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<document id="1D32E8A5403BB40B51B14A95A40274F8" ID-CLB-Dataset="6318" ID-DOI="10.11646/zootaxa.4970.2.1" ID-GBIF-Dataset="89d3c795-198b-4162-8903-9c6cf3f1e665" ID-ISSN="1175-5326" ID-Zenodo-Dep="4761544" ID-ZooBank="6DBA0C02-9E52-4FD0-8B48-8C7B405713A8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620984897660" checkinUser="plazi" docAuthor="Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M. &amp; Brescovit, Antonio Domingos" docDate="2021" docId="052487CFFFC6E52E46C6B2C6896FFB76" docLanguage="en" docName="zootaxa.4970.2.1.pdf" docOrigin="Zootaxa 4970 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Arthrosolaenomeris chapadensis Schubart 1943" docType="treatment" docVersion="6" lastPageNumber="213" masterDocId="F91DFFB7FFCAE5224651B2518D76FFF7" masterDocTitle="Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae)" masterLastPageNumber="256" masterPageNumber="201" pageNumber="213" updateTime="1698939836712" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="BAC989EFA24191C8CADFF5D6805FF89D">Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae)</mods:title>
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<mods:namePart id="74466F56FCF3434DC2440896DC022F08">Bouzan, Rodrigo Salvador</mods:namePart>
<mods:affiliation id="6CE527E74EB8FE15CCF9E2DCE28948A0">Pós-graduação em Zoologia, Instituto de Biociências, Universidade de São Paulo, São Paulo, Brasil.</mods:affiliation>
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<mods:namePart id="551AD6652773916578E8148178BDA6A8">Iniesta, Luiz Felipe M.</mods:namePart>
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<mods:affiliation id="E5F55435B2E570C4711B4F7B415C5FB5">Pós-graduação em Zoologia, Instituto de Biociências, Universidade de São Paulo, São Paulo, Brasil. &amp; luiz-moretti @ hotmail. com; https: // orcid. org / 0000 - 0002 - 0529 - 4162</mods:affiliation>
<mods:nameIdentifier id="63F199EC6736E572E37B372E84B88EC5" type="email">luiz-moretti@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="F019F052CD1F7A885C2348C6A2F6CEE2">Brescovit, Antonio Domingos</mods:namePart>
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<mods:affiliation id="AFD66E58420EE53CBB8396F786FCDAB5">antonio. brescovit @ butantan. gov. br; https: // orcid. org / 0000 - 0002 - 1511 - 5324</mods:affiliation>
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<treatment id="052487CFFFC6E52E46C6B2C6896FFB76" ID-DOI="http://doi.org/10.5281/zenodo.4761557" ID-GBIF-Taxon="182486297" ID-Zenodo-Dep="4761557" LSID="urn:lsid:plazi:treatment:052487CFFFC6E52E46C6B2C6896FFB76" httpUri="http://treatment.plazi.org/id/052487CFFFC6E52E46C6B2C6896FFB76" lastPageNumber="213" pageId="12" pageNumber="213">
<subSubSection id="C5976552FFC6E52E46C6B2C68F9FFF46" box="[151,745,151,178]" pageId="12" pageNumber="213" type="nomenclature">
<paragraph id="8D3236D9FFC6E52E46C6B2C68F9FFF46" blockId="12.[151,745,151,213]" box="[151,745,151,178]" pageId="12" pageNumber="213">
<heading id="D67A81B5FFC6E52E46C6B2C68F9FFF46" bold="true" box="[151,745,151,178]" fontSize="11" level="1" pageId="12" pageNumber="213" reason="1">
<taxonomicName id="4A8D4D5AFFC6E52E46C6B2C68F9FFF46" ID-CoL="67SD4" authority="Schubart, 1943" authorityName="Schubart" authorityYear="1943" box="[151,745,151,178]" class="Diplopoda" family="Chelodesmidae" genus="Arthrosolaenomeris" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="chapadensis">
<emphasis id="BFF9EACBFFC6E52E46C6B2C68F9FFF46" bold="true" box="[151,745,151,178]" pageId="12" pageNumber="213">
<emphasis id="BFF9EACBFFC6E52E46C6B2C68F52FF45" bold="true" box="[151,548,151,178]" italics="true" pageId="12" pageNumber="213">Arthrosolaenomeris chapadensis</emphasis>
<bibRefCitation id="E91C4B28FFC6E52E447AB2C68F9FFF46" author="Schubart, O." box="[555,745,151,177]" pageId="12" pageNumber="213" pagination="127 - 164" refId="ref33096" refString="Schubart, O. (1943) Especies novas das familias Strongylosomidae e Leptodesmidae da ordem Proterospermophora do interior dos Estados de Sao Paulo e Mato Grosso. Papeis Avulsos de Zoologia, 3, 127 - 164." type="journal article" year="1943">Schubart, 1943</bibRefCitation>
</emphasis>
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<subSubSection id="C5976552FFC6E52E46C6B2EA8CFEFF22" box="[151,392,187,213]" pageId="12" pageNumber="213" type="description">
<paragraph id="8D3236D9FFC6E52E46C6B2EA8CFEFF22" blockId="12.[151,745,151,213]" box="[151,392,187,213]" pageId="12" pageNumber="213">
<figureCitation id="15B62A5CFFC6E52E46C6B2EA8C74FF22" box="[151,258,187,213]" captionStart="FIGURE 4" captionStartId="13.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-20@13.[161,1426,181,1680]" captionTargetPageId="13" captionText="FIGURE 4. Arthrosolaenomeris chapadensis Schubart, 1943 (holotype, MZSP 1092), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm. Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere; SPr, secondary process of the prefemoral process." figureDoi="http://doi.org/10.5281/zenodo.4761555" httpUri="https://zenodo.org/record/4761555/files/figure.png" pageId="12" pageNumber="213">Figures 4</figureCitation>
,
<figureCitation id="15B62A5CFFC6E52E475FB2EA8C17FF22" box="[270,353,187,213]" captionStart="FIGURE 10" captionStartId="21.[151,250,1798,1822]" captionTargetBox="[314,1273,181,1773]" captionTargetId="figure-20@21.[314,1273,181,1773]" captionTargetPageId="21" captionText="FIGURE 10. Left vulva. Arthrosolaenomeris chapadensis Schubart, 1943 (IBSP 1049): (A), ventral view; (B), lateral view. Arthrosolaenomeris pantanalensis Schubart, 1943 (CZUFMT 847): (C), ventral view; (D), lateral view. Arthrosolaenomeris saci n. sp. (paratype, IBSP 2577): (E), ventral view; (F), lateral view.Arthrosolaenomeris iara n. sp. (paratype, IBSP 7537). (G), ventral view; (H), lateral view. Scale bars: 0,5 mm. Abbreviations: EV, external valve; IV, internal valve; Op, operculum." figureDoi="http://doi.org/10.5281/zenodo.4761577" httpUri="https://zenodo.org/record/4761577/files/figure.png" pageId="12" pageNumber="213">10AB</figureCitation>
,
<figureCitation id="15B62A5CFFC6E52E473CB2EA8CFEFF22" box="[365,392,187,213]" captionStart="FIGURE 11" captionStartId="22.[151,250,1361,1385]" captionTargetBox="[151,1436,181,1337]" captionTargetId="figure-24@22.[151,1436,181,1337]" captionTargetPageId="22" captionText="FIGURE 11. Distribution map of Arthrosolaenomeris Schubart, 1943 species." figureDoi="http://doi.org/10.5281/zenodo.4761582" httpUri="https://zenodo.org/record/4761582/files/figure.png" pageId="12" pageNumber="213">11</figureCitation>
</paragraph>
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<subSubSection id="C5976552FFC6E52E46C6B35189A3FE46" pageId="12" pageNumber="213" type="reference_group">
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<treatmentCitation id="0C2C10C8FFC6E52E46C6B3518FB7FEEF" author="Schubart, O." box="[151,705,256,280]" page="144" pageId="12" pageNumber="213" year="1943">
<taxonomicName id="4A8D4D5AFFC6E52E46C6B3518FB7FEEF" ID-CoL="67SD4" authority="Schubart, 1943: 144" authorityName="Schubart" authorityPageNumber="144" authorityYear="1943" box="[151,705,256,280]" class="Diplopoda" family="Chelodesmidae" genus="Arthrosolaenomeris" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="chapadensis">
<emphasis id="BFF9EACBFFC6E52E46C6B3518C90FEEF" box="[151,486,256,280]" italics="true" pageId="12" pageNumber="213">Arthrosolaenomeris chapadensis</emphasis>
<bibRefCitation id="E91C4B28FFC6E52E47BEB3518FB7FEEF" author="Schubart, O." box="[495,705,256,280]" pageId="12" pageNumber="213" pagination="127 - 164" refId="ref33096" refString="Schubart, O. (1943) Especies novas das familias Strongylosomidae e Leptodesmidae da ordem Proterospermophora do interior dos Estados de Sao Paulo e Mato Grosso. Papeis Avulsos de Zoologia, 3, 127 - 164." type="journal article" year="1943">Schubart, 1943: 144</bibRefCitation>
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, 160, 161, figs 2838. Male
<typeStatus id="5236887BFFC6E52E45A0B351893CFEEF" box="[1009,1098,256,280]" pageId="12" pageNumber="213" type="holotype">holotype</typeStatus>
from Indubrasil, Campo Grande (
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,
<geoCoordinate id="E8B9501EFFC6E52E4702B34F8CAEFEC1" box="[339,472,286,310]" degrees="54" direction="west" minutes="38" orientation="longitude" pageId="12" pageNumber="213" precision="15" seconds="47" value="-54.64639">54°3847”W</geoCoordinate>
),
<collectingRegion id="4F49F83BFFC6E52E47B8B34E8FC5FEC1" box="[489,691,286,310]" country="Brazil" name="Mato Grosso do Sul" pageId="12" pageNumber="213">Mato Grosso do Sul</collectingRegion>
,
<collectingCountry id="F59A7649FFC6E52E44EDB34F8F8AFEC1" box="[700,764,286,310]" name="Brazil" pageId="12" pageNumber="213">Brazil</collectingCountry>
,
<date id="F9331019FFC6E52E4557B34F8E05FEC1" box="[774,883,286,310]" pageId="12" pageNumber="213" value="1938-10-17">17.X.1938</date>
, Instituto Osvaldo Cruz coll., deposited in MZSP 1092, examined.
<typeStatus id="5236887BFFC6E52E4767B36C8CEBFEA2" box="[310,413,317,341]" pageId="12" pageNumber="213" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9B8BFD50FFC6E52E47F7B36C8F77FEA2" box="[422,513,317,341]" pageId="12" pageNumber="213" type="male">one male</specimenCount>
from Camapuã (
<geoCoordinate id="E8B9501EFFC6E52E44FDB36C8E50FEA2" box="[684,806,317,341]" degrees="19" direction="south" minutes="31" orientation="latitude" pageId="12" pageNumber="213" precision="15" seconds="54" value="-19.531666">19°3154”S</geoCoordinate>
,
<geoCoordinate id="E8B9501EFFC6E52E4560B36C8EC3FEA2" box="[817,949,317,341]" degrees="54" direction="west" minutes="02" orientation="longitude" pageId="12" pageNumber="213" precision="15" seconds="37" value="-54.04361">54°0237”W</geoCoordinate>
),
<collectingRegion id="4F49F83BFFC6E52E4596B36C89E2FEA2" box="[967,1172,317,341]" country="Brazil" name="Mato Grosso do Sul" pageId="12" pageNumber="213">Mato Grosso do Sul</collectingRegion>
,
<collectingCountry id="F59A7649FFC6E52E42CFB36C89A8FEA2" box="[1182,1246,317,341]" name="Brazil" pageId="12" pageNumber="213">Brazil</collectingCountry>
,
<date id="F9331019FFC6E52E42B9B36C8840FEA2" box="[1256,1334,317,341]" pageId="12" pageNumber="213" value="1938-10">X.1938</date>
, Camargo coll., deposited in MZSP 1090 and
<specimenCount id="9B8BFD50FFC6E52E446EB30D8FDCFE83" box="[575,682,348,372]" pageId="12" pageNumber="213" type="male">two males</specimenCount>
from Porto Murtinho (
<geoCoordinate id="E8B9501EFFC6E52E45CEB30A896CFE84" box="[927,1050,347,371]" degrees="21" direction="south" minutes="41" orientation="latitude" pageId="12" pageNumber="213" precision="15" seconds="56" value="-21.698889">21°4156”S</geoCoordinate>
,
<geoCoordinate id="E8B9501EFFC6E52E4276B30A89DAFE84" box="[1063,1196,347,371]" degrees="57" direction="west" minutes="52" orientation="longitude" pageId="12" pageNumber="213" precision="15" seconds="56" value="-57.88222">57°5256”W</geoCoordinate>
),
<collectingRegion id="4F49F83BFFC6E52E4290B30D88EEFE83" box="[1217,1432,348,372]" country="Brazil" name="Mato Grosso do Sul" pageId="12" pageNumber="213">Mato Grosso do Sul</collectingRegion>
,
<collectingCountry id="F59A7649FFC6E52E4696B32B8C71FE65" box="[199,263,378,402]" name="Brazil" pageId="12" pageNumber="213">Brazil</collectingCountry>
,
<date id="F9331019FFC6E52E4742B32B8C04FE65" box="[275,370,378,402]" pageId="12" pageNumber="213" value="1929-12">XII.1929</date>
, R. Spitz coll. deposited in MZSP 522 and 536, respectively, examined;
<treatmentCitation id="0C2C10C8FFC6E52E420CB32B8859FE65" author="Schubart, O." box="[1117,1327,378,402]" page="238" pageId="12" pageNumber="213" year="1958">
<bibRefCitation id="E91C4B28FFC6E52E420CB32B8859FE65" author="Schubart, O." box="[1117,1327,378,402]" pageId="12" pageNumber="213" pagination="203 - 252" refId="ref33204" refString="Schubart, O. (1958) Sobre alguns Diplopoda de Mato Grosso e Goias, Brasil e a familia Spirostreptidae. Arquivos do Museu Nacional, 46, 203 - 252." type="journal article" year="1958">Schubart, 1958: 238</bibRefCitation>
</treatmentCitation>
;
<bibRefCitation id="E91C4B28FFC6E52E436AB32B8D8BFE46" author="Hoffman, R. L." pageId="12" pageNumber="213" pagination="171 - 183" refId="ref32248" refString="Hoffman, R. L. (1976) Chelodesmid studies IX. A synopsis of the new Brazilian tribe Arthrosolaenomeridini (Diplopoda; Polydesmida). Papeis Avulsos de Zoologia, 30, 171 - 183." type="journal article" year="1976">Hoffman, 1976</bibRefCitation>
; 177;
<treatmentCitation id="0C2C10C8FFC6E52E476AB3C889A7FE46" author="Golovatch, S. I. &amp; Hoffman, R. L. &amp; Adis, J. &amp; Marques, M. I. &amp; Raizer, J. &amp; Silva, F. H. O. &amp; Ribeiro, R. A. K. &amp; Silva, J. L. &amp; Pinheiro, T. G." box="[315,1233,409,433]" page="276" pageId="12" pageNumber="213" year="2005">
<bibRefCitation id="E91C4B28FFC6E52E476AB3C889A7FE46" author="Golovatch, S. I. &amp; Hoffman, R. L. &amp; Adis, J. &amp; Marques, M. I. &amp; Raizer, J. &amp; Silva, F. H. O. &amp; Ribeiro, R. A. K. &amp; Silva, J. L. &amp; Pinheiro, T. G." box="[315,1233,409,433]" pageId="12" pageNumber="213" pagination="273 - 288" refId="ref32110" refString="Golovatch, S. I., Hoffman, R. L., Adis, J., Marques, M. I., Raizer, J., Silva, F. H. O., Ribeiro, R. A. K., Silva, J. L. &amp; Pinheiro, T. G. (2005) Millipedes of the Brazilian Pantanal. Amazoniana, 18 (3 / 4), 273 - 288." type="journal article" year="2005">Golovatch, Hoffman, Adis, Marques, Raizer, Silva, Ribeiro, Silva and Pinheiro, 2005: 276</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C5976552FFC6E52E46C6B38E8832FDEA" pageId="12" pageNumber="213" type="materials_examined">
<paragraph id="8D3236D9FFC6E52E46C6B38E8832FDEA" blockId="12.[151,1437,479,1154]" pageId="12" pageNumber="213">
<emphasis id="BFF9EACBFFC6E52E46C6B38E8CFBFE0E" bold="true" box="[151,397,479,505]" pageId="12" pageNumber="213">Additional material.</emphasis>
<materialsCitation id="3DE53C84FFC6E52E47C9B38E8849FDEA" ID-GBIF-Occurrence="3125622303" collectionCode="IBSP" collectorName="Equipe Butantan" country="Brazil" latitude="-22.579723" location="Rosana" longLatPrecision="21" longitude="-53.058887" pageId="12" pageNumber="213" specimenCode="IBSP 1049" specimenCount="3" specimenCount-female="1" specimenCount-juvenile="1" specimenCount-male="1" stateProvince="Sao Paulo">
<collectingCountry id="F59A7649FFC6E52E47C9B38E8F74FE0E" box="[408,514,479,505]" name="Brazil" pageId="12" pageNumber="213">BRAZIL</collectingCountry>
:
<emphasis id="BFF9EACBFFC6E52E445EB38E8FF1FE0E" box="[527,647,479,505]" italics="true" pageId="12" pageNumber="213">
<collectingRegion id="4F49F83BFFC6E52E445EB38E8FF1FE0E" box="[527,647,479,505]" country="Brazil" name="Sao Paulo" pageId="12" pageNumber="213">São Paulo</collectingRegion>
</emphasis>
:
<location id="88526002FFC6E52E44C8B3B18F9DFE0D" LSID="urn:lsid:plazi:treatment:052487CFFFC6E52E46C6B2C6896FFB76:88526002FFC6E52E44C8B3B18F9DFE0D" box="[665,747,480,506]" country="Brazil" latitude="-22.579723" longLatPrecision="21" longitude="-53.058887" name="Rosana" pageId="12" pageNumber="213" stateProvince="Sao Paulo">Rosana</location>
(
<geoCoordinate id="E8B9501EFFC6E52E44AFB38E8EF0FE0E" box="[766,902,479,505]" degrees="22" direction="south" minutes="34" orientation="latitude" pageId="12" pageNumber="213" precision="15" seconds="47" value="-22.579723">22°3447”S</geoCoordinate>
,
<geoCoordinate id="E8B9501EFFC6E52E45C4B38E895EFE0D" box="[917,1064,479,506]" degrees="53" direction="west" minutes="03" orientation="longitude" pageId="12" pageNumber="213" precision="15" seconds="32" value="-53.058887">53°0332”W</geoCoordinate>
), Usina Hidrelétrica Engenheiro Sérgio Motta (Porto Primavera),
<specimenCount id="9B8BFD50FFC6E52E4459B0528F5AFDEA" box="[520,556,515,541]" pageId="12" pageNumber="213" type="male">1♂</specimenCount>
,
<specimenCount id="9B8BFD50FFC6E52E4468B0528F2AFDEA" box="[569,604,515,541]" pageId="12" pageNumber="213" type="female">1♀</specimenCount>
and
<specimenCount id="9B8BFD50FFC6E52E44C2B0558E69FDEA" box="[659,799,515,542]" pageId="12" pageNumber="213" type="juvenile">one juvenile</specimenCount>
, 19992000,
<collectorName id="2078530FFFC6E52E45E7B0528907FDE9" box="[950,1137,515,542]" pageId="12" pageNumber="213">Equipe Butantan</collectorName>
coll. (
<specimenCode id="DD2B9EA2FFC6E52E42E9B0528841FDEA" box="[1208,1335,515,541]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="12" pageNumber="213">IBSP 1049</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C5976552FFC6E52E4696B0768C95FD7E" pageId="12" pageNumber="213" type="diagnosis">
<paragraph id="8D3236D9FFC6E52E4696B0768C95FD7E" blockId="12.[151,1437,479,1154]" pageId="12" pageNumber="213">
<emphasis id="BFF9EACBFFC6E52E4696B0768C34FDB6" bold="true" box="[199,322,551,577]" pageId="12" pageNumber="213">Diagnosis.</emphasis>
Adult males differ from other species of the genus by the combination of the following characters: presence of indentations in the prefemoral process (
<figureCitation id="15B62A5CFFC6E52E4485B01A8E5AFD92" box="[724,812,587,613]" captionStart="FIGURE 4" captionStartId="13.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-20@13.[161,1426,181,1680]" captionTargetPageId="13" captionText="FIGURE 4. Arthrosolaenomeris chapadensis Schubart, 1943 (holotype, MZSP 1092), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm. Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere; SPr, secondary process of the prefemoral process." figureDoi="http://doi.org/10.5281/zenodo.4761555" httpUri="https://zenodo.org/record/4761555/files/figure.png" pageId="12" pageNumber="213">Fig. 4D</figureCitation>
) and absence of a conspicuous groove in the apex of the prefemoral process (
<figureCitation id="15B62A5CFFC6E52E472DB03E8CA1FD7E" box="[380,471,623,649]" captionStart="FIGURE 4" captionStartId="13.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-20@13.[161,1426,181,1680]" captionTargetPageId="13" captionText="FIGURE 4. Arthrosolaenomeris chapadensis Schubart, 1943 (holotype, MZSP 1092), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm. Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere; SPr, secondary process of the prefemoral process." figureDoi="http://doi.org/10.5281/zenodo.4761555" httpUri="https://zenodo.org/record/4761555/files/figure.png" pageId="12" pageNumber="213">Fig. 4D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C5976552FFC6E52E4696B0C2895CFBA9" pageId="12" pageNumber="213" type="description">
<paragraph id="8D3236D9FFC6E52E4696B0C2894CFC3A" blockId="12.[151,1437,479,1154]" pageId="12" pageNumber="213">
<emphasis id="BFF9EACBFFC6E52E4696B0C28C2FFD5A" bold="true" box="[199,345,659,685]" pageId="12" pageNumber="213">Description.</emphasis>
Male (
<typeStatus id="5236887BFFC6E52E47FAB0C28F6EFD5A" box="[427,536,659,685]" pageId="12" pageNumber="213" type="holotype">Holotype</typeStatus>
, MZSP 1092). Coloration (long-preserved in 70% ethanol): Head and antennae ocher. Body ocher and paranota tip with the same color as the body (
<figureCitation id="15B62A5CFFC6E52E45F7B0E6895EFD26" box="[934,1064,695,721]" captionStart="FIGURE 4" captionStartId="13.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-20@13.[161,1426,181,1680]" captionTargetPageId="13" captionText="FIGURE 4. Arthrosolaenomeris chapadensis Schubart, 1943 (holotype, MZSP 1092), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm. Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere; SPr, secondary process of the prefemoral process." figureDoi="http://doi.org/10.5281/zenodo.4761555" httpUri="https://zenodo.org/record/4761555/files/figure.png" pageId="12" pageNumber="213">Figs 4AC</figureCitation>
). Legs ocher. Telson ocher. Total length: 65.98. Total width: 10.02. Collum, length 3.10, width 9.33. Antennomere lengths (1&gt;7): 0.71; 1.91; 1.82; 1.62; 1.56; 1.51; 0.39. Podomeres lengths (1&gt;7): 1.03; 1.60; 2.78; 1.52; 1.96; 1.97; 0.68. Gonopod aperture, length 1.98, width 3.35. Telson, length 0.77. Gonopod: length 2.86, width 2.07. Gonocoxae: length 1.11, width 1.02. Telopodite: length 2.77, width 1.00. Prefemoral process long (greater than or equal to the solenomere) and narrow (
<figureCitation id="15B62A5CFFC6E52E46CEB13A8C69FC72" box="[159,287,875,901]" captionStart="FIGURE 4" captionStartId="13.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-20@13.[161,1426,181,1680]" captionTargetPageId="13" captionText="FIGURE 4. Arthrosolaenomeris chapadensis Schubart, 1943 (holotype, MZSP 1092), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm. Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere; SPr, secondary process of the prefemoral process." figureDoi="http://doi.org/10.5281/zenodo.4761555" httpUri="https://zenodo.org/record/4761555/files/figure.png" pageId="12" pageNumber="213">Figs 4DF</figureCitation>
). Presence of a large secondary process positioned in its medial portion (
<figureCitation id="15B62A5CFFC6E52E423BB13A89B1FC72" box="[1130,1223,875,901]" captionStart="FIGURE 4" captionStartId="13.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-20@13.[161,1426,181,1680]" captionTargetPageId="13" captionText="FIGURE 4. Arthrosolaenomeris chapadensis Schubart, 1943 (holotype, MZSP 1092), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm. Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere; SPr, secondary process of the prefemoral process." figureDoi="http://doi.org/10.5281/zenodo.4761555" httpUri="https://zenodo.org/record/4761555/files/figure.png" pageId="12" pageNumber="213">Fig. 4D</figureCitation>
). Posterior portion to the secondary process with several indentations. Solenomere long and narrow (
<figureCitation id="15B62A5CFFC6E52E4211B1DE89ECFC5E" box="[1088,1178,911,937]" captionStart="FIGURE 4" captionStartId="13.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-20@13.[161,1426,181,1680]" captionTargetPageId="13" captionText="FIGURE 4. Arthrosolaenomeris chapadensis Schubart, 1943 (holotype, MZSP 1092), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm. Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere; SPr, secondary process of the prefemoral process." figureDoi="http://doi.org/10.5281/zenodo.4761555" httpUri="https://zenodo.org/record/4761555/files/figure.png" pageId="12" pageNumber="213">Fig. 4E</figureCitation>
). Cingulum positioned centroposteriorly (
<figureCitation id="15B62A5CFFC6E52E4737B1E28CCAFC3A" box="[358,444,947,973]" captionStart="FIGURE 4" captionStartId="13.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-20@13.[161,1426,181,1680]" captionTargetPageId="13" captionText="FIGURE 4. Arthrosolaenomeris chapadensis Schubart, 1943 (holotype, MZSP 1092), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm. Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere; SPr, secondary process of the prefemoral process." figureDoi="http://doi.org/10.5281/zenodo.4761555" httpUri="https://zenodo.org/record/4761555/files/figure.png" pageId="12" pageNumber="213">Fig. 4F</figureCitation>
). Apex of the solenomere abruptly descending (
<figureCitation id="15B62A5CFFC6E52E4583B1E2895BFC3A" box="[978,1069,947,973]" captionStart="FIGURE 4" captionStartId="13.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-20@13.[161,1426,181,1680]" captionTargetPageId="13" captionText="FIGURE 4. Arthrosolaenomeris chapadensis Schubart, 1943 (holotype, MZSP 1092), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 1 mm. Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere; SPr, secondary process of the prefemoral process." figureDoi="http://doi.org/10.5281/zenodo.4761555" httpUri="https://zenodo.org/record/4761555/files/figure.png" pageId="12" pageNumber="213">Fig. 4D</figureCitation>
).
</paragraph>
<paragraph id="8D3236D9FFC6E52E4696B186895CFBA9" blockId="12.[151,1437,479,1154]" pageId="12" pageNumber="213">
Female (IBSP 1049). Body as in male. Total length: 57.60. Total width: 9.35. Antennomere lengths (1&gt;7): 0.87; 1.72; 1.54; 1.36; 1.37; 1.38; 0.33. Podomeres lengths (1&gt;7): 0.82; 1.26; 2.62; 1.04; 1.12; 1.63; 0.55. Telson, length 1.38. Epigyne rectangular shaped. Vulvae (
<figureCitation id="15B62A5CFFC6E52E4425B64E8E74FBCE" box="[628,770,1055,1081]" captionStart="FIGURE 10" captionStartId="21.[151,250,1798,1822]" captionTargetBox="[314,1273,181,1773]" captionTargetId="figure-20@21.[314,1273,181,1773]" captionTargetPageId="21" captionText="FIGURE 10. Left vulva. Arthrosolaenomeris chapadensis Schubart, 1943 (IBSP 1049): (A), ventral view; (B), lateral view. Arthrosolaenomeris pantanalensis Schubart, 1943 (CZUFMT 847): (C), ventral view; (D), lateral view. Arthrosolaenomeris saci n. sp. (paratype, IBSP 2577): (E), ventral view; (F), lateral view.Arthrosolaenomeris iara n. sp. (paratype, IBSP 7537). (G), ventral view; (H), lateral view. Scale bars: 0,5 mm. Abbreviations: EV, external valve; IV, internal valve; Op, operculum." figureDoi="http://doi.org/10.5281/zenodo.4761577" httpUri="https://zenodo.org/record/4761577/files/figure.png" pageId="12" pageNumber="213">Figs 10AB</figureCitation>
): length 1.34, width 0.68. External valve: length 0.98, width 0.34. Internal valve: length 0.94, width 0.28. Operculum: length 0.39, width.0.56.
</paragraph>
</subSubSection>
<subSubSection id="C5976552FFC6E52E4696B636896FFB76" box="[199,1049,1127,1154]" pageId="12" pageNumber="213" type="distribution">
<paragraph id="8D3236D9FFC6E52E4696B636896FFB76" blockId="12.[151,1437,479,1154]" box="[199,1049,1127,1154]" pageId="12" pageNumber="213">
<emphasis id="BFF9EACBFFC6E52E4696B6368C17FB76" bold="true" box="[199,353,1127,1153]" pageId="12" pageNumber="213">Distribution.</emphasis>
States of
<collectingRegion id="4F49F83BFFC6E52E4781B6398FC6FB76" box="[464,688,1127,1154]" country="Brazil" name="Mato Grosso do Sul" pageId="12" pageNumber="213">Mato Grosso do Sul</collectingRegion>
and
<collectingRegion id="4F49F83BFFC6E52E44B6B6368E2DFB76" box="[743,859,1127,1153]" country="Brazil" name="Sao Paulo" pageId="12" pageNumber="213">São Paulo</collectingRegion>
,
<collectingCountry id="F59A7649FFC6E52E4537B6368EDCFB76" box="[870,938,1127,1153]" name="Brazil" pageId="12" pageNumber="213">Brazil</collectingCountry>
(
<figureCitation id="15B62A5CFFC6E52E45E8B636897DFB76" box="[953,1035,1127,1153]" captionStart="FIGURE 11" captionStartId="22.[151,250,1361,1385]" captionTargetBox="[151,1436,181,1337]" captionTargetId="figure-24@22.[151,1436,181,1337]" captionTargetPageId="22" captionText="FIGURE 11. Distribution map of Arthrosolaenomeris Schubart, 1943 species." figureDoi="http://doi.org/10.5281/zenodo.4761582" httpUri="https://zenodo.org/record/4761582/files/figure.png" pageId="12" pageNumber="213">Fig. 11</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>