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<document id="AA5877BB08D202110EE856F1D8654644" ID-DOI="10.5852/ejt.2020.621" ID-GBIF-Dataset="d13c2f0d-7f44-4b5f-aa68-6b4d69c05d2e" ID-Zenodo-Dep="3750251" ID-ZooBank="CF9060EF-84DA-4195-81AE-25B06B69D0DF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1586809403066" checkinUser="valdenar" docAuthor="Ranasinghe, Sasanka, Eberle, Jonas, Benjamin, Suresh P. &amp; Ahrens, Dirk" docDate="2020" docId="C46987D93F44FF9B9D2C03D3FE32FEE3" docLanguage="en" docName="ejt-621_ranasinghe_eberle_benjamin.pdf.imf" docOrigin="European Journal of Taxonomy 621" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maladera cervicornis Ranasinghe &amp; Eberle &amp; Benjamin &amp; Ahrens 2020, sp. nov." docType="treatment" docUuid="8B6DAFF6-F615-4D2E-A776-937A80C0924E" docUuidSource="ZooBank" docVersion="6" lastPageNumber="15" masterDocId="3850FFA13F4EFF959F44015BFFC7FFC6" masterDocTitle="New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae)" masterLastPageNumber="20" masterPageNumber="1" pageNumber="11" updateTime="1698737313009" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="CF79889DDA74E7E3008C602955CD19EB">New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae)</mods:title>
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<mods:namePart id="8D8218BBDF51BB559596C3A3769E61E1">Ranasinghe, Sasanka</mods:namePart>
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<mods:nameIdentifier id="5CA5D02CC39787324A76C88E06729044" type="email">lakmalisanky@gmail.com</mods:nameIdentifier>
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<mods:namePart id="29EAE4ECF274D8473AAEC95661DCE32B">Eberle, Jonas</mods:namePart>
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<mods:namePart id="822187708E9465B65DECF435723B6E27">Benjamin, Suresh P.</mods:namePart>
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<mods:nameIdentifier id="88CE7EE4656800781D49B333CA46F643" type="email">suresh.benjamin@gmail.com</mods:nameIdentifier>
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<mods:namePart id="697C46E8412574C08FA2772F79592E7E">Ahrens, Dirk</mods:namePart>
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<mods:nameIdentifier id="BE0AFD6574C7D9E4029A7C953A7DE430" type="email">ahrens.dirk_col@gmx.de;d.ahrens@leibniz-zfmk.de</mods:nameIdentifier>
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<mods:title id="07709EC128C62FF310D46211D9245F4E">European Journal of Taxonomy</mods:title>
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<treatment id="C46987D93F44FF9B9D2C03D3FE32FEE3" ID-DOI="http://doi.org/10.5281/zenodo.3808779" ID-GBIF-Taxon="163609934" ID-Zenodo-Dep="3808779" LSID="urn:lsid:zoobank.org:act:8B6DAFF6-F615-4D2E-A776-937A80C0924E" httpUri="http://treatment.plazi.org/id/C46987D93F44FF9B9D2C03D3FE32FEE3" lastPageId="14" lastPageNumber="15" pageId="10" pageNumber="11">
<subSubSection id="04DA65443F44FF9F9D2C03D3FB01FD00" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="4C7F36CF3F44FF9F9D2C03D3FC0BFD65" blockId="10.[366,1222,648,745]" box="[616,972,648,675]" pageId="10" pageNumber="11">
<heading id="173781A33F44FF9F9D2C03D3FC0BFD65" box="[616,972,648,675]" centered="true" fontSize="11" level="2" pageId="10" pageNumber="11" reason="2">
<taxonomicName id="8BC04D4C3F44FF9F9D2C03D3FCADFD64" authority="Ranasinghe &amp; Eberle &amp; Benjamin &amp; Ahrens, 2020" authorityName="Ranasinghe &amp; Eberle &amp; Benjamin &amp; Ahrens" authorityYear="2020" box="[616,874,648,675]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="cervicornis" status="sp. nov.">
<emphasis id="7EB4EADD3F44FF9F9D2C03D3FCADFD64" bold="true" box="[616,874,648,675]" italics="true" pageId="10" pageNumber="11">Maladera cervicornis</emphasis>
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<taxonomicNameLabel id="658757A63F44FF9F9C3503D2FC0BFD65" box="[881,972,649,675]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
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<uri id="38513ACD3F44FF9F9E2A03F0FB01FD00" box="[366,1222,683,710]" pageId="10" pageNumber="11">
urn:lsid:zoobank.org:act:
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</paragraph>
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<subSubSection id="04DA65443F44FF9F9DEC0395FC4BFD2E" box="[680,908,718,745]" pageId="10" pageNumber="11" type="description">
<paragraph id="4C7F36CF3F44FF9F9DEC0395FC4BFD2E" blockId="10.[366,1222,648,745]" box="[680,908,718,745]" pageId="10" pageNumber="11">
<figureCitation id="D4FB2A4A3F44FF9F9DEC0395FD36FD2F" box="[680,753,718,745]" captionStart="Fig" captionStartId="9.[189,232,1931,1957]" captionTargetBox="[209,1372,940,1880]" captionTargetId="figure@9.[193,1394,919,1894]" captionTargetPageId="9" captionText="Fig. 4. A–D. Maladera galdaththana sp. nov. (holotype). E–H. M. cervicornis sp. nov. (holotype). A, E. Aedeagus, left side lateral view. C, G. Aedeagus, right side lateral view. B, F. Parameres, dorsal view. D, H. Habitus (not to scale). Scales: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3750259" httpUri="https://zenodo.org/record/3750259/files/figure.png" pageId="10" pageNumber="11">Figs 4</figureCitation>
E–H, 5D, 6D
</paragraph>
</subSubSection>
<subSubSection id="04DA65443F44FF9F9FF9024BFE9AFC59" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="4C7F36CF3F44FF9F9FF9024BFEF1FCEC" blockId="10.[189,310,784,810]" box="[189,310,784,810]" pageId="10" pageNumber="11">
<heading id="173781A33F44FF9F9FF9024BFEF1FCEC" bold="true" box="[189,310,784,810]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="7EB4EADD3F44FF9F9FF9024BFEF1FCEC" bold="true" box="[189,310,784,810]" pageId="10" pageNumber="11">Diagnosis</emphasis>
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<taxonomicName id="8BC04D4C3F44FF9F9FF90265FE06FC9E" authority="Ranasinghe &amp; Eberle &amp; Benjamin &amp; Ahrens, 2020" authorityName="Ranasinghe &amp; Eberle &amp; Benjamin &amp; Ahrens" authorityYear="2020" box="[189,449,830,856]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="cervicornis" status="sp. nov.">
<emphasis id="7EB4EADD3F44FF9F9FF90265FE06FC9E" box="[189,449,830,856]" italics="true" pageId="10" pageNumber="11">Maladera cervicornis</emphasis>
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<taxonomicNameLabel id="658757A63F44FF9F9E8A0264FDE9FC9F" box="[462,558,831,857]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
is in external morphology very similar to
<taxonomicName id="8BC04D4C3F44FF9F9B020264FE98FCBA" authority="Fabrizi &amp; Ahrens, 2014" authorityName="Fabrizi &amp; Ahrens" authorityYear="2014" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="kandyensis">
<emphasis id="7EB4EADD3F44FF9F9B020264FB99FC9E" box="[1094,1118,831,856]" italics="true" pageId="10" pageNumber="11">M</emphasis>
.
<emphasis id="7EB4EADD3F44FF9F9B370265FB33FC9E" box="[1139,1268,830,856]" italics="true" pageId="10" pageNumber="11">kandyensis</emphasis>
<bibRefCitation id="28514B3E3F44FF9F9A440265FE98FCBA" author="Fabrizi S. &amp; Ahrens D." pageId="10" pageNumber="11" pagination="1 - 124" refId="ref9492" refString="Fabrizi S. &amp; Ahrens D. 2014. A Monograph of the Sericini of Sri Lanka (Coleoptera: Scarabaeidae). Bonn Zoological Bulletin, Supplements 61: 1 - 124." type="journal article" year="2014">Fabrizi &amp; Ahrens, 2014</bibRefCitation>
</taxonomicName>
. The new species differs by the tubercle on 3
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abdominal sternite as well as by the shape of parameres.
</paragraph>
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<subSubSection id="04DA65443F44FF9F9FF9029DFAB0FBF5" pageId="10" pageNumber="11" type="etymology">
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<heading id="173781A33F44FF9F9FF9029DFE83FC26" bold="true" box="[189,324,966,992]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="7EB4EADD3F44FF9F9FF9029DFE83FC26" bold="true" box="[189,324,966,992]" pageId="10" pageNumber="11">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="4C7F36CF3F44FF9F9FF902AEFAB0FBF5" blockId="10.[189,1399,1013,1075]" pageId="10" pageNumber="11">The name of the new species is derived from the combined Latin nouns cornu (horn) and cervus (deer), with reference to the shape of the parameres, resembling the horns of a deer (noun in nominative case).</paragraph>
</subSubSection>
<subSubSection id="04DA65443F44FF9F9FF90501FB9FFAB7" pageId="10" pageNumber="11" type="materials_examined">
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<emphasis id="7EB4EADD3F44FF9F9FF90501FE2BFBB5" bold="true" box="[189,492,1113,1140]" pageId="10" pageNumber="11">Type material examined</emphasis>
</heading>
</paragraph>
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<typeStatus id="937B886D3F44FF9F9FAE05D3FE9BFB64" box="[234,348,1160,1186]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
</emphasis>
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•
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; “
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,
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,
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,
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&amp;
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”;
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.
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<paragraph id="4C7F36CF3F44FF9F9FF90468FB9FFAB7" blockId="10.[189,1398,1296,1393]" pageId="10" pageNumber="11">
<materialsCitation id="FCA83C923F44FF9F9FF90468FB94FAB7" ID-GBIF-Occurrence="2597609332" collectingDate="2019-02-15" collectingMethod="Black light" collectionCode="ZFMK" collectorName="Eberle &amp; Ranasinghe" country="Sri Lanka" county="Pitawala Pathana" elevation="902" latitude="7.549767" location="Sri Lanka" longLatPrecision="1" longitude="80.75212" municipality="Riverston" pageId="10" pageNumber="11" specimenCode="X-SR0189" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<collectingCountry id="34D7765F3F44FF9F9FF90468FE91FA8B" box="[189,342,1331,1357]" name="Sri Lanka" pageId="10" pageNumber="11">SRI LANKA</collectingCountry>
•
<specimenCount id="5AC6FD463F44FF9F9E280468FE5EFA88" box="[364,409,1331,1358]" pageId="10" pageNumber="11">1 ♂</specimenCount>
; “
<specimenCode id="1C669EB43F44FF9F9EF40468FDF4FA8B" box="[432,563,1331,1357]" pageId="10" pageNumber="11">X-SR0189</specimenCode>
,
<location id="491F60143F44FF9F9D790468FD75FA8B" LSID="urn:lsid:plazi:treatment:C46987D93F44FF9B9D2C03D3FE32FEE3:491F60143F44FF9F9D790468FD75FA8B" box="[573,690,1331,1357]" country="Sri Lanka" county="Pitawala Pathana" latitude="7.549767" longLatPrecision="1" longitude="80.75212" municipality="Riverston" name="Sri Lanka" pageId="10" pageNumber="11">Sri Lanka</location>
,
<location id="491F60143F44FF9F9DF90468FCA8FA8B" LSID="urn:lsid:plazi:treatment:C46987D93F44FF9B9D2C03D3FE32FEE3:491F60143F44FF9F9DF90468FCA8FA8B" box="[701,879,1331,1357]" country="Sri Lanka" county="Pitawala Pathana" latitude="7.549767" longLatPrecision="1" longitude="80.75212" municipality="Riverston" name="Matale District" pageId="10" pageNumber="11">Matale District</location>
,
<collectingMunicipality id="AC1BACB53F44FF9F9C3E0468FC28FA8B" box="[890,1007,1331,1357]" pageId="10" pageNumber="11">Riverston</collectingMunicipality>
,
<collectingCounty id="A51E4E433F44FF9F9CBD0468FB03FA8B" box="[1017,1220,1331,1357]" pageId="10" pageNumber="11">Pitawala Pathana</collectingCounty>
,
<geoCoordinate id="29F450083F44FF9F9B8B0468FAB5FA88" box="[1231,1394,1331,1358]" degrees="7,54976718" direction="north" orientation="latitude" pageId="10" pageNumber="11" precision="1" value="7.549767">7,54976718N</geoCoordinate>
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<geoCoordinate id="29F450083F44FF9F9FF9040DFEABFAB7" box="[189,364,1366,1393]" degrees="80,75212294" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="1" value="80.75212">80,75212294E</geoCoordinate>
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<quantity id="8B389B2A3F44FF9F9E3C040DFE7AFAB6" box="[376,445,1366,1392]" metricMagnitude="2" metricUnit="m" metricValue="9.02" pageId="10" pageNumber="11" unit="m" value="902.0">
<elevation id="C7EDD1FC3F44FF9F9E3C040DFE7AFAB6" box="[376,445,1366,1392]" metricMagnitude="2" metricUnit="m" metricValue="9.02" pageId="10" pageNumber="11" unit="m" value="902.0">902m</elevation>
</quantity>
,
<collectingMethod id="95814ED83F44FF9F9E8D040DFD89FAB6" box="[457,590,1366,1392]" pageId="10" pageNumber="11">Black light</collectingMethod>
,
<date id="387E100F3F44FF9F9D1D040DFD1CFAB7" box="[601,731,1366,1393]" pageId="10" pageNumber="11" value="2019-02-15">
<collectingDate id="283AE9E73F44FF9F9D1D040DFD1CFAB7" box="[601,731,1366,1393]" pageId="10" pageNumber="11" value="2019-02-15">15-II-2019</collectingDate>
</date>
,
<collectorName id="E13553193F44FF9F9DA3040DFCF3FAB6" box="[743,820,1366,1392]" pageId="10" pageNumber="11">Eberle</collectorName>
&amp;
<collectorName id="E13553193F44FF9F9C1D040DFC2FFAB6" box="[857,1000,1366,1392]" pageId="10" pageNumber="11">Ranasinghe</collectorName>
”;
<collectionCode id="2AD1AE0A3F44FF9F9CBA040CFB94FAB7" box="[1022,1107,1367,1393]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="10" pageNumber="11" type="Museum">ZFMK</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="04DA65443F44FF9E9FF904CCFEFEF8A7" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="description">
<paragraph id="4C7F36CF3F44FF9F9FF904CCFE89FA77" blockId="10.[189,334,1431,1457]" box="[189,334,1431,1457]" pageId="10" pageNumber="11">
<heading id="173781A33F44FF9F9FF904CCFE89FA77" bold="true" box="[189,334,1431,1457]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="7EB4EADD3F44FF9F9FF904CCFE89FA77" bold="true" box="[189,334,1431,1457]" pageId="10" pageNumber="11">Description</emphasis>
</heading>
</paragraph>
<paragraph id="4C7F36CF3F44FF9F9FF9049CFBE2FA27" blockId="10.[189,1061,1478,1505]" box="[189,1061,1478,1505]" pageId="10" pageNumber="11">
MEASUREMENTS. Length:
<quantity id="8B389B2A3F44FF9F9E9A049DFDFCFA27" box="[478,571,1478,1505]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="10" pageNumber="11" unit="mm" value="7.2">7.2 mm</quantity>
, length of elytra:
<quantity id="8B389B2A3F44FF9F9C4D049DFCA1FA27" box="[777,870,1478,1505]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="10" pageNumber="11" unit="mm" value="5.4">5.4 mm</quantity>
, width:
<quantity id="8B389B2A3F44FF9F9C87049DFBE7FA27" box="[963,1056,1478,1505]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="10" pageNumber="11" unit="mm" value="4.4">4.4 mm</quantity>
.
</paragraph>
<paragraph id="4C7F36CF3F44FF9F9FF90753FB42F9E4" blockId="10.[189,1157,1543,1570]" box="[189,1157,1543,1570]" pageId="10" pageNumber="11">
VARIATION. Length:
<quantity id="8B389B2A3F44FF9F9EE00753FDF2F9E4" box="[420,565,1544,1570]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="7.8" metricValueMin="7.2" pageId="10" pageNumber="11" unit="mm" value="7.5" valueMax="7.8" valueMin="7.2">7.2–7.8 mm</quantity>
, length of elytra:
<quantity id="8B389B2A3F44FF9F9C47075CFC54F9E4" box="[771,915,1543,1570]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" metricValueMax="5.8" metricValueMin="5.4" pageId="10" pageNumber="11" unit="mm" value="5.6" valueMax="5.8" valueMin="5.4">5.4–5.8 mm</quantity>
, width:
<quantity id="8B389B2A3F44FF9F9CB5075CFB46F9E4" box="[1009,1153,1543,1570]" metricMagnitude="-3" metricUnit="m" metricValue="4.45" metricValueMax="4.5" metricValueMin="4.4" pageId="10" pageNumber="11" unit="mm" value="4.45" valueMax="4.5" valueMin="4.4">4.4–4.5 mm</quantity>
.
</paragraph>
<paragraph id="4C7F36CF3F44FF9F9FF90711FE73F940" blockId="10.[189,1398,1609,1670]" pageId="10" pageNumber="11">HABITUS AND COLORATION. Body short oval, dark brown, antenna yellow, dorsal surface shiny, nearly completely glabrous.</paragraph>
<paragraph id="4C7F36CF3F44FF9E9FF907F5FBB3FE8E" blockId="10.[189,1399,1710,1983]" lastBlockId="11.[189,1398,267,329]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">HEAD. Labroclypeus short and rectangular, wider than long, widest at base, lateral margins moderately convex and convergent to broadly rounded anterior angles, lateral border and ocular canthus producing an indistinct blunt angle, margins weakly reflexed, anterior margin almost feebly sinuate medially; surface slightly convex, finely and densely punctate, distance between punctures smaller than their diameter, with numerous erect setae in larger punctures; frontoclypeal suture indistinctly incised and bluntly bent medially; smooth area in front of eye approximately three times as wide as long; ocular canthus long and narrow, minutely and superficially punctate, without a single short terminal seta. Frons with fine, dense punctures, with a few long erect setae in larger punctures. Eyes very large, ratio of diameter/ interocular width: 0.97. Antenna yellow, with ten antennomeres; club with three antennomeres, as long as remaining antennomeres combined. Mentum elevated and anteriorly flattened.</paragraph>
<paragraph id="4C7F36CF3F45FF9E9FF9002BFDE1FDFC" blockId="11.[189,1399,368,570]" pageId="11" pageNumber="12">PRONOTUM. Wide, widest at base, lateral margins weakly convex and evenly narrowed to the anterior angles, anterior angles moderately produced and sharp, anterior marginal line very fine but complete medially, anterior margin straight; surface finely densely punctate, except a few short and fine setae laterally glabrous; anterior and lateral borders setose, basal margin without marginal line; hypomeron ventrobasally carinate and slightly produced ventrally. Scutellum short and broad, triangular, with fine and dense punctures, glabrous.</paragraph>
<paragraph id="4C7F36CF3F45FF9E9FF90339FB6AFD23" blockId="11.[189,1399,609,742]" pageId="11" pageNumber="12">ELYTRA. Short oval, widest shortly behind middle, striae distinctly impressed, finely and densely punctate, intervals weakly convex, with moderately fine, dense punctures and with dense, fine, short setae; epipleural edge fine, ending at the weakly convex external apical angle of elytra, epipleura densely setose; apical border narrowly membranous, apex covered with short microtrichomes.</paragraph>
<paragraph id="4C7F36CF3F45FF9E9FF90256FB71FBA2" blockId="11.[189,1399,780,1124]" pageId="11" pageNumber="12">
VENTRAL SURFACE. Shiny, thorax and metacoxa with large and dense punctures, sparsely setose, metacoxa with minute adjacent setae in the punctures except for numerous long setae laterally, apical margin weakly convex, without a broad rim of long white microtrichomes; each abdominal sternite, in addition to evenly distributed fine and dense punctures bearing each a fine seta, with a distinct transverse row of coarse punctures each bearing a long and more robust seta, 3
<superScript id="BBB59B873F45FF9E9CE802C3FC7DFC60" attach="left" box="[940,954,920,934]" fontSize="6" pageId="11" pageNumber="12">rd</superScript>
sternite with a sharp median tubercle being half as high as sternite length, penultimate sternite apically with a shiny smooth chitinous border, which is a quarter as long as the sternite. Mesosternum between mesocoxae as wide as mesofemur, with a semi-circular ridge bearing robust setae. Ratio of length of metepisternum/ metacoxa: 1/ 1.89. Pygidium moderately convex, moderately finely and densely punctate, without smooth midline, punctures with short and moderately dense, adjacent setae or with moderately dense, long, erect setae.
</paragraph>
<paragraph id="4C7F36CF3F45FF9E9FF905D0FCFAF9A8" blockId="11.[189,1399,1163,1646]" pageId="11" pageNumber="12">LEGS. Short and wide; femur with two longitudinal rows of setae, finely and densely punctate; metafemur shiny, anterior edge acute, lacking an adjacent serrated line, ventral surface densely punctate and setose, posterior ventral margin straight, only little widened in apical half, finely serrate apically, dorsally not serrated, glabrous. Metatibia short, widest at middle, posteriorly only very little narrowed, ratio width/ length: 1/ 2.87, dorsally sharply carinate, with two groups of spines, basal one at middle, apical one at three quarters of metatibial length, basally beside dorsal margin with two single punctures with serrated margins, each bearing a single robust spine and beside them a longitudinal serrated line; lateral face almost flat, with dense, large but superficial punctures and with minute setae in the punctures; ventral margin with four strong spines equidistant from each other, medial face smooth, apex interiorly near tarsal articulation shallowly concave. Meso- and metatarsomeres finely and sparsely punctate but glabrous dorsally, ventrally with sparse, short setae; metatarsomeres ventrally with a strongly serrated ridge, beside which is a strong longitudinal carina; first metatarsomere as long as following two tarsomeres combined and as long as dorsal tibial spur. Protibia short, bidentate. All claws symmetrical, feebly curved and long, with normally developed basal tooth.</paragraph>
<paragraph id="4C7F36CF3F45FF9E9FF907CDFE05F976" blockId="11.[189,450,1686,1712]" box="[189,450,1686,1712]" pageId="11" pageNumber="12">
AEDEAGUS.
<figureCitation id="D4FB2A4A3F45FF9E9E0507CDFE05F976" box="[321,450,1686,1712]" captionStart="Fig" captionStartId="9.[189,232,1931,1957]" captionTargetBox="[209,1372,940,1880]" captionTargetId="figure@9.[193,1394,919,1894]" captionTargetPageId="9" captionText="Fig. 4. A–D. Maladera galdaththana sp. nov. (holotype). E–H. M. cervicornis sp. nov. (holotype). A, E. Aedeagus, left side lateral view. C, G. Aedeagus, right side lateral view. B, F. Parameres, dorsal view. D, H. Habitus (not to scale). Scales: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3750259" httpUri="https://zenodo.org/record/3750259/files/figure.png" pageId="11" pageNumber="12">
Fig. 4E
<emphasis id="7EB4EADD3F45FF9E9ED307CDFE61F976" bold="true" box="[407,422,1686,1712]" pageId="11" pageNumber="12">–</emphasis>
G.
</figureCitation>
</paragraph>
<paragraph id="4C7F36CF3F45FF9E9FF90783FE4BF937" blockId="11.[189,396,1751,1778]" box="[189,396,1751,1778]" pageId="11" pageNumber="12">
HABITUS.
<figureCitation id="D4FB2A4A3F45FF9E9E6F078CFE4FF937" box="[299,392,1751,1777]" captionStart="Fig" captionStartId="9.[189,232,1931,1957]" captionTargetBox="[209,1372,940,1880]" captionTargetId="figure@9.[193,1394,919,1894]" captionTargetPageId="9" captionText="Fig. 4. A–D. Maladera galdaththana sp. nov. (holotype). E–H. M. cervicornis sp. nov. (holotype). A, E. Aedeagus, left side lateral view. C, G. Aedeagus, right side lateral view. B, F. Parameres, dorsal view. D, H. Habitus (not to scale). Scales: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3750259" httpUri="https://zenodo.org/record/3750259/files/figure.png" pageId="11" pageNumber="12">Fig. 4H</figureCitation>
.
</paragraph>
<paragraph id="4C7F36CF3F45FF9E9FF90642FEDFF8F5" blockId="11.[189,280,1817,1843]" box="[189,280,1817,1843]" pageId="11" pageNumber="12">
<heading id="173781A33F45FF9E9FF90642FEDFF8F5" bold="true" box="[189,280,1817,1843]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="7EB4EADD3F45FF9E9FF90642FEDFF8F5" bold="true" box="[189,280,1817,1843]" pageId="11" pageNumber="12">Female</emphasis>
</heading>
</paragraph>
<paragraph id="4C7F36CF3F45FF9E9FF9061CFEFEF8A7" blockId="11.[189,313,1863,1889]" box="[189,313,1863,1889]" pageId="11" pageNumber="12">Unknown.</paragraph>
</subSubSection>
<subSubSection id="04DA65443F45FF9B9FF906D3FE32FEE3" lastPageId="14" lastPageNumber="15" pageId="11" pageNumber="12" type="distribution">
<paragraph id="4C7F36CF3F45FF9E9FF906D3FE91F864" blockId="11.[189,342,1928,1954]" box="[189,342,1928,1954]" pageId="11" pageNumber="12">
<heading id="173781A33F45FF9E9FF906D3FE91F864" bold="true" box="[189,342,1928,1954]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="7EB4EADD3F45FF9E9FF906D3FE91F864" bold="true" box="[189,342,1928,1954]" pageId="11" pageNumber="12">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="4C7F36CF3F45FF9E9FF906ECFE97F814" blockId="11.[189,336,1975,2002]" box="[189,336,1975,2002]" pageId="11" pageNumber="12">
See
<figureCitation id="D4FB2A4A3F45FF9E9FAB06ECFE8CF814" box="[239,331,1975,2002]" captionStart="Fig" captionStartId="12.[189,232,1965,1991]" captionTargetBox="[189,1398,324,1918]" captionTargetId="figure@12.[189,1398,324,1918]" captionTargetPageId="12" captionText="Fig. 5. Distribution of the new species. A. Selaserica athukoralai sp. nov. B. Neoserica dharmapriyai sp. nov. C. Maladera galdaththana sp. nov. D. M. cervicornis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3750261" httpUri="https://zenodo.org/record/3750261/files/figure.png" pageId="11" pageNumber="12">Fig. 5D</figureCitation>
.
</paragraph>
<caption id="18BF66473F42FF999FF906F6FC33F82D" ID-DOI="http://doi.org/10.5281/zenodo.3750261" ID-Zenodo-Dep="3750261" httpUri="https://zenodo.org/record/3750261/files/figure.png" pageId="12" pageNumber="13" startId="12.[189,232,1965,1991]" targetBox="[189,1398,324,1918]" targetPageId="12">
<paragraph id="4C7F36CF3F42FF999FF906F6FC33F82D" blockId="12.[189,1398,1965,2027]" pageId="12" pageNumber="13">
<emphasis id="7EB4EADD3F42FF999FF906F6FECBF80E" bold="true" box="[189,268,1965,1992]" pageId="12" pageNumber="13">Fig. 5.</emphasis>
Distribution of the new species.
<emphasis id="7EB4EADD3F42FF999DD206F6FD6CF801" bold="true" box="[662,683,1965,1991]" pageId="12" pageNumber="13">A</emphasis>
.
<taxonomicName id="8BC04D4C3F42FF999DFF06F6FC01F801" authority="Ranasinghe &amp; Eberle &amp; Benjamin &amp; Ahrens, 2020" authorityName="Ranasinghe &amp; Eberle &amp; Benjamin &amp; Ahrens" authorityYear="2020" box="[699,966,1965,1991]" class="Insecta" family="Scarabaeidae" genus="Selaserica" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="athukoralai" status="sp. nov.">
<emphasis id="7EB4EADD3F42FF999DFF06F6FC01F801" box="[699,966,1965,1991]" italics="true" pageId="12" pageNumber="13">Selaserica athukoralai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="658757A63F42FF999C8B06F5FBEDF80E" box="[975,1066,1966,1992]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="7EB4EADD3F42FF999B7606F5FB81F80E" bold="true" box="[1074,1094,1966,1992]" pageId="12" pageNumber="13">B</emphasis>
.
<taxonomicName id="8BC04D4C3F42FF999B1206F5FAB1F801" authority="Ranasinghe &amp; Eberle &amp; Benjamin &amp; Ahrens, 2020" authorityName="Ranasinghe &amp; Eberle &amp; Benjamin &amp; Ahrens" authorityYear="2020" box="[1110,1398,1965,1991]" class="Insecta" family="Melolonthidae" genus="Neoserica" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="dharmapriyai" status="sp. nov.">
<emphasis id="7EB4EADD3F42FF999B1206F5FAB1F801" box="[1110,1398,1965,1991]" italics="true" pageId="12" pageNumber="13">Neoserica dharmapriyai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="658757A63F42FF999FF9068AFEDFF82D" box="[189,280,2001,2027]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="7EB4EADD3F42FF999E5B068BFEF3F82C" bold="true" box="[287,308,2000,2026]" pageId="12" pageNumber="13">C</emphasis>
.
<taxonomicName id="8BC04D4C3F42FF999E07068AFD9DF82D" authority="Ranasinghe &amp; Eberle &amp; Benjamin &amp; Ahrens, 2020" authorityName="Ranasinghe &amp; Eberle &amp; Benjamin &amp; Ahrens" authorityYear="2020" box="[323,602,2001,2027]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="galdaththana" status="sp. nov.">
<emphasis id="7EB4EADD3F42FF999E07068AFD9DF82D" box="[323,602,2001,2027]" italics="true" pageId="12" pageNumber="13">Maladera galdaththana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="658757A63F42FF999D25068AFD7BF82D" box="[609,700,2001,2027]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="7EB4EADD3F42FF999D87068AFD1FF82D" bold="true" box="[707,728,2001,2027]" pageId="12" pageNumber="13">D</emphasis>
.
<taxonomicName id="8BC04D4C3F42FF999DA30689FC55F82D" authority="Ranasinghe &amp; Eberle &amp; Benjamin &amp; Ahrens, 2020" authorityName="Ranasinghe &amp; Eberle &amp; Benjamin &amp; Ahrens" authorityYear="2020" box="[743,914,2002,2027]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cervicornis" status="sp. nov.">
<emphasis id="7EB4EADD3F42FF999DA30689FC55F82D" box="[743,914,2002,2027]" italics="true" pageId="12" pageNumber="13">M. cervicornis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="658757A63F42FF999CDD068AFC33F82D" box="[921,1012,2001,2027]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<caption id="18BF66473F43FF989FF906F6FB78F82D" ID-DOI="http://doi.org/10.5281/zenodo.3750263" ID-Zenodo-Dep="3750263" httpUri="https://zenodo.org/record/3750263/files/figure.png" pageId="13" pageNumber="14" startId="13.[189,232,1965,1991]" targetBox="[224,1364,318,1924]" targetPageId="13">
<paragraph id="4C7F36CF3F43FF989FF906F6FB78F82D" blockId="13.[189,1399,1965,2027]" pageId="13" pageNumber="14">
<emphasis id="7EB4EADD3F43FF989FF906F6FEC8F801" bold="true" box="[189,271,1965,1991]" pageId="13" pageNumber="14">Fig. 6.</emphasis>
Photos of the habitats of the new species.
<emphasis id="7EB4EADD3F43FF989C6F06F6FC87F801" bold="true" box="[811,832,1965,1991]" pageId="13" pageNumber="14">A</emphasis>
.
<taxonomicName id="8BC04D4C3F43FF989C1006F6FBA4F801" authority="Ranasinghe &amp; Eberle &amp; Benjamin &amp; Ahrens, 2020" authorityName="Ranasinghe &amp; Eberle &amp; Benjamin &amp; Ahrens" authorityYear="2020" box="[852,1123,1965,1991]" class="Insecta" family="Scarabaeidae" genus="Selaserica" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="athukoralai" status="sp. nov.">
<emphasis id="7EB4EADD3F43FF989C1006F6FBA4F801" box="[852,1123,1965,1991]" italics="true" pageId="13" pageNumber="14">Selaserica athukoralai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="658757A63F43FF989B2A06F5FB09F80E" box="[1134,1230,1966,1992]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="7EB4EADD3F43FF989B9D06F5FB2AF80E" bold="true" box="[1241,1261,1966,1992]" pageId="13" pageNumber="14">B</emphasis>
.
<taxonomicName id="8BC04D4C3F43FF989A4406F5FE99F82D" authority="Ranasinghe &amp; Eberle &amp; Benjamin &amp; Ahrens, 2020" authorityName="Ranasinghe &amp; Eberle &amp; Benjamin &amp; Ahrens" authorityYear="2020" class="Insecta" family="Melolonthidae" genus="Neoserica" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="dharmapriyai" status="sp. nov.">
<emphasis id="7EB4EADD3F43FF989A4406F5FE99F82D" italics="true" pageId="13" pageNumber="14">Neoserica dharmapriyai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="658757A63F43FF989E22068AFE06F82D" box="[358,449,2001,2027]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="7EB4EADD3F43FF989E8C068BFE38F82C" bold="true" box="[456,511,2000,2026]" pageId="13" pageNumber="14">B–C</emphasis>
.
<taxonomicName id="8BC04D4C3F43FF989D4A068AFCE2F82D" authority="Ranasinghe &amp; Eberle &amp; Benjamin &amp; Ahrens, 2020" authorityName="Ranasinghe &amp; Eberle &amp; Benjamin &amp; Ahrens" authorityYear="2020" box="[526,805,2001,2027]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="galdaththana" status="sp. nov.">
<emphasis id="7EB4EADD3F43FF989D4A068AFCE2F82D" box="[526,805,2001,2027]" italics="true" pageId="13" pageNumber="14">Maladera galdaththana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="658757A63F43FF989C68068AFC40F82D" box="[812,903,2001,2027]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="7EB4EADD3F43FF989CCA068AFC64F82D" bold="true" box="[910,931,2001,2027]" pageId="13" pageNumber="14">D</emphasis>
.
<taxonomicName id="8BC04D4C3F43FF989CF60689FB9AF82D" authority="Ranasinghe &amp; Eberle &amp; Benjamin &amp; Ahrens, 2020" authorityName="Ranasinghe &amp; Eberle &amp; Benjamin &amp; Ahrens" authorityYear="2020" box="[946,1117,2002,2027]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="cervicornis" status="sp. nov.">
<emphasis id="7EB4EADD3F43FF989CF60689FB9AF82D" box="[946,1117,2002,2027]" italics="true" pageId="13" pageNumber="14">M. cervicornis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="658757A63F43FF989B20068AFB78F82D" box="[1124,1215,2001,2027]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="4C7F36CF3F40FF9B9FF90050FE32FEE3" blockId="14.[189,501,267,293]" box="[189,501,267,293]" pageId="14" pageNumber="15">
<heading id="173781A33F40FF9B9FF90050FE32FEE3" bold="true" box="[189,501,267,293]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="7EB4EADD3F40FF9B9FF90050FE32FEE3" bold="true" box="[189,501,267,293]" pageId="14" pageNumber="15">New distribution records</emphasis>
</heading>
</paragraph>
</subSubSection>
</treatment>
</document>
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