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<document id="29285505DFD5E43B52DF372CFB1E8417" ID-ISSN="0024-4082" ID-ZooBank="C1BDE9AB-5360-48B1-8689-E16F497A417A" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630096132686" checkinUser="felipe" docAuthor="Kolibáč, Jiří, Bocakova, Milada, Liebherr, James K., Ramage, Thiboult &amp; Porch, Nick" docDate="2021" docId="F047DF66FFCEFFFE8486F95FFAC5F97A" docLanguage="en" docName="zlaa064.pdf" docOrigin="Zoological Journal of the Linnean Society 191" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Tenebroides (Parallelodera)" docType="treatment" docVersion="2" masterDocId="0C7EA71EFFC0FFF08463FFE4FFBDFFCF" masterDocTitle="Extinct and extant Pacific Trogossitidae and the evolution of Cleroidea (Coleoptera) after the Late Triassic biotic crisis" updateTime="1699143628773" updateUser="plazi">
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<mods:title id="E3CC4138DE82ED29C148A762523CADF5">Extinct and extant Pacific Trogossitidae and the evolution of Cleroidea (Coleoptera) after the Late Triassic biotic crisis</mods:title>
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<mods:namePart id="B7734A73476CB32F0B0D32A603AB23DE">Kolibáč, Jiří</mods:namePart>
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<mods:namePart id="3AAAFE83408CE83EFD915C3F6FA3502A">Bocakova, Milada</mods:namePart>
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<mods:namePart id="1688A5278444829DE70EBFB15FD4606F">Liebherr, James K.</mods:namePart>
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<mods:namePart id="B4364567D24350CBCF8E48036189C8E3">Ramage, Thiboult</mods:namePart>
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<mods:title id="1A44CCDF96F0960926B68F447F162950">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="D25F6757D00CDB9DE2DB24FA84D7D9D1">2021</mods:date>
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<treatment id="F047DF66FFCEFFFE8486F95FFAC5F97A" LSID="urn:lsid:plazi:treatment:F047DF66FFCEFFFE8486F95FFAC5F97A" httpUri="http://treatment.plazi.org/id/F047DF66FFCEFFFE8486F95FFAC5F97A" pageId="14">
<subSubSection id="30F43DFBFFCEFFFE8486F95FFD75F91C" box="[229,712,1722,1747]" pageId="14" type="nomenclature">
<paragraph id="78516E70FFCEFFFE8486F95FFD75F91C" blockId="14.[229,712,1722,1747]" box="[229,712,1722,1747]" pageId="14">
<heading id="2319D91CFFCEFFFE8486F95FFD75F91C" box="[229,712,1722,1747]" centered="true" fontSize="9" level="2" pageId="14" reason="2">
<taxonomicName id="BFEE15F3FFCEFFFE8486F95FFDE1F91C" authority="" box="[229,604,1722,1747]" class="Insecta" family="Trogossitidae" genus="Tenebroides" kingdom="Animalia" order="Coleoptera" pageId="14" phylum="Arthropoda" rank="subGenus" subGenus="Parallelodera">
<emphasis id="4A9AB262FFCEFFFE8486F95FFE38F91E" box="[229,389,1723,1746]" italics="true" pageId="14">TENEBROIDES</emphasis>
(
<emphasis id="4A9AB262FFCEFFFE85F6F95FFDE9F91E" box="[405,596,1723,1746]" italics="true" pageId="14">PARALLELODERA</emphasis>
)
</taxonomicName>
SP. G038
</heading>
</paragraph>
</subSubSection>
<subSubSection id="30F43DFBFFCEFFFE84A6F90CFAC5F97A" pageId="14" type="description">
<paragraph id="78516E70FFCEFFFE84A6F90CFD55F8CE" blockId="14.[197,744,1768,1793]" box="[197,744,1768,1793]" pageId="14">
(
<figureCitation id="E0D572F5FFCEFFFE84AEF90CFE62F8CF" box="[205,479,1768,1793]" captionStart="Fig" captionStartId="16.[163,204,887,909]" captionTargetBox="[164,1436,213,846]" captionTargetId="figure-487@16.[163,1446,195,847]" captionTargetPageId="16" captionText="Fig. 7. Tenebroides sp. G038 (Panama): A, ovipositor ventrally; B, detail of coxitae and styli; C, apex of mesotibia and mesotarsus. D, Tenebroides (Parallelodera) quadraticollis, mesotarsus. E, Tenebroides (Tenebroides) mauritanicus, mesotarsus." pageId="14">FIG. 7AC; SUPPORTING</figureCitation>
INFORMATION,
<figureCitation id="E0D572F5FFCEFFFE86EDF90CFD5DF8CE" box="[654,736,1768,1793]" captionStart="Fig" captionStartId="13.[145,185,1734,1756]" captionTargetBox="[274,1290,198,1692]" captionTargetId="figure-73@13.[271,1298,195,1694]" captionTargetPageId="13" captionText="Fig. 5. A, Tenebroides luteicornis. BH, T. fairmairei: B, habitus; C, protibia and tarsus; D, middle leg; E, hind leg;F, wing; G, male abdominal segment IX; H, tegmen dorsally; I, penis. J, T. quadraticollis." pageId="14">FIG. S5</figureCitation>
)
</paragraph>
<paragraph id="78516E70FFCEFFFE84C0F8C3FCB6F8B6" blockId="14.[163,779,1830,1914]" pageId="14">
<emphasis id="4A9AB262FFCEFFFE84C0F8C3FE07F8F3" box="[163,442,1830,1852]" italics="true" pageId="14">M a t e r i a l e x a m i n e d:</emphasis>
<materialsCitation id="C886642DFFCEFFFE85AFF8C2FD56F8B5" collectingDate="2009-06-06" country="Panama" elevation="75" latitude="8.218467" location="Santiago" longLatPrecision="1" longitude="-80.96412" pageId="14" specimenCount="1" stateProvince="Veraguas">
O n e s p e c i m e n (G 0 3 8):
<collectingCountry id="00F92EE0FFCEFFFE84C4F8A1FEAEF895" box="[167,275,1861,1882]" name="Panama" pageId="14">Panama</collectingCountry>
:
<collectingRegion id="BA2AA092FFCEFFFE8540F8A1FE5BF895" box="[291,486,1861,1882]" country="Panama" name="Veraguas" pageId="14">Veraguas Prov.</collectingRegion>
/
<locationDeviation id="69E37752FFCEFFFE858CF8A1FD60F894" box="[495,733,1861,1883]" location="El Espino" pageId="14">
<quantity id="BF16C395FFCEFFFE858CF8A1FD8DF894" box="[495,560,1861,1883]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="14" unit="km" value="2.0">2 km</quantity>
N El Espino
</locationDeviation>
nr.
<location id="7D3138ABFFCEFFFE84C0F880FEB4F8B5" LSID="urn:lsid:plazi:treatment:F047DF66FFCEFFFE8486F95FFAC5F97A:7D3138ABFFCEFFFE84C0F880FEB4F8B5" box="[163,265,1892,1914]" country="Panama" latitude="8.218467" longLatPrecision="1" longitude="-80.96412" name="Santiago" pageId="14" stateProvince="Veraguas">Santiago</location>
/
<geoCoordinate id="1DDA08B7FFCEFFFE8571F880FE37F8B5" box="[274,394,1892,1914]" degrees="8" direction="north" minutes="13.108" orientation="latitude" pageId="14" precision="1" value="8.218467">8°13.108N</geoCoordinate>
<geoCoordinate id="1DDA08B7FFCEFFFE85F0F887FDA0F8B6" box="[403,541,1891,1913]" degrees="80" direction="west" minutes="57.847" orientation="longitude" pageId="14" precision="1" value="-80.96412">80°57.847W</geoCoordinate>
,
<quantity id="BF16C395FFCEFFFE864AF887FDD9F8B5" box="[553,612,1891,1914]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="14" unit="m" value="75.0">
<elevation id="F3C38943FFCEFFFE864AF887FDD9F8B5" box="[553,612,1891,1914]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="14" unit="m" value="75.0">75 m</elevation>
</quantity>
/
<date id="0C5048B0FFCEFFFE860FF880FD56F8B5" box="[620,747,1892,1914]" pageId="14" value="2009-06-06">
<collectingDate id="1C14B158FFCEFFFE860FF880FD56F8B5" box="[620,747,1892,1914]" pageId="14" value="2009-06-06">06Jun2009</collectingDate>
</date>
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N.
</paragraph>
<paragraph id="78516E70FFCEFFFE8758FCE8FAC2FC8E" blockId="14.[827,1443,780,833]" pageId="14">
Lord, K. Miller/E. Nearns, peeling bark (UNMM). Two specimens
<collectingCountry id="00F92EE0FFCEFFFE87D9FCCFFB95FC8E" box="[954,1064,811,833]" name="Colombia" pageId="14">Colombia</collectingCountry>
, Mamatoco, 1912, I. (MMBC).
</paragraph>
<paragraph id="78516E70FFCEFFFE8758FC8EFA91FAC1" blockId="14.[827,1444,874,1294]" pageId="14">
<emphasis id="4A9AB262FFCEFFFE8758FC8EFC06FCB0" box="[827,955,874,895]" italics="true" pageId="14">Diagnosis:</emphasis>
Black species with fine punctures in head and pronotum, without pubescence. Frons with medial longitudinal depression and distinct furrow in centre reaching to anterior margin of head. Ctenidium absent. Antenna 11-segmented, club three-segmented, asymmetric. Eye flat, not exceeding contour of head; finely facetted. Pronotum subparallel-sided, with small posterior angles; elytron with denticle at humerus; elytral rim meets fifth stria from suture; elytron with fine transverse wrinkles. Tarsal formula 4-4-4. Tibiae with two spines at apex, one of them large, hooked in protibia; all tibiae with three to five denticles along the outer margin. Elytron subparallel, weakly widened backwards, widest at apical third; wings developed. Body size:
<quantity id="BF16C395FFCEFFFE80C4FB1CFA95FAC1" box="[1191,1320,1272,1294]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="8.5" metricValueMin="7.5" pageId="14" unit="mm" value="8.0" valueMax="8.5" valueMin="7.5">7.58.5 mm</quantity>
.
</paragraph>
<paragraph id="78516E70FFCEFFFE8758FAD3FB0BFA82" blockId="14.[827,1206,1335,1357]" box="[827,1206,1335,1357]" pageId="14">
<emphasis id="4A9AB262FFCEFFFE8758FAD3FC71FA83" box="[827,972,1335,1356]" italics="true" pageId="14">Distribution:</emphasis>
<collectingCountry id="00F92EE0FFCEFFFE87BAFAD3FBFBFA82" box="[985,1094,1335,1357]" name="Colombia" pageId="14">Colombia</collectingCountry>
,
<collectingCountry id="00F92EE0FFCEFFFE8032FADCFB0FFA82" box="[1105,1202,1336,1357]" name="Panama" pageId="14">Panama</collectingCountry>
.
</paragraph>
<paragraph id="78516E70FFCEFFFE8758FA92FC62F9AD" blockId="14.[827,1443,1398,1634]" pageId="14">
<emphasis id="4A9AB262FFCEFFFE8758FA92FC15FA44" box="[827,936,1398,1419]" italics="true" pageId="14">Remarks:</emphasis>
Two specimens from
<collectingCountry id="00F92EE0FFCEFFFE80C9FA92FAAAFA43" box="[1194,1303,1398,1420]" name="Colombia" pageId="14">Colombia</collectingCountry>
agree in all details with the voucher specimen G038 from
<collectingCountry id="00F92EE0FFCEFFFE815CFA71FA23FA65" box="[1343,1438,1429,1450]" name="Panama" pageId="14">Panama</collectingCountry>
, which was used in the analysis by
<bibRefCitation id="1C7F1381FFCEFFFE80DDFA57FA1FFA06" author="Gimmel ML &amp; Bocakova M &amp; Gunter N &amp; Leschen R" box="[1214,1442,1459,1481]" pageId="14" pagination="527 - 558" refId="ref20958" refString="Gimmel ML, Bocakova M, Gunter N, Leschen R. 2019. Comprehensive phylogeny of the Cleroidea (Coleoptera: Cucujiformia). Systematic Entomology 44: 527 - 558." type="journal article" year="2019">
Gimmel
<emphasis id="4A9AB262FFCEFFFE817CFA50FAEBFA07" box="[1311,1366,1459,1481]" italics="true" pageId="14">et al.</emphasis>
(2019)
</bibRefCitation>
and in the present analysis. The species is denoted as
<taxonomicName id="BFEE15F3FFCEFFFE8723FA14FC4EF9C9" box="[832,1011,1520,1542]" class="Insecta" family="Trogossitidae" genus="Tenebroides" kingdom="Animalia" order="Coleoptera" pageId="14" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="4A9AB262FFCEFFFE8723FA14FC76F9CA" box="[832,971,1520,1541]" italics="true" pageId="14">Tenebroides</emphasis>
sp.
</taxonomicName>
Panama in tree diagrams. A formal description of the species is impossible without a systematic revision of
<taxonomicName id="BFEE15F3FFCEFFFE805CF9CAFB76F98B" authorityName="Piller &amp; Mitterpacher" authorityYear="1783" box="[1087,1227,1582,1604]" class="Insecta" family="Trogossitidae" genus="Tenebroides" kingdom="Animalia" order="Coleoptera" pageId="14" phylum="Arthropoda" rank="genus">Tenebroides</taxonomicName>
south of
<collectingCountry id="00F92EE0FFCEFFFE815BF9CAFAD3F98C" box="[1336,1390,1582,1603]" name="United States of America" pageId="14">USA</collectingCountry>
(see
<bibRefCitation id="1C7F1381FFCEFFFE8758F9A9FC72F9AE" author="Barron JR" box="[827,975,1612,1634]" pageId="14" pagination="1 - 143" refId="ref20295" refString="Barron JR. 1971. A revision of the Trogositidae of America North of Mexico (Coleoptera: Cleroidea). Memoirs of the Entomological Society of Canada 75: 1 - 143." type="journal article" year="1971">Barron, 1971</bibRefCitation>
).
</paragraph>
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<emphasis id="4A9AB262FFCEFFFE8706F97BFB2EF97A" bold="true" box="[869,1171,1695,1718]" pageId="14">
<taxonomicName id="BFEE15F3FFCEFFFE8706F97BFBB3F97A" authorityName="Piller &amp; Mitterpacher" authorityYear="1783" box="[869,1038,1695,1718]" class="Insecta" family="Trogossitidae" genus="Tenebroides" kingdom="Animalia" order="Coleoptera" pageId="14" phylum="Arthropoda" rank="genus">
<emphasis id="4A9AB262FFCEFFFE8706F97BFBB3F97A" bold="true" box="[869,1038,1695,1718]" italics="true" pageId="14">TENEBROIDES</emphasis>
</taxonomicName>
SUBGENUS
</emphasis>
<taxonomicName id="BFEE15F3FFCEFFFE80F9F97BFAC5F97A" authorityName="Kolibáč &amp; Bocakova &amp; Liebherr &amp; Ramage &amp; Porch" authorityYear="2021" box="[1178,1400,1695,1718]" class="Insecta" family="Trogossitidae" genus="Tenebroides" kingdom="Animalia" order="Coleoptera" pageId="14" phylum="Arthropoda" rank="subGenus" subGenus="Polynesibroides">
<emphasis id="4A9AB262FFCEFFFE80F9F97BFAC5F97A" bold="true" box="[1178,1400,1695,1718]" italics="true" pageId="14">POLYNESIBROIDES</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
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