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<document id="91FF8CAFBB208CB1BD1514E9A94F78F3" ID-CLB-Dataset="50947" ID-DOI="10.5281/zenodo.170373" ID-GBIF-Dataset="0a55d84c-3a76-4137-91e8-a1917d9eb3b3" ID-ISSN="1175­5326" ID-Zenodo-Dep="170373" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459747732027" checkinUser="plazi" docAuthor="Lewis, Julian J." docDate="2005" docId="11396778FFB1FFD1FEB173339C8E1866" docLanguage="en" docName="zt01080p031.pdf" docOrigin="Zootaxa 1080" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Pseudotremia wallaceae Lewis, 2005, new species" docType="treatment" docVersion="10" lastPageNumber="28" masterDocId="ED001F00FFBAFFDDFFB9722F9D781938" masterDocTitle="Cumberland Plateau (Diplopoda: Chordeumatida: Cleidogonidae)" masterLastPageNumber="31" masterPageNumber="17" pageNumber="28" updateTime="1698214545952" updateUser="ExternalLinkService">
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<mods:title id="DE1C76541CAD077AD66BD77F3BD3B150">Cumberland Plateau (Diplopoda: Chordeumatida: Cleidogonidae)</mods:title>
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<mods:namePart id="5E47D8D073AE6AC43BE4BF5F26C3C0C5">Lewis, Julian J.</mods:namePart>
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<mods:date id="349C44A72F8D689E31C8387E21328B00">2005</mods:date>
<mods:detail id="EEE9FB5786190F5BF39E46B5FC171D81" type="volume">
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<mods:identifier id="7BD1ECCF2CCFF10C7C84121D6FBACC5C" type="CLB-Dataset">50947</mods:identifier>
<mods:identifier id="FE59FEB35F5560D65A97A888191D9BCE" type="DOI">10.5281/zenodo.170373</mods:identifier>
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<mods:identifier id="463AC0C628C71C2A5A2DCA89991470BC" type="ISSN">1175­5326</mods:identifier>
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<treatment id="11396778FFB1FFD1FEB173339C8E1866" ID-DOI="http://doi.org/10.5281/zenodo.6264409" ID-GBIF-Taxon="119336856" ID-Zenodo-Dep="6264409" LSID="urn:lsid:plazi:treatment:11396778FFB1FFD1FEB173339C8E1866" httpUri="http://treatment.plazi.org/id/11396778FFB1FFD1FEB173339C8E1866" lastPageId="12" lastPageNumber="28" pageId="11" pageNumber="28">
<subSubSection id="D18A85E5FFB1FFD6FEB173339CE91861" pageId="11" pageNumber="28" type="nomenclature">
<paragraph id="992FD66EFFB1FFD6FEB173339FBC180D" blockId="11.[264,708,283,345]" box="[264,708,283,310]" pageId="11" pageNumber="28">
<heading id="C2676102FFB1FFD6FEB173339FBC180D" bold="true" box="[264,708,283,310]" fontSize="11" level="1" pageId="11" pageNumber="28" reason="1">
<emphasis id="ABE40A7CFFB1FFD6FEB173339FBC180D" bold="true" box="[264,708,283,310]" pageId="11" pageNumber="28">
<taxonomicName id="5E90ADEDFFB1FFD6FEB173339F54180E" ID-CoL="78HKS" box="[264,556,284,310]" class="Diplopoda" family="Cleidogonidae" genus="Pseudotremia" kingdom="Animalia" order="Chordeumatida" pageId="11" pageNumber="28" phylum="Arthropoda" rank="species" species="wallaceae" status="sp. nov.">
<emphasis id="ABE40A7CFFB1FFD6FEB173339F54180E" bold="true" box="[264,556,284,310]" italics="true" pageId="11" pageNumber="28">Pseudotremia wallaceae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B0D7B707FFB1FFD6FD8A73349FBC180D" box="[563,708,283,309]" pageId="11" pageNumber="28" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="992FD66EFFB1FFD6FEB173109CE91861" blockId="11.[264,708,283,345]" box="[264,401,319,345]" pageId="11" pageNumber="28">
<figureCitation id="01ABCAEBFFB1FFD6FEB173109CE91861" box="[264,401,319,345]" captionStart="FIGURES 16 23" captionStartId="10.[264,382,1695,1719]" captionTargetBox="[301,1267,334,1645]" captionTargetId="figure@10.[292,1296,321,1669]" captionTargetPageId="10" captionText="FIGURES 16 23. Pseudotremia roebuckorum, n. sp., Fallen Entrance Cave, Fentress Co., Tennessee: (16) anterior gonopods, anterior aspect, (17) anterior gonopods, lateral aspect, (18) leg 9, (19) cyphopod. Pseudotremia wallaceae, n. sp., Frog Cave, Pickett Co., Tennessee: (20) anterior gonopods, anterior aspect, (21) anterior gonopods, lateral aspect, (22) leg 9, (23) cyphopod." httpUri="https://zenodo.org/record/170376/files/figure.png" pageId="11" pageNumber="28">Figs. 2023</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="D18A85E5FFB1FFD6FEB173A09F181B19" pageId="11" pageNumber="28" type="materials_examined">
<paragraph id="992FD66EFFB1FFD6FEB173A09F181B19" blockId="11.[264,1325,398,1870]" pageId="11" pageNumber="28">
<emphasis id="ABE40A7CFFB1FFD6FEB173A09C971890" box="[264,495,399,424]" italics="true" pageId="11" pageNumber="28">Material examined:</emphasis>
<emphasis id="ABE40A7CFFB1FFD6FE4E73A19E6D1890" bold="true" box="[503,789,398,424]" pageId="11" pageNumber="28">Tennessee: Pickett Co.,</emphasis>
Cordell Hull State Park, Frog Cave,
<typeStatus id="462B68CCFFB1FFD6FB7A73A098521891" box="[1219,1322,399,425]" pageId="11" pageNumber="28" type="holotype">holotype</typeStatus>
male, 2
<typeStatus id="462B68CCFFB1FFD6FED073989CB618E9" box="[361,462,439,465]" pageId="11" pageNumber="28" type="paratype">paratype</typeStatus>
males,
<typeStatus id="462B68CCFFB1FFD6FD9573989FE918E9" box="[556,657,439,465]" pageId="11" pageNumber="28" type="paratype">paratype</typeStatus>
female,
<specimenCount id="8F961DE7FFB1FFD6FD4373989E0618E9" box="[762,894,439,465]" pageId="11" pageNumber="28" type="juvenile">3 juveniles</specimenCount>
, J. Lewis, S. Lewis, H. Garland,
<date id="ED2EF0AEFFB1FFD6FAB473989CA818C1" pageId="11" pageNumber="28" value="2003-09-12">12 September 2003</date>
; Bunkum Cave, 3
<typeStatus id="462B68CCFFB1FFD6FD0673F09E5C18C1" box="[703,804,479,505]" pageId="11" pageNumber="28" type="paratype">paratype</typeStatus>
males,
<specimenCount id="8F961DE7FFB1FFD6FC3173F0997A18C1" box="[904,1026,479,505]" pageId="11" pageNumber="28" type="female">6 females</specimenCount>
, J. Lewis, S. Lewis, H. Garland,
<date id="ED2EF0AEFFB1FFD6FECC70289F241B19" box="[373,604,519,545]" pageId="11" pageNumber="28" value="2003-09-12">12 September 2003</date>
.
</paragraph>
</subSubSection>
<subSubSection id="D18A85E5FFB1FFD6FE8170009C8E1BD1" pageId="11" pageNumber="28" type="diagnosis">
<paragraph id="992FD66EFFB1FFD6FE8170009C8E1BD1" blockId="11.[264,1325,398,1870]" pageId="11" pageNumber="28">
<emphasis id="ABE40A7CFFB1FFD6FE8170009CC11B70" box="[312,441,559,584]" italics="true" pageId="11" pageNumber="28">Diagnosis:</emphasis>
Within the
<taxonomicName id="5E90ADEDFFB1FFD6FDE870009FCE1B70" box="[593,694,559,584]" class="Diplopoda" family="Cleidogonidae" genus="Pseudotremia" kingdom="Animalia" order="Chordeumatida" pageId="11" pageNumber="29" phylum="Arthropoda" rank="species" species="eburnea">
<emphasis id="ABE40A7CFFB1FFD6FDE870009FCE1B70" box="[593,694,559,584]" italics="true" pageId="11" pageNumber="28">Eburnea</emphasis>
</taxonomicName>
Group of
<bibRefCitation id="FD01AB9FFFB1FFD6FC8470009EA51B71" author="Shear" box="[829,989,559,585]" pageId="11" pageNumber="31" refString="Shear, W. A. (1972) Studies in the milliped Order Chordeumida (Diplopoda): A revision of the Family Cleidogonidae and a reclassification of the Order Chordeumida in the New World. Bulletin of the Museum of Comparative Zoology, 144, 151 - 352." type="journal article" year="1972">Shear (1972)</bibRefCitation>
,
<emphasis id="ABE40A7CFFB1FFD6FC57700099FA1B70" box="[1006,1154,559,584]" italics="true" pageId="11" pageNumber="28">P. w a l l a c e a e</emphasis>
most closely resembles
<emphasis id="ABE40A7CFFB1FFD6FE3D70789C841B48" box="[388,508,599,624]" italics="true" pageId="11" pageNumber="28">P. a c h e ro n</emphasis>
Shear (Warren and Van Buren counties, Tennessee) and
<taxonomicName id="5E90ADEDFFB1FFD6FB2C707898531B48" box="[1173,1323,599,624]" class="Diplopoda" family="Cleidogonidae" genus="Pseudotremia" kingdom="Animalia" order="Chordeumatida" pageId="11" pageNumber="28" phylum="Arthropoda" rank="species" species="garlandae" status="sp. nov.">
<emphasis id="ABE40A7CFFB1FFD6FB2C707898531B48" box="[1173,1323,599,624]" italics="true" pageId="11" pageNumber="28">P. garlandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B0D7B707FFB1FFD6FEB170509C471BA1" box="[264,319,639,665]" pageId="11" pageNumber="28" rank="species">n.sp.</taxonomicNameLabel>
(Fentress County), from which
<emphasis id="ABE40A7CFFB1FFD6FD0F70509E3D1BA0" box="[694,837,639,664]" italics="true" pageId="11" pageNumber="28">P. w a l l a c e a e</emphasis>
can be separated by the pigmented body, the presence of a spine on the median angiocoxite and a syncolpocoxite process distally entire, not bifurcate.
</paragraph>
</subSubSection>
<subSubSection id="D18A85E5FFB1FFD6FE8170D89C371CDE" pageId="11" pageNumber="28" type="description">
<paragraph id="992FD66EFFB1FFD6FE8170D89E371D16" blockId="11.[264,1325,398,1870]" pageId="11" pageNumber="28">
<emphasis id="ABE40A7CFFB1FFD6FE8170D89F4A1A28" box="[312,562,759,784]" italics="true" pageId="11" pageNumber="28">Description of male:</emphasis>
Longest approximately
<quantity id="5E687B8BFFB1FFD6FCE570D89EDF1A29" box="[860,935,759,785]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="11" pageNumber="28" unit="mm" value="26.0">26mm</quantity>
(coiled), width increasing from
<quantity id="5E687B8BFFB1FFD6FEB1710D9C211A04" box="[264,345,802,828]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="28" unit="mm" value="1.2">1.2mm</quantity>
(collum) to
<quantity id="5E687B8BFFB1FFD6FDB8710D9F2B1A04" box="[513,595,802,828]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="11" pageNumber="28" unit="mm" value="2.3">2.3mm</quantity>
(7th segment); body light to medium brown with whitish infusion, metaterga uniform brown with about 20 darker, slightly elongated tubercles, dorsal midline stripe light brown, purplish infusion in pregonopodal segments; sterna and legs pale brown. Eyes with 1617 clear, irregularly shaped, unpigmented ocelli within subtriangular purple ocellaria. Antenna about
<quantity id="5E687B8BFFB1FFD6FCFE71EB9EE11AE6" box="[839,921,964,990]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="11" pageNumber="28" unit="mm" value="4.2">4.2mm</quantity>
long, slender, 3rd segment about
<quantity id="5E687B8BFFB1FFD6FEB171C39C221D3E" box="[264,346,1004,1030]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="11" pageNumber="28" unit="mm" value="1.4">1.4mm</quantity>
long. Segmental paranota mildly produced, becoming indistinct from lateral striae about segment 2224. Lateral striae about 1113.
</paragraph>
<paragraph id="992FD66EFFB1FFD6FE8176139C1C1C56" blockId="11.[264,1325,398,1870]" pageId="11" pageNumber="28">
Gonopods with syncolpocoxites mitten­shaped, separated by U­shaped cleft, process class I (
<bibRefCitation id="FD01AB9FFFB1FFD6FEDD764B9F321D46" author="Shear" box="[356,586,1124,1150]" pageId="11" pageNumber="31" refString="Shear, W. A. (1972) Studies in the milliped Order Chordeumida (Diplopoda): A revision of the Family Cleidogonidae and a reclassification of the Order Chordeumida in the New World. Bulletin of the Museum of Comparative Zoology, 144, 151 - 352." type="journal article" year="1972" yearSuffix="p">Shear 1972, p. 167</bibRefCitation>
), arises from notch between syncolpocoxites, curving dorsad between t angiocoxites, in anterior view broadened distally, tapering to single point best seen in lateral aspect. Median angiocoxite with single small mesial spine midlength. Lateral angiocoxite separated from median by deep, U­shaped cleft, sigmoidally curved, divided, mesial branch shorter, slightly sigmoid, extending obliquely mesiad in anterior view. Leg 9 with distal 3 segments well formed and apparently functional, single claw present.
</paragraph>
<paragraph id="992FD66EFFB1FFD6FE8177529C371CDE" blockId="11.[264,1325,398,1870]" pageId="11" pageNumber="28">
<emphasis id="ABE40A7CFFB1FFD6FE8177529CE81CAE" box="[312,400,1405,1430]" italics="true" pageId="11" pageNumber="28">Female</emphasis>
: Approximately
<quantity id="5E687B8BFFB1FFD6FDE277539FDF1CAE" box="[603,679,1404,1430]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="11" pageNumber="28" unit="mm" value="24.0">24mm</quantity>
(coiled), non­sexual characters similar to male, ocelli 1415. Cyphopod valves elongate in dorsal aspect, mesal valve about 1.5X length lateral valve.
</paragraph>
</subSubSection>
<subSubSection id="D18A85E5FFB1FFD6FE8177DA9E401FBE" pageId="11" pageNumber="28" type="etymology">
<paragraph id="992FD66EFFB1FFD6FE8177DA9C1E1F66" blockId="11.[264,1325,398,1870]" pageId="11" pageNumber="28">
<emphasis id="ABE40A7CFFB1FFD6FE8177DA9CC61F36" box="[312,446,1525,1550]" italics="true" pageId="11" pageNumber="28">Etymology:</emphasis>
This species is named in honor of Linda Wallace­Gray, philanthropist and benefactor of The Nature Conservancys biological inventory of caves of the Cumberland Plateau.
</paragraph>
<paragraph id="992FD66EFFB1FFD6FE8174429E401FBE" blockId="11.[264,1325,398,1870]" box="[312,824,1644,1670]" pageId="11" pageNumber="28">
<emphasis id="ABE40A7CFFB1FFD6FE8174429F731FBE" box="[312,523,1645,1670]" italics="true" pageId="11" pageNumber="28">Vernacular name:</emphasis>
Wallaces cave milliped.
</paragraph>
</subSubSection>
<subSubSection id="D18A85E5FFB1FFD1FE8174BA9C8E1866" lastPageId="12" lastPageNumber="29" pageId="11" pageNumber="28" type="description">
<paragraph id="992FD66EFFB1FFD1FE8174BA9C8E1866" blockId="11.[264,1325,398,1870]" lastBlockId="12.[264,1322,284,350]" lastPageId="12" lastPageNumber="29" pageId="11" pageNumber="28">
<emphasis id="ABE40A7CFFB1FFD6FE8174BA9F511F96" box="[312,553,1685,1710]" italics="true" pageId="11" pageNumber="28">Habitat and Range:</emphasis>
Frog and Bunkum caves essentially comprise a single locality, since the entrances are less than 50 meters apart. Bunkum Cave was described by
<bibRefCitation id="FD01AB9FFFB1FFD6FB4E74939C2E1FC6" author="Barr" pageId="11" pageNumber="30" refString="Barr, T. C., Jr. (1961) Caves of Tennessee. State of Tennessee Department of Conservation and Commerce, Division of Geology, Bulletin 64, 1 - 567." type="journal article" year="1961">Barr (1961)</bibRefCitation>
as having one of the largest and most scenic entrances in Tennessee, with a main stream passage
<quantity id="5E687B8BFFB1FFD6FE7875239F581E1E" box="[449,544,1804,1830]" metricMagnitude="2" metricUnit="m" metricValue="2.286" pageId="11" pageNumber="28" unit="ft" value="750.0">750 feet</quantity>
(ca. 250 meters) in length.
<taxonomicName id="5E90ADEDFFB1FFD6FCD2752299F31E1E" box="[875,1163,1805,1830]" class="Diplopoda" family="Cleidogonidae" genus="Pseudotremia" kingdom="Animalia" order="Chordeumatida" pageId="11" pageNumber="28" phylum="Arthropoda" rank="species" species="wallaceae">
<emphasis id="ABE40A7CFFB1FFD6FCD2752299F31E1E" box="[875,1163,1805,1830]" italics="true" pageId="11" pageNumber="28">Pseudotremia wallaceae</emphasis>
</taxonomicName>
was found in Frog and Bunkum caves on riparian mud banks. Although
<taxonomicName id="5E90ADEDFFB1FFD6FC00751A99351E76" box="[953,1101,1845,1870]" class="Diplopoda" family="Cleidogonidae" genus="Pseudotremia" kingdom="Animalia" order="Chordeumatida" pageId="11" pageNumber="28" phylum="Arthropoda" rank="species" species="wallaceae">
<emphasis id="ABE40A7CFFB1FFD6FC00751A99351E76" box="[953,1101,1845,1870]" italics="true" pageId="11" pageNumber="28">P. wallaceae</emphasis>
</taxonomicName>
has been collected only from caves, in contrast to the unpigmented troglobite
<taxonomicName id="5E90ADEDFFB6FFD1FC0473339942180D" box="[957,1082,284,309]" class="Diplopoda" family="Cleidogonidae" genus="Pseudotremia" kingdom="Animalia" order="Chordeumatida" pageId="12" pageNumber="29" phylum="Arthropoda" rank="species" species="acheron">
<emphasis id="ABE40A7CFFB6FFD1FC0473339942180D" box="[957,1082,284,309]" italics="true" pageId="12" pageNumber="29">P. acheron</emphasis>
</taxonomicName>
, it is pigmented and may be troglophilic.
</paragraph>
</subSubSection>
</treatment>
</document>