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<document id="63B61C42ED3270B055A62C8C7C0BC0C0" ID-DOI="10.5852/ejt.2024.921.2429" ID-ISSN="2118-9773" ID-Zenodo-Dep="10663874" ID-ZooBank="02633F29-4CDF-4027-BEBF-07AD2F925B42" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1707990651991" checkinUser="plazi" docAuthor="Bellvert, Adrià, Dimitrov, Dragomir, Zamani, Alireza &amp; Arnedo, Miquel A." docDate="2024" docId="03DA7943B728D84371A28F58412DFC30" docLanguage="en" docName="EJT.2024.921.210-235.pdf" docOrigin="European Journal of Taxonomy 921 (210)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2429/10759" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dysdera sancticedri Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo 2024, comb. nov." docType="treatment" docVersion="3" lastPageNumber="228" masterDocId="FFE3013BB738D85170598D5B445AFFC5" masterDocTitle="Integrating museum collections and molecules reveals genus-level synonymy and new species in red devil spiders (Araneae, Dysderidae) from the Middle East and Central Asia" masterLastPageNumber="235" masterPageNumber="210" pageNumber="226" updateTime="1708090743251" updateUser="tatiana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="EDE10045803F37E0FD804794C7A9FDC9">
<mods:title id="8AFC8BAF9AE4B4D58132E2E18669DCCC">Integrating museum collections and molecules reveals genus-level synonymy and new species in red devil spiders (Araneae, Dysderidae) from the Middle East and Central Asia</mods:title>
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<mods:namePart id="02FC8C6E21EB1E9DBBCAD9FB3E6D101F">Bellvert, Adrià</mods:namePart>
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<mods:affiliation id="51330ED8A76DE4C222667A25B634F211">Departament de Biologia Evolutiva, Ecologia i Ciències Ambientals, Universitat de Barcelona (UB), Av. Diagonal, 643, 08028 Barcelona, Spain. &amp; Institut de Recerca de la Biodiversitat (IRBio), Universitat de Barcelona (UB) Barcelona, Spain.</mods:affiliation>
<mods:nameIdentifier id="8B8B57D08623BC60AC4B35FDF398FC35" type="email">abellvertba@gmail.com</mods:nameIdentifier>
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<mods:namePart id="124137F2EC54ADFF10418C3516E6F478">Dimitrov, Dragomir</mods:namePart>
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<mods:affiliation id="B091C2E922FB5A437BFAA8F9575F89AB">Departament de Biologia Evolutiva, Ecologia i Ciències Ambientals, Universitat de Barcelona (UB), Av. Diagonal, 643, 08028 Barcelona, Spain. &amp; Institut de Recerca de la Biodiversitat (IRBio), Universitat de Barcelona (UB) Barcelona, Spain. &amp; National Museum of Natural Sciences, Bulgarian Academy of Sciences, 1 Tsar Osvoboditel Blvd, 1000 Sofia, Bulgaria.</mods:affiliation>
<mods:nameIdentifier id="96927FCBC0BB1154A14D6FC8577EA27E" type="email">dimitrov.drk@gmail.com</mods:nameIdentifier>
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<mods:namePart id="11571F8815C71056A46ECD0127E60672">Zamani, Alireza</mods:namePart>
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<mods:affiliation id="06C0B976D539E00CD5E7E3B156667B2A">Zoological Museum, Biodiversity Unit, FI- 20014 University of Turku, Finland.</mods:affiliation>
<mods:nameIdentifier id="E5C16AA40BD3D09654033600849767BD" type="email">zamani.alireza5@gmail.com</mods:nameIdentifier>
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<mods:namePart id="7C847287F8B6E1FC97A60E358A67C640">Arnedo, Miquel A.</mods:namePart>
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<mods:affiliation id="806105E386F47AEB6548337C4C233D09">Departament de Biologia Evolutiva, Ecologia i Ciències Ambientals, Universitat de Barcelona (UB), Av. Diagonal, 643, 08028 Barcelona, Spain. &amp; Institut de Recerca de la Biodiversitat (IRBio), Universitat de Barcelona (UB) Barcelona, Spain.</mods:affiliation>
<mods:nameIdentifier id="85EBEDCD9108A5EC27660B23C451B3FA" type="email">marnedo@gmail.com</mods:nameIdentifier>
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<mods:title id="6E965BB171F7D00E644784DFE0666EA5">European Journal of Taxonomy</mods:title>
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<mods:date id="69E77075EA6DF703CF0C13C0513FD3C7">2024</mods:date>
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<mods:number id="58B5D339014410D776E406087B54630C">2024-02-14</mods:number>
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<mods:number id="76660B73B840E2F87C81805B8BBD2170">921</mods:number>
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<treatment id="03DA7943B728D84371A28F58412DFC30" ID-DOI="http://doi.org/10.5281/zenodo.10663898" ID-Zenodo-Dep="10663898" LSID="urn:lsid:plazi:treatment:03DA7943B728D84371A28F58412DFC30" httpUri="http://treatment.plazi.org/id/03DA7943B728D84371A28F58412DFC30" lastPageId="18" lastPageNumber="228" pageId="16" pageNumber="226">
<subSubSection id="C3699BDEB728D84171A28F584063FDDB" box="[507,1081,515,542]" pageId="16" pageNumber="226" type="nomenclature">
<paragraph id="8BCCC855B728D84171A28F584063FDDB" blockId="16.[507,1081,515,578]" box="[507,1081,515,542]" pageId="16" pageNumber="226">
<heading id="D0847F39B728D84171A28F584063FDDB" box="[507,1081,515,542]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="226" reason="2">
<taxonomicName id="4C73B3D6B728D84171A28F5847F5FDD8" authority="(Brignoli, 1978)" authorityName="Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo" authorityYear="2024" baseAuthorityName="Brignoli" baseAuthorityYear="1978" box="[507,943,515,542]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="226" phylum="Arthropoda" rank="species" species="sancticedri" status="comb. nov.">
<emphasis id="B9071447B728D84171A28F5846B0FDDB" bold="true" box="[507,746,515,542]" italics="true" pageId="16" pageNumber="226">Dysdera sancticedri</emphasis>
(
<bibRefCitation id="EFE2B5A4B728D84172A08F5847FDFDD8" author="Brignoli" box="[761,935,515,541]" firstAuthor="Brignoli" pageId="16" pageNumber="226" pagination="172 - 175" refId="ref11294" refString="Brignoli P. M. 1978. Araignees du Liban IV. Notes sur quelques Dysderidae (Araneae). Bulletin de la Societe d'Histoire Naturelle de Toulouse 114: 172 - 175." type="journal article" year="1978">Brignoli, 1978</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A234A93CB728D84173EE8F584063FDDB" box="[951,1081,515,542]" pageId="16" pageNumber="226" rank="species">comb. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3699BDEB728D84172818F7C4701FD87" box="[728,859,551,578]" pageId="16" pageNumber="226" type="description">
<paragraph id="8BCCC855B728D84172818F7C4701FD87" blockId="16.[507,1081,515,578]" box="[728,859,551,578]" pageId="16" pageNumber="226">
<figureCitation id="1348D4D0B728D84172818F7C4701FD87" box="[728,859,551,578]" captionStart="Figs 21–25" captionStartId="17.[189,241,1929,1955]" captionTargetBox="[278,1309,265,1891]" captionTargetId="figure-16@17.[278,1309,265,1891]" captionTargetPageId="17" captionText="Figs 21–25. Male Dysdera sancticedri (Brignoli, 1978) comb. nov. (MCSN). 21–22. Carapace. 21. Dorsal view. 22. Ventral view. 23–25. Left palp. 23. Prolateral view. 24. Anterior view. 25. Retrolateral view; arrowhead showing a small knob on the anterior side of T. Scale bars: 21–22 = 1.5 mm; 23–25 =0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663894" httpUri="https://zenodo.org/record/10663894/files/figure.png" pageId="16" pageNumber="226">Figs 21–25</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3699BDEB728D84170E48F304748FD41" box="[189,786,618,644]" pageId="16" pageNumber="226" type="reference_group">
<paragraph id="8BCCC855B728D84170E48F304748FD41" blockId="16.[189,786,618,644]" box="[189,786,618,644]" pageId="16" pageNumber="226">
<treatmentCitationGroup id="AB63EF7BB728D84170E48F304748FD41" box="[189,786,618,644]" pageId="16" pageNumber="226">
<treatmentCitation id="0AD2EE44B728D84170E48F3046F8FD41" author="Brignoli P. M." box="[189,674,618,644]" page="174" pageId="16" pageNumber="226" year="1978">
<taxonomicName id="4C73B3D6B728D84170E48F3046F8FD41" ID-CoL="3479F" authority="Brignoli, 1978: 174" authorityName="Brignoli" authorityPageNumber="174" authorityYear="1978" box="[189,674,618,644]" class="Arachnida" family="Dysderidae" genus="Dasumia" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="226" phylum="Arthropoda" rank="species" species="sancticedri">
<emphasis id="B9071447B728D84170E48F3045EAFD41" box="[189,432,618,644]" italics="true" pageId="16" pageNumber="226">Dasumia sancticedri</emphasis>
<bibRefCitation id="EFE2B5A4B728D84171E18F3146F8FD41" author="Brignoli" box="[440,674,618,644]" firstAuthor="Brignoli" page="174" pageId="16" pageNumber="226" pagination="172 - 175" refId="ref11294" refString="Brignoli P. M. 1978. Araignees du Liban IV. Notes sur quelques Dysderidae (Araneae). Bulletin de la Societe d'Histoire Naturelle de Toulouse 114: 172 - 175." type="journal article" year="1978">Brignoli, 1978: 174</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 1–2.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3699BDEB728D84170E48FF7456CFD03" box="[189,310,684,710]" pageId="16" pageNumber="226" type="diagnosis">
<paragraph id="8BCCC855B728D84170E48FF7456CFD03" blockId="16.[189,310,684,710]" box="[189,310,684,710]" pageId="16" pageNumber="226">
<heading id="D0847F39B728D84170E48FF7456CFD03" bold="true" box="[189,310,684,710]" fontSize="11" level="3" pageId="16" pageNumber="226" reason="3">
<emphasis id="B9071447B728D84170E48FF7456CFD03" bold="true" box="[189,310,684,710]" pageId="16" pageNumber="226">Diagnosis</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3699BDEB728D84170E48F874501FC36" pageId="16" pageNumber="226" type="reference_group">
<paragraph id="8BCCC855B728D84170E48F874501FC36" blockId="16.[188,1400,732,1011]" pageId="16" pageNumber="226">
<taxonomicName id="4C73B3D6B728D84170E48F8745F2FD33" authorityName="Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo" authorityYear="2024" baseAuthorityName="Brignoli" baseAuthorityYear="1978" box="[189,424,732,758]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="226" phylum="Arthropoda" rank="species" species="sancticedri" status="comb. nov.">
<emphasis id="B9071447B728D84170E48F8745F2FD33" box="[189,424,732,758]" italics="true" pageId="16" pageNumber="226">Dysdera sancticedri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A234A93CB728D84171F48F874677FD33" box="[429,557,732,758]" pageId="16" pageNumber="226" rank="species">comb. nov.</taxonomicNameLabel>
resembles
<taxonomicName id="4C73B3D6B728D84172F48F874064FD33" authority="(Caporiacco, 1929)" baseAuthorityName="Caporiacco" baseAuthorityYear="1929" box="[685,1086,732,758]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="226" phylum="Arthropoda" rank="species" species="festai">
<emphasis id="B9071447B728D84172F48F87470CFD33" box="[685,854,732,758]" italics="true" pageId="16" pageNumber="226">Dysdera festai</emphasis>
(
<bibRefCitation id="EFE2B5A4B728D841733C8F87406CFD33" author="Caporiacco" box="[869,1078,732,758]" firstAuthor="Caporiacco" pageId="16" pageNumber="226" pagination="221 - 242" refId="ref11328" refString="Caporiacco L. di 1929. Aracnidi. In: Ricerche faunitische nelle isole italiane dell'Egeo. Archivio Zoologico Italiano 13: 221 - 242." type="journal article" year="1929">Caporiacco, 1929</bibRefCitation>
)
</taxonomicName>
by the conformation of the male palp, having a similar external sclerite. The two species differ by (1) the lack of a posterior apophysis in
<taxonomicName id="4C73B3D6B728D84170828E7E45DBFCFB" authorityName="Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo" authorityYear="2024" baseAuthorityName="Brignoli" baseAuthorityYear="1978" box="[219,385,804,830]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="226" phylum="Arthropoda" rank="species" species="sancticedri">
<emphasis id="B9071447B728D84170828E7E45DBFCFB" box="[219,385,804,830]" italics="true" pageId="16" pageNumber="226">D. sancticedri</emphasis>
</taxonomicName>
(
<figureCitation id="1348D4D0B728D84171C88E7F4643FCFA" box="[401,537,804,831]" captionStart="Figs 21–25" captionStartId="17.[189,241,1929,1955]" captionTargetBox="[278,1309,265,1891]" captionTargetId="figure-16@17.[278,1309,265,1891]" captionTargetPageId="17" captionText="Figs 21–25. Male Dysdera sancticedri (Brignoli, 1978) comb. nov. (MCSN). 21–22. Carapace. 21. Dorsal view. 22. Ventral view. 23–25. Left palp. 23. Prolateral view. 24. Anterior view. 25. Retrolateral view; arrowhead showing a small knob on the anterior side of T. Scale bars: 21–22 = 1.5 mm; 23–25 =0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663894" httpUri="https://zenodo.org/record/10663894/files/figure.png" pageId="16" pageNumber="226">Figs 23, 25</figureCitation>
), vs present in
<taxonomicName id="4C73B3D6B728D84172938E7E4774FCFB" baseAuthorityName="Caporiacco" baseAuthorityYear="1929" box="[714,814,804,830]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="226" phylum="Arthropoda" rank="species" species="festai">
<emphasis id="B9071447B728D84172938E7E4774FCFB" box="[714,814,804,830]" italics="true" pageId="16" pageNumber="226">D. festai</emphasis>
</taxonomicName>
(
<figureCitation id="1348D4D0B728D84173678E7F47CFFCFB" box="[830,917,804,830]" captionStart="Figs 26–33" captionStartId="19.[189,241,1928,1954]" captionTargetBox="[381,1207,271,1890]" captionTargetId="figure-15@19.[381,1207,265,1890]" captionTargetPageId="19" captionText="Figs 26–33. Male Dysdera festai Caporiacco, 1929 (26, 28–30 = specimen from Turkey (NHMW); 27, 31–33 = specimen from Greece (MZSUT)). 26–27. Cephalic region. 28–33. Left palp. 28, 31. Prolateral view. 29, 32. Anterior view. 30, 33. Retrolateral view. Scale bars: 26–27 = 2 mm; 28–33 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663896" httpUri="https://zenodo.org/record/10663896/files/figure.png" pageId="16" pageNumber="226">Fig. 28</figureCitation>
), (2) the reduced, hollow interior sclerite with a tubular shape in
<taxonomicName id="4C73B3D6B728D84171948E124629FCA7" authorityName="Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo" authorityYear="2024" baseAuthorityName="Brignoli" baseAuthorityYear="1978" box="[461,627,840,866]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="226" phylum="Arthropoda" rank="species" species="sancticedri">
<emphasis id="B9071447B728D84171948E124629FCA7" box="[461,627,840,866]" italics="true" pageId="16" pageNumber="226">D. sancticedri</emphasis>
</taxonomicName>
(
<figureCitation id="1348D4D0B728D84172DA8E134681FCA6" box="[643,731,840,867]" captionStart="Figs 21–25" captionStartId="17.[189,241,1929,1955]" captionTargetBox="[278,1309,265,1891]" captionTargetId="figure-16@17.[278,1309,265,1891]" captionTargetPageId="17" captionText="Figs 21–25. Male Dysdera sancticedri (Brignoli, 1978) comb. nov. (MCSN). 21–22. Carapace. 21. Dorsal view. 22. Ventral view. 23–25. Left palp. 23. Prolateral view. 24. Anterior view. 25. Retrolateral view; arrowhead showing a small knob on the anterior side of T. Scale bars: 21–22 = 1.5 mm; 23–25 =0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663894" httpUri="https://zenodo.org/record/10663894/files/figure.png" pageId="16" pageNumber="226">Fig. 24</figureCitation>
), vs well developed and connected basally to the exterior one in
<taxonomicName id="4C73B3D6B728D84171488E35452CFC42" baseAuthorityName="Caporiacco" baseAuthorityYear="1929" box="[273,374,877,903]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="226" phylum="Arthropoda" rank="species" species="festai">
<emphasis id="B9071447B728D84171488E35452CFC42" box="[273,374,877,903]" italics="true" pageId="16" pageNumber="226">D. festai</emphasis>
</taxonomicName>
(
<figureCitation id="1348D4D0B728D84171DF8E364585FC42" box="[390,479,877,903]" captionStart="Figs 26–33" captionStartId="19.[189,241,1928,1954]" captionTargetBox="[381,1207,271,1890]" captionTargetId="figure-15@19.[381,1207,265,1890]" captionTargetPageId="19" captionText="Figs 26–33. Male Dysdera festai Caporiacco, 1929 (26, 28–30 = specimen from Turkey (NHMW); 27, 31–33 = specimen from Greece (MZSUT)). 26–27. Cephalic region. 28–33. Left palp. 28, 31. Prolateral view. 29, 32. Anterior view. 30, 33. Retrolateral view. Scale bars: 26–27 = 2 mm; 28–33 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663896" httpUri="https://zenodo.org/record/10663896/files/figure.png" pageId="16" pageNumber="226">Fig. 29</figureCitation>
), (3) the short, rounded tegulum, wider than long in
<taxonomicName id="4C73B3D6B728D841740C8E3540A6FC42" authorityName="Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo" authorityYear="2024" baseAuthorityName="Brignoli" baseAuthorityYear="1978" box="[1109,1276,877,903]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="226" phylum="Arthropoda" rank="species" species="sancticedri">
<emphasis id="B9071447B728D841740C8E3540A6FC42" box="[1109,1276,877,903]" italics="true" pageId="16" pageNumber="226">D. sancticedri</emphasis>
</taxonomicName>
(
<figureCitation id="1348D4D0B728D841754F8E364484FC69" captionStart="Figs 21–25" captionStartId="17.[189,241,1929,1955]" captionTargetBox="[278,1309,265,1891]" captionTargetId="figure-16@17.[278,1309,265,1891]" captionTargetPageId="17" captionText="Figs 21–25. Male Dysdera sancticedri (Brignoli, 1978) comb. nov. (MCSN). 21–22. Carapace. 21. Dorsal view. 22. Ventral view. 23–25. Left palp. 23. Prolateral view. 24. Anterior view. 25. Retrolateral view; arrowhead showing a small knob on the anterior side of T. Scale bars: 21–22 = 1.5 mm; 23–25 =0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663894" httpUri="https://zenodo.org/record/10663894/files/figure.png" pageId="16" pageNumber="226">Figs 23, 25</figureCitation>
), vs more elongated and longer than wide in
<taxonomicName id="4C73B3D6B728D84172BC8EC94712FC6E" baseAuthorityName="Caporiacco" baseAuthorityYear="1929" box="[741,840,913,939]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="226" phylum="Arthropoda" rank="species" species="festai">
<emphasis id="B9071447B728D84172BC8EC94712FC6E" box="[741,840,913,939]" italics="true" pageId="16" pageNumber="226">D. festai</emphasis>
</taxonomicName>
(
<figureCitation id="1348D4D0B728D841730E8ECA4785FC6E" box="[855,991,913,939]" captionStart="Figs 26–33" captionStartId="19.[189,241,1928,1954]" captionTargetBox="[381,1207,271,1890]" captionTargetId="figure-15@19.[381,1207,265,1890]" captionTargetPageId="19" captionText="Figs 26–33. Male Dysdera festai Caporiacco, 1929 (26, 28–30 = specimen from Turkey (NHMW); 27, 31–33 = specimen from Greece (MZSUT)). 26–27. Cephalic region. 28–33. Left palp. 28, 31. Prolateral view. 29, 32. Anterior view. 30, 33. Retrolateral view. Scale bars: 26–27 = 2 mm; 28–33 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663896" httpUri="https://zenodo.org/record/10663896/files/figure.png" pageId="16" pageNumber="226">Figs 28–30</figureCitation>
), and (4) the long chelicerae, about ⅗ of carapace length in dorsal view in
<taxonomicName id="4C73B3D6B728D84172DE8EED4777FC0A" authorityName="Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo" authorityYear="2024" baseAuthorityName="Brignoli" baseAuthorityYear="1978" box="[647,813,949,975]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="226" phylum="Arthropoda" rank="species" species="sancticedri">
<emphasis id="B9071447B728D84172DE8EED4777FC0A" box="[647,813,949,975]" italics="true" pageId="16" pageNumber="226">D. sancticedri</emphasis>
</taxonomicName>
(
<figureCitation id="1348D4D0B728D84173678EEE479DFC0A" box="[830,967,949,975]" captionStart="Figs 21–25" captionStartId="17.[189,241,1929,1955]" captionTargetBox="[278,1309,265,1891]" captionTargetId="figure-16@17.[278,1309,265,1891]" captionTargetPageId="17" captionText="Figs 21–25. Male Dysdera sancticedri (Brignoli, 1978) comb. nov. (MCSN). 21–22. Carapace. 21. Dorsal view. 22. Ventral view. 23–25. Left palp. 23. Prolateral view. 24. Anterior view. 25. Retrolateral view; arrowhead showing a small knob on the anterior side of T. Scale bars: 21–22 = 1.5 mm; 23–25 =0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663894" httpUri="https://zenodo.org/record/10663894/files/figure.png" pageId="16" pageNumber="226">Figs 21–22</figureCitation>
), vs significantly smaller in
<taxonomicName id="4C73B3D6B728D841754A8EED412DFC0A" baseAuthorityName="Caporiacco" baseAuthorityYear="1929" box="[1299,1399,949,975]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="226" phylum="Arthropoda" rank="species" species="festai">
<emphasis id="B9071447B728D841754A8EED412DFC0A" box="[1299,1399,949,975]" italics="true" pageId="16" pageNumber="226">D. festai</emphasis>
</taxonomicName>
(
<figureCitation id="1348D4D0B728D841709C8E824515FC36" box="[197,335,985,1011]" captionStart="Figs 26–33" captionStartId="19.[189,241,1928,1954]" captionTargetBox="[381,1207,271,1890]" captionTargetId="figure-15@19.[381,1207,265,1890]" captionTargetPageId="19" captionText="Figs 26–33. Male Dysdera festai Caporiacco, 1929 (26, 28–30 = specimen from Turkey (NHMW); 27, 31–33 = specimen from Greece (MZSUT)). 26–27. Cephalic region. 28–33. Left palp. 28, 31. Prolateral view. 29, 32. Anterior view. 30, 33. Retrolateral view. Scale bars: 26–27 = 2 mm; 28–33 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663896" httpUri="https://zenodo.org/record/10663896/files/figure.png" pageId="16" pageNumber="226">Figs 26–27</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3699BDEB728D84170E489474678FB6B" pageId="16" pageNumber="226" type="materials_examined">
<paragraph id="8BCCC855B728D84170E489474536FBF3" blockId="16.[189,364,1052,1078]" box="[189,364,1052,1078]" pageId="16" pageNumber="226">
<heading id="D0847F39B728D84170E489474536FBF3" bold="true" box="[189,364,1052,1078]" fontSize="11" level="3" pageId="16" pageNumber="226" reason="3">
<emphasis id="B9071447B728D84170E489474536FBF3" bold="true" box="[189,364,1052,1078]" pageId="16" pageNumber="226">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BCCC855B728D84170B389174506FBA3" blockId="16.[189,1399,1100,1198]" box="[234,348,1100,1126]" pageId="16" pageNumber="226">
<emphasis id="B9071447B728D84170B389174506FBA3" bold="true" box="[234,348,1100,1126]" pageId="16" pageNumber="226">
<typeStatus id="54C876F7B728D84170B389174506FBA3" box="[234,348,1100,1126]" pageId="16" pageNumber="226" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BCCC855B728D84170E4892B4678FB6B" blockId="16.[189,1399,1100,1198]" pageId="16" pageNumber="226">
<materialsCitation id="3B1BC208B728D84170E4892B4678FB6B" collectingDate="1972-06-05" collectionCode="MCSN" collectorName="de Bacharre &amp; P. Brignoli" country="Lebanon" elevation="1950" location="North Gorvernorate" municipality="Cedres de Bacharre" pageId="16" pageNumber="226" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectingCountry id="F36488C5B728D84170E4892B4510FB4F" box="[189,330,1136,1162]" name="Lebanon" pageId="16" pageNumber="226">LEBANON</collectingCountry>
•
<specimenCount id="9D7503DCB728D841713F892B45CDFB4F" box="[358,407,1136,1162]" count="1" pageId="16" pageNumber="226" type="male">1 ♂</specimenCount>
;
<location id="8EAC9E8EB728D84171FC892B46C8FB4F" LSID="urn:lsid:plazi:treatment:03DA7943B728D84371A28F58412DFC30:8EAC9E8EB728D84171FC892B46C8FB4F" box="[421,658,1136,1162]" country="Lebanon" municipality="Cedres de Bacharre" name="North Gorvernorate" pageId="16" pageNumber="226">North Gorvernorate</location>
, des
<collectingMunicipality id="6BA8522FB728D8417289892B47E0FB4F" box="[720,954,1136,1162]" pageId="16" pageNumber="226">
Cèdres
<collectorName id="2686AD83B728D8417372892B47E0FB4F" box="[811,954,1136,1162]" pageId="16" pageNumber="226">de Bacharré</collectorName>
</collectingMunicipality>
(caza Bcharré);
<elevation id="005E2F66B728D84174DA89344145FB4F" box="[1155,1311,1135,1162]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="16" pageNumber="226" unit="m" value="1950.0">
<quantity id="4C8B65B0B728D84174DA89344084FB4F" box="[1155,1246,1135,1162]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="16" pageNumber="226" unit="m" value="1950.0">1950 m</quantity>
a.s.l.
</elevation>
;
<date id="FFCDEE95B728D8417576893444A1FB6B" pageId="16" pageNumber="226" value="1972-06-05">
<collectingDate id="EF89177DB728D8417576893444A1FB6B" pageId="16" pageNumber="226" value="1972-06-05">5 Jun. 1972</collectingDate>
</date>
;
<collectorName id="2686AD83B728D841715E89CF45DFFB6B" box="[263,389,1172,1198]" pageId="16" pageNumber="226">P. Brignoli</collectorName>
leg.;
<collectionCode id="ED625090B728D841719E89CF4644FB6B" box="[455,542,1172,1198]" country="Italy" httpUri="http://grbio.org/cool/353t-cj9k" name="Museo Civico di Storia Naturale, Verona" pageId="16" pageNumber="226">MCSN</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3699BDEB728D84370E4898D4566FD62" lastPageId="18" lastPageNumber="228" pageId="16" pageNumber="226" type="description">
<paragraph id="8BCCC855B728D84170E4898D4514FB35" blockId="16.[189,334,1238,1264]" box="[189,334,1238,1264]" pageId="16" pageNumber="226">
<heading id="D0847F39B728D84170E4898D4514FB35" bold="true" box="[189,334,1238,1264]" fontSize="11" level="3" pageId="16" pageNumber="226" reason="3">
<emphasis id="B9071447B728D84170E4898D4514FB35" bold="true" box="[189,334,1238,1264]" pageId="16" pageNumber="226">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BCCC855B728D84170B3885D469CF987" blockId="16.[187,1400,1286,1602]" pageId="16" pageNumber="226">
<emphasis id="B9071447B728D84170B3885D4570FAE5" bold="true" box="[234,298,1286,1312]" pageId="16" pageNumber="226">Male</emphasis>
PROSOMA. 2.70 long; maximum width 2.14; minimum width 1.53. Uniformly red; cephalic region smooth, thorax heavily covered with circular depressions. Anterior border roughly round, about ⅗ of carapace length; anterior lateral borders divergent; rounded at maximum dorsal width, posterior lateral borders rounded; posterior margin narrow, slightly bilobulated. Eye diameters: AME 0.167; PLE 0.129; PME 0.122; AMEs on edge of the anterior border, separated from one another by about ¾ diameter, touching PLEs; PMEs very close to one another, about ⅓ of PME diameter from PLEs. Labium trapezoid-shaped, base wider than the distal part; as long as wide at base; triangular groove at the tip. Sternum dark orange, darkened on borders; very slightly wrinkled.
</paragraph>
<paragraph id="8BCCC855B728D84170E48B314518F909" blockId="16.[189,1398,1641,1741]" pageId="16" pageNumber="226">
CHELICERAE. 1.62 long, about ⅗ of carapace length in dorsal view; fang 1.865 long; paturon smooth, with no granulations (
<figureCitation id="1348D4D0B728D841719B8BD54614F96D" box="[450,590,1678,1704]" captionStart="Figs 21–25" captionStartId="17.[189,241,1929,1955]" captionTargetBox="[278,1309,265,1891]" captionTargetId="figure-16@17.[278,1309,265,1891]" captionTargetPageId="17" captionText="Figs 21–25. Male Dysdera sancticedri (Brignoli, 1978) comb. nov. (MCSN). 21–22. Carapace. 21. Dorsal view. 22. Ventral view. 23–25. Left palp. 23. Prolateral view. 24. Anterior view. 25. Retrolateral view; arrowhead showing a small knob on the anterior side of T. Scale bars: 21–22 = 1.5 mm; 23–25 =0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663894" httpUri="https://zenodo.org/record/10663894/files/figure.png" pageId="16" pageNumber="226">Figs 21–22</figureCitation>
). Cheliceral inner groove not visible; armed with slender long setae at the base.
</paragraph>
<paragraph id="8BCCC855B728D84370E48BAE4785FE82" blockId="16.[187,1400,1781,2025]" lastBlockId="18.[189,1399,265,327]" lastPageId="18" lastPageNumber="228" pageId="16" pageNumber="226">LEGS. Orange. Lengths of leg segments: fe1 2; pa1 1.29; ti1 1.73; me1 1.72; ta1 0.71; total 8.45; fe2 1.70; pa2 1.14; ti2 1.40; me2 1.44; ta2 0.63; total 6.31; fe3 1.37; pa3 0.85; ti3 0.91; me3 1.12; ta3 0.44; total 4.69; fe4 1.83; pa4 1.07; ti4 1.41; me4 1.58; ta4 0.49; total 6.38; fe Pdp 1.27; pa Pdp 0.76; ti Pdp 0.64; ta Pdp 0.81; total 3.48; leg formula: 1&gt;4&gt; 2&gt;3. Leg2 spineless; leg4 spineless. Fe3d spineless; pa3 spineless; tb3d spines arranged in one band; proximal 0; medial-proximal 0; medial-distal 0; distal 1.0.0; tb3v spines spineless; without terminal spines. Dorsal side of frontal legs covered with setae, lacking small grains; ventral side of pedipalp smooth. Claws with less than 5 teeth; hardly larger than claw width. OPISTHOSOMA. 3.57 long; cream-colored; cylindrical. Abdominal dorsal setae 0.03 long; thin, roughly straight, not compressed, pointed; uniformly and thickly distributed.</paragraph>
<caption id="DF0C98DDB729D84070E48AD24122F82F" ID-DOI="http://doi.org/10.5281/zenodo.10663894" ID-Zenodo-Dep="10663894" httpUri="https://zenodo.org/record/10663894/files/figure.png" pageId="17" pageNumber="227" startId="17.[189,241,1929,1955]" targetBox="[278,1309,265,1891]" targetPageId="17">
<paragraph id="8BCCC855B729D84070E48AD24122F82F" blockId="17.[187,1400,1929,2026]" pageId="17" pageNumber="227">
<emphasis id="B9071447B729D84070E48AD2451CF866" bold="true" box="[189,326,1929,1955]" pageId="17" pageNumber="227">Figs 21–25.</emphasis>
Male
<taxonomicName id="4C73B3D6B729D84071D48AD2476FF866" authority="(Brignoli, 1978)" authorityName="Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo" authorityYear="2024" baseAuthorityName="Brignoli" baseAuthorityYear="1978" box="[397,821,1929,1955]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="227" phylum="Arthropoda" rank="species" species="sancticedri" status="comb. nov.">
<emphasis id="B9071447B729D84071D48AD2462CF866" box="[397,630,1929,1955]" italics="true" pageId="17" pageNumber="227">Dysdera sancticedri</emphasis>
(
<bibRefCitation id="EFE2B5A4B729D84072DA8AD24777F866" author="Brignoli" box="[643,813,1929,1955]" firstAuthor="Brignoli" pageId="17" pageNumber="227" pagination="172 - 175" refId="ref11294" refString="Brignoli P. M. 1978. Araignees du Liban IV. Notes sur quelques Dysderidae (Araneae). Bulletin de la Societe d'Histoire Naturelle de Toulouse 114: 172 - 175." type="journal article" year="1978">Brignoli, 1978</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A234A93CB729D84073628AD247E0F861" box="[827,954,1929,1956]" pageId="17" pageNumber="227" rank="species">comb. nov.</taxonomicNameLabel>
(MCSN).
<emphasis id="B9071447B729D84074698AD24022F866" bold="true" box="[1072,1144,1929,1955]" pageId="17" pageNumber="227">21–22</emphasis>
. Carapace.
<emphasis id="B9071447B729D84074A68AD24146F866" bold="true" box="[1279,1308,1929,1955]" pageId="17" pageNumber="227">21</emphasis>
. Dorsal view.
<emphasis id="B9071447B729D840715D8AF7457BF803" bold="true" box="[260,289,1964,1990]" pageId="17" pageNumber="227">22</emphasis>
. Ventral view.
<emphasis id="B9071447B729D840718C8AF74644F803" bold="true" box="[469,542,1964,1990]" pageId="17" pageNumber="227">23–25</emphasis>
. Left palp.
<emphasis id="B9071447B729D84072F38AF7469CF803" bold="true" box="[682,710,1964,1990]" pageId="17" pageNumber="227">23</emphasis>
. Prolateral view.
<emphasis id="B9071447B729D84073C18AF747EEF803" bold="true" box="[920,948,1964,1990]" pageId="17" pageNumber="227">24</emphasis>
. Anterior view.
<emphasis id="B9071447B729D840742C8AF740C8F803" bold="true" box="[1141,1170,1964,1990]" pageId="17" pageNumber="227">25</emphasis>
. Retrolateral view; arrowhead showing a small knob on the anterior side of T. Scale bars: 21–22 = 1.5 mm; 23–25 =0.5 mm.
</paragraph>
</caption>
<paragraph id="8BCCC855B72AD84370E48C344566FD62" blockId="18.[187,1399,366,679]" pageId="18" pageNumber="228">
PALP (
<figureCitation id="1348D4D0B72AD843715C8C3445D7FE4C" box="[261,397,366,393]" captionStart="Figs 21–25" captionStartId="17.[189,241,1929,1955]" captionTargetBox="[278,1309,265,1891]" captionTargetId="figure-16@17.[278,1309,265,1891]" captionTargetPageId="17" captionText="Figs 21–25. Male Dysdera sancticedri (Brignoli, 1978) comb. nov. (MCSN). 21–22. Carapace. 21. Dorsal view. 22. Ventral view. 23–25. Left palp. 23. Prolateral view. 24. Anterior view. 25. Retrolateral view; arrowhead showing a small knob on the anterior side of T. Scale bars: 21–22 = 1.5 mm; 23–25 =0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663894" httpUri="https://zenodo.org/record/10663894/files/figure.png" pageId="18" pageNumber="228">Figs 23–25</figureCitation>
). T short and rounded, slightly longer than ED, bearing small knob on the anterior side (arrowed in
<figureCitation id="1348D4D0B72AD84371118CC945C7FE68" box="[328,413,402,429]" captionStart="Figs 21–25" captionStartId="17.[189,241,1929,1955]" captionTargetBox="[278,1309,265,1891]" captionTargetId="figure-16@17.[278,1309,265,1891]" captionTargetPageId="17" captionText="Figs 21–25. Male Dysdera sancticedri (Brignoli, 1978) comb. nov. (MCSN). 21–22. Carapace. 21. Dorsal view. 22. Ventral view. 23–25. Left palp. 23. Prolateral view. 24. Anterior view. 25. Retrolateral view; arrowhead showing a small knob on the anterior side of T. Scale bars: 21–22 = 1.5 mm; 23–25 =0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663894" httpUri="https://zenodo.org/record/10663894/files/figure.png" pageId="18" pageNumber="228">Fig. 25</figureCitation>
); external distal border sloped forwards; internal sloped backward. ED not bent, same T axis in lateral view, internal distal border not expanded. ES wide markedly sclerotized, IS reduced to whip-like sclerotization (
<figureCitation id="1348D4D0B72AD84371BC8C814667FE31" box="[485,573,474,500]" captionStart="Figs 21–25" captionStartId="17.[189,241,1929,1955]" captionTargetBox="[278,1309,265,1891]" captionTargetId="figure-16@17.[278,1309,265,1891]" captionTargetPageId="17" captionText="Figs 21–25. Male Dysdera sancticedri (Brignoli, 1978) comb. nov. (MCSN). 21–22. Carapace. 21. Dorsal view. 22. Ventral view. 23–25. Left palp. 23. Prolateral view. 24. Anterior view. 25. Retrolateral view; arrowhead showing a small knob on the anterior side of T. Scale bars: 21–22 = 1.5 mm; 23–25 =0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663894" httpUri="https://zenodo.org/record/10663894/files/figure.png" pageId="18" pageNumber="228">Fig. 24</figureCitation>
). ED tip reduced, only the sclerotized structures are visible. C absent. AC absent. L lateral margin distally projected, split into two spine-like apophyses (
<figureCitation id="1348D4D0B72AD84374D88CA54082FDDD" box="[1153,1240,510,536]" captionStart="Figs 21–25" captionStartId="17.[189,241,1929,1955]" captionTargetBox="[278,1309,265,1891]" captionTargetId="figure-16@17.[278,1309,265,1891]" captionTargetPageId="17" captionText="Figs 21–25. Male Dysdera sancticedri (Brignoli, 1978) comb. nov. (MCSN). 21–22. Carapace. 21. Dorsal view. 22. Ventral view. 23–25. Left palp. 23. Prolateral view. 24. Anterior view. 25. Retrolateral view; arrowhead showing a small knob on the anterior side of T. Scale bars: 21–22 = 1.5 mm; 23–25 =0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663894" httpUri="https://zenodo.org/record/10663894/files/figure.png" pageId="18" pageNumber="228">Fig. 24</figureCitation>
). L is a single well-developed apophysis, curved mesally. LA absent. F absent. AL absent. P absent (
<figureCitation id="1348D4D0B72AD84374F58F7A416FFDF9" box="[1196,1333,545,572]" captionStart="Figs 21–25" captionStartId="17.[189,241,1929,1955]" captionTargetBox="[278,1309,265,1891]" captionTargetId="figure-16@17.[278,1309,265,1891]" captionTargetPageId="17" captionText="Figs 21–25. Male Dysdera sancticedri (Brignoli, 1978) comb. nov. (MCSN). 21–22. Carapace. 21. Dorsal view. 22. Ventral view. 23–25. Left palp. 23. Prolateral view. 24. Anterior view. 25. Retrolateral view; arrowhead showing a small knob on the anterior side of T. Scale bars: 21–22 = 1.5 mm; 23–25 =0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663894" httpUri="https://zenodo.org/record/10663894/files/figure.png" pageId="18" pageNumber="228">Figs 23, 25</figureCitation>
); DH sloped, forming an angle of ca 135° to T in lateral view; lateral length ca ¼ of T width; ridge ca 45° to T in lateral view, not sclerotized; not expanded, upper margin smooth; not distally projected; back margin not folded.
</paragraph>
</subSubSection>
<subSubSection id="C3699BDEB72AD84370E48F954030FCDD" box="[189,1130,718,792]" pageId="18" pageNumber="228" type="distribution">
<paragraph id="8BCCC855B72AD84370E48F95450CFD2D" blockId="18.[189,342,718,744]" box="[189,342,718,744]" pageId="18" pageNumber="228">
<heading id="D0847F39B72AD84370E48F95450CFD2D" bold="true" box="[189,342,718,744]" fontSize="11" level="3" pageId="18" pageNumber="228" reason="3">
<emphasis id="B9071447B72AD84370E48F95450CFD2D" bold="true" box="[189,342,718,744]" pageId="18" pageNumber="228">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BCCC855B72AD84370E48FA54030FCDD" blockId="18.[189,1130,766,792]" box="[189,1130,766,792]" pageId="18" pageNumber="228">
Known only from the
<typeStatus id="54C876F7B72AD843719B8FA545AFFCDD" box="[450,501,766,792]" pageId="18" pageNumber="228">type</typeStatus>
locality in the
<collectingRegion id="49B706B7B72AD84372FD8FA547DCFCDD" box="[676,902,766,792]" country="Lebanon" name="Liban-Nord" pageId="18" pageNumber="228">North Governorate</collectingRegion>
, northern
<collectingCountry id="F36488C5B72AD84373A58FA5403DFCDD" box="[1020,1127,766,792]" name="Lebanon" pageId="18" pageNumber="228">Lebanon</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3699BDEB72AD84370E48E1B412DFC30" pageId="18" pageNumber="228" type="discussion">
<paragraph id="8BCCC855B72AD84370E48E1B4575FC9F" blockId="18.[189,303,832,858]" box="[189,303,832,858]" pageId="18" pageNumber="228">
<heading id="D0847F39B72AD84370E48E1B4575FC9F" bold="true" box="[189,303,832,858]" fontSize="11" level="3" pageId="18" pageNumber="228" reason="3">
<emphasis id="B9071447B72AD84370E48E1B4575FC9F" bold="true" box="[189,303,832,858]" pageId="18" pageNumber="228">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BCCC855B72AD84370E48E34412DFC30" blockId="18.[188,1400,879,1013]" pageId="18" pageNumber="228">
The complete absence of a posterior apophysis is a unique feature of this species. Because of the overall resemblance of the male palp to that of
<taxonomicName id="4C73B3D6B72AD84372C58ECF475AFC68" baseAuthorityName="Caporiacco" baseAuthorityYear="1929" box="[668,768,915,941]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="228" phylum="Arthropoda" rank="species" species="festai">
<emphasis id="B9071447B72AD84372C58ECF46EAFC68" box="[668,688,916,941]" italics="true" pageId="18" pageNumber="228">D</emphasis>
.
<emphasis id="B9071447B72AD84372998EC8475AFC68" box="[704,768,915,941]" italics="true" pageId="18" pageNumber="228">festai</emphasis>
</taxonomicName>
, it most likely belongs to the
<taxonomicName id="4C73B3D6B72AD84374308EC840F3FC68" baseAuthorityName="Caporiacco" baseAuthorityYear="1929" box="[1129,1193,915,941]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="228" phylum="Arthropoda" rank="species" species="festai">
<emphasis id="B9071447B72AD84374308EC840F3FC68" box="[1129,1193,915,941]" italics="true" pageId="18" pageNumber="228">festai</emphasis>
</taxonomicName>
group. However, it differs from the other two species in this group by the elongated chelicerae. To determine its correct position in the phylogeny of
<taxonomicName id="4C73B3D6B72AD843725C8E814632FC31" baseAuthorityName="sensu Deeleman-Reinhold &amp; Deeleman" baseAuthorityYear="1988" box="[517,616,986,1012]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="228" phylum="Arthropoda" rank="genus">
<emphasis id="B9071447B72AD843725C8E814632FC31" box="[517,616,986,1012]" italics="true" pageId="18" pageNumber="228">Dysdera</emphasis>
</taxonomicName>
, it is necessary to collect fresh material and describe the female too.
</paragraph>
</subSubSection>
</treatment>
</document>
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