168 lines
19 KiB
XML
168 lines
19 KiB
XML
<document id="FC559DF443C51E77A5BFA52A048D02A1" ID-DOI="10.33910/2686-9519-2023-15-2-325-354" ID-ISSN="2686-9519" ID-Zenodo-Dep="12816849" ID-ZooBank="912125ED-10E5-4A6C-86E7-548CCD69F364" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721879379315" checkinUser="felipe" docAuthor="Kosterin, Oleg E." docDate="2023" docId="05368791C523FFB12962FF23FBF9EACD" docLanguage="en" docName="AmurianZoolJ.XV.2.325-354.pdf" docOrigin="Amurian Zoological Journal XV (2)" docSource="http://dx.doi.org/10.33910/2686-9519-2023-15-2-325-354" docStyle="DocumentStyle:1BE7E8521B83181C3ED08D600C3F2F53.3:AmurianZoolJ.2019-.journal_article.english" docStyleId="1BE7E8521B83181C3ED08D600C3F2F53" docStyleName="AmurianZoolJ.2019-.journal_article.english" docStyleVersion="3" docTitle="Plebejus idas subsp. subsolanus" docType="treatment" docVersion="2" lastPageNumber="344" masterDocId="F90FFFE9C530FFA22A78FFBBFFBCED24" masterDocTitle="Early summer aspect of butterflies (Lepidoptera: Papilionoidea) of Republic of Khakassia as examined in 2000, with some additional data" masterLastPageNumber="354" masterPageNumber="325" pageNumber="344" updateTime="1721879893216" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="1D72352744383B5E8948B57218D8E13A">Early summer aspect of butterflies (Lepidoptera: Papilionoidea) of Republic of Khakassia as examined in 2000, with some additional data</mods:title>
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<mods:namePart id="1DF5FFD262D23B336CFFEBE034B76B56">Kosterin, Oleg E.</mods:namePart>
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<mods:title id="5DEA339F3E9439D895E6FAAAE699E646">Amurian Zoological Journal</mods:title>
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<mods:date id="AD030DC57B1F42E01FD84881FAC19546">2023</mods:date>
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<mods:number id="AFA2D3772A1C199339AB638641A6EA89">XV</mods:number>
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<mods:identifier id="1D906F455A0767D47296233A14D57ED9" type="Zenodo-Dep">12816849</mods:identifier>
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<treatment id="05368791C523FFB12962FF23FBF9EACD" LSID="urn:lsid:plazi:treatment:05368791C523FFB12962FF23FBF9EACD" httpUri="http://treatment.plazi.org/id/05368791C523FFB12962FF23FBF9EACD" lastPageNumber="344" pageId="19" pageNumber="344">
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<subSubSection id="C585650CC523FFB12962FF23FB7BEC52" pageId="19" pageNumber="344" type="nomenclature">
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<paragraph id="8D203687C523FFB12962FF23FACAED92" blockId="19.[791,1401,152,2025]" box="[794,1398,152,182]" pageId="19" pageNumber="344">
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<heading id="D66881EBC523FFB12962FF23FACAED92" box="[794,1398,152,182]" centered="true" fontSize="12" level="2" pageId="19" pageNumber="344" reason="4">
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<taxonomicName id="4A9F4D04C523FFB12962FF23FACAED92" authority="(Eversmann, 1851)" baseAuthorityName="Eversmann" baseAuthorityYear="1851" box="[794,1398,152,182]" class="Insecta" family="Lycaenidae" genus="Plebejus" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="344" phylum="Arthropoda" rank="subSpecies" species="idas" subSpecies="subsolanus">
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<emphasis id="BFEBEA95C523FFB12962FF23FBCAED91" bold="true" box="[794,1142,152,181]" italics="true" pageId="19" pageNumber="344">Plebejus idas subsolanus</emphasis>
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(Eversmann, 1851)
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8D203687C523FFB12962FF04FB7BEC52" blockId="19.[791,1401,152,2025]" pageId="19" pageNumber="344">
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<emphasis id="BFEBEA95C523FFB12962FF04FC09EDF8" bold="true" box="[794,949,191,220]" pageId="19" pageNumber="344">Specimens:</emphasis>
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1\NNE Parnaya,
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<specimenCount id="9B99FD0EC523FFB12ED2FF04FB5DEDFB" box="[1194,1249,191,223]" count="11" pageId="19" pageNumber="344" type="male">11♂</specimenCount>
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,
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<specimenCount id="9B99FD0EC523FFB12E8EFF04FAA1EDFB" box="[1270,1309,191,223]" count="3" pageId="19" pageNumber="344" type="female">3♀</specimenCount>
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; 2\NE Kopyovo,
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<specimenCount id="9B99FD0EC523FFB129D8FF5DFC64EC21" box="[928,984,230,261]" count="18" pageId="19" pageNumber="344" type="male">18♂</specimenCount>
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,
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<specimenCount id="9B99FD0EC523FFB12990FF5DFBACEC21" box="[1000,1040,230,261]" count="2" pageId="19" pageNumber="344" type="female">2♀</specimenCount>
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; 3\NNE Kopyovo,
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<specimenCount id="9B99FD0EC523FFB12F77FF5DFA8BEC21" box="[1295,1335,230,261]" count="9" pageId="19" pageNumber="344" type="male">9♂</specimenCount>
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,
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<specimenCount id="9B99FD0EC523FFB12F3FFF5DFAD3EC21" box="[1351,1391,230,261]" count="2" pageId="19" pageNumber="344" type="female">2♀</specimenCount>
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; 4\WNW Shira,
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<specimenCount id="9B99FD0EC523FFB12988FEB7FBA4EC08" box="[1008,1048,268,300]" count="2" pageId="19" pageNumber="344" type="male">2♂</specimenCount>
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; 7\Sonskiy,
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<specimenCount id="9B99FD0EC523FFB12EC2FEB7FB5EEC08" box="[1210,1250,268,300]" count="3" pageId="19" pageNumber="344" type="male">3♂</specimenCount>
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,
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<specimenCount id="9B99FD0EC523FFB12E8CFEB7FAA0EC08" box="[1268,1308,268,300]" count="2" pageId="19" pageNumber="344" type="female">2♀</specimenCount>
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; 8\Uybat,
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<specimenCount id="9B99FD0EC523FFB12934FE88FCCFEC76" box="[844,883,307,338]" count="7" pageId="19" pageNumber="344" type="male">7♂</specimenCount>
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,
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<specimenCount id="9B99FD0EC523FFB12905FE88FC18EC76" box="[893,932,307,338]" count="1" pageId="19" pageNumber="344" type="female">1♀</specimenCount>
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; 11\Tashtyp,
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<specimenCount id="9B99FD0EC523FFB12E3CFE89FBC7EC76" box="[1092,1147,306,338]" count="25" pageId="19" pageNumber="344" type="male">25♂</specimenCount>
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,
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<specimenCount id="9B99FD0EC523FFB12EFCFE88FB07EC76" box="[1156,1211,307,338]" count="10" pageId="19" pageNumber="344" type="female">10♀</specimenCount>
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;12\Dzhabash,
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<specimenCount id="9B99FD0EC523FFB1296FFEE2FCEEEC5D" box="[791,850,345,377]" count="74" pageId="19" pageNumber="344" type="male">74♂</specimenCount>
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(from a large congregation).
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</paragraph>
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</subSubSection>
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<subSubSection id="C585650CC523FFB12962FEC4FA9DECB8" box="[794,1313,383,415]" pageId="19" pageNumber="344" type="description">
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<paragraph id="8D203687C523FFB12962FEC4FA9DECB8" blockId="19.[791,1401,152,2025]" box="[794,1313,383,415]" pageId="19" pageNumber="344">
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<emphasis id="BFEBEA95C523FFB12962FEC4FB8AECB8" bold="true" box="[794,1078,383,412]" pageId="19" pageNumber="344">Visual registrations:</emphasis>
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10\Terensug,
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<specimenCount id="9B99FD0EC523FFB12E8AFEC4FAA6ECBB" box="[1266,1306,383,415]" count="1" pageId="19" pageNumber="344" type="male">1♂</specimenCount>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C585650CC523FFB12962FE1DFBF9EACD" pageId="19" pageNumber="344" type="discussion">
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<paragraph id="8D203687C523FFB12962FE1DFB86E9B8" blockId="19.[791,1401,152,2025]" pageId="19" pageNumber="344">
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<emphasis id="BFEBEA95C523FFB12962FE1DFC1CECE6" bold="true" box="[794,928,422,450]" pageId="19" pageNumber="344">Remarks:</emphasis>
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This species was the main target of our expedition. Most male specimens had the UPS black border occupying not less than half of the wing lengths, so that in extreme cases the deep blue occupies about one third of the UPS total area (
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<figureCitation id="15A42A02C523FFB12E44FDDDFB2BEFA7" box="[1084,1175,614,643]" captionStart="Fig" captionStartId="20.[162,209,848,877]" captionTargetBox="[152,1398,151,937]" captionTargetId="figure-393@20.[162,1389,157,828]" captionTargetPageId="20" captionText="Fig. 13. A male of Plebejus idas subsolanus at Tashtyp Village, 6.07.2000 Рис. 13. Самец Plebejus idas subsolanus в окрестностях с. Таштып, 6.07.2000" figureDoi="http://doi.org/10.5281/zenodo.12816867" httpUri="https://zenodo.org/record/12816867/files/figure.png" pageId="19" pageNumber="344">Fig. 13</figureCitation>
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), hence well representing the subspecies
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<taxonomicName id="4A9F4D04C523FFB12E02FD36FACAEF8D" baseAuthorityName="Eversmann" baseAuthorityYear="1851" box="[1146,1398,652,681]" class="Insecta" family="Lycaenidae" genus="Plebejus" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="344" phylum="Arthropoda" rank="subSpecies" species="idas" subSpecies="subsolanus">
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<emphasis id="BFEBEA95C523FFB12E02FD36FACAEF8D" box="[1146,1398,652,681]" italics="true" pageId="19" pageNumber="344">P. idas subsolanus</emphasis>
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</taxonomicName>
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(
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<bibRefCitation id="E90E4B76C523FFB1295BFD09FBD0EFF4" author="Churkin, S. V. & Zhdanko, A. B." box="[803,1132,690,720]" pageId="19" pageNumber="344" refId="ref15527" refString="Churkin, S. V., Zhdanko, A. B. (2003) A review of the Plebejus idas-subsolanus complex of the Asian part of Russia and Mongolia, with the description of new taxa (Lepidoptera, Lycaenidae). In: Helios. Collection of lepidopterological articles. Vol. 4. Moscow: Chronos-Press, pp. 37 - 74. (In English)" type="book" year="2003">Churkin, Zhdanko 2003</bibRefCitation>
|
||
;
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<bibRefCitation id="E90E4B76C523FFB12E02FD09FC3FEFD2" author="Gorbunov, P. Y. & Kosterin, O. E." pageId="19" pageNumber="344" refId="ref15754" refString="Gorbunov, P. Y., Kosterin, O. E. (2003) The butterflies (Hesperioidea and Papilionoidea) of North Asia (Asian part of Russia) in nature. Vol. 1. Moscow: Rodina & Fodio Publ., 392 p. (In English)" type="book" year="2003">Gorbunov, Kosterin 2003</bibRefCitation>
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). Yet that border was variable and in some specimens (e.g. in males from 7\Sonskiy and the only male seen at 10\Terensug) narrowed to
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<quantity id="4A679B62C523FFB129EFFCF7FBB6EE4D" box="[919,1034,844,873]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="19" pageNumber="344" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
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, as in the subspecies
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<taxonomicName id="4A9F4D04C523FFB12F59FCF6FB8AEEB4" authority="(Tutt, 1909)" baseAuthorityName="Tutt" baseAuthorityYear="1909" class="Insecta" family="Lycaenidae" genus="Plebejus" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="344" phylum="Arthropoda" rank="subSpecies" species="idas" subSpecies="ongodai">
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<emphasis id="BFEBEA95C523FFB12F59FCF6FC3FEEAB" italics="true" pageId="19" pageNumber="344">P. idas ongodai</emphasis>
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(Tutt, 1909)
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</taxonomicName>
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widespread in the
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<collectingRegion id="4F5BF865C523FFB12F34FCC9FC80EE92" country="Russia" name="Altay" pageId="19" pageNumber="344">Altai</collectingRegion>
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Mts (
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<bibRefCitation id="E90E4B76C523FFB129EBFC22FB4EEE92" author="Gorbunov, P. Y. & Kosterin, O. E." box="[915,1266,921,950]" pageId="19" pageNumber="344" refId="ref15754" refString="Gorbunov, P. Y., Kosterin, O. E. (2003) The butterflies (Hesperioidea and Papilionoidea) of North Asia (Asian part of Russia) in nature. Vol. 1. Moscow: Rodina & Fodio Publ., 392 p. (In English)" type="book" year="2003">Gorbunov, Kosterin 2003</bibRefCitation>
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;
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<bibRefCitation id="E90E4B76C523FFB12F7CFC22FCE3EEF8" author="Kosterin, O. E." pageId="19" pageNumber="344" pagination="24 - 34" refId="ref16194" refString="Kosterin, O. E. (2007) New data on butterfly fauna (Lepidoptera, Diurna) of the Katunskii mountain range (Central Altai). Altaiskij Zoologicheskij Zhurnal - Altai Zoological Journal, vol. 1, pp. 24 - 34. (In English)" type="journal article" year="2007">Kosterin 2007</bibRefCitation>
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). (Curiously, the
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<typeStatus id="52248825C523FFB12E3FFC04FB01EEF8" box="[1095,1213,959,988]" pageId="19" pageNumber="344" type="holotype">holotype</typeStatus>
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of that taxon had the wings as in
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<taxonomicName id="4A9F4D04C523FFB12E54FC5CFAA0E927" baseAuthorityName="Eversmann" baseAuthorityYear="1851" box="[1068,1308,998,1027]" class="Insecta" family="Lycaenidae" genus="Plebejus" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="344" phylum="Arthropoda" rank="subSpecies" species="idas" subSpecies="subsolanus">
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<emphasis id="BFEBEA95C523FFB12E54FC5CFAA0E927" box="[1068,1308,998,1027]" italics="true" pageId="19" pageNumber="344">P. idas subsolanus</emphasis>
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</taxonomicName>
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, being an aberration among males with the black border as narrow as ca
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<quantity id="4A679B62C523FFB12E1DFB89FB6DE96B" box="[1125,1233,1074,1103]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="19" pageNumber="344" unit="mm" value="1.5">1.5 mm</quantity>
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, which predominate overwhelmingly in
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<collectingRegion id="4F5BF865C523FFB12EDBFBE2FB5AE952" box="[1187,1254,1113,1142]" country="Russia" name="Altay" pageId="19" pageNumber="344">Altai</collectingRegion>
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, see
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<bibRefCitation id="E90E4B76C523FFB12F5EFBE2FB8AE9B8" author="Churkin, S. V. & Zhdanko, A. B." pageId="19" pageNumber="344" refId="ref15527" refString="Churkin, S. V., Zhdanko, A. B. (2003) A review of the Plebejus idas-subsolanus complex of the Asian part of Russia and Mongolia, with the description of new taxa (Lepidoptera, Lycaenidae). In: Helios. Collection of lepidopterological articles. Vol. 4. Moscow: Chronos-Press, pp. 37 - 74. (In English)" type="book" year="2003">Churkin, Zhdanko (2003))</bibRefCitation>
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.
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</paragraph>
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<paragraph id="8D203687C523FFB12962FB1DFB62E88D" blockId="19.[791,1401,152,2025]" pageId="19" pageNumber="344">
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The most variable male UPS, with the mentioned versions of the black border development, and intermediates, of comparable frequences, was observed by me in the Central Tuvinian Depression, on sands at Lake Khadyn; that population looked intermediate between subspecies (
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<bibRefCitation id="E90E4B76C523FFB12E68FA37FB6DE88D" author="Kosterin, O. E." box="[1040,1233,1420,1449]" pageId="19" pageNumber="344" pagination="212 - 225" refId="ref16311" refString="Kosterin, O. E. (2009) Materialy ekspeditsij 1999 and 2000 gg. po faune dnevnykh babochek (Lepidoptera, Diurna) Ubsunurskoj kotloviny v predelakh Respubliki Tyva [Data of 1990 and 2000 expeditions on butterfly (Lepidoptera, Diurna) fauna of the Ubsu-Nur intermontane hollow within Tyva Republic]. In: Priroda zapovednika " Ubsubnurskaya Kotlovina ". Trudy gosudarstvennogo prirodnogo biosfernogo zapovednika " Ubsunurskaya Kotlovina ". Vyp. 1 [Nature of Ubsu-Nur Hollow Nature Reserve. Transactions of " Ubsunur Hollow " State Biosphere Nature Reserve. Vol. 1]. Krasnoyarsk: Darma Press, pp. 212 - 225 (In Russian)" type="book chapter" year="2009">Kosterin 2009</bibRefCitation>
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).
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</paragraph>
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<paragraph id="8D203687C523FFB12962FA09FBF9EACD" blockId="19.[791,1401,152,2025]" pageId="19" pageNumber="344">
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<emphasis id="BFEBEA95C523FFB12962FA09FC5CE8EB" bold="true" box="[794,992,1458,1487]" pageId="19" pageNumber="344">Observations:</emphasis>
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Usually occurs together with the previous species, mostly on meadows with dominance of
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<taxonomicName id="4A9F4D04C523FFB12E59F9BBFB66EB38" box="[1057,1242,1536,1564]" class="Magnoliopsida" family="Fabaceae" genus="Vicia" kingdom="Plantae" order="Fabales" pageId="19" pageNumber="344" phylum="Tracheophyta" rank="species" species="amoena">
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<emphasis id="BFEBEA95C523FFB12E59F9BBFB66EB38" box="[1057,1242,1536,1564]" italics="true" pageId="19" pageNumber="344">Vicia amoena</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4A9F4D04C523FFB12F64F9BBFCEFEB67" class="Magnoliopsida" family="Fabaceae" genus="Vicia" kingdom="Plantae" order="Fabales" pageId="19" pageNumber="344" phylum="Tracheophyta" rank="species" species="unijuga">
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<emphasis id="BFEBEA95C523FFB12F64F9BBFCEFEB67" italics="true" pageId="19" pageNumber="344">V. unijuga</emphasis>
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</taxonomicName>
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(but at 11\Tashtyp the imagines seemed to associate with
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<taxonomicName id="4A9F4D04C523FFB12982F9F7FB4CEB4D" box="[1018,1264,1612,1641]" class="Magnoliopsida" family="Fabaceae" genus="Lathyrus" kingdom="Plantae" order="Fabales" pageId="19" pageNumber="344" phylum="Tracheophyta" rank="species" species="pratensis">
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<emphasis id="BFEBEA95C523FFB12982F9F7FB4CEB4D" box="[1018,1264,1612,1641]" italics="true" pageId="19" pageNumber="344">Lathyrus pratensis</emphasis>
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</taxonomicName>
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L.), and is locally very abundant but less widespread, as found at fewer localities. For instance,
|
||
<taxonomicName id="4A9F4D04C523FFB12F59F921FACBEB92" baseAuthorityName="Linnaeus" baseAuthorityYear="1761" box="[1313,1399,1689,1718]" class="Insecta" family="Lycaenidae" genus="Plebejus" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="344" phylum="Arthropoda" rank="species" species="idas">
|
||
<emphasis id="BFEBEA95C523FFB12F59F921FACBEB92" box="[1313,1399,1689,1718]" italics="true" pageId="19" pageNumber="344">P. idas</emphasis>
|
||
</taxonomicName>
|
||
was absent from the Iyus River valley (6\SE Efremkino), where
|
||
<taxonomicName id="4A9F4D04C523FFB12E66F95CFB3AEA27" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1054,1158,1767,1795]" class="Insecta" family="Lycaenidae" genus="Plebejus" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="344" phylum="Arthropoda" rank="species" species="argus">
|
||
<emphasis id="BFEBEA95C523FFB12E66F95CFB3AEA27" box="[1054,1158,1767,1795]" italics="true" pageId="19" pageNumber="344">P. argus</emphasis>
|
||
</taxonomicName>
|
||
was so abundant. At 12\Dzhabash,
|
||
<specimenCount id="9B99FD0EC523FFB12985F8B7FBD3EA0D" box="[1021,1135,1804,1833]" count="74" pageId="19" pageNumber="344" type="male">74 males</specimenCount>
|
||
were collected from the same congregation, joint with
|
||
<taxonomicName id="4A9F4D04C523FFB12EA6F888FC70EA52" authorityName="Fabricius" authorityYear="1787" class="Insecta" family="Nymphalidae" genus="Coenonympha" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="344" phylum="Arthropoda" rank="species" species="oedippus">
|
||
<emphasis id="BFEBEA95C523FFB12EA6F888FC70EA52" italics="true" pageId="19" pageNumber="344">Coenonympha oedippus</emphasis>
|
||
</taxonomicName>
|
||
, on mud, in two attempts: after the first net stroke not less number of those re-gathered again after a while, probably attracted by cattle urine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |