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<document id="72CA37DB86348299E03FB068AA92964C" ID-DOI="10.5281/zenodo.196847" ID-GBIF-Dataset="5e5a206b-9cfe-4bde-bc1d-4de509a3a4e9" ID-ISSN="1175-5326" ID-Zenodo-Dep="196847" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460254280575" checkinUser="plazi" docAuthor="Pitts, James P., Wilson, Joseph S., Williams, Kevin A. &amp; Boehme, Nicole F." docDate="2010" docId="504687FBC744FFBAFF65FE133AA2FB73" docLanguage="en" docName="zt02553p034.pdf" docOrigin="Zootaxa 2553" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Sphaeropthalma megagnathos Schuster" docType="treatment" docVersion="10" lastPageNumber="26" masterDocId="AC7FFF83C75DFFA3FFF2FFF4397FFF89" masterDocTitle="Nocturnal velvet ant males (Hymenoptera: Mutillidae) of Deep Canyon, California including four new species and a fifth new species from Owens Lake Valley, California" masterLastPageNumber="34" masterPageNumber="1" pageNumber="26" updateTime="1698243197414" updateUser="plazi">
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<mods:title id="E1D0122213A1A7A56076FFF29F7CE47A">Nocturnal velvet ant males (Hymenoptera: Mutillidae) of Deep Canyon, California including four new species and a fifth new species from Owens Lake Valley, California</mods:title>
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<mods:namePart id="F8616E8F37879973D8A1E27AFF5A61E2">Pitts, James P.</mods:namePart>
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<mods:namePart id="AE7F8FE0088E6B58C4D637F5F52E95D6">Wilson, Joseph S.</mods:namePart>
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<mods:namePart id="C6D33FD20BC60C55BF6F5F2DE2F4AA7A">Williams, Kevin A.</mods:namePart>
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<mods:namePart id="6FE52BAA311C7EC065F26DD3A8B57BE8">Boehme, Nicole F.</mods:namePart>
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<mods:date id="1AD753448363E7050BE43B14D7D2F15C">2010</mods:date>
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<treatment id="504687FBC744FFBAFF65FE133AA2FB73" ID-DOI="http://doi.org/10.5281/zenodo.5670804" ID-GBIF-Taxon="119392248" ID-Zenodo-Dep="5670804" LSID="urn:lsid:plazi:treatment:504687FBC744FFBAFF65FE133AA2FB73" httpUri="http://treatment.plazi.org/id/504687FBC744FFBAFF65FE133AA2FB73" lastPageNumber="26" pageId="25" pageNumber="26">
<subSubSection id="90F56566C744FFBAFF65FE133B03FD88" box="[151,636,487,513]" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="D85036EDC744FFBAFF65FE133B03FD88" blockId="25.[151,636,487,513]" box="[151,636,487,513]" pageId="25" pageNumber="26">
<heading id="83188181C744FFBAFF65FE133B03FD88" bold="true" box="[151,636,487,513]" fontSize="11" level="1" pageId="25" pageNumber="26" reason="1">
<taxonomicName id="1FEF4D6EC744FFBAFF65FE133B03FD88" authority="Schuster" authorityName="Schuster" box="[151,636,487,513]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="megagnathos">
<emphasis id="EA9BEAFFC744FFBAFF65FE133B03FD88" bold="true" box="[151,636,487,513]" pageId="25" pageNumber="26">
<emphasis id="EA9BEAFFC744FFBAFF65FE133B78FD88" bold="true" box="[151,519,487,513]" italics="true" pageId="25" pageNumber="26">Sphaeropthalma megagnathos</emphasis>
Schuster
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="90F56566C744FFBAFF65FDC43A42FD2E" pageId="25" pageNumber="26" type="reference_group">
<paragraph id="D85036EDC744FFBAFF65FDC43A2FFDEE" blockId="25.[151,1437,560,679]" pageId="25" pageNumber="26">
<taxonomicName id="1FEF4D6EC744FFBAFF65FDC43AD0FDCE" authority="Schuster, 1958" authorityName="Schuster" authorityYear="1958" box="[151,943,560,583]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="subSpecies" species="megagnathos" subGenus="Photopsis" subSpecies="megagnathos">
<emphasis id="EA9BEAFFC744FFBAFF65FDC43839FDCE" box="[151,326,560,583]" italics="true" pageId="25" pageNumber="26">Sphaeropthalma</emphasis>
(
<emphasis id="EA9BEAFFC744FFBAFEA8FDC438BCFDCE" box="[346,451,560,583]" italics="true" pageId="25" pageNumber="26">Photopsis</emphasis>
)
<emphasis id="EA9BEAFFC744FFBAFE25FDC43B80FDCE" box="[471,767,560,583]" italics="true" pageId="25" pageNumber="26">megagnathos megagnathos</emphasis>
<bibRefCitation id="BC7E4B1CC744FFBAFCF8FDC43AD0FDCE" author="Schuster" box="[778,943,560,583]" pageId="25" pageNumber="34" refString="Schuster, R. M. (1958) A revision of the sphaeropthalmine Mutillidae of America north of Mexico. II. Entomologica Americana, 37, 1 - 130." type="journal article" year="1958">Schuster, 1958</bibRefCitation>
</taxonomicName>
. Ent. Amer. (n. s.) 37: 36. Male.
<typeStatus id="0754884FC744FFBAFAC3FDC43CE7FDCE" box="[1329,1432,560,583]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
: Arizona, Ehrenberg,
<date id="AC51102DC744FFBAFE54FDA43B52FDEE" box="[422,557,592,615]" pageId="25" pageNumber="26" value="1939-04-27">27.Apr.1939</date>
, coll. F.H. Parker (UMSP).
</paragraph>
<paragraph id="D85036EDC744FFBAFF65FD843A42FD2E" blockId="25.[151,1437,560,679]" pageId="25" pageNumber="26">
<taxonomicName id="1FEF4D6EC744FFBAFF65FD843A19FD0E" authority="Schuster, 1958" authorityName="Schuster" authorityYear="1958" box="[151,870,624,647]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="subSpecies" species="megagnathos" subGenus="Photopsis" subSpecies="aurifera">
<emphasis id="EA9BEAFFC744FFBAFF65FD843839FD0E" box="[151,326,624,647]" italics="true" pageId="25" pageNumber="26">Sphaeropthalma</emphasis>
(
<emphasis id="EA9BEAFFC744FFBAFEA4FD8438C0FD0E" box="[342,447,624,647]" italics="true" pageId="25" pageNumber="26">Photopsis</emphasis>
)
<emphasis id="EA9BEAFFC744FFBAFE22FD843BC1FD0E" box="[464,702,624,647]" italics="true" pageId="25" pageNumber="26">megagnathos aurifera</emphasis>
<bibRefCitation id="BC7E4B1CC744FFBAFD37FD843A19FD0E" author="Schuster" box="[709,870,624,647]" pageId="25" pageNumber="34" refString="Schuster, R. M. (1958) A revision of the sphaeropthalmine Mutillidae of America north of Mexico. II. Entomologica Americana, 37, 1 - 130." type="journal article" year="1958">Schuster, 1958</bibRefCitation>
</taxonomicName>
. Ent. Amer. (n. s.) 37: 36. Male.
<typeStatus id="0754884FC744FFBAFB3FFD843C4BFD0E" box="[1229,1332,624,647]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
: Arizona, Tinajas Atlas Mountains, 1905, coll. W.J. McGee (UMSP).
</paragraph>
</subSubSection>
<subSubSection id="90F56566C744FFBAFF65FD213ACAFC23" pageId="25" pageNumber="26" type="diagnosis">
<paragraph id="D85036EDC744FFBAFF65FD213ACAFC23" blockId="25.[151,1437,725,1274]" pageId="25" pageNumber="26">
<emphasis id="EA9BEAFFC744FFBAFF65FD213806FD66" bold="true" box="[151,377,725,751]" pageId="25" pageNumber="26">Diagnosis of male.</emphasis>
The male of this species can be recognized by mandibular morphology (
<figureCitation id="40D42A68C744FFBAFB3BFD213C60FD66" box="[1225,1311,725,751]" captionStart="FIGURES 25 31" captionStartId="13.[151,274,1360,1384]" captionTargetBox="[151,1435,194,1326]" captionTargetId="figure@13.[151,1436,194,1338]" captionTargetPageId="13" captionText="FIGURES 25 31. Head, anterior view. 25. Sphaeropthalma. megagnathos; 26. S. mendica; 27. S. pallida; 28. S. sublobata; 29. S. tetracuspis; 30. S. triangularis; and 31. S. yumaella." httpUri="https://zenodo.org/record/196850/files/figure.png" pageId="25" pageNumber="26">Fig. 25</figureCitation>
; see
<figureCitation id="40D42A68C744FFBAFAAAFD213CE3FD66" box="[1368,1436,725,751]" captionStart="FIGURES 1 12" captionStartId="4.[151,278,1762,1786]" captionTargetBox="[151,1436,197,1724]" captionTargetId="figure@4.[151,1436,197,1739]" captionTargetPageId="4" captionText="FIGURES 1 12. Head, anterior view. 1. Acanthophotopsis falciformis; 2. Acrophotopsis campylognatha; 3. Odontophotopsis acmaea; 4. O. armata; 5. O. aufidia; 6. O. bellona; 7. O. biramosa; 8. O. clypeata; 9. O. inconspicua; 10. O. mamata; 11. O. melicausa; and 12. O. quadrispinosa." httpUri="https://zenodo.org/record/196848/files/figure.png" pageId="25" pageNumber="26">
Fig.
<quantity id="1F179B08C744FFBAFA7FFD213CE3FD66" box="[1421,1436,725,751]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="25" pageNumber="26" unit="in" value="7.0">7</quantity>
</figureCitation>
in
<bibRefCitation id="BC7E4B1CC744FFBAFF44FD0E3845FC9D" author="Pitts" box="[182,314,762,788]" pageId="25" pageNumber="34" refString="Pitts, J. P. (2006) Review of the Sphaeropthalma imperialis species-group (Hymenoptera: Mutillidae), with descriptions of females and taxonomic notes. Zootaxa, 1248, 1 - 20." type="journal article" year="2006">Pitts, 2006</bibRefCitation>
). The mandibles are broadly dilated, especially the ventral portion apically, and is much wider distally than its width at the ventral angulation, the ventral basal tooth of the mandible is small, and the apex is vertical. Additionally, the head is long and parallel posteriorly, the clypeus is deeply depressed below the dorsal mandibular margin, the mesosternum lacks tubercles, the wings are yellowish-hyaline and a sternal felt line is absent. The genitalic morphology also is diagnostic (Fig. 59).
</paragraph>
</subSubSection>
<subSubSection id="90F56566C744FFBAFF37FC4138D8FC46" box="[197,423,949,975]" pageId="25" pageNumber="26" type="description">
<paragraph id="D85036EDC744FFBAFF37FC4138D8FC46" blockId="25.[151,1437,725,1274]" box="[197,423,949,975]" pageId="25" pageNumber="26">
<emphasis id="EA9BEAFFC744FFBAFF37FC41385AFC47" box="[197,293,949,974]" italics="true" pageId="25" pageNumber="26">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="90F56566C744FFBAFF37FC2E3A4BFB93" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="D85036EDC744FFBAFF37FC2E3A4BFB93" blockId="25.[151,1437,725,1274]" pageId="25" pageNumber="26">
<emphasis id="EA9BEAFFC744FFBAFF37FC2E3B76FC7D" bold="true" box="[197,521,986,1012]" pageId="25" pageNumber="26">
Material examined.
<collectingCountry id="A0F8767DC744FFBAFE3FFC2E3B76FC7D" box="[461,521,986,1012]" name="United States of America" pageId="25" pageNumber="26">USA</collectingCountry>
</emphasis>
:
<emphasis id="EA9BEAFFC744FFBAFDEEFC2E3BD4FC7D" bold="true" box="[540,683,986,1012]" pageId="25" pageNumber="26">California,</emphasis>
<emphasis id="EA9BEAFFC744FFBAFD41FC2E3A2BFC7A" box="[691,852,986,1011]" italics="true" pageId="25" pageNumber="26">Riverside Co</emphasis>
.: Deep Canyon,
<specimenCount id="CEE9FD64C744FFBAFBD7FC2E3D03FC7D" box="[1061,1148,986,1012]" pageId="25" pageNumber="26" type="male">1 male</specimenCount>
,
<date id="AC51102DC744FFBAFB7BFC2E3C52FC7D" box="[1161,1325,986,1012]" pageId="25" pageNumber="26" value="1973-05-19">19.May.1973</date>
, coll. M. Quillman,
<specimenCount id="CEE9FD64C744FFBAFEE6FBF43819FB93" box="[276,358,1024,1050]" pageId="25" pageNumber="26" type="male">1 male</specimenCount>
, 2223.
<date id="AC51102DC744FFBAFE33FBF43B29FB93" box="[449,598,1024,1050]" pageId="25" pageNumber="26" value="2007-05-01">
May.2007,
<specimenCount id="CEE9FD64C744FFBAFDB5FBF43B29FB93" box="[583,598,1024,1050]" pageId="25" pageNumber="26" type="male">1</specimenCount>
</date>
male,
<date id="AC51102DC744FFBAFD56FBF43A4FFB93" box="[676,816,1024,1050]" pageId="25" pageNumber="26" value="1969-07-23">23.Jul.1969</date>
.
</paragraph>
</subSubSection>
<subSubSection id="90F56566C744FFBAFF37FBD13A26FBB6" box="[197,857,1061,1087]" pageId="25" pageNumber="26" type="distribution">
<paragraph id="D85036EDC744FFBAFF37FBD13A26FBB6" blockId="25.[151,1437,725,1274]" box="[197,857,1061,1087]" pageId="25" pageNumber="26">
<emphasis id="EA9BEAFFC744FFBAFF37FBD1381AFBB6" bold="true" box="[197,357,1061,1087]" pageId="25" pageNumber="26">Distribution.</emphasis>
The western Sonoran and Mojave deserts.
</paragraph>
</subSubSection>
<subSubSection id="90F56566C744FFBAFF34FBBE3AA2FB73" pageId="25" pageNumber="26" type="discussion">
<paragraph id="D85036EDC744FFBAFF34FBBE3AA2FB73" blockId="25.[151,1437,725,1274]" pageId="25" pageNumber="26">
<emphasis id="EA9BEAFFC744FFBAFF34FBBE383FFBED" bold="true" box="[198,320,1098,1124]" pageId="25" pageNumber="26">Remarks.</emphasis>
In some specimens the coloration of the setae is bright orange, although setal coloration varies from orange to white (see
<figureCitation id="40D42A68C744FFBAFE3DFB843B5CFB03" box="[463,547,1136,1162]" captionStart="FIGURES 25 31" captionStartId="13.[151,274,1360,1384]" captionTargetBox="[151,1435,194,1326]" captionTargetId="figure@13.[151,1436,194,1338]" captionTargetPageId="13" captionText="FIGURES 25 31. Head, anterior view. 25. Sphaeropthalma. megagnathos; 26. S. mendica; 27. S. pallida; 28. S. sublobata; 29. S. tetracuspis; 30. S. triangularis; and 31. S. yumaella." httpUri="https://zenodo.org/record/196850/files/figure.png" pageId="25" pageNumber="26">
Fig.
<quantity id="1F179B08C744FFBAFDF4FB843B5CFB03" box="[518,547,1136,1162]" metricMagnitude="-1" metricUnit="m" metricValue="7.366" pageId="25" pageNumber="26" unit="in" value="29.0">29</quantity>
</figureCitation>
in
<bibRefCitation id="BC7E4B1CC744FFBAFDB8FB843BB1FB03" author="Pitts" box="[586,718,1136,1162]" pageId="25" pageNumber="34" refString="Pitts, J. P. (2006) Review of the Sphaeropthalma imperialis species-group (Hymenoptera: Mutillidae), with descriptions of females and taxonomic notes. Zootaxa, 1248, 1 - 20." type="journal article" year="2006">Pitts, 2006</bibRefCitation>
). The orangish specimens of
<taxonomicName id="1FEF4D6EC744FFBAFBDAFB843D9DFB00" box="[1064,1250,1136,1161]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="megagnathos">
<emphasis id="EA9BEAFFC744FFBAFBDAFB843D9DFB00" box="[1064,1250,1136,1161]" italics="true" pageId="25" pageNumber="26">S. megagnathos</emphasis>
</taxonomicName>
could easily be confused with
<taxonomicName id="1FEF4D6EC744FFBAFEB4FB6138C5FB27" box="[326,442,1173,1198]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="luiseno">
<emphasis id="EA9BEAFFC744FFBAFEB4FB6138C5FB27" box="[326,442,1173,1198]" italics="true" pageId="25" pageNumber="26">S. luiseno</emphasis>
</taxonomicName>
or
<taxonomicName id="1FEF4D6EC744FFBAFE11FB613B0CFB27" box="[483,627,1173,1198]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="contracta">
<emphasis id="EA9BEAFFC744FFBAFE11FB613B0CFB27" box="[483,627,1173,1198]" italics="true" pageId="25" pageNumber="26">S. contracta</emphasis>
</taxonomicName>
based on coloration. They do not occur in Deep Canyon;
<taxonomicName id="1FEF4D6EC744FFBAFADAFB613CE4FB27" box="[1320,1435,1173,1198]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="luiseno">
<emphasis id="EA9BEAFFC744FFBAFADAFB613CE4FB27" box="[1320,1435,1173,1198]" italics="true" pageId="25" pageNumber="26">S. luiseno</emphasis>
</taxonomicName>
has large projections bearing long, curved setae on the middle and hind coxae, while
<taxonomicName id="1FEF4D6EC744FFBAFB60FB4F3C5DFB5D" box="[1170,1314,1211,1236]" class="Insecta" family="Mutillidae" genus="Sphaeropthalma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="contracta">
<emphasis id="EA9BEAFFC744FFBAFB60FB4F3C5DFB5D" box="[1170,1314,1211,1236]" italics="true" pageId="25" pageNumber="26">S. contracta</emphasis>
</taxonomicName>
has much smaller ocelli and different genitalic morphology (see
<bibRefCitation id="BC7E4B1CC744FFBAFCE6FB143AAEFB73" author="Pitts" box="[788,977,1248,1274]" pageId="25" pageNumber="34" refString="Pitts, J. P., Parker, F. D. &amp; Pitts-Singer, T. L. (2004) A review of the Sphaeropthalma uro species-group (Hymenoptera: Mutillidae), with taxonomic changes. Journal of the Kansas Entomological Society, 77, 223 - 234." type="journal article" year="2004">
Pitts
<emphasis id="EA9BEAFFC744FFBAFCBDFB143AF5FB70" box="[847,906,1248,1273]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
2004
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>