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<document id="6FF5DE9E0A788AE844F71C20F0ED1A44" ID-DOI="10.5852/ejt.2024.921.2429" ID-ISSN="2118-9773" ID-Zenodo-Dep="10663874" ID-ZooBank="02633F29-4CDF-4027-BEBF-07AD2F925B42" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1707990651991" checkinUser="plazi" docAuthor="Bellvert, Adrià, Dimitrov, Dragomir, Zamani, Alireza &amp; Arnedo, Miquel A." docDate="2024" docId="03DA7943B730D85B72568F934016FC48" docLanguage="en" docName="EJT.2024.921.210-235.pdf" docOrigin="European Journal of Taxonomy 921 (210)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2429/10759" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dysdera jaegeri Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo 2024, sp. nov." docType="treatment" docUuid="579D5ABE-98BC-4B60-96DC-721AA2C027FE" docUuidSource="ZooBank" docVersion="2" lastPageNumber="220" masterDocId="FFE3013BB738D85170598D5B445AFFC5" masterDocTitle="Integrating museum collections and molecules reveals genus-level synonymy and new species in red devil spiders (Araneae, Dysderidae) from the Middle East and Central Asia" masterLastPageNumber="235" masterPageNumber="210" pageNumber="218" updateTime="1707991084405" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="CFABF15AEFA9D38AEE05F15B00FF8707" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5EAD6AF6D01004E5D90E37DF39DE6889">
<mods:title id="2270C2A60F43D6CE6B36A06C61D0C801">Integrating museum collections and molecules reveals genus-level synonymy and new species in red devil spiders (Araneae, Dysderidae) from the Middle East and Central Asia</mods:title>
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<mods:namePart id="5908D7E963C2FC40214DB8092F2AF6D7">Bellvert, Adrià</mods:namePart>
<mods:nameIdentifier id="C86078C63891F070EF254AC2FD2C27BC" type="LSID">0ABC3DFE-E982-47F5-A5DA-732688BCDEA9</mods:nameIdentifier>
<mods:affiliation id="25470E42681FB69750C6F88CC444EFDF">Departament de Biologia Evolutiva, Ecologia i Ciències Ambientals, Universitat de Barcelona (UB), Av. Diagonal, 643, 08028 Barcelona, Spain. &amp; Institut de Recerca de la Biodiversitat (IRBio), Universitat de Barcelona (UB) Barcelona, Spain.</mods:affiliation>
<mods:nameIdentifier id="23F2EEA280C4DF8F76DFA7CB4DF68109" type="email">abellvertba@gmail.com</mods:nameIdentifier>
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<mods:namePart id="621A826108AD03525708F24F28080352">Dimitrov, Dragomir</mods:namePart>
<mods:nameIdentifier id="D798A8AB13CED98A4EBB8C2449D97A65" type="LSID">37EC22F1-6348-4A61-B55F-3E74498AE5E9</mods:nameIdentifier>
<mods:affiliation id="12F7CB7612514CB455F4E8F3C66CD7D3">Departament de Biologia Evolutiva, Ecologia i Ciències Ambientals, Universitat de Barcelona (UB), Av. Diagonal, 643, 08028 Barcelona, Spain. &amp; Institut de Recerca de la Biodiversitat (IRBio), Universitat de Barcelona (UB) Barcelona, Spain. &amp; National Museum of Natural Sciences, Bulgarian Academy of Sciences, 1 Tsar Osvoboditel Blvd, 1000 Sofia, Bulgaria.</mods:affiliation>
<mods:nameIdentifier id="68413AD80EE026BCD7A0AA3C806CCD23" type="email">dimitrov.drk@gmail.com</mods:nameIdentifier>
</mods:name>
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<mods:namePart id="4CB194E7D01BCA6F9108B77203C18BB7">Zamani, Alireza</mods:namePart>
<mods:nameIdentifier id="4090E46690DFEEDC5FAC7AA3A3F02860" type="LSID">A21C0B82-E2D9-4402-9408-D6160843DAF4</mods:nameIdentifier>
<mods:affiliation id="7E1799E65DDFAEBBA4A8225D12EA9601">Zoological Museum, Biodiversity Unit, FI- 20014 University of Turku, Finland.</mods:affiliation>
<mods:nameIdentifier id="6F7B61AB79F4F5AAB116167EE7041334" type="email">zamani.alireza5@gmail.com</mods:nameIdentifier>
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<mods:namePart id="29A4BBFE2250AA87E7085A7470F6FC5A">Arnedo, Miquel A.</mods:namePart>
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<mods:affiliation id="4755735528681D8A5C65FC4E9C87377D">Departament de Biologia Evolutiva, Ecologia i Ciències Ambientals, Universitat de Barcelona (UB), Av. Diagonal, 643, 08028 Barcelona, Spain. &amp; Institut de Recerca de la Biodiversitat (IRBio), Universitat de Barcelona (UB) Barcelona, Spain.</mods:affiliation>
<mods:nameIdentifier id="89B88B52A904E4112299E0976F59EAF6" type="email">marnedo@gmail.com</mods:nameIdentifier>
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<mods:title id="37A83F44709B9E03C9D760CDC2D67EE3">European Journal of Taxonomy</mods:title>
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<mods:date id="941C87BCE3EEF4E1FD2DC416645BF0C8">2024</mods:date>
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<mods:number id="9CDDCBB2BF97DDE8680BB93569CEB341">2024-02-14</mods:number>
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<mods:number id="A47D12C5537D6BFD596BEAA9593FF71B">921</mods:number>
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<mods:number id="11A1029BDED9069152B8FD217A360393">210</mods:number>
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<mods:start id="5F4688D9AD3DDDFC36561DB42222FCFD">210</mods:start>
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<mods:identifier id="62F00D2F17CE2BA659918F00B2290418" type="DOI">10.5852/ejt.2024.921.2429</mods:identifier>
<mods:identifier id="7057D6BBBE04C0BB1CE67E1612CE507C" type="ISSN">2118-9773</mods:identifier>
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<treatment id="03DA7943B730D85B72568F934016FC48" LSID="urn:lsid:zoobank.org:act:579D5ABE-98BC-4B60-96DC-721AA2C027FE" httpUri="http://treatment.plazi.org/id/03DA7943B730D85B72568F934016FC48" lastPageId="10" lastPageNumber="220" pageId="8" pageNumber="218">
<subSubSection id="C3699BDEB730D85972568F934094FCCD" pageId="8" pageNumber="218" type="nomenclature">
<paragraph id="8BCCC855B730D85972568F93407FFD26" blockId="8.[358,1230,712,811]" box="[527,1061,712,739]" pageId="8" pageNumber="218">
<heading id="D0847F39B730D85972568F93407FFD26" box="[527,1061,712,739]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="218" reason="2">
<taxonomicName id="4C73B3D6B730D85972568F93479EFD26" authority="Bellvert &amp; Dimitrov" authorityName="Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo" authorityYear="2024" box="[527,964,712,739]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="218" phylum="Arthropoda" rank="species" species="jaegeri" status="sp. nov.">
<emphasis id="B9071447B730D85972568F934696FD26" bold="true" box="[527,716,712,739]" italics="true" pageId="8" pageNumber="218">Dysdera jaegeri</emphasis>
Bellvert &amp; Dimitrov
</taxonomicName>
<taxonomicNameLabel id="A234A93CB730D85973928F92407FFD26" box="[971,1061,713,739]" pageId="8" pageNumber="218" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="C3699BDEB730D85972BE8E4A4717FCEE" box="[743,845,785,811]" pageId="8" pageNumber="218" type="description">
<paragraph id="8BCCC855B730D85972BE8E4A4717FCEE" blockId="8.[358,1230,712,811]" box="[743,845,785,811]" pageId="8" pageNumber="218">
<figureCitation id="1348D4D0B730D85972BE8E4A4717FCEE" box="[743,845,785,811]" captionStart="Figs 6–8" captionStartId="9.[191,244,1963,1989]" captionTargetBox="[189,1398,1041,1924]" captionTargetId="figure-312@9.[189,1398,1041,1924]" captionTargetPageId="9" captionText="Figs 6–8. Male Dysdera jaegeri Bellvert &amp; Dimitrov sp. nov. (SMF), left palp. 6. Prolateral view. 7. Anterior view. 8. Retrolateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663886" httpUri="https://zenodo.org/record/10663886/files/figure.png" pageId="8" pageNumber="218">Figs 6–8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3699BDEB730D85970E48E0F4057FB05" pageId="8" pageNumber="218" type="diagnosis">
<paragraph id="8BCCC855B730D85970E48E0F456CFCAB" blockId="8.[189,310,852,878]" box="[189,310,852,878]" pageId="8" pageNumber="218">
<heading id="D0847F39B730D85970E48E0F456CFCAB" bold="true" box="[189,310,852,878]" fontSize="11" level="3" pageId="8" pageNumber="218" reason="3">
<emphasis id="B9071447B730D85970E48E0F456CFCAB" bold="true" box="[189,310,852,878]" pageId="8" pageNumber="218">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BCCC855B730D85970E48EDF4057FB05" blockId="8.[188,1399,900,1216]" pageId="8" pageNumber="218">
Male resembles that of
<taxonomicName id="4C73B3D6B730D859718B8EDF4032FC5B" authority="Deeleman-Reinhold &amp; Deeleman, 1988" authorityName="Deeleman-Reinhold &amp; Deeleman" authorityYear="1988" box="[466,1128,900,926]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="218" phylum="Arthropoda" rank="species" species="sultani">
<emphasis id="B9071447B730D859718B8EDF46D4FC5B" box="[466,654,900,926]" italics="true" pageId="8" pageNumber="218">Dysdera sultani</emphasis>
<bibRefCitation id="EFE2B5A4B730D85972CC8EDF4032FC5B" author="Deeleman-Reinhold C. L. &amp; Deeleman P. R." box="[661,1128,900,926]" pageId="8" pageNumber="218" pagination="141 - 269" refId="ref11600" refString="Deeleman-Reinhold C. L. &amp; Deeleman P. R. 1988. Revision des Dysderinae (Araneae, Dysderidae), les especes mediterraneennes occidentales exceptees. Tijdschrift voor Entomologie 131: 141 - 269." type="journal article" year="1988">Deeleman-Reinhold &amp; Deeleman, 1988</bibRefCitation>
</taxonomicName>
by having the external and internal sclerites fused in a single anterior sclerite, surrounding the lateral sheet (
<figureCitation id="1348D4D0B730D85974D68EF3408CFC07" box="[1167,1238,936,962]" captionStart="Figs 6–8" captionStartId="9.[191,244,1963,1989]" captionTargetBox="[189,1398,1041,1924]" captionTargetId="figure-312@9.[189,1398,1041,1924]" captionTargetPageId="9" captionText="Figs 6–8. Male Dysdera jaegeri Bellvert &amp; Dimitrov sp. nov. (SMF), left palp. 6. Prolateral view. 7. Anterior view. 8. Retrolateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663886" httpUri="https://zenodo.org/record/10663886/files/figure.png" pageId="8" pageNumber="218">Fig. 7</figureCitation>
), and the bent embolic division (
<figureCitation id="1348D4D0B730D85971D78E9745A2FC23" box="[398,504,972,998]" captionStart="Figs 6–8" captionStartId="9.[191,244,1963,1989]" captionTargetBox="[189,1398,1041,1924]" captionTargetId="figure-312@9.[189,1398,1041,1924]" captionTargetPageId="9" captionText="Figs 6–8. Male Dysdera jaegeri Bellvert &amp; Dimitrov sp. nov. (SMF), left palp. 6. Prolateral view. 7. Anterior view. 8. Retrolateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663886" httpUri="https://zenodo.org/record/10663886/files/figure.png" pageId="8" pageNumber="218">Figs 7–8</figureCitation>
). The two species can be distinguished by (1) the short and stout lateral sheet with transparent lamellar extension in
<taxonomicName id="4C73B3D6B730D85972DC8EAA46A7FBCF" authority="Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo, 2024" authorityName="Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo" authorityYear="2024" box="[645,765,1009,1034]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="218" phylum="Arthropoda" rank="species" species="jaegeri" status="sp. nov.">
<emphasis id="B9071447B730D85972DC8EAA46A7FBCF" box="[645,765,1009,1034]" italics="true" pageId="8" pageNumber="218">D. jaegeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A234A93CB730D859735D8EAA473AFBCE" box="[772,864,1009,1035]" pageId="8" pageNumber="218" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="1348D4D0B730D85973298EAB47E2FBCE" box="[880,952,1008,1035]" captionStart="Figs 6–8" captionStartId="9.[191,244,1963,1989]" captionTargetBox="[189,1398,1041,1924]" captionTargetId="figure-312@9.[189,1398,1041,1924]" captionTargetPageId="9" captionText="Figs 6–8. Male Dysdera jaegeri Bellvert &amp; Dimitrov sp. nov. (SMF), left palp. 6. Prolateral view. 7. Anterior view. 8. Retrolateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663886" httpUri="https://zenodo.org/record/10663886/files/figure.png" pageId="8" pageNumber="218">Fig. 8</figureCitation>
), vs thin and long single processus in
<taxonomicName id="4C73B3D6B730D85970E7894D456BFBEA" authorityName="Deeleman-Reinhold &amp; Deeleman" authorityYear="1988" box="[190,305,1045,1071]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="218" phylum="Arthropoda" rank="species" species="sultani">
<emphasis id="B9071447B730D85970E7894D456BFBEA" box="[190,305,1045,1071]" italics="true" pageId="8" pageNumber="218">D. sultani</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFE2B5A4B730D8597167894E4751FBEA" author="Deeleman-Reinhold C. L. &amp; Deeleman P. R." box="[318,779,1045,1071]" pageId="8" pageNumber="218" pagination="141 - 269" refId="ref11600" refString="Deeleman-Reinhold C. L. &amp; Deeleman P. R. 1988. Revision des Dysderinae (Araneae, Dysderidae), les especes mediterraneennes occidentales exceptees. Tijdschrift voor Entomologie 131: 141 - 269." type="journal article" year="1988">Deeleman-Reinhold &amp; Deeleman 1988</bibRefCitation>
: figs 236–237), (2) less bent embolic division (i.e., ca 60°) in
<taxonomicName id="4C73B3D6B730D859714F896145D7FB96" authorityName="Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo" authorityYear="2024" box="[278,397,1082,1107]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="218" phylum="Arthropoda" rank="species" species="jaegeri">
<emphasis id="B9071447B730D859714F896145D7FB96" box="[278,397,1082,1107]" italics="true" pageId="8" pageNumber="218">D. jaegeri</emphasis>
</taxonomicName>
, vs ca 80° in
<taxonomicName id="4C73B3D6B730D8597277896146F8FB96" authorityName="Deeleman-Reinhold &amp; Deeleman" authorityYear="1988" box="[558,674,1081,1107]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="218" phylum="Arthropoda" rank="species" species="sultani">
<emphasis id="B9071447B730D8597277896146F8FB96" box="[558,674,1081,1107]" italics="true" pageId="8" pageNumber="218">D. sultani</emphasis>
</taxonomicName>
, (3) the apical part of the embolic division is a whole plate in
<taxonomicName id="4C73B3D6B730D85970E48905456BFBB2" authorityName="Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo" authorityYear="2024" box="[189,305,1118,1143]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="218" phylum="Arthropoda" rank="species" species="jaegeri">
<emphasis id="B9071447B730D85970E48905456BFBB2" box="[189,305,1118,1143]" italics="true" pageId="8" pageNumber="218">D. jaegeri</emphasis>
</taxonomicName>
(
<figureCitation id="1348D4D0B730D8597166890645FEFBBD" box="[319,420,1117,1144]" captionStart="Figs 6–8" captionStartId="9.[191,244,1963,1989]" captionTargetBox="[189,1398,1041,1924]" captionTargetId="figure-312@9.[189,1398,1041,1924]" captionTargetPageId="9" captionText="Figs 6–8. Male Dysdera jaegeri Bellvert &amp; Dimitrov sp. nov. (SMF), left palp. 6. Prolateral view. 7. Anterior view. 8. Retrolateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663886" httpUri="https://zenodo.org/record/10663886/files/figure.png" pageId="8" pageNumber="218">Figs 6, 8</figureCitation>
), while in
<taxonomicName id="4C73B3D6B730D8597243890546D7FBB2" authorityName="Deeleman-Reinhold &amp; Deeleman" authorityYear="1988" box="[538,653,1117,1143]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="218" phylum="Arthropoda" rank="species" species="sultani">
<emphasis id="B9071447B730D8597243890546D7FBB2" box="[538,653,1117,1143]" italics="true" pageId="8" pageNumber="218">D. sultani</emphasis>
</taxonomicName>
it is sulcate apically, and (4) the posterior apophysis in
<taxonomicName id="4C73B3D6B730D85975588905412CFBB2" authorityName="Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo" authorityYear="2024" box="[1281,1398,1118,1143]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="218" phylum="Arthropoda" rank="species" species="jaegeri">
<emphasis id="B9071447B730D85975588905412CFBB2" box="[1281,1398,1118,1143]" italics="true" pageId="8" pageNumber="218">D. jaegeri</emphasis>
</taxonomicName>
is more or less straight, perpendicular to the tegulum (
<figureCitation id="1348D4D0B730D859731489D947E0FB59" box="[845,954,1154,1180]" captionStart="Figs 6–8" captionStartId="9.[191,244,1963,1989]" captionTargetBox="[189,1398,1041,1924]" captionTargetId="figure-312@9.[189,1398,1041,1924]" captionTargetPageId="9" captionText="Figs 6–8. Male Dysdera jaegeri Bellvert &amp; Dimitrov sp. nov. (SMF), left palp. 6. Prolateral view. 7. Anterior view. 8. Retrolateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663886" httpUri="https://zenodo.org/record/10663886/files/figure.png" pageId="8" pageNumber="218">Figs 6, 8</figureCitation>
), vs long and more bent towards the tegulum in
<taxonomicName id="4C73B3D6B730D859711A89FC45ECFB05" authorityName="Deeleman-Reinhold &amp; Deeleman" authorityYear="1988" box="[323,438,1190,1216]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="218" phylum="Arthropoda" rank="species" species="sultani">
<emphasis id="B9071447B730D859711A89FC45ECFB05" box="[323,438,1190,1216]" italics="true" pageId="8" pageNumber="218">D. sultani</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFE2B5A4B730D859719C89FD47CCFB05" author="Deeleman-Reinhold C. L. &amp; Deeleman P. R." box="[453,918,1190,1216]" pageId="8" pageNumber="218" pagination="141 - 269" refId="ref11600" refString="Deeleman-Reinhold C. L. &amp; Deeleman P. R. 1988. Revision des Dysderinae (Araneae, Dysderidae), les especes mediterraneennes occidentales exceptees. Tijdschrift voor Entomologie 131: 141 - 269." type="journal article" year="1988">Deeleman-Reinhold &amp; Deeleman 1988</bibRefCitation>
: fig. 236).
</paragraph>
</subSubSection>
<subSubSection id="C3699BDEB730D85970E489B24096FAF6" box="[189,1228,1257,1331]" pageId="8" pageNumber="218" type="etymology">
<paragraph id="8BCCC855B730D85970E489B2451EFAC6" blockId="8.[189,324,1257,1283]" box="[189,324,1257,1283]" pageId="8" pageNumber="218">
<heading id="D0847F39B730D85970E489B2451EFAC6" bold="true" box="[189,324,1257,1283]" fontSize="11" level="3" pageId="8" pageNumber="218" reason="3">
<emphasis id="B9071447B730D85970E489B2451EFAC6" bold="true" box="[189,324,1257,1283]" pageId="8" pageNumber="218">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BCCC855B730D85970E488434096FAF6" blockId="8.[189,1228,1304,1331]" box="[189,1228,1304,1331]" pageId="8" pageNumber="218">This species is named after our colleague and reputed German arachnologist Peter Jäger.</paragraph>
</subSubSection>
<subSubSection id="C3699BDEB730D85970E48800466BFA28" pageId="8" pageNumber="218" type="materials_examined">
<paragraph id="8BCCC855B730D85970E488004536FAB1" blockId="8.[189,364,1370,1397]" box="[189,364,1370,1397]" pageId="8" pageNumber="218">
<heading id="D0847F39B730D85970E488004536FAB1" bold="true" box="[189,364,1370,1397]" fontSize="11" level="3" pageId="8" pageNumber="218" reason="3">
<emphasis id="B9071447B730D85970E488004536FAB1" bold="true" box="[189,364,1370,1397]" pageId="8" pageNumber="218">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BCCC855B730D85970B388D14506FA61" blockId="8.[189,1399,1418,1517]" box="[234,348,1418,1444]" pageId="8" pageNumber="218">
<emphasis id="B9071447B730D85970B388D14506FA61" bold="true" box="[234,348,1418,1444]" pageId="8" pageNumber="218">
<typeStatus id="54C876F7B730D85970B388D14506FA61" box="[234,348,1418,1444]" pageId="8" pageNumber="218" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BCCC855B730D85970E488F4466BFA28" blockId="8.[189,1399,1418,1517]" pageId="8" pageNumber="218">
<materialsCitation id="3B1BC208B730D85970E488F4466BFA28" collectingDate="1973-03-18" collectionCode="R, SMF" collectorName="Kinzelbach" country="Syria" location="Homs Province" pageId="8" pageNumber="218" specimenCount="1" specimenCount-male="1" stateProvince="Hims" typeStatus="holotype">
<collectingCountry id="F36488C5B730D85970E488F44549FA0C" box="[189,275,1455,1481]" name="Syria" pageId="8" pageNumber="218">SYRIA</collectingCountry>
•
<specimenCount id="9D7503DCB730D859717588F44501FA0C" box="[300,347,1455,1481]" count="1" pageId="8" pageNumber="218" type="male">1 ♂</specimenCount>
;
<collectingRegion id="49B706B7B730D859713E88F44645FA0C" box="[359,543,1455,1481]" country="Syria" name="Hims" pageId="8" pageNumber="218">Homs Province</collectingRegion>
, west of
<collectingRegion id="49B706B7B730D85972DE88F4468AFA0C" box="[647,720,1455,1481]" country="Syria" name="Hims" pageId="8" pageNumber="218">Homs</collectingRegion>
, ruins and abandoned cemetery NW of the castle;
<date id="FFCDEE95B730D859754288F444A1FA28" pageId="8" pageNumber="218" value="1973-03-18">
<collectingDate id="EF89177DB730D859754288F444A1FA28" pageId="8" pageNumber="218" value="1973-03-18">18 Mar. 1973</collectingDate>
</date>
;
<collectionCode id="ED625090B730D859715E88884544FA28" box="[263,286,1491,1517]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="8" pageNumber="218">R</collectionCode>
.
<collectorName id="2686AD83B730D8597170888845F5FA28" box="[297,431,1491,1517]" pageId="8" pageNumber="218">Kinzelbach</collectorName>
leg.;
<collectionCode id="ED625090B730D85971A888884677FA28" box="[497,557,1491,1517]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="8" pageNumber="218">SMF</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3699BDEB730D85B70E48B4E4563FDE8" lastPageId="10" lastPageNumber="220" pageId="8" pageNumber="218" type="description">
<paragraph id="8BCCC855B730D85970E48B4E4514F9EA" blockId="8.[189,334,1557,1583]" box="[189,334,1557,1583]" pageId="8" pageNumber="218">
<heading id="D0847F39B730D85970E48B4E4514F9EA" bold="true" box="[189,334,1557,1583]" fontSize="11" level="3" pageId="8" pageNumber="218" reason="3">
<emphasis id="B9071447B730D85970E48B4E4514F9EA" bold="true" box="[189,334,1557,1583]" pageId="8" pageNumber="218">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BCCC855B730D85970B38B1E45F1F99A" blockId="8.[187,1401,1605,1922]" box="[234,427,1605,1631]" pageId="8" pageNumber="218">
<emphasis id="B9071447B730D85970B38B1E4570F99A" bold="true" box="[234,298,1605,1631]" pageId="8" pageNumber="218">Male</emphasis>
(
<figureCitation id="1348D4D0B730D85971608B1E45F8F99A" box="[313,418,1605,1631]" captionStart="Figs 6–8" captionStartId="9.[191,244,1963,1989]" captionTargetBox="[189,1398,1041,1924]" captionTargetId="figure-312@9.[189,1398,1041,1924]" captionTargetPageId="9" captionText="Figs 6–8. Male Dysdera jaegeri Bellvert &amp; Dimitrov sp. nov. (SMF), left palp. 6. Prolateral view. 7. Anterior view. 8. Retrolateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663886" httpUri="https://zenodo.org/record/10663886/files/figure.png" pageId="8" pageNumber="218">Figs 6–8</figureCitation>
)
</paragraph>
<paragraph id="8BCCC855B730D85970E48B31405CF847" blockId="8.[187,1401,1605,1922]" pageId="8" pageNumber="218">PROSOMA. 3.13 long; maximum width 2.32; minimum width 1.61. Reddish, uniformly; with small dark grains mainly in frontal part; frontally covered with small white setae. Frontal border roughly round, ca ½ of carapace length; anterior lateral borders parallel; rounded at maximum dorsal width, back lateral borders rounded; back margin narrow, bilobulated. Eye diameters: AME 0.17; PLE 0.15; PME 0.14; AMEs on edge of frontal border, separated from one another by ca ½ diameter, close to PLEs; PMEs very close to one another, ca ⅓ of PME diameter from PLEs. Labium trapezoid-shaped, base wider than distal part; as long as wide at base; semicircular groove at tip. Sternum orange-red, frontally darker, becoming lighter posteriorly; smooth; uniformly covered with slender black setae.</paragraph>
<paragraph id="8BCCC855B730D85970E48AF1412CF82D" blockId="8.[189,1399,1961,2025]" pageId="8" pageNumber="218">CHELICERAE. 1.13 long, ca ⅓ of carapace length in dorsal view; fang medium-sized, 0.88 long; paturon dorsal and ventral side completely covered with piligerous granulations. Cheliceral inner groove short,</paragraph>
<paragraph id="8BCCC855B731D85870E48C524793FE82" blockId="9.[189,1398,263,327]" pageId="9" pageNumber="219">ca ⅓ of cheliceral length; armed with three teeth and lamina at base; B&gt; M&gt;D; D triangular, located roughly at centre of groove; B close to basal lamina; M close to B.</paragraph>
<paragraph id="8BCCC855B731D85870E48C354119FD2D" blockId="9.[187,1400,365,744]" pageId="9" pageNumber="219">LEGS. Anterior legs orange, posterior legs yellow. Lengths of leg segments: fe1 2.36; pa1 1.49; ti1 1.87; me1 2.06; ta1 0.58; total 8.36; fe2 2.14; pa2 1.38; ti2 1.68; me2 1.95; ta2 0.56; total 7.72; fe3 1.65; pa3 0.94; ti3 1.00; me3 1.45; ta3 0.51; total 5.59; fe4 2.04; pa4 1.48; ti4 1.46; me4 1.97; ta4 0.62; total 7.58; fe Pdp 1.50; pa Pdp 0.88; ti Pdp 0.57; ta Pdp 0.91; total 3.86; leg formula: 1&gt;2&gt; 4&gt;3. Leg2 spineless. Fe3d spineless; pa3 spineless; tb3d spines arranged in two bands; proximal 1.0.0-1; medial-proximal 0; medial-distal 0; distal 1.0.1; tb3v spines arranged in two bands; proximal 1-2.0.0; medial-proximal 0; medial-distal 0; distal 1-0.0.0; with one terminal spine on forward margin. Fe4d spines in two rows; forward 1; backward 2; pa4 spineless; tb4d spines arranged in two bands; proximal 1.0.1; medial-proximal 0; medial-distal 0; distal 0.0.1; tb4v spines arranged in two bands; proximal 1.1.0; medial-proximal 0; medial-distal 0; distal 0-1.0.0; with one terminal spine at the distal border. Dorsal side of frontal legs smooth; ventral side of pedipalp smooth. Claws with 8 teeth or less; hardly larger than claw width.</paragraph>
<paragraph id="8BCCC855B731D85870E48E544743FC89" blockId="9.[189,1398,783,844]" pageId="9" pageNumber="219">OPISTHOSOMA. 3.87 long; cream-colored; cylindrical. Abdominal dorsal setae 0.04 long; thin, almost straight, pointed; uniformly and thickly distributed.</paragraph>
<paragraph id="8BCCC855B731D85B70E48E2F410FFE0E" blockId="9.[189,1399,884,981]" lastBlockId="10.[187,1398,265,460]" lastPageId="10" lastPageNumber="220" pageId="9" pageNumber="219">
PALP (
<figureCitation id="1348D4D0B731D858715E8E2F452BFC4B" box="[263,369,884,910]" captionStart="Figs 6–8" captionStartId="9.[191,244,1963,1989]" captionTargetBox="[189,1398,1041,1924]" captionTargetId="figure-312@9.[189,1398,1041,1924]" captionTargetPageId="9" captionText="Figs 6–8. Male Dysdera jaegeri Bellvert &amp; Dimitrov sp. nov. (SMF), left palp. 6. Prolateral view. 7. Anterior view. 8. Retrolateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663886" httpUri="https://zenodo.org/record/10663886/files/figure.png" pageId="9" pageNumber="219">Figs 6–8</figureCitation>
). T slightly longer than ED; external distal border straight, internal projected at middle. ED bent ca 60 degrees in lateral view, internal distal border markedly expanded. IS and ES completely fused, forming a single anterior sclerite (
<figureCitation id="1348D4D0B731D85872F38EE146AFFC11" box="[682,757,954,980]" captionStart="Figs 6–8" captionStartId="9.[191,244,1963,1989]" captionTargetBox="[189,1398,1041,1924]" captionTargetId="figure-312@9.[189,1398,1041,1924]" captionTargetPageId="9" captionText="Figs 6–8. Male Dysdera jaegeri Bellvert &amp; Dimitrov sp. nov. (SMF), left palp. 6. Prolateral view. 7. Anterior view. 8. Retrolateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663886" httpUri="https://zenodo.org/record/10663886/files/figure.png" pageId="9" pageNumber="219">Fig. 7</figureCitation>
). ED tip straight in lateral view. C present, short and poorly developed, located in distal end on ED internal tip, proximal border of C continuously decreasing. LF absent. L well developed, anteriorly not projected with a spine-like basal projection. External border poorly sclerotized, forming a transparent ridge between spine-like apophysis and distal part, not folded.AL present, well developed; proximal border in posterior view smooth, not fused with distal haematodocha. P short, fused to T; lateral length of P from ⅖ to ½ of T width; ridge present, perpendicular to T; distinctly expanded, rounded, upper margin smooth; distally ridge-like expanded; posterior margin not folded.
</paragraph>
<caption id="DF0C98DDB731D85870E68AF04731F82C" ID-DOI="http://doi.org/10.5281/zenodo.10663886" ID-Zenodo-Dep="10663886" httpUri="https://zenodo.org/record/10663886/files/figure.png" pageId="9" pageNumber="219" startId="9.[191,244,1963,1989]" targetBox="[189,1398,1041,1924]" targetPageId="9">
<paragraph id="8BCCC855B731D85870E68AF04731F82C" blockId="9.[191,1399,1963,2025]" pageId="9" pageNumber="219">
<emphasis id="B9071447B731D85870E68AF04569F800" bold="true" box="[191,307,1963,1989]" pageId="9" pageNumber="219">Figs 6–8.</emphasis>
Male
<taxonomicName id="4C73B3D6B731D85871DB8AF04711F800" authority="Bellvert &amp; Dimitrov" authorityName="Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo" authorityYear="2024" box="[386,843,1963,1989]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="219" phylum="Arthropoda" rank="species" species="jaegeri" status="sp. nov.">
<emphasis id="B9071447B731D85871DB8AF0461DF800" box="[386,583,1963,1989]" italics="true" pageId="9" pageNumber="219">Dysdera jaegeri</emphasis>
Bellvert &amp; Dimitrov
</taxonomicName>
<taxonomicNameLabel id="A234A93CB731D858730C8AF747EEF803" box="[853,948,1964,1990]" pageId="9" pageNumber="219" rank="species">sp. nov.</taxonomicNameLabel>
(SMF), left palp.
<emphasis id="B9071447B731D85874C08AF040FCF800" bold="true" box="[1177,1190,1963,1989]" pageId="9" pageNumber="219">6</emphasis>
. Prolateral view.
<emphasis id="B9071447B731D85870E68A954491F82D" bold="true" box="[191,203,1998,2024]" pageId="9" pageNumber="219">7</emphasis>
. Anterior view.
<emphasis id="B9071447B731D85871D18A9545CDF82D" bold="true" box="[392,407,1998,2024]" pageId="9" pageNumber="219">8</emphasis>
. Retrolateral view. Scale bar = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BCCC855B732D85B70B38CAA451FFDCE" blockId="10.[189,325,497,557]" box="[234,325,497,523]" pageId="10" pageNumber="220">
<emphasis id="B9071447B732D85B70B38CAA451FFDCE" bold="true" box="[234,325,497,523]" pageId="10" pageNumber="220">Female</emphasis>
</paragraph>
<paragraph id="8BCCC855B732D85B70E48F484563FDE8" blockId="10.[189,325,497,557]" box="[189,313,531,557]" pageId="10" pageNumber="220">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3699BDEB732D85B70E48F094780FD5F" box="[189,986,594,666]" pageId="10" pageNumber="220" type="distribution">
<paragraph id="8BCCC855B732D85B70E48F09450CFDA9" blockId="10.[189,342,594,620]" box="[189,342,594,620]" pageId="10" pageNumber="220">
<heading id="D0847F39B732D85B70E48F09450CFDA9" bold="true" box="[189,342,594,620]" fontSize="11" level="3" pageId="10" pageNumber="220" reason="3">
<emphasis id="B9071447B732D85B70E48F09450CFDA9" bold="true" box="[189,342,594,620]" pageId="10" pageNumber="220">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BCCC855B732D85B70E48FDB4780FD5F" blockId="10.[189,986,640,666]" box="[189,986,640,666]" pageId="10" pageNumber="220">
Known only from the
<typeStatus id="54C876F7B732D85B719B8FDB45AFFD5F" box="[450,501,640,666]" pageId="10" pageNumber="220">type</typeStatus>
locality in
<collectingRegion id="49B706B7B732D85B72208FDB476BFD5F" box="[633,817,640,666]" country="Syria" name="Hims" pageId="10" pageNumber="220">Homs Province</collectingRegion>
, central
<collectingCountry id="F36488C5B732D85B73CC8FDB478DFD5F" box="[917,983,640,666]" name="Syria" pageId="10" pageNumber="220">Syria</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3699BDEB732D85B70E48F9B4016FC48" pageId="10" pageNumber="220" type="discussion">
<paragraph id="8BCCC855B732D85B70E48F9B4575FD1F" blockId="10.[189,303,704,730]" box="[189,303,704,730]" pageId="10" pageNumber="220">
<heading id="D0847F39B732D85B70E48F9B4575FD1F" bold="true" box="[189,303,704,730]" fontSize="11" level="3" pageId="10" pageNumber="220" reason="3">
<emphasis id="B9071447B732D85B70E48F9B4575FD1F" bold="true" box="[189,303,704,730]" pageId="10" pageNumber="220">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BCCC855B732D85B70E48FB74016FC48" blockId="10.[187,1399,748,910]" pageId="10" pageNumber="220">
<taxonomicName id="4C73B3D6B732D85B70E48FB74520FCC3" authority="Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo, 2024" authorityName="Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo" authorityYear="2024" box="[189,378,748,774]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="220" phylum="Arthropoda" rank="species" species="jaegeri" status="sp. nov.">
<emphasis id="B9071447B732D85B70E48FB74520FCC3" box="[189,378,748,774]" italics="true" pageId="10" pageNumber="220">Dysdera jaegeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A234A93CB732D85B71D98FB64580FCC2" box="[384,474,749,775]" pageId="10" pageNumber="220" rank="species">sp. nov.</taxonomicNameLabel>
along with
<taxonomicName id="4C73B3D6B732D85B72388FB647DDFCC2" authority="Nosek, 1905" authorityName="Nosek" authorityYear="1905" box="[609,903,748,775]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="220" phylum="Arthropoda" rank="species" species="argaeica">
<emphasis id="B9071447B732D85B72388FB646B1FCC3" box="[609,747,749,774]" italics="true" pageId="10" pageNumber="220">D. argaeica</emphasis>
<bibRefCitation id="EFE2B5A4B732D85B72AB8FB747DDFCC2" author="Nosek A." box="[754,903,748,775]" pageId="10" pageNumber="220" pagination="114 - 154" refId="ref12754" refString="Nosek A. 1905. Araneiden, Opilionen und Chernetiden. In: Penther A. &amp; Zederbauer E. (eds) Ergebnisse einer naturwissenschaftlichen Reise zum Erdschias-Dagh (Kleinasien). Annalen des Kaiserlich- Koniglichen Naturhistorischen Hofmuseums in Wien: 20: 114 - 154." type="journal article" year="1905">Nosek, 1905</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C73B3D6B732D85B73C88FB6405EFCC3" authorityName="Deeleman-Reinhold &amp; Deeleman" authorityYear="1988" box="[913,1028,748,774]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="220" phylum="Arthropoda" rank="species" species="sultani">
<emphasis id="B9071447B732D85B73C88FB6405EFCC3" box="[913,1028,748,774]" italics="true" pageId="10" pageNumber="220">D. sultani</emphasis>
</taxonomicName>
,
<taxonomicName id="4C73B3D6B732D85B744A8FB6411CFCC2" authority="Dimitrov, 2018" authorityName="Dimitrov" authorityYear="2018" box="[1043,1350,748,775]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="220" phylum="Arthropoda" rank="species" species="galinae">
<emphasis id="B9071447B732D85B744A8FB640D4FCC3" box="[1043,1166,748,774]" italics="true" pageId="10" pageNumber="220">D. galinae</emphasis>
<bibRefCitation id="EFE2B5A4B732D85B74CD8FB7411CFCC2" author="Dimitrov D." box="[1172,1350,748,775]" pageId="10" pageNumber="220" pagination="457 - 462" refId="ref11637" refString="Dimitrov D. 2018. A Contribution to the study of Turkish Dysderidae with description of a new Dysdera (Araneae, Dysderidae). Arachnology 17 (9): 457 - 462. https: // doi. org / 10.13156 / arac. 2018.17.9.457" type="journal article" year="2018">Dimitrov, 2018</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C73B3D6B732D85B70E58E544639FCED" authority="Deeleman-Reinhold, 1988" authorityName="Deeleman-Reinhold" authorityYear="1988" box="[188,611,782,808]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="220" phylum="Arthropoda" rank="species" species="yozgat">
<emphasis id="B9071447B732D85B70E58E544571FCED" box="[188,299,783,808]" italics="true" pageId="10" pageNumber="220">D. yozgat</emphasis>
<bibRefCitation id="EFE2B5A4B732D85B71688E554639FCED" author="Deeleman-Reinhold C. L. &amp; Deeleman P. R." box="[305,611,782,808]" pageId="10" pageNumber="220" pagination="141 - 269" refId="ref11600" refString="Deeleman-Reinhold C. L. &amp; Deeleman P. R. 1988. Revision des Dysderinae (Araneae, Dysderidae), les especes mediterraneennes occidentales exceptees. Tijdschrift voor Entomologie 131: 141 - 269." type="journal article" year="1988">Deeleman-Reinhold, 1988</bibRefCitation>
</taxonomicName>
form a complex of species within the diverse
<emphasis id="B9071447B732D85B74368E544091FCED" box="[1135,1227,783,808]" italics="true" pageId="10" pageNumber="220">asiatica</emphasis>
group that can be characterized by the long cylindrical T, entirely fused IS and ES, and the sclerotized processus-like lateral sheet. This complex is distributed in
<collectingCountry id="F36488C5B732D85B72EB8E0A475FFCAE" box="[690,773,849,875]" name="Turkey" pageId="10" pageNumber="220">Turkey</collectingCountry>
and
<collectingCountry id="F36488C5B732D85B73628E0A4726FCAE" box="[827,892,849,875]" name="Syria" pageId="10" pageNumber="220">Syria</collectingCountry>
. Unfortunately, we were unable to sequence any of these species and their position in the
<taxonomicName id="4C73B3D6B732D85B72928E28470BFC48" box="[715,849,883,909]" pageId="10" pageNumber="211" rank="subFamily" subFamily="Dysderinae">Dysderinae</taxonomicName>
tree remains unclear.
</paragraph>
</subSubSection>
</treatment>
</document>
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