506 lines
72 KiB
XML
506 lines
72 KiB
XML
<document id="9F35CACBBE9FA67966A93CA3AC38D401" ID-DOI="10.11646/zootaxa.4748.2.2" ID-GBIF-Dataset="62a26005-245c-4298-ae96-2de02d77897b" ID-ISSN="1175-5326" ID-Zenodo-Dep="3698755" ID-ZooBank="90DAFF0F-6ABD-402F-8F37-85AF23CA83BF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1583482566449" checkinUser="plazi" docAuthor="Vidanapathirana, L. J. Mendis Wickramasinghe Dulan Ranga, Pushpamal, Vishan & Wickramasinghe, Nethu" docDate="2020" docId="263B94701738FFC28EE0FA504CBBCA23" docLanguage="en" docName="zootaxa.4748.2.2.pdf" docOrigin="Zootaxa 4748 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dryocalamus chithrasekarai Vidanapathirana & Pushpamal & Wickramasinghe 2020, sp. nov." docType="treatment" docVersion="8" lastPageNumber="254" masterDocId="DA02EC081739FFC48E77FFBC4E44CC3B" masterDocTitle="A new species of Dryocalamus (Serpentes: Colubridae) endemic to the rainforests of southwestern Sri Lanka" masterLastPageNumber="260" masterPageNumber="248" pageNumber="249" updateTime="1702002991662" updateUser="plazi" zenodo-license-document="CLOSED">
|
||
<mods:mods id="CD398DD711EE00EEEE3FDFA11DB293E4" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="BE86E4EF32CD1C3ABC1891B4ACB9428B">
|
||
<mods:title id="F00E9FCA0EA90F3AEB6C140AF8E7D874">A new species of Dryocalamus (Serpentes: Colubridae) endemic to the rainforests of southwestern Sri Lanka</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="43464E958DBC908C5A599F2D630B138C" type="personal">
|
||
<mods:role id="D3DA1D05AAFB3D93C5DF7220107FEDA2">
|
||
<mods:roleTerm id="E709F83E7B91EFE9537C89A0DF8A1176">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="C6D59E4AA39A2C45043C17629D29C79C">Vidanapathirana, L. J. Mendis Wickramasinghe Dulan Ranga</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="A89699FCE54354086E56B957AFF01680" type="personal">
|
||
<mods:role id="7398DA8F4E896902E3ADA15376E81B3A">
|
||
<mods:roleTerm id="59C88A550924E12EDCD8D59DCE9BF30E">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B9A9043FF7683E5D1980734489B1E7B9">Pushpamal, Vishan</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="AE044C24182BDC24683463492DC8F2F5" type="personal">
|
||
<mods:role id="051C39FC22EFAC79711143E5F07E17CD">
|
||
<mods:roleTerm id="16499C81ECE969D42FD05F91C3BE82A7">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="124EF3EAEE9F1BD003136B913BD7DB25">Wickramasinghe, Nethu</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="F7CD09AEB135A984581A701D3CF1B542">text</mods:typeOfResource>
|
||
<mods:relatedItem id="4AA7968001AA9B270322AC6FA8CF9947" type="host">
|
||
<mods:titleInfo id="0A305C0287901AB74624815FB916AC83">
|
||
<mods:title id="277F93D2C40FB9A060B06D424D6D9EDA">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="EA4859E68D367FB7451CEC333830239E">
|
||
<mods:date id="480AC471D0E64DF3B843A0107F97E648">2020</mods:date>
|
||
<mods:detail id="2E78AFC0C9E7ADBCB2907BBB9C869DF7" type="pubDate">
|
||
<mods:number id="2763E0BAD8E2BBC75208CD7CFB629CD3">2020-03-06</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="89A204FAE262FBBD0E91EC86669DFAC2" type="volume">
|
||
<mods:number id="9FA243BB6A0988392ABAFDF88DE34500">4748</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="C9AC86FAA29A40DE8CA0BDF790156E88" type="issue">
|
||
<mods:number id="A70A6091519C4E7843DC79A2496D976E">2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="BE95732AB495938802736EF62A8B238A" unit="page">
|
||
<mods:start id="F44F934F8D396593B84997FF88428312">248</mods:start>
|
||
<mods:end id="572368FC19073A25518F7EDBA75D0211">260</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="76E636680029C8615218186642BBEDCF">journal article</mods:classification>
|
||
<mods:identifier id="80238CF9369D7E7BA8B3CBA4ED736198" type="DOI">10.11646/zootaxa.4748.2.2</mods:identifier>
|
||
<mods:identifier id="D072259861EC10939ECEBC377DE6167A" type="GBIF-Dataset">62a26005-245c-4298-ae96-2de02d77897b</mods:identifier>
|
||
<mods:identifier id="933BC5CD950154B13A1B2671A1F0A3E4" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="3C42175E1AEA95EABEEF1BDBD794B93F" type="Zenodo-Dep">3698755</mods:identifier>
|
||
<mods:identifier id="62539C5F988606F14395373C48DDFAF5" type="ZooBank">90DAFF0F-6ABD-402F-8F37-85AF23CA83BF</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="263B94701738FFC28EE0FA504CBBCA23" ID-DOI="http://doi.org/10.5281/zenodo.3704868" ID-GBIF-Taxon="162462230" ID-Zenodo-Dep="3704868" LSID="urn:lsid:plazi:treatment:263B94701738FFC28EE0FA504CBBCA23" httpUri="http://treatment.plazi.org/id/263B94701738FFC28EE0FA504CBBCA23" lastPageId="6" lastPageNumber="254" pageId="1" pageNumber="249">
|
||
<subSubSection id="E68876ED1738FFC58EE0FA504C18CA3D" box="[151,604,1516,1543]" pageId="1" pageNumber="249" type="nomenclature">
|
||
<paragraph id="AE2D25661738FFC58EE0FA504C18CA3D" blockId="1.[151,604,1516,1579]" box="[151,604,1516,1543]" pageId="1" pageNumber="249">
|
||
<heading id="F565920A1738FFC58EE0FA504C18CA3D" bold="true" box="[151,604,1516,1543]" fontSize="11" level="1" pageId="1" pageNumber="249" reason="1">
|
||
<emphasis id="9CE6F9741738FFC58EE0FA504C18CA3D" bold="true" box="[151,604,1516,1543]" pageId="1" pageNumber="249">
|
||
<taxonomicName id="69925EE51738FFC58EE0FA504FB4CA3C" authority="Vidanapathirana & Pushpamal & Wickramasinghe, 2020" authorityName="Vidanapathirana & Pushpamal & Wickramasinghe" authorityYear="2020" box="[151,496,1516,1543]" class="Reptilia" family="Colubridae" genus="Dryocalamus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="249" phylum="Chordata" rank="species" species="chithrasekarai" status="sp. nov.">
|
||
<emphasis id="9CE6F9741738FFC58EE0FA504FB4CA3C" bold="true" box="[151,496,1516,1543]" italics="true" pageId="1" pageNumber="249">Dryocalamus chithrasekarai</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="87D5440F1738FFC58F88FA504C18CA3D" box="[511,604,1516,1542]" pageId="1" pageNumber="249" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E68876ED1738FFC58EE0F9AC4FC3CA11" box="[151,391,1552,1579]" pageId="1" pageNumber="249" type="description">
|
||
<paragraph id="AE2D25661738FFC58EE0F9AC4FC3CA11" blockId="1.[151,604,1516,1579]" box="[151,391,1552,1579]" pageId="1" pageNumber="249">
|
||
(
|
||
<figureCitation id="36A939E31738FFC58EE8F9AC4F47CA11" box="[159,259,1552,1578]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,250,1243,1268]" captionStartId-1="3.[151,250,1955,1980]" captionStartId-2="4.[151,250,1088,1113]" captionStartId-3="4.[151,250,1980,2005]" captionTargetBox-0="[176,1412,259,1194]" captionTargetBox-1="[161,1427,1111,1921]" captionTargetBox-2="[220,1365,203,1042]" captionTargetBox-3="[191,1397,1179,1946]" captionTargetId-0="figure@2.[151,1436,237,1218]" captionTargetId-1="figure@3.[151,1436,1101,1931]" captionTargetId-2="figure@4.[185,1403,181,1063]" captionTargetId-3="figure@4.[181,1406,1171,1954]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="4" captionText-0="FIGURE 1. Distribution of Dryocalamus chithrasekarai sp. nov., holotype (solid white dot), paratype (white circle), specimens illustrated in figures (black solid dots), sight-only records (black circle). Black solid square indicates the type locality (Jaffna) of Odontomus fergusonii; the black square indicates Dambulla, and the white square indicates Kurunegala, the only localities from which the only confirmed localities for Dryocalamus gracilis has been recorded by previous authors." captionText-1="FIGURE 2. The holotype of Dryocalamus chithrasekarai sp. nov., anterior body (A) dorsal, (B) lateral, (C) ventral aspects; midbody (D) dorsal, (E) lateral, and (F) ventral aspects, respectively; and (G) ventral aspect of tail." captionText-2="FIGURE 3. Dryocalamus chithrasekarai sp. nov., holotype, NMSL-NH 2019.26.01, 328 mm SVL, head scalation: (A) lateral, (B) dorsal, (C) ventral aspects, respectively." captionText-3="FIGURE 4. The paratype, of Dryocalamus chithrasekarai sp. nov., in life (not collected) showing (A) dorsal, (B) dorsolateral (laterally inverted), and (C) ventral aspects, respectively." figureDoi-0="http://doi.org/10.5281/zenodo.3698757" figureDoi-1="http://doi.org/10.5281/zenodo.3698761" figureDoi-2="http://doi.org/10.5281/zenodo.3698763" figureDoi-3="http://doi.org/10.5281/zenodo.3698765" httpUri-0="https://zenodo.org/record/3698757/files/figure.png" httpUri-1="https://zenodo.org/record/3698761/files/figure.png" httpUri-2="https://zenodo.org/record/3698763/files/figure.png" httpUri-3="https://zenodo.org/record/3698765/files/figure.png" pageId="1" pageNumber="249">Figs 1–4</figureCitation>
|
||
,
|
||
<figureCitation id="36A939E31738FFC58F67F9AD4F64CA10" box="[272,288,1553,1579]" captionStart="FIGURE 7" captionStartId="7.[151,250,1728,1753]" captionTargetBox="[162,1425,464,1693]" captionTargetId="figure@7.[151,1436,453,1704]" captionTargetPageId="7" captionText="FIGURE 7. Dryocalamus chithrasekarai sp. nov., in life (not collected) from (A-C) Runakanda Forest Reserve, (D) from Sinharaja World Heritage Site." figureDoi="http://doi.org/10.5281/zenodo.3698771" httpUri="https://zenodo.org/record/3698771/files/figure.png" pageId="1" pageNumber="249">7</figureCitation>
|
||
,
|
||
<tableCitation id="E31010DD1738FFC58F5CF9AC4F38CA11" box="[299,380,1552,1578]" captionStart="TABLE 1" captionStartId="9.[151,239,586,610]" captionTargetPageId="9" captionText="TABLE 1. Meristic and morphometric data for 40 characters of the holotype (*) and paratype (**) of Dryocalamus chithrasekarai sp. nov. Bilateral scale are counts given in left, right order. Character numbers in parentheses, as follows: (1) CL, (2) DBE, (3) DFE, (4) ED, (5) HD, (6) HL, (7) HW, (8) IN, (9) IO, (10)NE, (11) ND, (12) SE, (13) SG, (14) SN, (15) SVL, (16) TL, (17) relative tail length (Tol/TL), (18) anal scute, (19) anterior chinshield, (20) dorsal scale rows at midbody, (21) coastals with apical pits, (22) supralabials in contact with eye, (23) frontal, (24) infralabials, (25) internasals, (26) lorials, (27) nasal, (28) parietals, (29) posterior chinshield, (30) post-occulars, (31) preanal, (32) prefrontals, (33) preventrals, (34) subcaudals, (35) subcaudals, (36) supralabials, (37) supraocculars, (38) temporals, (39) ventral marking, (40) ventrals." httpUri="http://table.plazi.org/id/FAED75EE1730FFCD8EE0FDF64FC8CF64" pageId="1" pageNumber="249" tableUuid="FAED75EE1730FFCD8EE0FDF64FC8CF64">Table 1</tableCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E68876ED1738FFC58EE0F9EF4F28CB1A" pageId="1" pageNumber="249" type="materials_examined">
|
||
<paragraph id="AE2D25661738FFC58EE0F9EF4B42CA8E" blockId="1.[151,1437,1619,2042]" pageId="1" pageNumber="249">
|
||
<materialsCitation id="1EFA2F3B1738FFC58EE0F9EF4B42CA8E" ID-GBIF-Occurrence="2574624215" collectingDate="2009-11-05" collectionCode="NMSL-NH" collectorName="Mendis Wickramasinghe & Dulan Ranga Vidanapathirana" country="Sri Lanka" county="Ratnapura District" elevation="655" latitude="6.7950835" location="Uda Pawanalla Police Post" longLatPrecision="1" longitude="80.46122" municipality="Sripada" pageId="1" pageNumber="249" specimenCode="NMSL-NH 2019.26" specimenCount="1" stateProvince="Sabaragamuwa" typeStatus="holotype">
|
||
<emphasis id="9CE6F9741738FFC58EE0F9EF4F4FCA56" bold="true" box="[151,267,1619,1645]" pageId="1" pageNumber="249">
|
||
<typeStatus id="71299BC41738FFC58EE0F9EF4F42CA56" box="[151,262,1619,1645]" pageId="1" pageNumber="249" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<specimenCode id="FE348D1D1738FFC58F62F9EF4FBACA56" box="[277,510,1619,1645]" collectionCode="NMSL-NH" pageId="1" pageNumber="249">NMSL-NH 2019.26</specimenCode>
|
||
.01 (
|
||
<figureCitation id="36A939E31738FFC58C59F9EF4CC4CA56" box="[558,640,1619,1645]" captionStart="FIGURE 2" captionStartId="3.[151,250,1955,1980]" captionTargetBox="[161,1427,1111,1921]" captionTargetId="figure@3.[151,1436,1101,1931]" captionTargetPageId="3" captionText="FIGURE 2. The holotype of Dryocalamus chithrasekarai sp. nov., anterior body (A) dorsal, (B) lateral, (C) ventral aspects; midbody (D) dorsal, (E) lateral, and (F) ventral aspects, respectively; and (G) ventral aspect of tail." figureDoi="http://doi.org/10.5281/zenodo.3698761" httpUri="https://zenodo.org/record/3698761/files/figure.png" pageId="1" pageNumber="249">Figs. 2</figureCitation>
|
||
,
|
||
<figureCitation id="36A939E31738FFC58CF9F9EF4CD9CA56" box="[654,669,1619,1645]" captionStart="FIGURE 3" captionStartId="4.[151,250,1088,1113]" captionTargetBox="[220,1365,203,1042]" captionTargetId="figure@4.[185,1403,181,1063]" captionTargetPageId="4" captionText="FIGURE 3. Dryocalamus chithrasekarai sp. nov., holotype, NMSL-NH 2019.26.01, 328 mm SVL, head scalation: (A) lateral, (B) dorsal, (C) ventral aspects, respectively." figureDoi="http://doi.org/10.5281/zenodo.3698763" httpUri="https://zenodo.org/record/3698763/files/figure.png" pageId="1" pageNumber="249">3</figureCitation>
|
||
), adult female, SVL
|
||
<quantity id="696A88831738FFC58DE5F9EF4DB7CA56" box="[914,1011,1619,1645]" metricMagnitude="-1" metricUnit="m" metricValue="3.28" pageId="1" pageNumber="249" unit="mm" value="328.0">328 mm</quantity>
|
||
,
|
||
<location id="AB4D73BD1738FFC58A76F9EF4B7DCA56" LSID="urn:lsid:plazi:treatment:263B94701738FFC28EE0FA504CBBCA23:AB4D73BD1738FFC58A76F9EF4B7DCA56" box="[1025,1337,1619,1645]" country="Sri Lanka" county="Ratnapura District" latitude="6.7950835" longLatPrecision="1" longitude="80.46122" municipality="Sripada" name="Uda Pawanalla Police Post" pageId="1" pageNumber="249" stateProvince="Sabaragamuwa">Uda Pawanalla Police Post</location>
|
||
,
|
||
<collectingMunicipality id="4E49BF1C1738FFC58B30F9EF4BD9CA56" box="[1351,1437,1619,1645]" pageId="1" pageNumber="249">Sripada</collectingMunicipality>
|
||
(Peak Wilderness),
|
||
<collectingCounty id="474C5DEA1738FFC58F06F9CB4C06CAAA" box="[369,578,1655,1681]" pageId="1" pageNumber="249">Ratnapura District</collectingCounty>
|
||
,
|
||
<collectingRegion id="6C56EB841738FFC58C3BF9CB4D20CAAA" box="[588,868,1655,1681]" country="Sri Lanka" name="Sabaragamuwa" pageId="1" pageNumber="249">Sabaragamuwa Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="D68565F61738FFC58D18F9CB4D98CAAA" box="[879,988,1655,1681]" name="Sri Lanka" pageId="1" pageNumber="249">Sri Lanka</collectingCountry>
|
||
(
|
||
<geoCoordinate id="CBA643A11738FFC58D9CF9CB4AD5CAAA" box="[1003,1169,1655,1681]" degrees="06" direction="north" minutes="47" orientation="latitude" pageId="1" pageNumber="249" precision="1" seconds="42.3" value="6.7950835">06°47’42.3’’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="CBA643A11738FFC58AEBF9CB4B09CAAA" box="[1180,1357,1655,1681]" degrees="080" direction="east" minutes="27" orientation="longitude" pageId="1" pageNumber="249" precision="1" seconds="40.4" value="80.46122">080°27’40.4’’E</geoCoordinate>
|
||
;
|
||
<elevation id="25BFC2551738FFC58B2FF9CB4F08CA8D" metricMagnitude="2" metricUnit="m" metricValue="6.55" pageId="1" pageNumber="249" unit="m" value="655.0">
|
||
elevation
|
||
<quantity id="696A88831738FFC58EBDF9264F55CA8E" box="[202,273,1690,1717]" metricMagnitude="2" metricUnit="m" metricValue="6.55" pageId="1" pageNumber="249" unit="m" value="655.0">655 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
), coll.
|
||
<collectorName id="036740B01738FFC58FECF9274CF6CA8E" box="[411,690,1691,1717]" pageId="1" pageNumber="249">Mendis Wickramasinghe</collectorName>
|
||
&
|
||
<collectorName id="036740B01738FFC58CA1F9274A6DCA8E" box="[726,1065,1691,1717]" pageId="1" pageNumber="249">Dulan Ranga Vidanapathirana</collectorName>
|
||
,
|
||
<date id="DA2C03A61738FFC58A42F9264B47CA8E" box="[1077,1283,1690,1717]" pageId="1" pageNumber="249" value="2009-11-05">
|
||
<collectingDate id="CA68FA4E1738FFC58A42F9264B47CA8E" box="[1077,1283,1690,1717]" pageId="1" pageNumber="249" value="2009-11-05">5 November 2009</collectingDate>
|
||
</date>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="AE2D25661738FFC58EB0F9034F28CB1A" blockId="1.[151,1437,1619,2042]" pageId="1" pageNumber="249">
|
||
<emphasis id="9CE6F9741738FFC58EB0F9034F7FCAE2" bold="true" box="[199,315,1727,1753]" pageId="1" pageNumber="249">
|
||
<typeStatus id="71299BC41738FFC58EB0F9034F72CAE2" box="[199,310,1727,1753]" pageId="1" pageNumber="249" type="paratype">Paratype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
DWC
|
||
<date id="DA2C03A61738FFC58FF9F9024C4FCAE2" box="[398,523,1726,1753]" pageId="1" pageNumber="249" value="2019-05-03">2019.05.03</date>
|
||
(
|
||
<figureCitation id="36A939E31738FFC58C6AF9034C27CAE2" box="[541,611,1727,1753]" captionStart="FIGURE 4" captionStartId="4.[151,250,1980,2005]" captionTargetBox="[191,1397,1179,1946]" captionTargetId="figure@4.[181,1406,1171,1954]" captionTargetPageId="4" captionText="FIGURE 4. The paratype, of Dryocalamus chithrasekarai sp. nov., in life (not collected) showing (A) dorsal, (B) dorsolateral (laterally inverted), and (C) ventral aspects, respectively." figureDoi="http://doi.org/10.5281/zenodo.3698765" httpUri="https://zenodo.org/record/3698765/files/figure.png" pageId="1" pageNumber="249">Fig. 4</figureCitation>
|
||
),
|
||
<materialsCitation id="1EFA2F3B1738FFC58C0DF9034F2DCB1A" ID-GBIF-Occurrence="2574624213" collectingDate="2017-06-18" collectorName="Vishan Pushpamal" country="Sri Lanka" county="Galle" elevation="45" latitude="6.247916" location="Kanneliya" longLatPrecision="1" longitude="80.33016" municipality="Panangala" pageId="1" pageNumber="249" specimenCount="1" stateProvince="Southern" typeStatus="paratype">
|
||
juvenile male
|
||
<quantity id="696A88831738FFC58D6BF9034D3FCAE2" box="[796,891,1727,1753]" metricMagnitude="-1" metricUnit="m" metricValue="1.9899999999999998" pageId="1" pageNumber="249" unit="mm" value="199.0">199 mm</quantity>
|
||
SVL, from
|
||
<location id="AB4D73BD1738FFC58A70F9034A39CAE2" LSID="urn:lsid:plazi:treatment:263B94701738FFC28EE0FA504CBBCA23:AB4D73BD1738FFC58A70F9034A39CAE2" box="[1031,1149,1727,1753]" country="Sri Lanka" county="Galle" latitude="6.247916" longLatPrecision="1" longitude="80.33016" municipality="Panangala" name="Kanneliya" pageId="1" pageNumber="249" stateProvince="Southern">Kanneliya</location>
|
||
,
|
||
<location id="AB4D73BD1738FFC58AFDF9034B52CAE2" LSID="urn:lsid:plazi:treatment:263B94701738FFC28EE0FA504CBBCA23:AB4D73BD1738FFC58AFDF9034B52CAE2" box="[1162,1302,1727,1753]" country="Sri Lanka" county="Galle" latitude="6.247916" longLatPrecision="1" longitude="80.33016" municipality="Panangala" name="Koralegama" pageId="1" pageNumber="249" stateProvince="Southern">Koralegama</location>
|
||
,
|
||
<collectingMunicipality id="4E49BF1C1738FFC58B55F9034BDCCAE2" box="[1314,1432,1727,1753]" pageId="1" pageNumber="249">Panangala</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="474C5DEA1738FFC58EE0F95F4E92CAC6" box="[151,214,1763,1789]" pageId="1" pageNumber="249">Galle</collectingCounty>
|
||
,
|
||
<collectingRegion id="6C56EB841738FFC58E94F95F4FF1CAC6" box="[227,437,1763,1789]" country="Sri Lanka" name="Southern" pageId="1" pageNumber="249">Southern Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="D68565F61738FFC58FB5F95F4C77CAC6" box="[450,563,1763,1789]" name="Sri Lanka" pageId="1" pageNumber="249">Sri Lanka</collectingCountry>
|
||
, (
|
||
<geoCoordinate id="CBA643A11738FFC58C30F95E4CABCAC6" box="[583,751,1762,1789]" degrees="06" direction="north" minutes="14" orientation="latitude" pageId="1" pageNumber="249" precision="1" seconds="52.5" value="6.247916">06º14’52.5” N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="CBA643A11738FFC58C8CF95F4DE8CAC6" box="[763,940,1763,1789]" degrees="080" direction="east" minutes="19" orientation="longitude" pageId="1" pageNumber="249" precision="1" seconds="48.6" value="80.33016">080º19’48.6” E</geoCoordinate>
|
||
;
|
||
<elevation id="25BFC2551738FFC58DCFF95F4AD9CAC5" box="[952,1181,1762,1790]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="249" unit="m" value="45.0">
|
||
elevation
|
||
<quantity id="696A88831738FFC58A50F95E4A25CAC6" box="[1063,1121,1762,1789]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="249" unit="m" value="45.0">45 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
), coll.
|
||
<collectorName id="036740B01738FFC58A9AF95F4E80CB1A" pageId="1" pageNumber="249">Vishan Pushpamal</collectorName>
|
||
,
|
||
<date id="DA2C03A61738FFC58EB8F8BB4F2DCB1A" box="[207,361,1799,1825]" pageId="1" pageNumber="249" value="2017-06-18">
|
||
<collectingDate id="CA68FA4E1738FFC58EB8F8BB4F2DCB1A" box="[207,361,1799,1825]" pageId="1" pageNumber="249" value="2017-06-18">18 June 2017</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E68876ED1738FFC68EB0F8964D4ECCEC" lastPageId="2" lastPageNumber="250" pageId="1" pageNumber="249" type="diagnosis">
|
||
<paragraph id="AE2D25661738FFC68EB0F8964D4ECCEC" blockId="1.[151,1437,1619,2042]" lastBlockId="2.[151,1436,153,215]" lastPageId="2" lastPageNumber="250" pageId="1" pageNumber="249">
|
||
<emphasis id="9CE6F9741738FFC58EB0F8964F06CB7E" bold="true" box="[199,322,1834,1861]" pageId="1" pageNumber="249">Diagnosis.</emphasis>
|
||
We assign the new species to the genus
|
||
<taxonomicName id="69925EE51738FFC58D7FF8974DDBCB7E" authorityName="Gunther" authorityYear="1858" box="[776,927,1835,1861]" class="Reptilia" family="Colubridae" genus="Dryocalamus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="249" phylum="Chordata" rank="genus">
|
||
<emphasis id="9CE6F9741738FFC58D7FF8974DDBCB7E" box="[776,927,1835,1861]" italics="true" pageId="1" pageNumber="249">Dryocalamus</emphasis>
|
||
</taxonomicName>
|
||
because it possesses the following characteristics: costal scales at midbody 13–15; loreal touching eye; broad and keeled ventrals, which are raised on the sides; divided subcaudals, which are raised on the sides.
|
||
<taxonomicName id="69925EE51738FFC58CB2F8CF4A41CBB6" authority="Vidanapathirana & Pushpamal & Wickramasinghe, 2020" authorityName="Vidanapathirana & Pushpamal & Wickramasinghe" authorityYear="2020" box="[709,1029,1907,1933]" class="Reptilia" family="Colubridae" genus="Dryocalamus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="249" phylum="Chordata" rank="species" species="chithrasekarai" status="sp. nov.">
|
||
<emphasis id="9CE6F9741738FFC58CB2F8CF4A41CBB6" box="[709,1029,1907,1933]" italics="true" pageId="1" pageNumber="249">Dryocalamus chithrasekarai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9CE6F9741738FFC58A7BF8CF4A21CBB6" bold="true" box="[1036,1125,1907,1933]" pageId="1" pageNumber="249">
|
||
<taxonomicNameLabel id="87D5440F1738FFC58A7BF8CF4A21CBB6" box="[1036,1125,1907,1933]" pageId="1" pageNumber="249" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, can be easily distinguished from all its congeners by the combination of the following characters: (1) the absence of a preocular scale; (2) loreal in contact with supraocular; (3) costal scales in 15 straight rows; (4) posterior margin of nuchal band appears flat; (5) white solid bands on anterior body, which appear somewhat circular centrally; (6) checkered appearance on posterior lateral body where consecutive bands coalesce at the ventrolateral margin; (7) banding present throughout the anterior venter; and (8) tail region consistently dark.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="FAED75EE173BFFC68EE0FB674AD2C95B" ID-DOI="http://doi.org/10.5281/zenodo.3698757" ID-Zenodo-Dep="3698757" httpUri="https://zenodo.org/record/3698757/files/figure.png" pageId="2" pageNumber="250" startId="2.[151,250,1243,1268]" targetBox="[176,1412,259,1194]" targetPageId="2">
|
||
<paragraph id="AE2D2566173BFFC68EE0FB674AD2C95B" blockId="2.[151,1437,1243,1376]" pageId="2" pageNumber="250">
|
||
<emphasis id="9CE6F974173BFFC68EE0FB674F56C8CF" bold="true" box="[151,274,1243,1268]" pageId="2" pageNumber="250">FIGURE 1.</emphasis>
|
||
Distribution of
|
||
<taxonomicName id="69925EE5173BFFC68FC5FB604C90C8CF" authority="Vidanapathirana & Pushpamal & Wickramasinghe, 2020" authorityName="Vidanapathirana & Pushpamal & Wickramasinghe" authorityYear="2020" box="[434,724,1244,1268]" class="Reptilia" family="Colubridae" genus="Dryocalamus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="250" phylum="Chordata" rank="species" species="chithrasekarai" status="sp. nov.">
|
||
<emphasis id="9CE6F974173BFFC68FC5FB604C90C8CF" box="[434,724,1244,1268]" italics="true" pageId="2" pageNumber="250">Dryocalamus chithrasekarai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9CE6F974173BFFC68CAEFB604D6CC8CF" bold="true" box="[729,808,1244,1268]" pageId="2" pageNumber="250">
|
||
<taxonomicNameLabel id="87D5440F173BFFC68CAEFB604D6CC8CF" box="[729,808,1244,1268]" pageId="2" pageNumber="250" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype (solid white dot), paratype (white circle), specimens illustrated in figures (black solid dots), sight-only records (black circle). Black solid square indicates the type locality (Jaffna) of
|
||
<emphasis id="9CE6F974173BFFC68EE0FA984FC4C907" box="[151,384,1316,1340]" italics="true" pageId="2" pageNumber="250">
|
||
<taxonomicName id="69925EE5173BFFC68EE0FA984F3EC907" authorityName="Haly" authorityYear="1888" box="[151,378,1316,1340]" class="Squamata" family="Colubridae" genus="Odontomus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="2" pageNumber="250" phylum="Chordata" rank="species" species="fergusonii">Odontomus fergusonii</taxonomicName>
|
||
;
|
||
</emphasis>
|
||
the black square indicates Dambulla, and the white square indicates Kurunegala, the only localities from which the only confirmed localities for
|
||
<taxonomicName id="69925EE5173BFFC68C5FFAF44D40C95B" baseAuthorityName="Gunther" baseAuthorityYear="1864" box="[552,772,1352,1376]" class="Reptilia" family="Colubridae" genus="Dryocalamus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="250" phylum="Chordata" rank="species" species="gracilis">
|
||
<emphasis id="9CE6F974173BFFC68C5FFAF44D40C95B" box="[552,772,1352,1376]" italics="true" pageId="2" pageNumber="250">Dryocalamus gracilis</emphasis>
|
||
</taxonomicName>
|
||
has been recorded by previous authors.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="E68876ED173BFFC18EB0FA324F75CD78" lastPageId="5" lastPageNumber="253" pageId="2" pageNumber="250" type="description">
|
||
<paragraph id="AE2D2566173BFFC68EB0FA324A4CC9CB" blockId="2.[151,1437,1422,2025]" pageId="2" pageNumber="250">
|
||
<emphasis id="9CE6F974173BFFC68EB0FA324FA0C992" bold="true" box="[199,484,1422,1449]" pageId="2" pageNumber="250">
|
||
Description of
|
||
<typeStatus id="71299BC4173BFFC68F01FA334F9BC992" box="[374,479,1423,1449]" pageId="2" pageNumber="250" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
The
|
||
<typeStatus id="71299BC4173BFFC68C6AFA324C3BC993" box="[541,639,1422,1448]" pageId="2" pageNumber="250" type="holotype">holotype</typeStatus>
|
||
is an adult female with a snout-vent length of
|
||
<quantity id="696A8883173BFFC68AF1FA334AA7C992" box="[1158,1251,1423,1449]" metricMagnitude="-1" metricUnit="m" metricValue="3.28" pageId="2" pageNumber="250" unit="mm" value="328.0">328 mm</quantity>
|
||
and a tail length of
|
||
<quantity id="696A8883173BFFC68EC2FA0F4F51C9F6" box="[181,277,1459,1485]" metricMagnitude="-1" metricUnit="m" metricValue="1.08" pageId="2" pageNumber="250" unit="mm" value="108.0">108 mm</quantity>
|
||
. Head oval in shape (
|
||
<figureCitation id="36A939E3173BFFC68C74FA0E4C19C9F6" box="[515,605,1458,1485]" captionStart="FIGURE 2" captionStartId="3.[151,250,1955,1980]" captionTargetBox="[161,1427,1111,1921]" captionTargetId="figure@3.[151,1436,1101,1931]" captionTargetPageId="3" captionText="FIGURE 2. The holotype of Dryocalamus chithrasekarai sp. nov., anterior body (A) dorsal, (B) lateral, (C) ventral aspects; midbody (D) dorsal, (E) lateral, and (F) ventral aspects, respectively; and (G) ventral aspect of tail." figureDoi="http://doi.org/10.5281/zenodo.3698761" httpUri="https://zenodo.org/record/3698761/files/figure.png" pageId="2" pageNumber="250">Fig. 2A</figureCitation>
|
||
), depressed (
|
||
<figureCitation id="36A939E3173BFFC68C99FA0E4D01C9F6" box="[750,837,1458,1485]" captionStart="FIGURE 2" captionStartId="3.[151,250,1955,1980]" captionTargetBox="[161,1427,1111,1921]" captionTargetId="figure@3.[151,1436,1101,1931]" captionTargetPageId="3" captionText="FIGURE 2. The holotype of Dryocalamus chithrasekarai sp. nov., anterior body (A) dorsal, (B) lateral, (C) ventral aspects; midbody (D) dorsal, (E) lateral, and (F) ventral aspects, respectively; and (G) ventral aspect of tail." figureDoi="http://doi.org/10.5281/zenodo.3698761" httpUri="https://zenodo.org/record/3698761/files/figure.png" pageId="2" pageNumber="250">Fig. 2B</figureCitation>
|
||
), distinct from neck (
|
||
<figureCitation id="36A939E3173BFFC68A44FA0E4A88C9F6" box="[1075,1228,1458,1485]" captionStart="FIGURE 2" captionStartId="3.[151,250,1955,1980]" captionTargetBox="[161,1427,1111,1921]" captionTargetId="figure@3.[151,1436,1101,1931]" captionTargetPageId="3" captionText="FIGURE 2. The holotype of Dryocalamus chithrasekarai sp. nov., anterior body (A) dorsal, (B) lateral, (C) ventral aspects; midbody (D) dorsal, (E) lateral, and (F) ventral aspects, respectively; and (G) ventral aspect of tail." figureDoi="http://doi.org/10.5281/zenodo.3698761" httpUri="https://zenodo.org/record/3698761/files/figure.png" pageId="2" pageNumber="250">Figs. 2A & C</figureCitation>
|
||
); snout rounded in dorsal, lateral and ventral views. Body slender, subcylindrical in cross-section.
|
||
</paragraph>
|
||
<paragraph id="AE2D2566173BFFC78EB0FA464CD6CD78" blockId="2.[151,1437,1422,2025]" lastBlockId="3.[151,1437,152,1079]" lastPageId="3" lastPageNumber="251" pageId="2" pageNumber="250">
|
||
Dorsals smooth, in 15/15/15 straight rows, of uniform shape (almost as long as wide, rhomboid); vertebral scale row (
|
||
<figureCitation id="36A939E3173BFFC68F64F9A24F2FCA02" box="[275,363,1566,1593]" captionStart="FIGURE 2" captionStartId="3.[151,250,1955,1980]" captionTargetBox="[161,1427,1111,1921]" captionTargetId="figure@3.[151,1436,1101,1931]" captionTargetPageId="3" captionText="FIGURE 2. The holotype of Dryocalamus chithrasekarai sp. nov., anterior body (A) dorsal, (B) lateral, (C) ventral aspects; midbody (D) dorsal, (E) lateral, and (F) ventral aspects, respectively; and (G) ventral aspect of tail." figureDoi="http://doi.org/10.5281/zenodo.3698761" httpUri="https://zenodo.org/record/3698761/files/figure.png" pageId="2" pageNumber="250">Fig. 2D</figureCitation>
|
||
) and outer row not enlarged (
|
||
<figureCitation id="36A939E3173BFFC68CCBF9A24D50CA02" box="[700,788,1566,1593]" captionStart="FIGURE 2" captionStartId="3.[151,250,1955,1980]" captionTargetBox="[161,1427,1111,1921]" captionTargetId="figure@3.[151,1436,1101,1931]" captionTargetPageId="3" captionText="FIGURE 2. The holotype of Dryocalamus chithrasekarai sp. nov., anterior body (A) dorsal, (B) lateral, (C) ventral aspects; midbody (D) dorsal, (E) lateral, and (F) ventral aspects, respectively; and (G) ventral aspect of tail." figureDoi="http://doi.org/10.5281/zenodo.3698761" httpUri="https://zenodo.org/record/3698761/files/figure.png" pageId="2" pageNumber="250">Fig. 2E</figureCitation>
|
||
); individual apical pits prominent on the three scale rows on mid-dorsum between second and third white crossbars from head (weaker towards posterior body); 1 preventral, 228 keeled ventrals (
|
||
<figureCitation id="36A939E3173BFFC68FF3F9DA4F9ECABA" box="[388,474,1638,1665]" captionStart="FIGURE 2" captionStartId="3.[151,250,1955,1980]" captionTargetBox="[161,1427,1111,1921]" captionTargetId="figure@3.[151,1436,1101,1931]" captionTargetPageId="3" captionText="FIGURE 2. The holotype of Dryocalamus chithrasekarai sp. nov., anterior body (A) dorsal, (B) lateral, (C) ventral aspects; midbody (D) dorsal, (E) lateral, and (F) ventral aspects, respectively; and (G) ventral aspect of tail." figureDoi="http://doi.org/10.5281/zenodo.3698761" httpUri="https://zenodo.org/record/3698761/files/figure.png" pageId="2" pageNumber="250">Fig. 2F</figureCitation>
|
||
), raised on the sides; cloacal shield divided; 99 divided subcaudals (Fig, 2G); caudal reduction: 11 to 7 (from the 2
|
||
<superScript id="59E7882E173BFFC68FABF9354FA8CAAC" attach="left" box="[476,492,1673,1687]" fontSize="6" pageId="2" pageNumber="250">nd</superScript>
|
||
SC), to 6 (from the 6
|
||
<superScript id="59E7882E173BFFC68CA0F9354CA0CAAC" attach="left" box="[727,740,1673,1687]" fontSize="6" pageId="2" pageNumber="250">th</superScript>
|
||
SC), to 5 (from the 22
|
||
<superScript id="59E7882E173BFFC68DAAF9354DA9CAAC" attach="left" box="[989,1005,1673,1687]" fontSize="6" pageId="2" pageNumber="250">nd</superScript>
|
||
(L) and 21
|
||
<superScript id="59E7882E173BFFC68A10F9364A36CAA3" attach="left" box="[1127,1138,1674,1688]" fontSize="6" pageId="2" pageNumber="250">st</superScript>
|
||
(R) SC) to 4 (from the 24
|
||
<superScript id="59E7882E173BFFC68BE7F9354BD9CAAC" attach="right" box="[1424,1437,1673,1687]" fontSize="6" pageId="2" pageNumber="250">th</superScript>
|
||
SC), to 3 (from the 65
|
||
<superScript id="59E7882E173BFFC68FE7F9114FD9CA80" attach="left" box="[400,413,1709,1723]" fontSize="6" pageId="2" pageNumber="250">th</superScript>
|
||
SC), to 2 (from the 67
|
||
<superScript id="59E7882E173BFFC68CE9F9114CEFCA80" attach="left" box="[670,683,1709,1723]" fontSize="6" pageId="2" pageNumber="250">th</superScript>
|
||
SC); 7/7 supralabials, the 3
|
||
<superScript id="59E7882E173BFFC68D93F9114DB6CA80" attach="left" box="[996,1010,1709,1723]" fontSize="6" pageId="2" pageNumber="250">rd</superScript>
|
||
and 4
|
||
<superScript id="59E7882E173BFFC68A41F9114A07CA80" attach="left" box="[1078,1091,1709,1723]" fontSize="6" pageId="2" pageNumber="250">th</superScript>
|
||
in contact with the eye, the 6
|
||
<superScript id="59E7882E173BFFC68BE7F9114BD9CA80" attach="none" box="[1424,1437,1709,1723]" fontSize="6" pageId="2" pageNumber="250">th</superScript>
|
||
being longest in longitudinal axis, the 7
|
||
<superScript id="59E7882E173BFFC68C27F96D4C19CAE4" attach="left" box="[592,605,1745,1759]" fontSize="6" pageId="2" pageNumber="250">th</superScript>
|
||
widest in the transverse axis; 1/1 loreals, length equal to height, in contact with nasal, prefrontal, supraocular, eye, and 2
|
||
<superScript id="59E7882E173BFFC68CF9F9494CDACB38" attach="left" box="[654,670,1781,1795]" fontSize="6" pageId="2" pageNumber="250">nd</superScript>
|
||
and 3
|
||
<superScript id="59E7882E173BFFC68C96F9494CABCB38" attach="left" box="[737,751,1781,1795]" fontSize="6" pageId="2" pageNumber="250">rd</superScript>
|
||
supralabials; no preocular; nasal divided, with suture meeting with first supralabial; naris in prenasal; 2/2 postoculars, lower one larger than upper one, which is in contact with supraocular, parietal and upper anterior temporal, while lower one touches the two anterior temporals and 4
|
||
<superScript id="59E7882E173BFFC68B28F8814B28CB70" attach="left" box="[1375,1388,1853,1867]" fontSize="6" pageId="2" pageNumber="250">th</superScript>
|
||
and 5
|
||
<superScript id="59E7882E173BFFC68ED2F8DD4EF6CB54" attach="left" box="[165,178,1889,1903]" fontSize="6" pageId="2" pageNumber="250">th</superScript>
|
||
supralabials; 2+2 / 2+2 temporals (
|
||
<figureCitation id="36A939E3173BFFC68C48F8DE4CDDCB46" box="[575,665,1890,1917]" captionStart="FIGURE 3" captionStartId="4.[151,250,1088,1113]" captionTargetBox="[220,1365,203,1042]" captionTargetId="figure@4.[185,1403,181,1063]" captionTargetPageId="4" captionText="FIGURE 3. Dryocalamus chithrasekarai sp. nov., holotype, NMSL-NH 2019.26.01, 328 mm SVL, head scalation: (A) lateral, (B) dorsal, (C) ventral aspects, respectively." figureDoi="http://doi.org/10.5281/zenodo.3698763" httpUri="https://zenodo.org/record/3698763/files/figure.png" pageId="2" pageNumber="250">Fig. 3A</figureCitation>
|
||
); rostral distinctly wider than high, visible in dorsal view; single pair of internasals, wider than long, shorter than prefrontals; single pair of prefrontals, wider than long, extending far onto sides of head to about level of center of eye; frontal pentagonal, longer than wide, longer than its distance from rostral, but shorter than parietals; supraoculars narrow, smaller than frontal; a single pair of parietals, longer than wide (
|
||
<figureCitation id="36A939E3173AFFC78EABFF254F77CC88" box="[220,307,153,179]" captionStart="FIGURE 3" captionStartId="4.[151,250,1088,1113]" captionTargetBox="[220,1365,203,1042]" captionTargetId="figure@4.[185,1403,181,1063]" captionTargetPageId="4" captionText="FIGURE 3. Dryocalamus chithrasekarai sp. nov., holotype, NMSL-NH 2019.26.01, 328 mm SVL, head scalation: (A) lateral, (B) dorsal, (C) ventral aspects, respectively." figureDoi="http://doi.org/10.5281/zenodo.3698763" httpUri="https://zenodo.org/record/3698763/files/figure.png" pageId="3" pageNumber="251">Fig. 3B</figureCitation>
|
||
); 8/8 infralabials, first pair in contact behind mental, first four touching anterior chinshield, 4
|
||
<superScript id="59E7882E173AFFC78B37FF244B09CC9D" attach="left" box="[1344,1357,152,166]" fontSize="6" pageId="3" pageNumber="251">th</superScript>
|
||
and 5
|
||
<superScript id="59E7882E173AFFC78BE7FF244BD9CC9D" attach="none" box="[1424,1437,152,166]" fontSize="6" pageId="3" pageNumber="251">th</superScript>
|
||
in contact with posterior chinshield, 5
|
||
<superScript id="59E7882E173AFFC78C33FF004C15CCF1" attach="left" box="[580,593,188,202]" fontSize="6" pageId="3" pageNumber="251">th</superScript>
|
||
largest; 2 pairs of chinshields, anterior pair longer and slightly wider than posterior pair; no invasive gulars between the chinshields (
|
||
<figureCitation id="36A939E3173AFFC78D52FF5D4D39CCC0" box="[805,893,225,251]" captionStart="FIGURE 3" captionStartId="4.[151,250,1088,1113]" captionTargetBox="[220,1365,203,1042]" captionTargetId="figure@4.[185,1403,181,1063]" captionTargetPageId="4" captionText="FIGURE 3. Dryocalamus chithrasekarai sp. nov., holotype, NMSL-NH 2019.26.01, 328 mm SVL, head scalation: (A) lateral, (B) dorsal, (C) ventral aspects, respectively." figureDoi="http://doi.org/10.5281/zenodo.3698763" httpUri="https://zenodo.org/record/3698763/files/figure.png" pageId="3" pageNumber="251">Fig. 3C</figureCitation>
|
||
); eye large, with slightly vertical-elliptical pupil, diameter of eye less than distance between anterior margin of eye and tip of snout, eye diameter slightly greater than distance from anterior margin of eye to naris.
|
||
</paragraph>
|
||
<paragraph id="AE2D2566173AFFC78EB0FEF14ABDCE90" blockId="3.[151,1437,152,1079]" pageId="3" pageNumber="251">
|
||
Colour in alcohol (
|
||
<typeStatus id="71299BC4173AFFC78FE8FEF14C4CCD53" box="[415,520,333,360]" pageId="3" pageNumber="251" type="holotype">holotype</typeStatus>
|
||
). Except nuchal band, which covers posterior third of parietals, entire head light brown dorsally. Ground colour light brown, with cream-coloured bands (32 on body, not counting nuchal band, 22 on tail). In anterior 18 bands, cream colour is solid, with outer margins scalloped and wider ventrolaterally, centrally with a somewhat circular appearance. Bands posterior to 19
|
||
<superScript id="59E7882E173AFFC78D4FFE044D01CDFD" attach="left" box="[824,837,440,454]" fontSize="6" pageId="3" pageNumber="251">th</superScript>
|
||
cream band exhibit a dark blotch within the widened base, where adjacent bands coalesce at ventrolateral margin, giving a checkered appearance laterally. The light bands are 2½ scales wide anteriorly, at level of vertebral scale row, decreasing to 2 scales wide towards base of tail. Width of light-brown interspaces decreases gradually caudad: 11 dorsals in vertebral row in the first dark interspace between nuchal band and first cream band, decreasing to 4 dorsals at tail base and from there to tip of tail. Dark areas reaching ventrals in anterior body: hence venter in that region appearing banded. Entire venter colouration cream with darker interspaces beginning in nuchal region; tail venter bears light brown speckles, no bands.
|
||
</paragraph>
|
||
<paragraph id="AE2D2566173AFFC78EB0FD094CBCC80C" blockId="3.[151,1437,152,1079]" pageId="3" pageNumber="251">
|
||
Colour in life (
|
||
<typeStatus id="71299BC4173AFFC78F04FD094F9ECEF4" box="[371,474,693,719]" pageId="3" pageNumber="251" type="paratype">paratype</typeStatus>
|
||
). Except for nuchal band, which covers posterior third of parietals, entire head dark brown dorsally. Ground colour jet black, with pure-white bands (32 on body omitting nuchal band, 21 on tail) (
|
||
<figureCitation id="36A939E3173AFFC78B18FD654EE2CF2C" captionStart="FIGURE 4" captionStartId="4.[151,250,1980,2005]" captionTargetBox="[191,1397,1179,1946]" captionTargetId="figure@4.[181,1406,1171,1954]" captionTargetPageId="4" captionText="FIGURE 4. The paratype, of Dryocalamus chithrasekarai sp. nov., in life (not collected) showing (A) dorsal, (B) dorsolateral (laterally inverted), and (C) ventral aspects, respectively." figureDoi="http://doi.org/10.5281/zenodo.3698765" httpUri="https://zenodo.org/record/3698765/files/figure.png" pageId="3" pageNumber="251">Fig. 4</figureCitation>
|
||
A–B). In anteriormost 18 white bands, the white band is solid, with band-margins scalloped and wider ventrolaterally (
|
||
<figureCitation id="36A939E3173AFFC78EA5FC9D4F68CF00" box="[210,300,801,827]" captionStart="FIGURE 4" captionStartId="4.[151,250,1980,2005]" captionTargetBox="[191,1397,1179,1946]" captionTargetId="figure@4.[181,1406,1171,1954]" captionTargetPageId="4" captionText="FIGURE 4. The paratype, of Dryocalamus chithrasekarai sp. nov., in life (not collected) showing (A) dorsal, (B) dorsolateral (laterally inverted), and (C) ventral aspects, respectively." figureDoi="http://doi.org/10.5281/zenodo.3698765" httpUri="https://zenodo.org/record/3698765/files/figure.png" pageId="3" pageNumber="251">Fig. 4B</figureCitation>
|
||
), centrally with a somewhat circular appearance. While posterior bands (caudad of 19
|
||
<superScript id="59E7882E173AFFC78B77FC9C4B49CF15" attach="left" box="[1280,1293,800,814]" fontSize="6" pageId="3" pageNumber="251">th</superScript>
|
||
white band) exhibit a dark blotch within the widened base, where consecutive bands coalesce at ventrolateral margin, resulting in a checkered appearance laterally. The light bands 2½ scales wide at level of vertebral scale row anteriorly, decreasing to 2 scales wide towards base of tail. Width of dark interspaces decreasing gradually caudad: 14 dorsals in vertebral row in first dark interspace between the nuchal band and first white band, declining to 4 dorsals from tail base to tip of tail. The dark interspaces reaching ventrals, giving venter a banded appearance. The entire venter colouration is greyish-white, producing a clouded appearance with darker interspaces beginning in nuchal region, while tail is consistently dark with no bands (
|
||
<figureCitation id="36A939E3173AFFC78CE4FBA14CA8C80C" box="[659,748,1053,1079]" captionStart="FIGURE 4" captionStartId="4.[151,250,1980,2005]" captionTargetBox="[191,1397,1179,1946]" captionTargetId="figure@4.[181,1406,1171,1954]" captionTargetPageId="4" captionText="FIGURE 4. The paratype, of Dryocalamus chithrasekarai sp. nov., in life (not collected) showing (A) dorsal, (B) dorsolateral (laterally inverted), and (C) ventral aspects, respectively." figureDoi="http://doi.org/10.5281/zenodo.3698765" httpUri="https://zenodo.org/record/3698765/files/figure.png" pageId="3" pageNumber="251">Fig. 4C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="FAED75EE173AFFC78EE0F81F4AC7CBDB" ID-DOI="http://doi.org/10.5281/zenodo.3698761" ID-Zenodo-Dep="3698761" httpUri="https://zenodo.org/record/3698761/files/figure.png" pageId="3" pageNumber="251" startId="3.[151,250,1955,1980]" targetBox="[161,1427,1111,1921]" targetPageId="3">
|
||
<paragraph id="AE2D2566173AFFC78EE0F81F4AC7CBDB" blockId="3.[151,1436,1955,2016]" pageId="3" pageNumber="251">
|
||
<emphasis id="9CE6F974173AFFC78EE0F81F4F50CB87" bold="true" box="[151,276,1955,1980]" pageId="3" pageNumber="251">FIGURE 2.</emphasis>
|
||
The holotype of
|
||
<taxonomicName id="69925EE5173AFFC78FBEF8184CA9CB87" authority="Vidanapathirana & Pushpamal & Wickramasinghe, 2020" authorityName="Vidanapathirana & Pushpamal & Wickramasinghe" authorityYear="2020" box="[457,749,1956,1980]" class="Reptilia" family="Colubridae" genus="Dryocalamus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="251" phylum="Chordata" rank="species" species="chithrasekarai" status="sp. nov.">
|
||
<emphasis id="9CE6F974173AFFC78FBEF8184CA9CB87" box="[457,749,1956,1980]" italics="true" pageId="3" pageNumber="251">Dryocalamus chithrasekarai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9CE6F974173AFFC78C82F8184D03CB87" bold="true" box="[757,839,1956,1980]" pageId="3" pageNumber="251">
|
||
<taxonomicNameLabel id="87D5440F173AFFC78C82F8184D03CB87" box="[757,839,1956,1980]" pageId="3" pageNumber="251" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, anterior body (A) dorsal, (B) lateral, (C) ventral aspects; midbody (D) dorsal, (E) lateral, and (F) ventral aspects, respectively; and (G) ventral aspect of tail.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="FAED75EE173DFFC08EE0FBFC4C11C846" ID-DOI="http://doi.org/10.5281/zenodo.3698763" ID-Zenodo-Dep="3698763" httpUri="https://zenodo.org/record/3698763/files/figure.png" pageId="4" pageNumber="252" startId="4.[151,250,1088,1113]" targetBox="[220,1365,203,1042]" targetPageId="4">
|
||
<paragraph id="AE2D2566173DFFC08EE0FBFC4C11C846" blockId="4.[151,1436,1088,1149]" pageId="4" pageNumber="252">
|
||
<emphasis id="9CE6F974173DFFC08EE0FBFC4F57C862" bold="true" box="[151,275,1088,1113]" pageId="4" pageNumber="252">FIGURE 3.</emphasis>
|
||
<taxonomicName id="69925EE5173DFFC08F6EFBFD4C7FC862" authority="Vidanapathirana & Pushpamal & Wickramasinghe, 2020" authorityName="Vidanapathirana & Pushpamal & Wickramasinghe" authorityYear="2020" box="[281,571,1089,1113]" class="Reptilia" family="Colubridae" genus="Dryocalamus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="252" phylum="Chordata" rank="species" species="chithrasekarai" status="sp. nov.">
|
||
<emphasis id="9CE6F974173DFFC08F6EFBFD4C7FC862" box="[281,571,1089,1113]" italics="true" pageId="4" pageNumber="252">Dryocalamus chithrasekarai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9CE6F974173DFFC08C36FBFD4CD5C862" bold="true" box="[577,657,1089,1113]" pageId="4" pageNumber="252">
|
||
<taxonomicNameLabel id="87D5440F173DFFC08C36FBFD4CD5C862" box="[577,657,1089,1113]" pageId="4" pageNumber="252" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, NMSL-NH 2019.26.01, 328 mm SVL, head scalation: (A) lateral, (B) dorsal, (C) ventral aspects, respectively.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="FAED75EE173DFFC08EE0F8004C93CBC2" ID-DOI="http://doi.org/10.5281/zenodo.3698765" ID-Zenodo-Dep="3698765" httpUri="https://zenodo.org/record/3698765/files/figure.png" pageId="4" pageNumber="252" startId="4.[151,250,1980,2005]" targetBox="[191,1397,1179,1946]" targetPageId="4">
|
||
<paragraph id="AE2D2566173DFFC08EE0F8004C93CBC2" blockId="4.[151,1437,1980,2041]" pageId="4" pageNumber="252">
|
||
<emphasis id="9CE6F974173DFFC08EE0F8004F57CBEE" bold="true" box="[151,275,1980,2005]" pageId="4" pageNumber="252">FIGURE 4.</emphasis>
|
||
The paratype, of
|
||
<taxonomicName id="69925EE5173DFFC08FBFF8004CAFCBEF" authority="Vidanapathirana & Pushpamal & Wickramasinghe, 2020" authorityName="Vidanapathirana & Pushpamal & Wickramasinghe" authorityYear="2020" box="[456,747,1980,2004]" class="Reptilia" family="Colubridae" genus="Dryocalamus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="252" phylum="Chordata" rank="species" species="chithrasekarai" status="sp. nov.">
|
||
<emphasis id="9CE6F974173DFFC08FBFF8004CAFCBEF" box="[456,747,1980,2004]" italics="true" pageId="4" pageNumber="252">Dryocalamus chithrasekarai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9CE6F974173DFFC08C86F8004D06CBEF" bold="true" box="[753,834,1980,2004]" pageId="4" pageNumber="252">
|
||
<taxonomicNameLabel id="87D5440F173DFFC08C86F8004D06CBEF" box="[753,834,1980,2004]" pageId="4" pageNumber="252" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, in life (not collected) showing (A) dorsal, (B) dorsolateral (laterally inverted), and (C) ventral aspects, respectively.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="AE2D2566173CFFC18EB0FF254F75CD78" blockId="5.[151,1437,153,828]" pageId="5" pageNumber="253">
|
||
<emphasis id="9CE6F974173CFFC18EB0FF254F75CC88" bold="true" box="[199,305,153,179]" pageId="5" pageNumber="253">Notes on</emphasis>
|
||
<materialsCitation id="1EFA2F3B173CFFC18F4EFF254F69CD78" ID-GBIF-Occurrence="2574624214" collectionCode="R" pageId="5" pageNumber="253" specimenCount="1" typeStatus="paratype">
|
||
<emphasis id="9CE6F974173CFFC18F4EFF254FEFCC88" bold="true" box="[313,427,153,179]" pageId="5" pageNumber="253">
|
||
<typeStatus id="71299BC4173CFFC18F4EFF254FE2CC88" box="[313,422,153,179]" pageId="5" pageNumber="253" type="paratype">paratype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
The right eye was absent in the live juvenile male specimen at the time of collection. A
|
||
<quantity id="696A8883173CFFC18EE0FF014EA3CCEC" box="[151,231,189,215]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="253" unit="mm" value="12.0">12 mm</quantity>
|
||
length of the tail was snipped off and preserved in ‘DNA-grade’ ethanol for future molecular studies. The preserved specimen contains 15 white bands on the tail, and 74 (L)/ 75 (
|
||
<collectionCode id="C883BDA3173CFFC18DBEFF5D4D9ACCC0" box="[969,990,225,251]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="253">R</collectionCode>
|
||
) SC; 8/8 supralabials with the 6
|
||
<superScript id="59E7882E173CFFC18B3DFF5C4B13CCD5" attach="left" box="[1354,1367,224,238]" fontSize="6" pageId="5" pageNumber="253">th</superScript>
|
||
being the longest, and 7
|
||
<superScript id="59E7882E173CFFC18F2BFEB84F2DCD29" attach="left" box="[348,361,260,274]" fontSize="6" pageId="5" pageNumber="253">th</superScript>
|
||
the widest; 8/7 infralabials where the 6
|
||
<superScript id="59E7882E173CFFC18D55FEB84D6BCD29" attach="left" box="[802,815,260,274]" fontSize="6" pageId="5" pageNumber="253">th</superScript>
|
||
and 7
|
||
<superScript id="59E7882E173CFFC18D04FEB84DC4CD29" attach="left" box="[883,896,260,274]" fontSize="6" pageId="5" pageNumber="253">th</superScript>
|
||
scales have been fused to form a single scale on the right side
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E68876ED173CFFC28EB0FEF14A19CD1B" lastPageId="6" lastPageNumber="254" pageId="5" pageNumber="253" type="etymology">
|
||
<paragraph id="AE2D2566173CFFC18EB0FEF14F74CD94" blockId="5.[151,1437,153,828]" pageId="5" pageNumber="253">
|
||
<emphasis id="9CE6F974173CFFC18EB0FEF14F0ACD5C" bold="true" box="[199,334,333,359]" pageId="5" pageNumber="253">Etymology.</emphasis>
|
||
The species is named for Nagamulla Hewage Chithrasekara, in recognition of his efforts to protect the Kanneliya Forest Reserve, where the
|
||
<typeStatus id="71299BC4173CFFC18C13FECD4C80CDB0" box="[612,708,369,395]" pageId="5" pageNumber="253" type="paratype">paratype</typeStatus>
|
||
was collected. The species epithet is a patronym Latinized in the genitive case.
|
||
</paragraph>
|
||
<paragraph id="AE2D2566173CFFC18EB0FE044B27CDCC" blockId="5.[151,1437,153,828]" pageId="5" pageNumber="253">
|
||
<emphasis id="9CE6F974173CFFC18EB0FE044C60CDE8" bold="true" box="[199,548,440,467]" pageId="5" pageNumber="253">Suggested vernacular names.</emphasis>
|
||
The vernacular names recommended for the species are ‘Chithrasekarage gata radanakaya’ and Chithrasekara’s bridle snake in the languages Sinhala (vernacular) and English, respectively.
|
||
</paragraph>
|
||
<paragraph id="AE2D2566173CFFC18EB0FDBC4FF4CF07" blockId="5.[151,1437,153,828]" pageId="5" pageNumber="253">
|
||
<emphasis id="9CE6F974173CFFC18EB0FDBC4F29CE20" bold="true" box="[199,365,512,539]" pageId="5" pageNumber="253">Comparisons.</emphasis>
|
||
<taxonomicName id="69925EE5173CFFC18F02FDBD4CF3CE20" authority="Vidanapathirana & Pushpamal & Wickramasinghe, 2020" authorityName="Vidanapathirana & Pushpamal & Wickramasinghe" authorityYear="2020" box="[373,695,513,539]" class="Reptilia" family="Colubridae" genus="Dryocalamus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="253" phylum="Chordata" rank="species" species="chithrasekarai" status="sp. nov.">
|
||
<emphasis id="9CE6F974173CFFC18F02FDBD4CF3CE20" box="[373,695,513,539]" italics="true" pageId="5" pageNumber="253">Dryocalamus chithrasekarai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9CE6F974173CFFC18CB7FDBD4D5FCE20" bold="true" box="[704,795,513,539]" pageId="5" pageNumber="253">
|
||
<taxonomicNameLabel id="87D5440F173CFFC18CB7FDBD4D5FCE20" box="[704,795,513,539]" pageId="5" pageNumber="253" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, is distinguished from its congeners in
|
||
<collectingCountry id="D68565F6173CFFC18AAAFDBD4B08CE20" box="[1245,1356,513,539]" name="Sri Lanka" pageId="5" pageNumber="253">Sri Lanka</collectingCountry>
|
||
as follows: It differs from
|
||
<taxonomicName id="69925EE5173CFFC18FF6FD9A4FBDCE04" baseAuthorityName="Gunther" baseAuthorityYear="1864" box="[385,505,549,575]" class="Reptilia" family="Colubridae" genus="Dryocalamus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="253" phylum="Chordata" rank="species" species="gracilis">
|
||
<emphasis id="9CE6F974173CFFC18FF6FD9A4FBDCE04" box="[385,505,549,575]" italics="true" pageId="5" pageNumber="253">D. gracilis</emphasis>
|
||
</taxonomicName>
|
||
by the absence (vs. presence) of a preocular scale; having the loreal in contact (vs. not in contact) with the supraocular (
|
||
<figureCitation id="36A939E3173CFFC18C40FDF54CDFCE58" box="[567,667,585,611]" captionStart="FIGURE 3" captionStartId="4.[151,250,1088,1113]" captionTargetBox="[220,1365,203,1042]" captionTargetId="figure@4.[185,1403,181,1063]" captionTargetPageId="4" captionText="FIGURE 3. Dryocalamus chithrasekarai sp. nov., holotype, NMSL-NH 2019.26.01, 328 mm SVL, head scalation: (A) lateral, (B) dorsal, (C) ventral aspects, respectively." figureDoi="http://doi.org/10.5281/zenodo.3698763" httpUri="https://zenodo.org/record/3698763/files/figure.png" pageId="5" pageNumber="253">Figs. 3A</figureCitation>
|
||
,
|
||
<figureCitation id="36A939E3173CFFC18CDFFDF44C8ACE58" box="[680,718,584,611]" captionStart="FIGURE 5" captionStartId="5.[151,250,1827,1852]" captionTargetBox="[162,1425,861,1790]" captionTargetId="figure@5.[151,1436,849,1802]" captionTargetPageId="5" captionText="FIGURE 5. Dryocalamus gracilis holotype, BMNH 1946.1.13.86, 403 mm SVL: (A) lateral head; (B) dorsal, and (C) ventral aspects, respectively." figureDoi="http://doi.org/10.5281/zenodo.3698767" httpUri="https://zenodo.org/record/3698767/files/figure.png" pageId="5" pageNumber="253">5A</figureCitation>
|
||
); the cloacal shield divided (vs. undivided); posterior margin of the nuchal band flat (vs. convex) banding present throughout the anterior venter (vs. not banded, uniform creamy white); and the tail region consistently dark (vs. creamy white) (
|
||
<figureCitation id="36A939E3173CFFC18D16FD2D4DF4CE90" box="[865,944,657,683]" captionStart="FIGURE 2" captionStartId="3.[151,250,1955,1980]" captionTargetBox="[161,1427,1111,1921]" captionTargetId="figure@3.[151,1436,1101,1931]" captionTargetPageId="3" captionText="FIGURE 2. The holotype of Dryocalamus chithrasekarai sp. nov., anterior body (A) dorsal, (B) lateral, (C) ventral aspects; midbody (D) dorsal, (E) lateral, and (F) ventral aspects, respectively; and (G) ventral aspect of tail." figureDoi="http://doi.org/10.5281/zenodo.3698761" httpUri="https://zenodo.org/record/3698761/files/figure.png" pageId="5" pageNumber="253">Figs. 2</figureCitation>
|
||
,
|
||
<figureCitation id="36A939E3173CFFC18DCBFD2D4D88CE90" box="[956,972,657,683]" captionStart="FIGURE 4" captionStartId="4.[151,250,1980,2005]" captionTargetBox="[191,1397,1179,1946]" captionTargetId="figure@4.[181,1406,1171,1954]" captionTargetPageId="4" captionText="FIGURE 4. The paratype, of Dryocalamus chithrasekarai sp. nov., in life (not collected) showing (A) dorsal, (B) dorsolateral (laterally inverted), and (C) ventral aspects, respectively." figureDoi="http://doi.org/10.5281/zenodo.3698765" httpUri="https://zenodo.org/record/3698765/files/figure.png" pageId="5" pageNumber="253">4</figureCitation>
|
||
,
|
||
<figureCitation id="36A939E3173CFFC18DA0FD2C4DA0CE90" box="[983,996,656,683]" captionStart="FIGURE 5" captionStartId="5.[151,250,1827,1852]" captionTargetBox="[162,1425,861,1790]" captionTargetId="figure@5.[151,1436,849,1802]" captionTargetPageId="5" captionText="FIGURE 5. Dryocalamus gracilis holotype, BMNH 1946.1.13.86, 403 mm SVL: (A) lateral head; (B) dorsal, and (C) ventral aspects, respectively." figureDoi="http://doi.org/10.5281/zenodo.3698767" httpUri="https://zenodo.org/record/3698767/files/figure.png" pageId="5" pageNumber="253">5</figureCitation>
|
||
B–C). The new species is distinguished from
|
||
<taxonomicName id="69925EE5173CFFC18EA2FD0A4F14CEF4" baseAuthorityName="Daudin" baseAuthorityYear="1803" box="[213,336,693,719]" class="Reptilia" family="Colubridae" genus="Dryocalamus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="253" phylum="Chordata" rank="species" species="nympha">
|
||
<emphasis id="9CE6F974173CFFC18EA2FD0A4F14CEF4" box="[213,336,693,719]" italics="true" pageId="5" pageNumber="253">D. nympha</emphasis>
|
||
</taxonomicName>
|
||
by the absence (vs. presence) of a preocular scale; having the loreal in contact (vs. not in contact) with the supraocular (
|
||
<figureCitation id="36A939E3173CFFC18FFCFD654FABCEC8" box="[395,495,729,755]" captionStart="FIGURE 3" captionStartId="4.[151,250,1088,1113]" captionTargetBox="[220,1365,203,1042]" captionTargetId="figure@4.[185,1403,181,1063]" captionTargetPageId="4" captionText="FIGURE 3. Dryocalamus chithrasekarai sp. nov., holotype, NMSL-NH 2019.26.01, 328 mm SVL, head scalation: (A) lateral, (B) dorsal, (C) ventral aspects, respectively." figureDoi="http://doi.org/10.5281/zenodo.3698763" httpUri="https://zenodo.org/record/3698763/files/figure.png" pageId="5" pageNumber="253">Figs. 3A</figureCitation>
|
||
,
|
||
<figureCitation id="36A939E3173CFFC18F8CFD654C65CECF" box="[507,545,729,756]" captionStart="FIGURE 6" captionStartId="6.[151,250,1247,1272]" captionTargetBox="[161,1426,317,1215]" captionTargetId="figure@6.[151,1436,309,1223]" captionTargetPageId="6" captionText="FIGURE 6. Dryocalamus nympha, lectotype, BMNH 1946.1.13.69, 338 mm SVL: (A) lateral head; (B) dorsal,and (C) ventral aspects, respectively." figureDoi="http://doi.org/10.5281/zenodo.3698769" httpUri="https://zenodo.org/record/3698769/files/figure.png" pageId="5" pageNumber="253">6A</figureCitation>
|
||
); having 15 (vs. 13) costals; banding present throughout the anterior venter (vs. not banded, uniform creamy white); and the tail region consistently dark, with no white bands (vs. uniformly creamy white) (
|
||
<figureCitation id="36A939E3173CFFC18E99FC9D4F79CF00" box="[238,317,801,827]" captionStart="FIGURE 2" captionStartId="3.[151,250,1955,1980]" captionTargetBox="[161,1427,1111,1921]" captionTargetId="figure@3.[151,1436,1101,1931]" captionTargetPageId="3" captionText="FIGURE 2. The holotype of Dryocalamus chithrasekarai sp. nov., anterior body (A) dorsal, (B) lateral, (C) ventral aspects; midbody (D) dorsal, (E) lateral, and (F) ventral aspects, respectively; and (G) ventral aspect of tail." figureDoi="http://doi.org/10.5281/zenodo.3698761" httpUri="https://zenodo.org/record/3698761/files/figure.png" pageId="5" pageNumber="253">Figs. 2</figureCitation>
|
||
,
|
||
<figureCitation id="36A939E3173CFFC18F35FC9D4F16CF00" box="[322,338,801,827]" captionStart="FIGURE 4" captionStartId="4.[151,250,1980,2005]" captionTargetBox="[191,1397,1179,1946]" captionTargetId="figure@4.[181,1406,1171,1954]" captionTargetPageId="4" captionText="FIGURE 4. The paratype, of Dryocalamus chithrasekarai sp. nov., in life (not collected) showing (A) dorsal, (B) dorsolateral (laterally inverted), and (C) ventral aspects, respectively." figureDoi="http://doi.org/10.5281/zenodo.3698765" httpUri="https://zenodo.org/record/3698765/files/figure.png" pageId="5" pageNumber="253">4</figureCitation>
|
||
,
|
||
<figureCitation id="36A939E3173CFFC18F29FC9D4F28CF07" box="[350,364,801,828]" captionStart="FIGURE 6" captionStartId="6.[151,250,1247,1272]" captionTargetBox="[161,1426,317,1215]" captionTargetId="figure@6.[151,1436,309,1223]" captionTargetPageId="6" captionText="FIGURE 6. Dryocalamus nympha, lectotype, BMNH 1946.1.13.69, 338 mm SVL: (A) lateral head; (B) dorsal,and (C) ventral aspects, respectively." figureDoi="http://doi.org/10.5281/zenodo.3698769" httpUri="https://zenodo.org/record/3698769/files/figure.png" pageId="5" pageNumber="253">6</figureCitation>
|
||
B–C).
|
||
</paragraph>
|
||
<caption id="FAED75EE173CFFC18EE0F89F4F29CB5B" ID-DOI="http://doi.org/10.5281/zenodo.3698767" ID-Zenodo-Dep="3698767" httpUri="https://zenodo.org/record/3698767/files/figure.png" pageId="5" pageNumber="253" startId="5.[151,250,1827,1852]" targetBox="[162,1425,861,1790]" targetPageId="5">
|
||
<paragraph id="AE2D2566173CFFC18EE0F89F4F29CB5B" blockId="5.[151,1436,1827,1888]" pageId="5" pageNumber="253">
|
||
<emphasis id="9CE6F974173CFFC18EE0F89F4F57CB07" bold="true" box="[151,275,1827,1852]" pageId="5" pageNumber="253">FIGURE 5.</emphasis>
|
||
<taxonomicName id="69925EE5173CFFC18F6DF89F4FB2CB00" baseAuthorityName="Gunther" baseAuthorityYear="1864" box="[282,502,1827,1851]" class="Reptilia" family="Colubridae" genus="Dryocalamus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="253" phylum="Chordata" rank="species" species="gracilis">
|
||
<emphasis id="9CE6F974173CFFC18F6DF89F4FB2CB00" box="[282,502,1827,1851]" italics="true" pageId="5" pageNumber="253">Dryocalamus gracilis</emphasis>
|
||
</taxonomicName>
|
||
holotype, BMNH 1946.1.13.86, 403 mm SVL: (A) lateral head; (B) dorsal, and (C) ventral aspects, respectively.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="AE2D2566173CFFC28EB0F8324A19CD1B" blockId="5.[151,1436,1934,2032]" lastBlockId="6.[151,1436,152,288]" lastPageId="6" lastPageNumber="254" pageId="5" pageNumber="253">
|
||
<taxonomicName id="69925EE5173CFFC18EB0F8324C4DCB93" authority="Vidanapathirana & Pushpamal & Wickramasinghe, 2020" authorityName="Vidanapathirana & Pushpamal & Wickramasinghe" authorityYear="2020" box="[199,521,1934,1960]" class="Reptilia" family="Colubridae" genus="Dryocalamus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="253" phylum="Chordata" rank="species" species="chithrasekarai" status="sp. nov.">
|
||
<emphasis id="9CE6F974173CFFC18EB0F8324C4DCB93" box="[199,521,1934,1960]" italics="true" pageId="5" pageNumber="253">Dryocalamus chithrasekarai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9CE6F974173CFFC18C65F8324C28CB93" bold="true" box="[530,620,1934,1960]" pageId="5" pageNumber="253">
|
||
<taxonomicNameLabel id="87D5440F173CFFC18C65F8324C28CB93" box="[530,620,1934,1960]" pageId="5" pageNumber="253" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is distinguished from
|
||
<taxonomicName id="69925EE5173CFFC18D1BF8334AF9CB93" authority="(Blanford, 1878)" baseAuthorityName="Blanford" baseAuthorityYear="1878" box="[876,1213,1934,1960]" class="Reptilia" family="Colubridae" genus="Dryocalamus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="253" phylum="Chordata" rank="species" species="davisonii">
|
||
<emphasis id="9CE6F974173CFFC18D1BF8334DB2CB93" box="[876,1014,1934,1960]" italics="true" pageId="5" pageNumber="253">D. davisonii</emphasis>
|
||
(
|
||
<bibRefCitation id="CA035897173CFFC18A72F8324AF1CB93" author="Blanford, W. T." box="[1029,1205,1934,1960]" pageId="5" pageNumber="253" pagination="125 - 131" refId="ref6891" refString="Blanford, W. T. (1878) Notes on some Reptilia from the Himalayas and Burma. Journal of the Asiatic Society of Bengal, 47 (2), 125 - 131." type="journal article" year="1878">Blanford, 1878</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, to which it bears a superficial resemblance, by the absence (vs. presence) of a preocular scale, having the loreal in contact (vs. not in contact) with the supraocular, and having 15 (vs. 13) costals. It is distinguished from
|
||
<taxonomicName id="69925EE5173CFFC18A3DF86B4BD3CBCB" authority="(Griffin, 1909)" baseAuthorityName="Griffin" baseAuthorityYear="1909" box="[1098,1431,2006,2032]" class="Reptilia" family="Colubridae" genus="Dryocalamus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="253" phylum="Chordata" rank="species" species="philippinus">
|
||
<emphasis id="9CE6F974173CFFC18A3DF86B4AADCBCB" box="[1098,1257,2006,2032]" italics="true" pageId="5" pageNumber="253">D. philippinus</emphasis>
|
||
(
|
||
<bibRefCitation id="CA035897173CFFC18A80F86A4BD4CBCB" author="Griffin, L. E." box="[1271,1424,2006,2032]" pageId="5" pageNumber="253" pagination="595 - 601" refId="ref7714" refString="Griffin, L. E. (1909) A list of snakes found in Palawan. Philippine Journal of Science, 4, 595 - 601." type="journal article" year="1909">Griffin, 1909</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="69925EE5173FFFC28EE0FF264CA8CC88" authority="(Dumeril, Bibron & Dumeril, 1854)" baseAuthorityName="Dumeril, Bibron & Dumeril" baseAuthorityYear="1854" box="[151,748,152,179]" class="Reptilia" family="Colubridae" genus="Dryocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="254" phylum="Chordata" rank="species" species="subannulatus">
|
||
<emphasis id="9CE6F974173FFFC28EE0FF264F0BCC88" box="[151,335,153,179]" italics="true" pageId="6" pageNumber="254">D. subannulatus</emphasis>
|
||
(
|
||
<bibRefCitation id="CA035897173FFFC28F28FF254CA0CC88" author="Dumeril, A. M. C. & Bibron, G. & Dumeril, A." box="[351,740,152,179]" pageId="6" pageNumber="254" refId="ref7512" refString="Dumeril, A. M. C., Bibron, G. & Dumeril, A. (1854) Erpetologie generale ou Histoire Naturelle complete des Reptiles. Vol. 9. Roret, Paris, 474 pp. https: // doi. org / 10.5962 / bhl. title. 100816" type="book" year="1854">Duméril, Bibron & Duméril, 1854</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="69925EE5173FFFC28D53FF264A3ACC88" authority="(Gunther, 1858)" baseAuthorityName="Gunther" baseAuthorityYear="1858" box="[804,1150,152,179]" class="Reptilia" family="Colubridae" genus="Dryocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="254" phylum="Chordata" rank="species" species="tristrigatus">
|
||
<emphasis id="9CE6F974173FFFC28D53FF264D86CC88" box="[804,962,154,179]" italics="true" pageId="6" pageNumber="254">D. tristrigatus</emphasis>
|
||
(
|
||
<bibRefCitation id="CA035897173FFFC28DA5FF254A32CC88" author="Gunther, A." box="[978,1142,152,179]" pageId="6" pageNumber="254" refId="ref7742" refString="Gunther, A. (1858) Catalogue of Colubrine snakes of the British Museum. Printed by order of the Trustees, London, xvi + 281 pp." type="book" year="1858">Günther, 1858</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
by the absence (vs. presence) of a preocular scale, having the loreal in contact (vs. not in contact) with the supraocular, the cloacal shield divided (vs. undivided), a banded appearance on the venter throughout the anterior body, and the tail region consistently dark, bearing no white bands (vs. entire venter and tail cream white, unbanded).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="FAED75EE173FFFC28EE0FB634F29C927" ID-DOI="http://doi.org/10.5281/zenodo.3698769" ID-Zenodo-Dep="3698769" httpUri="https://zenodo.org/record/3698769/files/figure.png" pageId="6" pageNumber="254" startId="6.[151,250,1247,1272]" targetBox="[161,1426,317,1215]" targetPageId="6">
|
||
<paragraph id="AE2D2566173FFFC28EE0FB634F29C927" blockId="6.[151,1436,1247,1308]" pageId="6" pageNumber="254">
|
||
<emphasis id="9CE6F974173FFFC28EE0FB634F57C8C3" bold="true" box="[151,275,1247,1272]" pageId="6" pageNumber="254">FIGURE 6.</emphasis>
|
||
<taxonomicName id="69925EE5173FFFC28F6EFB5C4FBCC8C3" baseAuthorityName="Daudin" baseAuthorityYear="1803" box="[281,504,1248,1272]" class="Reptilia" family="Colubridae" genus="Dryocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="254" phylum="Chordata" rank="species" species="nympha">
|
||
<emphasis id="9CE6F974173FFFC28F6EFB5C4FBCC8C3" box="[281,504,1248,1272]" italics="true" pageId="6" pageNumber="254">Dryocalamus nympha</emphasis>
|
||
</taxonomicName>
|
||
, lectotype, BMNH 1946.1.13.69, 338 mm SVL: (A) lateral head; (B) dorsal,and (C) ventral aspects, respectively.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="E68876ED173FFFC28EB0FAF74CBBCA23" pageId="6" pageNumber="254" type="discussion">
|
||
<paragraph id="AE2D2566173FFFC28EB0FAF74CBBCA23" blockId="6.[151,1437,1354,1561]" pageId="6" pageNumber="254">
|
||
<emphasis id="9CE6F974173FFFC28EB0FAF74F7FC95E" bold="true" box="[199,315,1355,1381]" pageId="6" pageNumber="254">Remarks.</emphasis>
|
||
Although
|
||
<taxonomicName id="69925EE5173FFFC28FC2FAF74C3FC95E" authority="Vidanapathirana & Pushpamal & Wickramasinghe, 2020" authorityName="Vidanapathirana & Pushpamal & Wickramasinghe" authorityYear="2020" box="[437,635,1355,1381]" class="Reptilia" family="Colubridae" genus="Dryocalamus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="254" phylum="Chordata" rank="species" species="chithrasekarai" status="sp. nov.">
|
||
<emphasis id="9CE6F974173FFFC28FC2FAF74C3FC95E" box="[437,635,1355,1381]" italics="true" pageId="6" pageNumber="254">D. chithrasekarai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9CE6F974173FFFC28CF4FAF74C9AC95E" bold="true" box="[643,734,1355,1381]" pageId="6" pageNumber="254">
|
||
<taxonomicNameLabel id="87D5440F173FFFC28CF4FAF74C9AC95E" box="[643,734,1355,1381]" pageId="6" pageNumber="254" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, is described from only two specimens, we have observed the species also at Kalubowitiyana (
|
||
<geoCoordinate id="CBA643A1173FFFC28C7CFAD24CE9C9B2" box="[523,685,1390,1417]" degrees="06" direction="north" minutes="19" orientation="latitude" pageId="6" pageNumber="254" precision="1" seconds="58.9" value="6.333028">06°19’58.9”N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="CBA643A1173FFFC28CCCFAD24D2CC9B2" box="[699,872,1390,1417]" degrees="080" direction="east" minutes="24" orientation="longitude" pageId="6" pageNumber="254" precision="1" seconds="09.5" value="80.40264">080°24’09.5”E</geoCoordinate>
|
||
), Morawaka (
|
||
<geoCoordinate id="CBA643A1173FFFC28A7FFAD24AEDC9B2" box="[1032,1193,1390,1417]" degrees="06" direction="north" minutes="15" orientation="latitude" pageId="6" pageNumber="254" precision="1" seconds="58.9" value="6.266361">06°15’58.9”N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="CBA643A1173FFFC28ACFFAD24B27C9B2" box="[1208,1379,1390,1417]" degrees="080" direction="east" minutes="28" orientation="longitude" pageId="6" pageNumber="254" precision="1" seconds="36.4" value="80.476776">080°28’36.4”E</geoCoordinate>
|
||
) and Runakanda (
|
||
<geoCoordinate id="CBA643A1173FFFC28F55FA2E4F87C996" box="[290,451,1426,1453]" degrees="06" direction="north" minutes="27" orientation="latitude" pageId="6" pageNumber="254" precision="1" seconds="15.1" value="6.454194">06°27’15.1”N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="CBA643A1173FFFC28FBBFA2F4C32C996" box="[460,630,1427,1453]" degrees="080" direction="east" minutes="19" orientation="longitude" pageId="6" pageNumber="254" precision="1" seconds="29.0" value="80.32472">080°19’29.0”E</geoCoordinate>
|
||
) (
|
||
<figureCitation id="36A939E3173FFFC28CFDFA2E4C90C996" box="[650,724,1426,1453]" captionStart="FIGURE 7" captionStartId="7.[151,250,1728,1753]" captionTargetBox="[162,1425,464,1693]" captionTargetId="figure@7.[151,1436,453,1704]" captionTargetPageId="7" captionText="FIGURE 7. Dryocalamus chithrasekarai sp. nov., in life (not collected) from (A-C) Runakanda Forest Reserve, (D) from Sinharaja World Heritage Site." figureDoi="http://doi.org/10.5281/zenodo.3698771" httpUri="https://zenodo.org/record/3698771/files/figure.png" pageId="6" pageNumber="254">Figs. 7</figureCitation>
|
||
A–C), in addition two individuals from Sinharaja [
|
||
<geoCoordinate id="CBA643A1173FFFC28A82FA2E4BD3C997" box="[1269,1431,1426,1452]" degrees="06" direction="north" minutes="26" orientation="latitude" pageId="6" pageNumber="254" precision="1" seconds="18.1" value="6.438361">06°26’18.1”N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="CBA643A1173FFFC28EE0FA0A4F04C9EA" box="[151,320,1462,1489]" degrees="080" direction="east" minutes="25" orientation="longitude" pageId="6" pageNumber="254" precision="1" seconds="22.3" value="80.42286">080°25’22.3”E</geoCoordinate>
|
||
(
|
||
<figureCitation id="36A939E3173FFFC28F25FA0A4FE8C9EA" box="[338,428,1462,1489]" captionStart="FIGURE 7" captionStartId="7.[151,250,1728,1753]" captionTargetBox="[162,1425,464,1693]" captionTargetId="figure@7.[151,1436,453,1704]" captionTargetPageId="7" captionText="FIGURE 7. Dryocalamus chithrasekarai sp. nov., in life (not collected) from (A-C) Runakanda Forest Reserve, (D) from Sinharaja World Heritage Site." figureDoi="http://doi.org/10.5281/zenodo.3698771" httpUri="https://zenodo.org/record/3698771/files/figure.png" pageId="6" pageNumber="254">Fig. 7D</figureCitation>
|
||
) and
|
||
<geoCoordinate id="CBA643A1173FFFC28F99FA0A4CCBC9EB" box="[494,655,1462,1488]" degrees="06" direction="north" minutes="26" orientation="latitude" pageId="6" pageNumber="254" precision="1" seconds="18.1" value="6.438361">06°26’18.1”N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="CBA643A1173FFFC28CEAFA0A4D0FC9EA" box="[669,843,1462,1489]" degrees="080" direction="east" minutes="26" orientation="longitude" pageId="6" pageNumber="254" precision="1" seconds="40.6" value="80.44461">080°26’40.6”E</geoCoordinate>
|
||
], all of which lie in the ‘rain’ forests of the island’s south-western wet zone quadrant. All five individuals were identified in the field on the basis of the unique banded ventral coloration and the absence of a preocular scale.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |