6AFA08" box="[896,1045,1384,1409]" italics="true" pageId="22" pageNumber="23">S. arnalduri</emphasis>
</taxonomicName>
(Fig. 65) are quite similar to
<emphasis id="EA9BEAFFC74BFFB5FA77FA9C3870FA2F" italics="true" pageId="22" pageNumber="23">
<taxonomicName id="1FEF4D6EC74BFFB5FA77FA9C3873FA2F" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="fergusoni">S. fergusoni</taxonomicName>
,
</emphasis>
which were not illustrated. There are, however, other characters that are distinct between these species making the distinction of the two species credible.
</paragraph>
<paragraph id="D85036EDC74BFFB5FF37FA2C3BF4F9B5" blockId="22.[151,1438,152,1708]" pageId="22" pageNumber="23">
This species has tubercles on the hind coxae similar to those of
<taxonomicName id="1FEF4D6EC74BFFB5FC5EFA2C3D66FA78" box="[940,1049,1496,1521]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="blakeii">
<emphasis id="EA9BEAFFC74BFFB5FC5EFA2C3D66FA78" box="[940,1049,1496,1521]" italics="true" pageId="22" pageNumber="23">S. blakeii</emphasis>
</taxonomicName>
. They are difficult to see in most cases being obscured by setae. The morphology of the mandibles coupled with that of the clypeus make this species distinctive enough to be identified.
</paragraph>
<paragraph id="D85036EDC74BFFB5FF37F9BC3A46F925" blockId="22.[151,1438,152,1708]" pageId="22" pageNumber="23">
In
<figureCitation id="40D42A68C74BFFB5FF16F9BC3845F9EB" box="[228,314,1608,1634]" captionStart="FIGURES 38 43" captionStartId="16.[151,273,1981,2005]" captionTargetBox="[189,1401,194,1939]" captionTargetId="figure@16.[189,1402,194,1950]" captionTargetPageId="16" captionText="FIGURES 38 43. Sphaeropthalma chandleri, sp. nov.: 38. Head, anterior view; and 39. Habitus. Sphaeropthalma fergusoni, sp. nov.: 40. Pygidium, dorsal view; 41. Head anterior view; 42. Head, dorsal view; and 43. Habitus." httpUri="https://zenodo.org/record/196852/files/figure.png" pageId="22" pageNumber="23">Fig. 43</figureCitation>
, the black coloration of the gena is caused by some kind of oily substance that leaks out of older specimens; under natural circumstances, the gena should be the same color as the remainder of the head and actually is on the other side of the head of this individual.
</paragraph>
</subSubSection>
</treatment>
</document>