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<document id="147B27E59CE6EACDCD3ED0F18979FA9A" ID-CLB-Dataset="6318" ID-DOI="10.11646/zootaxa.4970.2.1" ID-GBIF-Dataset="89d3c795-198b-4162-8903-9c6cf3f1e665" ID-ISSN="1175-5326" ID-Zenodo-Dep="4761544" ID-ZooBank="6DBA0C02-9E52-4FD0-8B48-8C7B405713A8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620984897660" checkinUser="plazi" docAuthor="Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M. &amp; Brescovit, Antonio Domingos" docDate="2021" docId="052487CFFFE1E50F46C6B36C8E2CFEB6" docLanguage="en" docName="zootaxa.4970.2.1.pdf" docOrigin="Zootaxa 4970 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Folcloredesmus Bouzan &amp; Iniesta &amp; Brescovit 2021, n. gen." docType="treatment" docUuid="DF201399-575A-4A47-8A79-A51DAFBEC925" docUuidSource="ZooBank" docVersion="6" lastPageNumber="246" masterDocId="F91DFFB7FFCAE5224651B2518D76FFF7" masterDocTitle="Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae)" masterLastPageNumber="256" masterPageNumber="201" pageNumber="244" updateTime="1698939836712" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="CADCAA3541BE5CFC57B01C6E71311763">Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae)</mods:title>
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<mods:namePart id="8DB82DF8D53EFF96A115984572427AF8">Bouzan, Rodrigo Salvador</mods:namePart>
<mods:affiliation id="9224C8916C9DE0D36971F4B3FAF0CC70">Pós-graduação em Zoologia, Instituto de Biociências, Universidade de São Paulo, São Paulo, Brasil.</mods:affiliation>
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<mods:namePart id="1289761F327D23931D349055D2166379">Iniesta, Luiz Felipe M.</mods:namePart>
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<mods:affiliation id="BCD55AADB22EA47EFDBAA0BBBF4A6305">Pós-graduação em Zoologia, Instituto de Biociências, Universidade de São Paulo, São Paulo, Brasil. &amp; luiz-moretti @ hotmail. com; https: // orcid. org / 0000 - 0002 - 0529 - 4162</mods:affiliation>
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<mods:namePart id="013D4DFC88101D619ECF02E4152F21A2">Brescovit, Antonio Domingos</mods:namePart>
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<mods:affiliation id="1A8F166083DD574BF9E5876C9AF7C418">antonio. brescovit @ butantan. gov. br; https: // orcid. org / 0000 - 0002 - 1511 - 5324</mods:affiliation>
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<treatment id="052487CFFFE1E50F46C6B36C8E2CFEB6" ID-DOI="http://doi.org/10.5281/zenodo.4761643" ID-GBIF-Taxon="182486284" ID-Zenodo-Dep="4761643" LSID="urn:lsid:zoobank.org:act:DF201399-575A-4A47-8A79-A51DAFBEC925" httpUri="http://treatment.plazi.org/id/052487CFFFE1E50F46C6B36C8E2CFEB6" lastPageId="45" lastPageNumber="246" pageId="43" pageNumber="244">
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<taxonomicName id="4A8D4D5AFFE1E50946C6B36C8C21FEAF" ID-CoL="8H7RP" authority="Bouzan &amp; Iniesta &amp; Brescovit, 2021" authorityName="Bouzan &amp; Iniesta &amp; Brescovit" authorityYear="2021" box="[151,343,317,344]" class="Diplopoda" family="Chelodesmidae" genus="Folcloredesmus" kingdom="Animalia" order="Polydesmida" pageId="43" pageNumber="244" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="BFF9EACBFFE1E50946C6B36C8C21FEAF" bold="true" box="[151,343,317,344]" italics="true" pageId="43" pageNumber="244">Folcloredesmus</emphasis>
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<typeStatus id="5236887BFFE1E50946C6B3CB8DBFFE45" box="[151,201,410,434]" pageId="43" pageNumber="244">Type</typeStatus>
species.
<taxonomicName id="4A8D4D5AFFE1E5094777B3CB8F6BFE45" authority="Bouzan &amp; Iniesta &amp; Brescovit, 2021" authorityName="Bouzan &amp; Iniesta &amp; Brescovit" authorityYear="2021" box="[294,541,410,434]" class="Diplopoda" family="Chelodesmidae" genus="Folcloredesmus" kingdom="Animalia" order="Polydesmida" pageId="43" pageNumber="244" phylum="Arthropoda" rank="species" species="thomasi" status="sp. nov.">
<emphasis id="BFF9EACBFFE1E5094777B3CB8F6BFE45" box="[294,541,410,434]" italics="true" pageId="43" pageNumber="244">Folcloredesmus thomasi</emphasis>
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<taxonomicNameLabel id="A4CA57B0FFE1E5094472B3CB8F2BFE45" box="[547,605,410,434]" pageId="43" pageNumber="244" rank="species">n. sp.</taxonomicNameLabel>
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<caption id="D9F26651FFE1E50946C6B5DC8FC0F81A" ID-DOI="http://doi.org/10.5281/zenodo.4761639" ID-Zenodo-Dep="4761639" httpUri="https://zenodo.org/record/4761639/files/figure.png" pageId="43" pageNumber="244" startId="43.[151,250,1933,1957]" targetBox="[181,1406,458,1909]" targetPageId="43">
<paragraph id="8D3236D9FFE1E50946C6B5DC8FC0F81A" blockId="43.[151,1437,1933,2029]" pageId="43" pageNumber="244">
<emphasis id="BFF9EACBFFE1E50946C6B5DC8C69F852" bold="true" box="[151,287,1933,1957]" pageId="43" pageNumber="244">FIGURE 26.</emphasis>
<taxonomicName id="4A8D4D5AFFE1E5094775B5DC8F6CF852" authority="Bouzan &amp; Iniesta &amp; Brescovit, 2021" authorityName="Bouzan &amp; Iniesta &amp; Brescovit" authorityYear="2021" box="[292,538,1933,1957]" class="Diplopoda" family="Chelodesmidae" genus="Folcloredesmus" kingdom="Animalia" order="Polydesmida" pageId="43" pageNumber="244" phylum="Arthropoda" rank="species" species="thomasi" status="sp. nov.">
<emphasis id="BFF9EACBFFE1E5094775B5DC8F6CF852" box="[292,538,1933,1957]" italics="true" pageId="43" pageNumber="244">Folcloredesmus thomasi</emphasis>
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<taxonomicNameLabel id="A4CA57B0FFE1E5094471B5DC8F2EF852" box="[544,600,1933,1957]" pageId="43" pageNumber="244" rank="species">n. sp.</taxonomicNameLabel>
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(holotype, IBSP 5452), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm. Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere.
</paragraph>
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<subSubSection id="C5976552FFE6E50E46C6B2C78E2AFF0E" pageId="44" pageNumber="245" type="diagnosis">
<paragraph id="8D3236D9FFE6E50E46C6B2C78E2AFF0E" blockId="44.[151,1437,150,789]" pageId="44" pageNumber="245">
<emphasis id="BFF9EACBFFE6E50E46C6B2C78C67FF46" bold="true" box="[151,273,150,177]" pageId="44" pageNumber="245">Diagnosis.</emphasis>
Males of
<taxonomicName id="4A8D4D5AFFE6E50E47C2B2C68F33FF46" authority="Bouzan &amp; Iniesta &amp; Brescovit, 2021" authorityName="Bouzan &amp; Iniesta &amp; Brescovit" authorityYear="2021" box="[403,581,151,177]" class="Diplopoda" family="Chelodesmidae" genus="Folcloredesmus" kingdom="Animalia" order="Polydesmida" pageId="44" pageNumber="245" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="BFF9EACBFFE6E50E47C2B2C68F33FF46" box="[403,581,151,177]" italics="true" pageId="44" pageNumber="245">Folcloredesmus</emphasis>
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<emphasis id="BFF9EACBFFE6E50E4400B2C68FD3FF46" bold="true" box="[593,677,151,177]" pageId="44" pageNumber="245">
<taxonomicNameLabel id="A4CA57B0FFE6E50E4400B2C68FD3FF46" box="[593,677,151,177]" pageId="44" pageNumber="245" rank="genus">n. gen.</taxonomicNameLabel>
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differ from those of other
<taxonomicName id="4A8D4D5AFFE6E50E45BEB2C68998FF46" authorityName="Hoffman" authorityYear="1976" box="[1007,1262,151,177]" class="Diplopoda" family="Chelodesmidae" kingdom="Animalia" order="Polydesmida" pageId="44" pageNumber="245" phylum="Arthropoda" rank="tribe" tribe="Arthrosolaenomeridini">Arthrosolaenomeridini</taxonomicName>
genera by the combination of the following characters: paranota sub-triangular (
<figureCitation id="15B62A5CFFE6E50E45DDB2EA8E83FF21" box="[908,1013,187,214]" captionStart="FIGURE 26" captionStartId="43.[151,250,1933,1957]" captionTargetBox="[181,1406,458,1909]" captionTargetId="figure-76@43.[181,1406,458,1909]" captionTargetPageId="43" captionText="FIGURE 26. Folcloredesmus thomasi n. sp. (holotype, IBSP 5452), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm.Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere." figureDoi="http://doi.org/10.5281/zenodo.4761639" httpUri="https://zenodo.org/record/4761639/files/figure.png" pageId="44" pageNumber="245">Fig. 26B</figureCitation>
); slight dorsal lobe on leg prefemur; presence of a spiniform process in gonocoxae (
<figureCitation id="15B62A5CFFE6E50E44F7B28E8E67FF0E" box="[678,785,223,249]" captionStart="FIGURE 26" captionStartId="43.[151,250,1933,1957]" captionTargetBox="[181,1406,458,1909]" captionTargetId="figure-76@43.[181,1406,458,1909]" captionTargetPageId="43" captionText="FIGURE 26. Folcloredesmus thomasi n. sp. (holotype, IBSP 5452), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm.Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere." figureDoi="http://doi.org/10.5281/zenodo.4761639" httpUri="https://zenodo.org/record/4761639/files/figure.png" pageId="44" pageNumber="245">Figs 26D</figureCitation>
,
<figureCitation id="15B62A5CFFE6E50E454CB28E8E39FF0E" box="[797,847,223,249]" captionStart="FIGURE 32" captionStartId="49.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-20@49.[161,1426,181,1680]" captionTargetPageId="49" captionText="FIGURE 32. Gonocoxae.A) Arthrosolaenomeris pantanalensis Schubart, 1943 (IBSP 530); B) Gangugia tapirapensis Schubart, 1947 (MNRJ 12008); C) Folcloredesmus thomasi n. sp. (CZUFMT 833); D) A. pantanalensis Schubart, 1943 (IBSP 530); E) F. thomasi n. sp. (CZUFMT 833); F) Abiliodesmus mapinguari n. sp. (CZUFMT 836). Scale bars: AB, DF) 1 mm.; C) 0,2 mm. Abbreviations: Cx, gonocoxae; RE, rounded extension; SP, spiniform process." figureDoi="http://doi.org/10.5281/zenodo.4761655" httpUri="https://zenodo.org/record/4761655/files/figure.png" pageId="44" pageNumber="245">32C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C5976552FFE6E50E4696B3538E93FCE2" pageId="44" pageNumber="245" type="description">
<paragraph id="8D3236D9FFE6E50E4696B3538EF3FDCA" blockId="44.[151,1437,150,789]" pageId="44" pageNumber="245">
<emphasis id="BFF9EACBFFE6E50E4696B3538C2FFEEB" bold="true" box="[199,345,258,284]" pageId="44" pageNumber="245">Description.</emphasis>
<emphasis id="BFF9EACBFFE6E50E470FB3528F41FEEA" box="[350,567,259,285]" italics="true" pageId="44" pageNumber="245">General characters</emphasis>
: Coloration (specimen long preserved in 70% ethanol) ocher with the paranota tip slightly whitish. Gnathochilarium: lingual plate covered by several setae; mentum only centrally covered by setae, with smooth edges and stipes completely covered by sparse setae. Body rings: cuticle slightly rough; alignment of paranota in posterior view straight. Sternite of the fourth segment with a pair of projections, sternite of the fifth segment with two pairs of projections covered with setae and sternite of sixth body ring with two pairs of ventral projections; seventh body ring with a pair of projections and post-gonopodal sternites with two pairs of rounded ventral projections (
<figureCitation id="15B62A5CFFE6E50E472CB38A8C9EFE02" box="[381,488,475,501]" captionStart="FIGURE 30" captionStartId="47.[151,250,1703,1727]" captionTargetBox="[161,1426,181,1679]" captionTargetId="figure-20@47.[161,1426,181,1679]" captionTargetPageId="47" captionText="FIGURE 30. Sternites, ventral projections. A) Arthrosolaenomeris pantanalensis Schubart, 1943 (IBSP 530), 6th body ring; B) Dioplosternus salvatrix Hoffman, 2005 (IBSP 1084), 6th body ring; C) Arthrosolaenomeris saci n. sp. (IBSP 1148), 7th body ring; D) Folcloredesmus thomasi n. sp. (CZUFMT 833), 9th body ring; E) A. saci n. sp., 9th body ring; F) Abiliodesmus planaltensis (Schubart, 1960) comb. nov. (IBSP 3570), 9th body ring. Scale bars: AC, 1 mm.; D, 0,5 mm.; E, 0,3 mm.; F, 0,3 mm." figureDoi="http://doi.org/10.5281/zenodo.4761651" httpUri="https://zenodo.org/record/4761651/files/figure.png" pageId="44" pageNumber="245">Fig. 30D</figureCitation>
). Paranota form: sub-triangular (
<figureCitation id="15B62A5CFFE6E50E450EB38A8EBEFE01" box="[863,968,475,502]" captionStart="FIGURE 26" captionStartId="43.[151,250,1933,1957]" captionTargetBox="[181,1406,458,1909]" captionTargetId="figure-76@43.[181,1406,458,1909]" captionTargetPageId="43" captionText="FIGURE 26. Folcloredesmus thomasi n. sp. (holotype, IBSP 5452), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm.Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere." figureDoi="http://doi.org/10.5281/zenodo.4761639" httpUri="https://zenodo.org/record/4761639/files/figure.png" pageId="44" pageNumber="245">Fig. 26B</figureCitation>
). Legs: slight dorsal lobe, a ventroapical projection in the prefemur of the anterior legs (
<figureCitation id="15B62A5CFFE6E50E44F3B3AE8E71FDEE" box="[674,775,511,537]" captionStart="FIGURE 31" captionStartId="48.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-24@48.[161,1426,181,1680]" captionTargetPageId="48" captionText="FIGURE 31. Legs. A) Arthrosolaenomeris pantanalensis Schubart, 1943 (IBSP 530); B) Gangugia boto n. sp. (holotype, IBSP 7539); CD) Abiliodesmus planaltensis (Schubart, 1960) comb. nov. (IBSP 3570); EF) Gangugia tapirapensis Schubart, 1947 (MNRJ 12008). Scale bars: A) 2 mm.; B) 1 mm.; C) 1 mm.; D) 1 mm.; E) 500 µm.; F)1 mm. Abbreviations: co, coxae; f, femur; prf, prefemur; pst, postfemur; t, tarsus; tb, tibia; tc; tarsal claw." figureDoi="http://doi.org/10.5281/zenodo.4761653" httpUri="https://zenodo.org/record/4761653/files/figure.png" pageId="44" pageNumber="245">Fig. 31E</figureCitation>
), and presence of granules in the tibia and tarsus; thin setae on the ventral part of the podomeres in the anterior legs (
<figureCitation id="15B62A5CFFE6E50E4544B0728E0EFDCA" box="[789,888,547,573]" captionStart="FIGURE 31" captionStartId="48.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-24@48.[161,1426,181,1680]" captionTargetPageId="48" captionText="FIGURE 31. Legs. A) Arthrosolaenomeris pantanalensis Schubart, 1943 (IBSP 530); B) Gangugia boto n. sp. (holotype, IBSP 7539); CD) Abiliodesmus planaltensis (Schubart, 1960) comb. nov. (IBSP 3570); EF) Gangugia tapirapensis Schubart, 1947 (MNRJ 12008). Scale bars: A) 2 mm.; B) 1 mm.; C) 1 mm.; D) 1 mm.; E) 500 µm.; F)1 mm. Abbreviations: co, coxae; f, femur; prf, prefemur; pst, postfemur; t, tarsus; tb, tibia; tc; tarsal claw." figureDoi="http://doi.org/10.5281/zenodo.4761653" httpUri="https://zenodo.org/record/4761653/files/figure.png" pageId="44" pageNumber="245">Fig. 31F</figureCitation>
).
</paragraph>
<paragraph id="8D3236D9FFE6E50E4696B0168E5EFD05" blockId="44.[151,1437,150,789]" pageId="44" pageNumber="245">
<emphasis id="BFF9EACBFFE6E50E4696B0168CF4FD96" box="[199,386,583,609]" italics="true" pageId="44" pageNumber="245">Male characters</emphasis>
: Gonopod aperture on seventh body ring with the posterior margin excavated, concave and transverse sulci; without projections. Gonopods: gonocoxae equivalent to about half the length of the telopodite, cylindrical, with a spiniform process (
<figureCitation id="15B62A5CFFE6E50E440DB0DE8FBDFD5D" box="[604,715,655,682]" captionStart="FIGURE 26" captionStartId="43.[151,250,1933,1957]" captionTargetBox="[181,1406,458,1909]" captionTargetId="figure-76@43.[181,1406,458,1909]" captionTargetPageId="43" captionText="FIGURE 26. Folcloredesmus thomasi n. sp. (holotype, IBSP 5452), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm.Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere." figureDoi="http://doi.org/10.5281/zenodo.4761639" httpUri="https://zenodo.org/record/4761639/files/figure.png" pageId="44" pageNumber="245">Fig. 26D</figureCitation>
). Prefemoral process: elongate and thin (
<figureCitation id="15B62A5CFFE6E50E42E4B0DE8857FD5D" box="[1205,1313,655,682]" captionStart="FIGURE 26" captionStartId="43.[151,250,1933,1957]" captionTargetBox="[181,1406,458,1909]" captionTargetId="figure-76@43.[181,1406,458,1909]" captionTargetPageId="43" captionText="FIGURE 26. Folcloredesmus thomasi n. sp. (holotype, IBSP 5452), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm.Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere." figureDoi="http://doi.org/10.5281/zenodo.4761639" httpUri="https://zenodo.org/record/4761639/files/figure.png" pageId="44" pageNumber="245">Fig. 26D</figureCitation>
) and with indentations in the apical portion, in ectal view (
<figureCitation id="15B62A5CFFE6E50E4495B0E28E5BFD3A" box="[708,813,691,717]" captionStart="FIGURE 34" captionStartId="51.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-20@51.[161,1426,181,1680]" captionTargetPageId="51" captionText="FIGURE 34. Gonopod. (AC) Ectal view. A) Abiliodesmus mapinguari n. sp. (CZUFMT 836); B) Folcloredesmus thomasi n. sp. (CZUFMT 833); C) Arthrosolaenomeris saci n. sp. (IBSP 1148). (DF) Ventral view. D) Arthrosolaenomeris saci n. sp. (IBSP 1148); E) Gangugia cuca n. sp. (holotype, IBSP 7600); F) Chondrodesmus sp. (IBSP 3823). Scale bars: AD) 1 mm.; E) 0,5 mm.; F) 0,3 mm. Arrows refer to cingulum. Abbreviations: PfP, prefemoral process; S, solenomere." figureDoi="http://doi.org/10.5281/zenodo.4761659" httpUri="https://zenodo.org/record/4761659/files/figure.png" pageId="44" pageNumber="245">Fig. 34B</figureCitation>
). Cingulum in medial position (
<figureCitation id="15B62A5CFFE6E50E42CAB0E28877FD39" box="[1179,1281,691,718]" captionStart="FIGURE 26" captionStartId="43.[151,250,1933,1957]" captionTargetBox="[181,1406,458,1909]" captionTargetId="figure-76@43.[181,1406,458,1909]" captionTargetPageId="43" captionText="FIGURE 26. Folcloredesmus thomasi n. sp. (holotype, IBSP 5452), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm.Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere." figureDoi="http://doi.org/10.5281/zenodo.4761639" httpUri="https://zenodo.org/record/4761639/files/figure.png" pageId="44" pageNumber="245">Fig. 26F</figureCitation>
). Solenomere also long and thin with apex slightly curved (
<figureCitation id="15B62A5CFFE6E50E44C0B0868E6AFD05" box="[657,796,727,754]" captionStart="FIGURE 26" captionStartId="43.[151,250,1933,1957]" captionTargetBox="[181,1406,458,1909]" captionTargetId="figure-76@43.[181,1406,458,1909]" captionTargetPageId="43" captionText="FIGURE 26. Folcloredesmus thomasi n. sp. (holotype, IBSP 5452), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm.Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere." figureDoi="http://doi.org/10.5281/zenodo.4761639" httpUri="https://zenodo.org/record/4761639/files/figure.png" pageId="44" pageNumber="245">Figs 26DF</figureCitation>
).
</paragraph>
<paragraph id="8D3236D9FFE6E50E4696B0AA8E93FCE2" blockId="44.[151,1437,150,789]" box="[199,997,763,789]" pageId="44" pageNumber="245">
<emphasis id="BFF9EACBFFE6E50E4696B0AA8CECFCE2" box="[199,410,763,789]" italics="true" pageId="44" pageNumber="245">Female characters</emphasis>
: Vulvae: oval-shaped, sub-triangular in lateral view.
</paragraph>
</subSubSection>
<caption id="D9F26651FFE6E50E46C6B52C8FCCF82A" ID-DOI="http://doi.org/10.5281/zenodo.4761641" ID-Zenodo-Dep="4761641" httpUri="https://zenodo.org/record/4761641/files/figure.png" pageId="44" pageNumber="245" startId="44.[151,250,1917,1941]" targetBox="[158,1430,813,1894]" targetPageId="44">
<paragraph id="8D3236D9FFE6E50E46C6B52C8FCCF82A" blockId="44.[151,1437,1917,2013]" pageId="44" pageNumber="245">
<emphasis id="BFF9EACBFFE6E50E46C6B52C8C52F862" bold="true" box="[151,292,1917,1941]" pageId="44" pageNumber="245">FIGURE 27.</emphasis>
Left vulva.
<taxonomicName id="4A8D4D5AFFE6E50E47FAB52C8FC5F862" authority="Bouzan &amp; Iniesta &amp; Brescovit, 2021" authorityName="Bouzan &amp; Iniesta &amp; Brescovit" authorityYear="2021" box="[427,691,1917,1941]" class="Diplopoda" family="Chelodesmidae" genus="Abiliodesmus" kingdom="Animalia" order="Polydesmida" pageId="44" pageNumber="245" phylum="Arthropoda" rank="species" species="mapinguari" status="sp. nov.">
<emphasis id="BFF9EACBFFE6E50E47FAB52C8FC5F862" box="[427,691,1917,1941]" italics="true" pageId="44" pageNumber="245">Abiliodesmus mapinguari</emphasis>
</taxonomicName>
<emphasis id="BFF9EACBFFE6E50E44ECB52C8F8CF862" bold="true" box="[701,762,1917,1941]" pageId="44" pageNumber="245">
<taxonomicNameLabel id="A4CA57B0FFE6E50E44ECB52C8F8CF862" box="[701,762,1917,1941]" pageId="44" pageNumber="245" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(paratype, CZUFMT 834): (A), ventral view; (B), lateral view.
<taxonomicName id="4A8D4D5AFFE6E50E46C6B5F08CF9F84E" authority="Bouzan &amp; Iniesta &amp; Brescovit, 2021" authorityName="Bouzan &amp; Iniesta &amp; Brescovit" authorityYear="2021" box="[151,399,1953,1977]" class="Diplopoda" family="Chelodesmidae" genus="Folcloredesmus" kingdom="Animalia" order="Polydesmida" pageId="44" pageNumber="245" phylum="Arthropoda" rank="species" species="thomasi" status="sp. nov.">
<emphasis id="BFF9EACBFFE6E50E46C6B5F08CF9F84E" box="[151,399,1953,1977]" italics="true" pageId="44" pageNumber="245">Folcloredesmus thomasi</emphasis>
</taxonomicName>
<emphasis id="BFF9EACBFFE6E50E47C7B5F08CA6F84E" bold="true" box="[406,464,1953,1977]" pageId="44" pageNumber="245">
<taxonomicNameLabel id="A4CA57B0FFE6E50E47C7B5F08CA6F84E" box="[406,464,1953,1977]" pageId="44" pageNumber="245" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(paratype, IBSP 5453): (C), ventral view; (D), lateral view. Scale bars: 0,5 mm. Abbreviations: EV, external valve; IV, internal valve; Op, operculum.
</paragraph>
</caption>
<subSubSection id="C5976552FFE7E50F4696B2C78FB0FF46" box="[199,710,150,177]" pageId="45" pageNumber="246" type="distribution">
<paragraph id="8D3236D9FFE7E50F4696B2C78FB0FF46" blockId="45.[151,1436,150,321]" box="[199,710,150,177]" pageId="45" pageNumber="246">
<emphasis id="BFF9EACBFFE7E50F4696B2C78C17FF47" bold="true" box="[199,353,150,176]" pageId="45" pageNumber="246">Distribution.</emphasis>
State of
<collectingRegion id="4F49F83BFFE7E50F4794B2C68F20FF46" box="[453,598,151,177]" country="Brazil" name="Mato Grosso" pageId="45" pageNumber="246">Mato Grosso</collectingRegion>
(
<figureCitation id="15B62A5CFFE7E50F4437B2C68FCCFF46" box="[614,698,151,177]" captionStart="FIGURE 28" captionStartId="45.[151,250,1524,1548]" captionTargetBox="[151,1436,345,1501]" captionTargetId="figure-76@45.[151,1436,345,1501]" captionTargetPageId="45" captionText="FIGURE 28. Distribution map of Angelodesmus Schubart, 1962 and Folcloredesmus n. gen. species." figureDoi="http://doi.org/10.5281/zenodo.4761645" httpUri="https://zenodo.org/record/4761645/files/figure.png" pageId="45" pageNumber="246">Fig. 28</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C5976552FFE7E50F4696B2EA8E2CFEB6" pageId="45" pageNumber="246" type="etymology">
<paragraph id="8D3236D9FFE7E50F4696B2EA8FBCFEEA" blockId="45.[151,1436,150,321]" pageId="45" pageNumber="246">
<emphasis id="BFF9EACBFFE7E50F4696B2EA8C38FF22" bold="true" box="[199,334,187,213]" pageId="45" pageNumber="246">Etymology.</emphasis>
The genus name comes from the Portuguese word for “Folclore” (culture shared by a particular group of people, as the tales characters used as species epithet in this paper) with the combining stem “-desmus”, commonly used in generic names in
<taxonomicName id="4A8D4D5AFFE7E50F4463B3528FB0FEEA" authorityName="Leach" authorityYear="1814" box="[562,710,259,285]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="45" pageNumber="246" phylum="Arthropoda" rank="order">Polydesmida</taxonomicName>
.
</paragraph>
<paragraph id="8D3236D9FFE7E50F4696B3778E2CFEB6" blockId="45.[151,1436,150,321]" box="[199,858,294,321]" pageId="45" pageNumber="246">
<emphasis id="BFF9EACBFFE7E50F4696B3778C11FEB7" bold="true" box="[199,359,294,320]" pageId="45" pageNumber="246">Composition.</emphasis>
One species:
<taxonomicName id="4A8D4D5AFFE7E50F4452B3768E63FEB6" authority="Bouzan &amp; Iniesta &amp; Brescovit, 2021" authorityName="Bouzan &amp; Iniesta &amp; Brescovit" authorityYear="2021" box="[515,789,295,321]" class="Diplopoda" family="Chelodesmidae" genus="Folcloredesmus" kingdom="Animalia" order="Polydesmida" pageId="45" pageNumber="246" phylum="Arthropoda" rank="species" species="thomasi" status="sp. nov.">
<emphasis id="BFF9EACBFFE7E50F4452B3768E63FEB6" box="[515,789,295,321]" italics="true" pageId="45" pageNumber="246">Folcloredesmus thomasi</emphasis>
</taxonomicName>
<emphasis id="BFF9EACBFFE7E50F454AB3768E2CFEB6" bold="true" box="[795,858,295,321]" pageId="45" pageNumber="246">
<taxonomicNameLabel id="A4CA57B0FFE7E50F454AB3768E2CFEB6" box="[795,858,295,321]" pageId="45" pageNumber="246" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>