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<document ID-DOI="10.11646/zootaxa.5254.3.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="7727597" ID-ZooBank="49C30D0E-C49A-4A03-9ACC-2BB9E2E0DA43" checkinTime="1678696635584" checkinUser="plazi" docAuthor="Zumbado Echavarria, Marco A., Barrantes Barrantes, Edwin A., Helmick, Ericka E., Bartlett, Charles R. &amp; Bahder, Brian W." docDate="2023" docId="03D187F0874CB028FF26BF30DEFFB65F" docLanguage="en" docName="zootaxa.5254.3.3.pdf" docOrigin="Zootaxa 5254 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5254.3.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Herpis Stal 1862" docType="treatment" docVersion="3" lastPageNumber="360" masterDocId="FFE8FF88874EB02BFFB1B802DF73B54A" masterDocTitle="A new species of planthopper from Costa Rica in the genus Herpis (Hemiptera: Derbidae) from palms in the cloud forest" masterLastPageNumber="369" masterPageNumber="357" pageNumber="359" updateTime="1678790628403" updateUser="tatiana" zenodo-license-document="CLOSED">
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<mods:title>A new species of planthopper from Costa Rica in the genus Herpis (Hemiptera: Derbidae) from palms in the cloud forest</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Zumbado Echavarria, Marco A.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-2591-7662</mods:nameIdentifier>
<mods:affiliation>Universidad de Costa Rica-Sede San Ramón, Departamento de Ciencias Naturales, de la Iglesia el Tremedal 400 mts al Oeste carretera hacia San Pedro, San Ramón, Alajuela, Costa Rica. nassua 75 @ gmail. com; https: // orcid. org / 0000 - 0002 - 2591 - 7662 Universidad de Costa Rica-Sede San Ramón, Departamento de Ciencias Naturales, de la Iglesia el Tremedal 400 mts al Oeste carretera hacia San Pedro, San Ramón, Alajuela, Costa Rica. edwbarrantes @ gmail. com; https: // orcid. org / 0000 - 0001 - 9565 - 2105</mods:affiliation>
<mods:nameIdentifier type="email">nassua75@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Barrantes Barrantes, Edwin A.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-9565-2105</mods:nameIdentifier>
<mods:nameIdentifier type="email">edwbarrantes@gmail.com</mods:nameIdentifier>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Helmick, Ericka E.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-5153-0891</mods:nameIdentifier>
<mods:affiliation>University of Florida, Department of Entomology and Nematology-Fort Lauderdale Research and Education Center; 3205 College Ave., Davie, FL 33314 - 7719, USA. ehelmick @ ufl. edu; https: // orcid. org / 0000 - 0002 - 5153 - 0891</mods:affiliation>
<mods:nameIdentifier type="email">ehelmick@ufl.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bartlett, Charles R.</mods:namePart>
<mods:affiliation>University of Delaware, Department of Entomology and Wildlife Ecology, 250 Townsend Hall, Newark, DE 19716 - 2160, USA.</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Bahder, Brian W.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-1118-4832</mods:nameIdentifier>
<mods:affiliation>University of Florida, Department of Entomology and Nematology-Fort Lauderdale Research and Education Center; 3205 College Ave., Davie, FL 33314 - 7719, USA. bbahder @ ufl. edu; https: // orcid. org / 0000 - 0002 - 1118 - 4832</mods:affiliation>
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<mods:title>Zootaxa</mods:title>
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<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-03-13</mods:number>
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<mods:detail type="volume">
<mods:number>5254</mods:number>
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<mods:detail type="issue">
<mods:number>3</mods:number>
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<mods:identifier type="DOI">10.11646/zootaxa.5254.3.3</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7727597</mods:identifier>
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<subSubSection box="[151,442,1841,1868]" pageId="2" pageNumber="359" type="nomenclature">
<paragraph blockId="2.[151,442,1841,1868]" box="[151,442,1841,1868]" pageId="2" pageNumber="359">
<heading bold="true" box="[151,442,1841,1868]" fontSize="11" level="1" pageId="2" pageNumber="359" reason="1">
<emphasis bold="true" box="[151,442,1841,1868]" pageId="2" pageNumber="359">
Genus
<taxonomicName ID-CoL="4VXD" ID-ENA="2710943" authority="Stal 1862" authorityName="Stal" authorityYear="1862" box="[238,442,1841,1868]" class="Insecta" family="Derbidae" genus="Herpis" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="359" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[238,319,1842,1868]" italics="true" pageId="2" pageNumber="359">Herpis</emphasis>
<bibRefCitation author="Stal, C." box="[327,442,1841,1868]" pageId="2" pageNumber="359" pagination="1 - 75" refId="ref7115" refString="Stal, C. (1862) Bidrag till Rio de Janeiro-tratkens Hemipterfauna. II. Handlingar. Kongliga Svenska Vetenskaps-Akademien, 3 (6), 1 - 75." type="journal article" year="1862">Stål 1862</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,1355,1911,1936]" pageId="2" pageNumber="359" type="materials_examined">
<paragraph blockId="2.[151,1437,1911,2033]" box="[151,1355,1911,1936]" pageId="2" pageNumber="359">
<typeStatus box="[151,201,1912,1936]" pageId="2" pageNumber="359">Type</typeStatus>
species
<taxonomicName authority="Stal, 1862" authorityName="Stal" authorityYear="1862" box="[287,592,1911,1935]" class="Insecta" family="Derbidae" genus="Herpis" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="359" phylum="Arthropoda" rank="species" species="fuscovittata">
<emphasis box="[287,481,1911,1935]" italics="true" pageId="2" pageNumber="359">Herpis fuscovittata</emphasis>
<bibRefCitation author="Stal, C." box="[487,592,1911,1935]" pageId="2" pageNumber="359" pagination="1 - 75" refId="ref7115" refString="Stal, C. (1862) Bidrag till Rio de Janeiro-tratkens Hemipterfauna. II. Handlingar. Kongliga Svenska Vetenskaps-Akademien, 3 (6), 1 - 75." type="journal article" year="1862">Stål, 1862</bibRefCitation>
</taxonomicName>
, by subsequent designation of
<bibRefCitation author="Muir F. A. G." box="[904,1050,1911,1935]" pageId="2" pageNumber="359" pagination="15 - 19" refId="ref6903" refString="Muir F. A. G. (1924) Notes on some genera of Derbidae (Hemip.). Proceedings of the Entomological Society of Washington, 26, 15 - 19." type="journal article" year="1924">Muir 1924: 16</bibRefCitation>
(see also
<bibRefCitation author="McAtee, W. L." box="[1151,1342,1911,1935]" pageId="2" pageNumber="359" pagination="175 - 186" refId="ref6647" refString="McAtee, W. L. (1924) Notes on Cenchrea Westwood and Cedusa Fowler in America (Homoptera: Fulgoroidea). Annals of the Entomological Society of America, 17, 175 - 186. https: // doi. org / 10.1093 / aesa / 17.2.175" type="journal article" year="1924">McAtee 1924: 175</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection box="[199,439,1942,1966]" pageId="2" pageNumber="359" type="reference_group">
<paragraph blockId="2.[151,1437,1911,2033]" box="[199,439,1942,1966]" pageId="2" pageNumber="359">
<treatmentCitationGroup box="[199,439,1942,1966]" pageId="2" pageNumber="359">
<taxonomicName authority="Fowler, 1905" authorityName="Fowler" authorityYear="1905" box="[199,439,1942,1966]" class="Insecta" family="Derbidae" genus="Syntames" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="359" phylum="Arthropoda" rank="genus">
<emphasis box="[199,295,1942,1965]" italics="true" pageId="2" pageNumber="359">Syntames</emphasis>
<bibRefCitation author="Fowler, W. W." box="[303,439,1942,1966]" pageId="2" pageNumber="359" pagination="125 - 139" refId="ref6565" refString="Fowler, W. W. (1905) Order Rhynchota. Suborder Hemiptera-Homoptera. (Continued). Biologia Centrali-Americana, 1, 125 - 139." type="journal article" year="1905">Fowler, 1905</bibRefCitation>
</taxonomicName>
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</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="359" type="multiple">
<paragraph blockId="2.[151,1437,1911,2033]" pageId="2" pageNumber="359">
(
<typeStatus box="[453,505,1942,1966]" pageId="2" pageNumber="359">Type</typeStatus>
species
<taxonomicName authority="Fowler 1905" authorityName="Fowler" authorityYear="1905" box="[594,930,1942,1966]" class="Insecta" family="Derbidae" genus="Syntames" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="359" phylum="Arthropoda" rank="species" species="delicatus">
<emphasis box="[594,789,1942,1966]" italics="true" pageId="2" pageNumber="359">Syntames delicatus</emphasis>
<bibRefCitation author="Fowler, W. W." box="[798,930,1942,1966]" pageId="2" pageNumber="359" pagination="125 - 139" refId="ref6565" refString="Fowler, W. W. (1905) Order Rhynchota. Suborder Hemiptera-Homoptera. (Continued). Biologia Centrali-Americana, 1, 125 - 139." type="journal article" year="1905">Fowler 1905</bibRefCitation>
</taxonomicName>
, by monotypy). Synonym with
<taxonomicName ID-CoL="4VXD" ID-ENA="2710943" authority="Stal 1862" authorityName="Stal" authorityYear="1862" box="[1259,1328,1942,1965]" class="Insecta" family="Derbidae" genus="Herpis" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="359" phylum="Arthropoda" rank="genus">
<emphasis box="[1259,1328,1942,1965]" italics="true" pageId="2" pageNumber="359">Herpis</emphasis>
</taxonomicName>
according to
<treatmentCitation author="Muir F. A. G." box="[180,334,1972,1997]" page="16" pageId="2" pageNumber="359" year="1924">
<bibRefCitation author="Muir F. A. G." box="[180,334,1972,1997]" pageId="2" pageNumber="359" pagination="15 - 19" refId="ref6903" refString="Muir F. A. G. (1924) Notes on some genera of Derbidae (Hemip.). Proceedings of the Entomological Society of Washington, 26, 15 - 19." type="journal article" year="1924">Muir 1924: 16</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Caldwell, J. S." box="[348,543,1972,1997]" page="99" pageId="2" pageNumber="359" year="1944">
<bibRefCitation author="Caldwell, J. S." box="[348,543,1972,1997]" pageId="2" pageNumber="359" pagination="99 - 110" refId="ref6469" refString="Caldwell, J. S. (1944) The tribe Cenchreini with special references to the Cenchrea complex (Homoptera: Derbidae). Bulletin of the Brooklyn Entomological Society, 39, 99 - 110." type="journal article" year="1944">Caldwell 1944: 99</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="O'Brien, L. B." box="[557,756,1972,1997]" page="383" pageId="2" pageNumber="359" year="1987">
<bibRefCitation author="O'Brien, L. B." box="[557,756,1972,1997]" pageId="2" pageNumber="359" pagination="379 - 390" refId="ref6938" refString="O'Brien, L. B. (1987) Corrections and additions to Metcalf's &quot; The Fulgorina of Barro Colorado and other parts of Panama &quot; (Homoptera: Fulgoroidea). Annals of the Entomological Society of America, 80 (3), 379 - 390. https: // doi. org / 10.1093 / aesa / 80.3.379" type="journal article" year="1987">OBrien 1987: 383</bibRefCitation>
</treatmentCitation>
; treated as valid by
<treatmentCitation author="Metcalf, Z. P." box="[972,1167,1972,1997]" page="324" pageId="2" pageNumber="359" year="1938">
<bibRefCitation author="Metcalf, Z. P." box="[972,1167,1972,1997]" pageId="2" pageNumber="359" pagination="277 - 423" refId="ref6697" refString="Metcalf, Z. P. (1938) The Fulgorina of Barro Colorado and other parts of Panama. Bulletin of the Museum of Comparative Zoology at Harvard College, 83 (5), 277 - 423." type="journal article" year="1938">Metcalf 1938: 324</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Metcalf, Z. P." box="[1180,1299,1972,1997]" page="101" pageId="2" pageNumber="359" year="1945">
<bibRefCitation author="Metcalf, Z. P." box="[1180,1299,1972,1997]" pageId="2" pageNumber="359" pagination="1 - 212" refId="ref6737" refString="Metcalf, Z. P. (1945 a) General Catalogue of the Hemiptera. Fascicle IV, Fulgoroidea. Part 4 Derbidae. Smith College, Northhampton, Massachusetts. pp. 1 - 212." type="book chapter" year="1945">1945a: 101</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Metcalf, Z. P." box="[1312,1433,1972,1996]" page="125" pageId="2" pageNumber="359" year="1945">
<bibRefCitation author="Metcalf, Z. P." box="[1312,1433,1972,1996]" pageId="2" pageNumber="359" pagination="125 - 143" refId="ref6775" refString="Metcalf, Z. P. (1945 b) Fulgoroidea (Homoptera) of Kartabo, Bartica District, British Guiana. Zoologica [Scientific Contributions of the New York Zoological Society], 30 (3), 125 - 143. https: // doi. org / 10.5962 / p. 210851" type="journal article" year="1945">1945b: 125</bibRefCitation>
</treatmentCitation>
. Synonymy of
<taxonomicName authorityName="Fowler" authorityYear="1905" box="[295,391,2009,2032]" class="Insecta" family="Derbidae" genus="Syntames" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="359" phylum="Arthropoda" rank="genus">
<emphasis box="[295,391,2009,2032]" italics="true" pageId="2" pageNumber="359">Syntames</emphasis>
</taxonomicName>
with
<taxonomicName ID-CoL="4VXD" ID-ENA="2710943" authority="Stal 1862" authorityName="Stal" authorityYear="1862" box="[448,517,2009,2032]" class="Insecta" family="Derbidae" genus="Herpis" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="359" phylum="Arthropoda" rank="genus">
<emphasis box="[448,517,2009,2032]" italics="true" pageId="2" pageNumber="359">Herpis</emphasis>
</taxonomicName>
confirmed in
<bibRefCitation author="Bahder, B. W. &amp; Zumbado Echavarria, M. A. &amp; Barrantes E. A. B. &amp; Helmick, E. E. &amp; Bartlett, C. R." box="[660,879,2008,2033]" pageId="2" pageNumber="359" pagination="121 - 135" refId="ref5979" refString="Bahder, B. W., Zumbado Echavarria, M. A., Barrantes E. A. B., Helmick, E. E. &amp; Bartlett, C. R. (2021 a) A new species of planthopper in genus Herpis (Hemiptera: Derbidae) from lowland tropical rainforest in Costa Rica. Zootaxa, 5032 (1), 121 - 135. https: // doi. org / 10.11646 / zootaxa. 4908.3.3" type="journal article" year="2021">
Bahder
<emphasis box="[739,791,2008,2032]" italics="true" pageId="2" pageNumber="359">et al.</emphasis>
(2021a)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="360" type="diagnosis">
<paragraph blockId="3.[151,1437,150,789]" pageId="3" pageNumber="360">
<emphasis bold="true" box="[151,392,150,176]" pageId="3" pageNumber="360">Amended Diagnosis</emphasis>
(Modified from
<bibRefCitation author="Bahder, B. W. &amp; Zumbado Echavarria, M. A. &amp; Barrantes E. A. B. &amp; Helmick, E. E. &amp; Bartlett, C. R." box="[591,818,151,177]" pageId="3" pageNumber="360" pagination="121 - 135" refId="ref5979" refString="Bahder, B. W., Zumbado Echavarria, M. A., Barrantes E. A. B., Helmick, E. E. &amp; Bartlett, C. R. (2021 a) A new species of planthopper in genus Herpis (Hemiptera: Derbidae) from lowland tropical rainforest in Costa Rica. Zootaxa, 5032 (1), 121 - 135. https: // doi. org / 10.11646 / zootaxa. 4908.3.3" type="journal article" year="2021">
Bahder
<emphasis box="[682,735,151,177]" italics="true" pageId="3" pageNumber="360">et al</emphasis>
2021a
</bibRefCitation>
). Relatively large, and robust cenchreines, measuring approximately
<quantity box="[321,410,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="3" pageNumber="360" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">5-8 mm</quantity>
in length, including wings. Wings held tectiform in repose, distinctly exceeding abdomen. Vertex broad, trapezoidal, just broader than long, nearly flat to weakly concave, transverse carinae present at fastigium. Pits present along lateral carinae of vertex and frons, varying in distinctness, appearing to be in 2+ rows. Frons (frontal view) broad, flat or nearly so (not concave), lateral margins parallel, medium carina of frons evident, frontoclypeal margin straight. Clypeus elongate-triangular, approximately in the same plane as frons, median carina present. Genae without subantennal processes. Antennae short, pedicle spheroid, about as long as wide. Lateral ocelli obscure or absent.
</paragraph>
<paragraph blockId="3.[151,1437,150,789]" pageId="3" pageNumber="360">Pronotum narrow, anterior convex, posterior concave, broadening laterally, in lateral view declinate anteriorly; paranotal region foliate, forming large foveae posterior to the antennae (a tribal feature). Mesonotum tricarinate, carinae approximately reaching posterior margin, scutellum contiguous with scutum (i.e., not separated by groove). Lateral teeth absent on tibiae.</paragraph>
<paragraph blockId="3.[151,1437,150,789]" pageId="3" pageNumber="360">Terminalia bilaterally symmetrical. Pygofer in lateral view narrow, irregular in outline; in ventral view, median process of pygofer present (longer than wide, apically conical). Gonostyli elongate, broad, bearing dorsal projection in proximal half, and (in ventral view) usually with a medial (ental) process near midlength. Aedeagal shaft with processes apically and near midlength; flagellum complex, with elongate retrorse processes. Anal tube elongate in lateral view (but usually shorter than gonostyli), distally expanded; in dorsal view appearing spatulate with caudal margin strongly bifid.</paragraph>
</subSubSection>
</treatment>
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