134 lines
10 KiB
XML
134 lines
10 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.675.13252" ID-GBIF-Dataset="724d6de0-f19a-457d-9339-84c836e19c38" ID-PMC="PMC5523889" ID-Pensoft-Pub="1313-2970-675-129" ID-PubMed="28769682" ID-ZBK="E5C8F163D61547B98418CEE8D71A7DAB" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-675-129" ModsDocOrigin="ZooKeys 675" ModsDocTitle="Conchological differentiation and genital anatomy of Nepalese Glessulinae (Gastropoda, Stylommatophora, Subulinidae), with descriptions of six new species" checkinTime="1495538522124" checkinUser="pensoft" docAuthor="Budha, Prem B., Naggs, Fred & Backeljau, Thierry" docDate="2017" docId="4F21CFEE76CEA348B1E9CE9A13C7C766" docLanguage="en" docName="ZooKeys 675: 129-156" docOrigin="ZooKeys 675" docSource="http://dx.doi.org/10.3897/zookeys.675.13252" docTitle="Rishetia nagarjunensis Budha & Naggs, sp. n." docType="treatment" docUuid="CB41BEB9-DCC9-47F5-9E13-7AC9A357BDDF" docUuidSource="ZooBank" docVersion="6" lastPageNumber="136" masterDocId="FF8BE96F4A658939FFA88167FF8E6E18" masterDocTitle="Conchological differentiation and genital anatomy of Nepalese Glessulinae (Gastropoda, Stylommatophora, Subulinidae), with descriptions of six new species" masterLastPageNumber="156" masterPageNumber="129" pageNumber="136" updateTime="1668164473028" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Conchological differentiation and genital anatomy of Nepalese Glessulinae (Gastropoda, Stylommatophora, Subulinidae), with descriptions of six new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Budha, Prem B.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Naggs, Fred</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Backeljau, Thierry</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>675</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>129</mods:start>
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<mods:end>156</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.675.13252</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.675.13252</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-675-129</mods:identifier>
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<mods:identifier type="ZBK">E5C8F163D61547B98418CEE8D71A7DAB</mods:identifier>
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<mods:identifier type="ZooBank">E5C8F163D61547B98418CEE8D71A7DAB</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="130844517" LSID="urn:lsid:zoobank.org:act:CB41BEB9-DCC9-47F5-9E13-7AC9A357BDDF" httpUri="http://treatment.plazi.org/id/4F21CFEE76CEA348B1E9CE9A13C7C766" lastPageNumber="136" pageId="7" pageNumber="136">
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<subSubSection pageId="7" pageNumber="136" type="nomenclature">
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<paragraph pageId="7" pageNumber="136">
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<taxonomicName LSID="http://zoobank.org/CB41BEB9-DCC9-47F5-9E13-7AC9A357BDDF" authority="Budha & Naggs" class="Gastropoda" family="Subulinidae" genus="Rishetia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rishetia nagarjunensis" order="Gastropoda" pageId="7" pageNumber="136" phylum="Mollusca" rank="species" species="nagarjunensis">
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<pageBreakToken pageId="7" pageNumber="136" start="start">Rishetia</pageBreakToken>
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nagarjunensis Budha & Naggs
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</taxonomicName>
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<taxonomicNameLabel pageId="7" pageNumber="136">sp. n.</taxonomicNameLabel>
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Figs 2G, 9, 13E
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="136" type="materials_examined">
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<paragraph pageId="7" pageNumber="136">Material examined.</paragraph>
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<paragraph pageId="7" pageNumber="136">
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Holotype: CDZMTU067P/1, Nagarjun Forest, Balaju-Jamacho trail, Nagarjun-Shivapuri National Park, Kathmandu, Nepal, 1850 m,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="27.745996">27.745997N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="85.28724">85.287240E</geoCoordinate>
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, 24.I.2009, leg. P.B. Budha, R. Devkota, S. Khatiwara and S. Khanal. Paratypes: CDZMTU067/9 shells and CDZMTU068P/1 specimen from the type locality (same data as holotype). Paratypes: CDZMTU069P/1 specimen (dissected), Phulchowki Hill, mixed
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<taxonomicName class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rhododendron" order="Ericales" pageId="7" pageNumber="136" phylum="Tracheophyta" rank="genus">Rhododendron</taxonomicName>
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forest, 2324 m,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="27.578318">27.578317N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="85.39689">85.396885E</geoCoordinate>
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, 03.V.2007, leg. P.B. Budha.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="136" type="distribution">
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<paragraph pageId="7" pageNumber="136">Distribution.</paragraph>
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<paragraph pageId="7" pageNumber="136">Nagarjun-Shivapuri National Park and Phulchowki Hill, Central Nepal.</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="136" type="etymology">
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<paragraph pageId="7" pageNumber="136">Etymology.</paragraph>
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<paragraph pageId="7" pageNumber="136">The name is derived from the type locality Nagarjun Forest.</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="136" type="shell">
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<paragraph pageId="7" pageNumber="136">Shell.</paragraph>
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<paragraph pageId="7" pageNumber="136">
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Measurements (n = 4): SH 33.2-38.4 mm, SW 10.6-11.9 mm, HA 10.2-10.8 mm, WA 5.5-6.2 mm, Wh 10-11.0; holotype: SH 38.0 mm, SW 12.0 mm, HA 10.5 mm, WA 6.3 mm, Wh 11; approx. 3.3
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<normalizedToken originalValue="×">x</normalizedToken>
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higher than wide, elongately turreted, thick, reddish-brown or dark chestnut colour. Surface striated with obliquely raised riblets on the first 2-3 whorls, later whorls with comparatively thin and dense sculpture. The first whorl smooth, rounded. Sides rather flat, suture shallow. Aperture ovate, 1.8
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<normalizedToken originalValue="×">x</normalizedToken>
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longer than wide, peristome thin, inner side of the aperture thickened and whitish, columella slightly truncate at the base.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="136" type="radula">
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<paragraph pageId="7" pageNumber="136">Radula</paragraph>
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<paragraph pageId="7" pageNumber="136">(Fig. 13E). Central tooth very small, lacks cusps, lateral teeth shouldered.</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="136" type="genitalia">
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<paragraph pageId="7" pageNumber="136">Genitalia</paragraph>
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<paragraph pageId="7" pageNumber="136">(n = 2) (Fig. 9). All specimens have a long vas deferens. Flagellum and epiphallic caecum are reduced to two very short knobs (Fig. 9A). Penis cylindrical, with a nearly constant diameter in its proximal half, distally it rapidly expands into a bulbous section before the diameter contracts to somewhat less than the proximal penis, narrowing slightly distally. The penial retractor muscle far apart from the flagellum. The inner surface contains weakly convoluted folds (Fig. 9B). Epiphallus length about 1/3 of penis length. Vagina nearly as long as the penis. Gametolytic duct cylindrical, ends into a simple gametolytic sac. Hermaphroditic duct is loosely convoluted and connected at the middle of the albumen gland.</paragraph>
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<caption pageId="7" pageNumber="136">
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<paragraph pageId="7" pageNumber="136">
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Figure 9. Genitalia of
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<taxonomicName lsidName="R. nagarjunensis" pageId="7" pageNumber="136" rank="species" species="nagarjunensis">R. nagarjunensis</taxonomicName>
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sp. n., holotype: CDZMTU067P, A General view B Interior of penis of the same, Nagarjun forest, Balaju-Jamacho trail Nagarjun-Shivapuri National Park, Kathmandu, Nepal.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="136" type="remarks">
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<paragraph pageId="7" pageNumber="136">Remarks.</paragraph>
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<paragraph pageId="7" pageNumber="136">
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The shell of
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<taxonomicName lsidName="R. nagarjunensis" pageId="7" pageNumber="136" rank="species" species="nagarjunensis">R. nagarjunensis</taxonomicName>
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sp. n. is similar in size and shape to that of sympatric
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<taxonomicName lsidName="R. kathmandica" pageId="7" pageNumber="136" rank="species" species="kathmandica">R. kathmandica</taxonomicName>
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sp. n. However, the shell of
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<taxonomicName lsidName="R. kathmandica" pageId="7" pageNumber="136" rank="species" species="kathmandica">R. kathmandica</taxonomicName>
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sp. n. is more slender and has more whorls than
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<taxonomicName lsidName="R. nagarjunensis" pageId="7" pageNumber="136" rank="species" species="nagarjunensis">R. nagarjunensis</taxonomicName>
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sp. n. Conversely,
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<taxonomicName lsidName="R. nagarjunensis" pageId="7" pageNumber="136" rank="species" species="nagarjunensis">R. nagarjunensis</taxonomicName>
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sp. n. has a wider body whorl, a comparatively more robust shell, and stronger radial ribs than
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<taxonomicName lsidName="R. kathmandica" pageId="7" pageNumber="136" rank="species" species="kathmandica">R. kathmandica</taxonomicName>
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sp. n. The genitalia of both species are consistently different due to the reduced knob-like flagellum and epiphalic caecum in
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<taxonomicName lsidName="R. nagarjunensis" pageId="7" pageNumber="136" rank="species" species="nagarjunensis">R. nagarjunensis</taxonomicName>
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sp. n., as well as by the inner surface of the penis, which in
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<taxonomicName lsidName="R. nagarjunensis" pageId="7" pageNumber="136" rank="species" species="nagarjunensis">R. nagarjunensis</taxonomicName>
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sp. n. shows three loosely convoluted folds, whereas in
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<taxonomicName lsidName="R. kathmandica" pageId="7" pageNumber="136" rank="species" species="kathmandica">R. kathmandica</taxonomicName>
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sp. n. it shows distinct rectangular zigzag folds (Fig. 7A1).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |