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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1053.67326" ID-GBIF-Dataset="6ab5131b-fa67-412c-ad75-6e75f8eac2f8" ID-PMC="PMC8352857" ID-Pensoft-Pub="1313-2970-1053-185" ID-Pensoft-UUID="4A8522D8A609541581BC3FB89AFFFCFE" ID-PubMed="34393559" ID-ZooBank="6C907A4B587440D1881E18BA380D9C21" ModsDocID="1313-2970-1053-185" checkinTime="1627926266140" checkinUser="pensoft" docAuthor="Ng, Peter K. L. &amp; Chan, Tin-Yam" docDate="2021" docId="BFBD7B01F0A15F6BBCBBEE3FB81EC14A" docLanguage="en" docName="ZooKeys 1053: 185-195" docOrigin="ZooKeys 1053" docPubDate="2021-08-02" docSource="http://dx.doi.org/10.3897/zookeys.1053.67326" docTitle="Orientotlos iishibai Sakai 1980" docType="treatment" docVersion="4" id="4A8522D8A609541581BC3FB89AFFFCFE" lastPageNumber="185" masterDocId="4A8522D8A609541581BC3FB89AFFFCFE" masterDocTitle="Rediscovery of Orientotlos iishibai Sakai, 1980 (Crustacea, Decapoda, Brachyura, Leucosiidae) in Taiwan" masterLastPageNumber="195" masterPageNumber="185" pageNumber="185" updateTime="1668150789084" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Rediscovery of Orientotlos iishibai Sakai, 1980 (Crustacea, Decapoda, Brachyura, Leucosiidae) in Taiwan</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ng, Peter K. L.</mods:namePart>
<mods:affiliation>Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Singapore</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chan, Tin-Yam</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8143-0007</mods:nameIdentifier>
<mods:affiliation>Institute of Marine Biology and Center of Excellence for the Oceans, National Taiwan Ocean University, Keelung 202301, Taiwan</mods:affiliation>
<mods:nameIdentifier type="email">tychan@mail.ntou.edu.tw</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-08-02</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1053</mods:number>
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<mods:start>185</mods:start>
<mods:end>195</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1053.67326</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1053.67326</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1053-185</mods:identifier>
<mods:identifier type="ZooBank">6C907A4B587440D1881E18BA380D9C21</mods:identifier>
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<subSection pageId="0" pageNumber="185" type="taxonomy">
<treatment ID-GBIF-Taxon="183907165" LSID="urn:lsid:plazi:treatment:BFBD7B01F0A15F6BBCBBEE3FB81EC14A" httpUri="http://treatment.plazi.org/id/BFBD7B01F0A15F6BBCBBEE3FB81EC14A" lastPageNumber="185" pageId="0" pageNumber="185">
<subSubSection pageId="0" pageNumber="185" type="nomenclature">
<paragraph pageId="0" pageNumber="185">
<taxonomicName LSID="BFBD7B01-F0A1-5F6B-BCBB-EE3FB81EC14A" authority="Sakai, 1980" authorityName="Sakai" authorityYear="1980" class="Malacostraca" family="Leucosiidae" genus="Orientotlos" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orientotlos iishibai" order="Decapoda" pageId="0" pageNumber="185" phylum="Arthropoda" rank="species" species="iishibai">Orientotlos iishibai Sakai, 1980</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="185" type="description">
<paragraph pageId="0" pageNumber="185">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Orientotlos iishibai Sakai, 1980 A holotype female (7.0 by 5.5 mm) (repository unknown), Japan [after Sakai 1980: text-fig. 1] B color in life, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan." figureDoi="10.3897/zookeys.1053.67326.figure1" httpUri="https://binary.pensoft.net/fig/572659" pageId="0" pageNumber="185">Figs 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A overall dorsal view B overall ventral view C, D dorsal view of carapace from different angles." figureDoi="10.3897/zookeys.1053.67326.figure2" httpUri="https://binary.pensoft.net/fig/572660" pageId="0" pageNumber="185">, 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A frontal view of cephalothorax B frontal view showing antennules, antennae and orbits C lateral view of cephalothorax D pterygostomial region, buccal cavity, third maxillipeds and anterior part of sternopleonal cavity E left third maxilliped F outer view of left chela G right fourth ambulatory leg." figureDoi="10.3897/zookeys.1053.67326.figure3" httpUri="https://binary.pensoft.net/fig/572661" pageId="0" pageNumber="185">, 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A intestinal region, posterior carapace lobes and pleonal somites 2 - 6 B posterior carapace lobes, pleonal somites 2 - 6, and telson C pleonal somites 4 - 6 and telson D anterior thoracic sternites and sternopleonal cavity E sternopleonal cavity and vulvae." figureDoi="10.3897/zookeys.1053.67326.figure4" httpUri="https://binary.pensoft.net/fig/572662" pageId="0" pageNumber="185">, 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="185" type="reference_group">
<paragraph pageId="0" pageNumber="185">
<taxonomicName authorityName="Sakai" authorityYear="1980" class="Malacostraca" family="Leucosiidae" genus="Orientotlos" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orientotlos iishibai" order="Decapoda" pageId="0" pageNumber="185" phylum="Arthropoda" rank="species" species="iishibai">Orientotlos iishibai</taxonomicName>
Sakai, 1980: 74, text-fig. 1. -
<bibRefCitation author="Ng, PKL" journalOrPublisher="Raffles Bulletin of Zoology, Supplement" pageId="0" pageNumber="185" pagination="1 - 286" refId="B15" refString="Ng, PKL, Guinot, D, Davie, PJF, 2008. Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17: 1 - 286" title="Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world." volume="17" year="2008">Ng et al. 2008</bibRefCitation>
: 92 (list).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="185" type="materials_examined">
<paragraph pageId="0" pageNumber="185">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="185">
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Taiwan" location="Taiwan" specimenCount="1" specimenCount-female="1">
<collectingCountry name="Taiwan">Taiwan</collectingCountry>
<specimenCount type="female">1 ♀</specimenCount>
ovigerous (7.8
<normalizedToken originalValue="×">x</normalizedToken>
<quantity metricMagnitude="-3" metricUnit="m" metricValue="6.5" unit="mm" value="6.5">6.5 mm</quantity>
); station CP4210, off southwestern
</materialsCitation>
<materialsCitation accessionNumber="B00125" collectingDate="2020-11-14" collectorName="T. - Y. Chan" country="Taiwan" elevation="138" latitude="22.315666" location="Taiwan" longLatPrecision="12" longitude="120.342834" specimenCount="1">
<collectingCountry name="Taiwan">Taiwan</collectingCountry>
coast;
<geoCoordinate degrees="22" direction="north" minutes="18.94" orientation="latitude" precision="9" value="22.315666">22°18.94'N</geoCoordinate>
,
<geoCoordinate degrees="120" direction="east" minutes="20.57" orientation="longitude" precision="9" value="120.342834">120°20.57'E</geoCoordinate>
; depth
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.375" metricValueMax="1.59" metricValueMin="1.16" unit="m" value="137.5" valueMax="159.0" valueMin="116.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.375" metricValueMax="1.59" metricValueMin="1.16" unit="m" value="137.5" valueMax="159.0" valueMin="116.0">116-159 m</elevation>
</quantity>
;
<collectingDate value="2020-11-14">14 Nov. 2020</collectingDate>
;
<collectorName>T.-Y. Chan</collectorName>
leg.; hard bottom substrate; NTOU
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/B00125">B00125</accessionNumber>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="185" type="diagnosis">
<paragraph pageId="0" pageNumber="185">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="185">As for genus.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="185" type="description">
<paragraph pageId="0" pageNumber="185">Description.</paragraph>
<paragraph pageId="0" pageNumber="185">
<emphasis bold="true" pageId="0" pageNumber="185">Female.</emphasis>
Carapace subhexagonal in outline, 1.2
<normalizedToken originalValue="×">x</normalizedToken>
as wide as long; upper surface (between plates and bosses) paved with numerous, well-spaced boletiform and rounded tubercles of varying sizes (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Orientotlos iishibai Sakai, 1980 A holotype female (7.0 by 5.5 mm) (repository unknown), Japan [after Sakai 1980: text-fig. 1] B color in life, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan." figureDoi="10.3897/zookeys.1053.67326.figure1" httpUri="https://binary.pensoft.net/fig/572659" pageId="0" pageNumber="185">1</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A overall dorsal view B overall ventral view C, D dorsal view of carapace from different angles." figureDoi="10.3897/zookeys.1053.67326.figure2" httpUri="https://binary.pensoft.net/fig/572660" pageId="0" pageNumber="185">2A-D</figureCitation>
). Front produced, upturned, margin weakly bilobed with shallow median concavity; frontal margin gently confluent with concave supraorbital margin; hepatic region plate-like, formed by coalesced granules; subhepatic region swollen, forming a distinct obtuse angle visible in dorsal view; hepatic plate separated from first anterolateral tooth by a wide, deep cleft; anterolateral margin with 3 large lobiform teeth increasing in size posteriorly, third tooth directed obliquely, surface paved with flattened granules, margins lined with low granules; posterolateral margin concave, with median triangular tooth, margin with rounded granules; posterior carapace margin with 2 large lozenge-shaped bosses, directed posteriorly, margin flattened, surface and margins lined with rounded and flattened granules (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Orientotlos iishibai Sakai, 1980 A holotype female (7.0 by 5.5 mm) (repository unknown), Japan [after Sakai 1980: text-fig. 1] B color in life, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan." figureDoi="10.3897/zookeys.1053.67326.figure1" httpUri="https://binary.pensoft.net/fig/572659" pageId="0" pageNumber="185">1</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A overall dorsal view B overall ventral view C, D dorsal view of carapace from different angles." figureDoi="10.3897/zookeys.1053.67326.figure2" httpUri="https://binary.pensoft.net/fig/572660" pageId="0" pageNumber="185">2A-D</figureCitation>
). Suborbital region substantially compressed; subhepatic region swollen, surface with large, flattened tubercles, margins of subhepatic and pterygostomial regions covered with numerous boletiform tubercles (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A frontal view of cephalothorax B frontal view showing antennules, antennae and orbits C lateral view of cephalothorax D pterygostomial region, buccal cavity, third maxillipeds and anterior part of sternopleonal cavity E left third maxilliped F outer view of left chela G right fourth ambulatory leg." figureDoi="10.3897/zookeys.1053.67326.figure3" httpUri="https://binary.pensoft.net/fig/572661" pageId="0" pageNumber="185">3D, E</figureCitation>
); subhepatic and hepatic regions separated by distinct groove lined with granules, inner edge leading to just before tip of efferent branchial channel, outer edge joining cleft between hepatic lobe and first anterolateral tooth (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A frontal view of cephalothorax B frontal view showing antennules, antennae and orbits C lateral view of cephalothorax D pterygostomial region, buccal cavity, third maxillipeds and anterior part of sternopleonal cavity E left third maxilliped F outer view of left chela G right fourth ambulatory leg." figureDoi="10.3897/zookeys.1053.67326.figure3" httpUri="https://binary.pensoft.net/fig/572661" pageId="0" pageNumber="185">3A, B</figureCitation>
). Postfrontal median keel on gastric region prominent, high, extending posteriorly to cardiac region as raised row of rounded tubercles; postorbital region gently concave, without deep depression; median part of carapace dome-shaped, protogastric region with ovate boletiform plate, obliquely positioned, margin granulated; cardiac region swollen, with raised, vaguely T-shaped ridge formed of fused granules surrounded by rounded granules; epibranchial region with a small, subovate, transversely positioned boletiform plate, margin lined with small granules; mesobranchial region with large ovate boletiform plate, margin lined with granules; intestinal region strongly inflated, with a large subtriangular boletiform plate, directed posteriorly, margin with distinct granules (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Orientotlos iishibai Sakai, 1980 A holotype female (7.0 by 5.5 mm) (repository unknown), Japan [after Sakai 1980: text-fig. 1] B color in life, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan." figureDoi="10.3897/zookeys.1053.67326.figure1" httpUri="https://binary.pensoft.net/fig/572659" pageId="0" pageNumber="185">1</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A overall dorsal view B overall ventral view C, D dorsal view of carapace from different angles." figureDoi="10.3897/zookeys.1053.67326.figure2" httpUri="https://binary.pensoft.net/fig/572660" pageId="0" pageNumber="185">2A, C, D</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A frontal view of cephalothorax B frontal view showing antennules, antennae and orbits C lateral view of cephalothorax D pterygostomial region, buccal cavity, third maxillipeds and anterior part of sternopleonal cavity E left third maxilliped F outer view of left chela G right fourth ambulatory leg." figureDoi="10.3897/zookeys.1053.67326.figure3" httpUri="https://binary.pensoft.net/fig/572661" pageId="0" pageNumber="185">3A, C</figureCitation>
). Orbital margin not clearly marked, gradually merging with granules from frontal margin and hepatic region; cornea visible in dorsal view, peduncle short with small sharp granules (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A overall dorsal view B overall ventral view C, D dorsal view of carapace from different angles." figureDoi="10.3897/zookeys.1053.67326.figure2" httpUri="https://binary.pensoft.net/fig/572660" pageId="0" pageNumber="185">2C, D</figureCitation>
). Antennule folded into an oblique fossa; basal segment finely granulate, occupying lower two-thirds of fossa (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A frontal view of cephalothorax B frontal view showing antennules, antennae and orbits C lateral view of cephalothorax D pterygostomial region, buccal cavity, third maxillipeds and anterior part of sternopleonal cavity E left third maxilliped F outer view of left chela G right fourth ambulatory leg." figureDoi="10.3897/zookeys.1053.67326.figure3" httpUri="https://binary.pensoft.net/fig/572661" pageId="0" pageNumber="185">3B</figureCitation>
). Basal antennal article large, subquadrate, fused with epistome, forming most of suborbital margin, covered with rounded tubercles; antennae small, slender, inserted in orbital hiatus (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A frontal view of cephalothorax B frontal view showing antennules, antennae and orbits C lateral view of cephalothorax D pterygostomial region, buccal cavity, third maxillipeds and anterior part of sternopleonal cavity E left third maxilliped F outer view of left chela G right fourth ambulatory leg." figureDoi="10.3897/zookeys.1053.67326.figure3" httpUri="https://binary.pensoft.net/fig/572661" pageId="0" pageNumber="185">3B</figureCitation>
). Anterior margin of efferent branchial channel produced, reaching to just before proepistome, slightly notched (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A frontal view of cephalothorax B frontal view showing antennules, antennae and orbits C lateral view of cephalothorax D pterygostomial region, buccal cavity, third maxillipeds and anterior part of sternopleonal cavity E left third maxilliped F outer view of left chela G right fourth ambulatory leg." figureDoi="10.3897/zookeys.1053.67326.figure3" httpUri="https://binary.pensoft.net/fig/572661" pageId="0" pageNumber="185">3B</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1053.67326.figure2" httpUri="https://binary.pensoft.net/fig/572660" pageId="0" pageNumber="185" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="185">
<emphasis bold="true" pageId="0" pageNumber="185">Figure 2.</emphasis>
<taxonomicName authorityName="Sakai" authorityYear="1980" class="Malacostraca" family="Leucosiidae" genus="Orientotlos" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orientotlos iishibai" order="Decapoda" pageId="0" pageNumber="185" phylum="Arthropoda" rank="species" species="iishibai">
<emphasis italics="true" pageId="0" pageNumber="185">Orientotlos iishibai</emphasis>
</taxonomicName>
Sakai, 1980, female (7.8
<normalizedToken originalValue="×">x</normalizedToken>
6.5 mm) (NTOU B00125), Taiwan
<emphasis bold="true" pageId="0" pageNumber="185">A</emphasis>
overall dorsal view
<emphasis bold="true" pageId="0" pageNumber="185">B</emphasis>
overall ventral view
<emphasis bold="true" pageId="0" pageNumber="185">C, D</emphasis>
dorsal view of carapace from different angles.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="185">
Third maxilliped with merus, ischium and exopod paved with numerous flattened, rounded tubercles of varying sizes, those on proximal parts generally larger; merus triangular, about half length of ischium; palp (carpus, propodus and dactylus) shorter than merus, inserted on inner surface; dactylus distinctly longer than propodus; ischium subrectangular, with no visible median sulcus; basis-ischium and coxa not expanded, covered with numerous large rounded tubercles and some boletiform tubercles; exopod stout, broad, reaching to about half length of merus, basal part with large rounded, boletiform tubercles (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A frontal view of cephalothorax B frontal view showing antennules, antennae and orbits C lateral view of cephalothorax D pterygostomial region, buccal cavity, third maxillipeds and anterior part of sternopleonal cavity E left third maxilliped F outer view of left chela G right fourth ambulatory leg." figureDoi="10.3897/zookeys.1053.67326.figure3" httpUri="https://binary.pensoft.net/fig/572661" pageId="0" pageNumber="185">3D, E</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1053.67326.figure3" httpUri="https://binary.pensoft.net/fig/572661" pageId="0" pageNumber="185" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="185">
<emphasis bold="true" pageId="0" pageNumber="185">Figure 3.</emphasis>
<taxonomicName authorityName="Sakai" authorityYear="1980" class="Malacostraca" family="Leucosiidae" genus="Orientotlos" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orientotlos iishibai" order="Decapoda" pageId="0" pageNumber="185" phylum="Arthropoda" rank="species" species="iishibai">
<emphasis italics="true" pageId="0" pageNumber="185">Orientotlos iishibai</emphasis>
</taxonomicName>
Sakai, 1980, female (7.8
<normalizedToken originalValue="×">x</normalizedToken>
6.5 mm) (NTOU B00125), Taiwan
<emphasis bold="true" pageId="0" pageNumber="185">A</emphasis>
frontal view of cephalothorax
<emphasis bold="true" pageId="0" pageNumber="185">B</emphasis>
frontal view showing antennules, antennae and orbits
<emphasis bold="true" pageId="0" pageNumber="185">C</emphasis>
lateral view of cephalothorax
<emphasis bold="true" pageId="0" pageNumber="185">D</emphasis>
pterygostomial region, buccal cavity, third maxillipeds and anterior part of sternopleonal cavity
<emphasis bold="true" pageId="0" pageNumber="185">E</emphasis>
left third maxilliped
<emphasis bold="true" pageId="0" pageNumber="185">F</emphasis>
outer view of left chela
<emphasis bold="true" pageId="0" pageNumber="185">G</emphasis>
right fourth ambulatory leg.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="185">
Chelipeds subequal, closely covered with tubercles of varying sizes and shapes; merus trigonal in cross-section, surface covered with closely-packed low, rounded granules, posterior margin with 3 or 4 large conical tubercles, inner margin with low, broad tubercles and granules; carpus rounded, surface covered with closely-packed low, rounded granules, inner distal angle with low tooth; palm short, stout, upper and outer surfaces with low, conical tubercles, relatively densely packed, inner surface prominently swollen, covered with large, rounded granules; fingers short, shorter than palm, dorsal margin of dactylus with row of low, short granules, outer surface with low, flattened granules, cutting edge with small sharp denticles, pollex relatively broader, outer surface with small rounded granules, submarginal ones arranged in approximate rows, cutting edge with low denticles (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Orientotlos iishibai Sakai, 1980 A holotype female (7.0 by 5.5 mm) (repository unknown), Japan [after Sakai 1980: text-fig. 1] B color in life, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan." figureDoi="10.3897/zookeys.1053.67326.figure1" httpUri="https://binary.pensoft.net/fig/572659" pageId="0" pageNumber="185">1</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A overall dorsal view B overall ventral view C, D dorsal view of carapace from different angles." figureDoi="10.3897/zookeys.1053.67326.figure2" httpUri="https://binary.pensoft.net/fig/572660" pageId="0" pageNumber="185">2A, B</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A frontal view of cephalothorax B frontal view showing antennules, antennae and orbits C lateral view of cephalothorax D pterygostomial region, buccal cavity, third maxillipeds and anterior part of sternopleonal cavity E left third maxilliped F outer view of left chela G right fourth ambulatory leg." figureDoi="10.3897/zookeys.1053.67326.figure3" httpUri="https://binary.pensoft.net/fig/572661" pageId="0" pageNumber="185">3A, F</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="185">
Ambulatory legs short, decreasing in size posteriorly; merus, carpus and propodus covered with slender and boletiform tubercles along upper and lower margins, outer surface with large, rounded granules and tubercles; dactylo-propodal lock present; dactylus slender, glabrous, lined with granules, tip curved, corneous (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A overall dorsal view B overall ventral view C, D dorsal view of carapace from different angles." figureDoi="10.3897/zookeys.1053.67326.figure2" httpUri="https://binary.pensoft.net/fig/572660" pageId="0" pageNumber="185">2A, B</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A frontal view of cephalothorax B frontal view showing antennules, antennae and orbits C lateral view of cephalothorax D pterygostomial region, buccal cavity, third maxillipeds and anterior part of sternopleonal cavity E left third maxilliped F outer view of left chela G right fourth ambulatory leg." figureDoi="10.3897/zookeys.1053.67326.figure3" httpUri="https://binary.pensoft.net/fig/572661" pageId="0" pageNumber="185">3G</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="185">
Anterior thoracic sternites (1-4) strongly compressed; sternites 1 and 2 completely fused to form small plate; sternites 3 and 4 appearing fused but vaguely demarcated by arrangement of granules, surface of sternite 3 with numerous boletiform tubercles, larger on lateral surfaces; surfaces of sternites 4-7 with flattened and low, rounded tubercles (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A overall dorsal view B overall ventral view C, D dorsal view of carapace from different angles." figureDoi="10.3897/zookeys.1053.67326.figure2" httpUri="https://binary.pensoft.net/fig/572660" pageId="0" pageNumber="185">2B</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A intestinal region, posterior carapace lobes and pleonal somites 2 - 6 B posterior carapace lobes, pleonal somites 2 - 6, and telson C pleonal somites 4 - 6 and telson D anterior thoracic sternites and sternopleonal cavity E sternopleonal cavity and vulvae." figureDoi="10.3897/zookeys.1053.67326.figure4" httpUri="https://binary.pensoft.net/fig/572662" pageId="0" pageNumber="185">4A-D</figureCitation>
). Sternite 4 forming keel around distal part of sternopleonal cavity, occupying anterior third of overall cavity (Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A frontal view of cephalothorax B frontal view showing antennules, antennae and orbits C lateral view of cephalothorax D pterygostomial region, buccal cavity, third maxillipeds and anterior part of sternopleonal cavity E left third maxilliped F outer view of left chela G right fourth ambulatory leg." figureDoi="10.3897/zookeys.1053.67326.figure3" httpUri="https://binary.pensoft.net/fig/572661" pageId="0" pageNumber="185">3D</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A intestinal region, posterior carapace lobes and pleonal somites 2 - 6 B posterior carapace lobes, pleonal somites 2 - 6, and telson C pleonal somites 4 - 6 and telson D anterior thoracic sternites and sternopleonal cavity E sternopleonal cavity and vulvae." figureDoi="10.3897/zookeys.1053.67326.figure4" httpUri="https://binary.pensoft.net/fig/572662" pageId="0" pageNumber="185">4D</figureCitation>
). Sternopleonal cavity subovate, deeply excavated, glabrous, reaching buccal cavity anteriorly; sutures between sternites 4/5, 5/6, 6/7, and 7/8 interrupted medially (Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A frontal view of cephalothorax B frontal view showing antennules, antennae and orbits C lateral view of cephalothorax D pterygostomial region, buccal cavity, third maxillipeds and anterior part of sternopleonal cavity E left third maxilliped F outer view of left chela G right fourth ambulatory leg." figureDoi="10.3897/zookeys.1053.67326.figure3" httpUri="https://binary.pensoft.net/fig/572661" pageId="0" pageNumber="185">3D</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A intestinal region, posterior carapace lobes and pleonal somites 2 - 6 B posterior carapace lobes, pleonal somites 2 - 6, and telson C pleonal somites 4 - 6 and telson D anterior thoracic sternites and sternopleonal cavity E sternopleonal cavity and vulvae." figureDoi="10.3897/zookeys.1053.67326.figure4" httpUri="https://binary.pensoft.net/fig/572662" pageId="0" pageNumber="185">4D, E</figureCitation>
). Vulvae relatively small, round, positioned distinctly apart, with opening directed laterally (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A intestinal region, posterior carapace lobes and pleonal somites 2 - 6 B posterior carapace lobes, pleonal somites 2 - 6, and telson C pleonal somites 4 - 6 and telson D anterior thoracic sternites and sternopleonal cavity E sternopleonal cavity and vulvae." figureDoi="10.3897/zookeys.1053.67326.figure4" httpUri="https://binary.pensoft.net/fig/572662" pageId="0" pageNumber="185">4E</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1053.67326.figure4" httpUri="https://binary.pensoft.net/fig/572662" pageId="0" pageNumber="185" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="185">
<emphasis bold="true" pageId="0" pageNumber="185">Figure 4.</emphasis>
<taxonomicName authorityName="Sakai" authorityYear="1980" class="Malacostraca" family="Leucosiidae" genus="Orientotlos" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orientotlos iishibai" order="Decapoda" pageId="0" pageNumber="185" phylum="Arthropoda" rank="species" species="iishibai">
<emphasis italics="true" pageId="0" pageNumber="185">Orientotlos iishibai</emphasis>
</taxonomicName>
Sakai, 1980, female (7.8
<normalizedToken originalValue="×">x</normalizedToken>
6.5 mm) (NTOU B00125), Taiwan
<emphasis bold="true" pageId="0" pageNumber="185">A</emphasis>
intestinal region, posterior carapace lobes and pleonal somites 2-6
<emphasis bold="true" pageId="0" pageNumber="185">B</emphasis>
posterior carapace lobes, pleonal somites 2-6, and telson
<emphasis bold="true" pageId="0" pageNumber="185">C</emphasis>
pleonal somites 4-6 and telson
<emphasis bold="true" pageId="0" pageNumber="185">D</emphasis>
anterior thoracic sternites and sternopleonal cavity
<emphasis bold="true" pageId="0" pageNumber="185">E</emphasis>
sternopleonal cavity and vulvae.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="185">
Pleon ovate, shield-like, entirely covered with closely packed, rounded tubercles of varying sizes, larger ones partially coalescing; somite 1 very narrow, girdle-like, not visible when pleon closed, free; somite 2 narrow, semicircular, free; somites 3-6 completely fused, sutures not clearly visible in dorsal view, distinct in ventral view, margins lined with rounded granules; telson narrowly triangular, with proximal part broad, distal part linguiform, lateral margins deeply concave (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A overall dorsal view B overall ventral view C, D dorsal view of carapace from different angles." figureDoi="10.3897/zookeys.1053.67326.figure2" httpUri="https://binary.pensoft.net/fig/572660" pageId="0" pageNumber="185">2D</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A intestinal region, posterior carapace lobes and pleonal somites 2 - 6 B posterior carapace lobes, pleonal somites 2 - 6, and telson C pleonal somites 4 - 6 and telson D anterior thoracic sternites and sternopleonal cavity E sternopleonal cavity and vulvae." figureDoi="10.3897/zookeys.1053.67326.figure4" httpUri="https://binary.pensoft.net/fig/572662" pageId="0" pageNumber="185">4A-C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="185" type="remarks">
<paragraph pageId="0" pageNumber="185">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="185">
<bibRefCitation DOI="https://doi.org/10.18353/rcustacea.10.0_73" author="Sakai, T" journalOrPublisher="Researches on Crustacea" pageId="0" pageNumber="185" pagination="73 - 84" refId="B19" refString="Sakai, T, 1980. On new and rare crabs taken from Japanese and Central Pacific waters. Researches on Crustacea 10: 73 - 84, DOI: https://doi.org/10.18353/rcustacea.10.0_73" title="On new and rare crabs taken from Japanese and Central Pacific waters." url="https://doi.org/10.18353/rcustacea.10.0_73" volume="10" year="1980">Sakai (1980</bibRefCitation>
: 74) described the genus and species on the basis of only one 7.0 by 5.5 mm female, and no depth information was indicated with the provenance data. The description is relatively short and only one figure was provided. The whereabouts of the holotype is not known. We have checked the various museums in Japan, Germany, Netherlands and the USA where Sakai is known to have deposited material, but we could not locate the holotype in any of these countries.
<bibRefCitation DOI="https://doi.org/10.18353/rcustacea.10.0_73" author="Sakai, T" journalOrPublisher="Researches on Crustacea" pageId="0" pageNumber="185" pagination="73 - 84" refId="B19" refString="Sakai, T, 1980. On new and rare crabs taken from Japanese and Central Pacific waters. Researches on Crustacea 10: 73 - 84, DOI: https://doi.org/10.18353/rcustacea.10.0_73" title="On new and rare crabs taken from Japanese and Central Pacific waters." url="https://doi.org/10.18353/rcustacea.10.0_73" volume="10" year="1980">Sakai (1980</bibRefCitation>
: 73) commented that the source of his material was from the collection of Eiji Iishiba, a member of the Japanese Carcinological Society, and it is likely that the material was returned to Iishiba after study. Where the holotype is today is not known.
</paragraph>
<paragraph pageId="0" pageNumber="185">
At 7.8 by 6.5 mm, the present ovigerous female from Taiwan is larger than the type but closely resembles it, except that the hepatic lobe is more plate-like (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Orientotlos iishibai Sakai, 1980 A holotype female (7.0 by 5.5 mm) (repository unknown), Japan [after Sakai 1980: text-fig. 1] B color in life, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan." figureDoi="10.3897/zookeys.1053.67326.figure1" httpUri="https://binary.pensoft.net/fig/572659" pageId="0" pageNumber="185">1B</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A overall dorsal view B overall ventral view C, D dorsal view of carapace from different angles." figureDoi="10.3897/zookeys.1053.67326.figure2" httpUri="https://binary.pensoft.net/fig/572660" pageId="0" pageNumber="185">2A, C, D</figureCitation>
) (versus prominently granuliform); and the anterolateral lobes are more dentiform, the margins of each lobe less distinctly granuliform (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Orientotlos iishibai Sakai, 1980 A holotype female (7.0 by 5.5 mm) (repository unknown), Japan [after Sakai 1980: text-fig. 1] B color in life, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan." figureDoi="10.3897/zookeys.1053.67326.figure1" httpUri="https://binary.pensoft.net/fig/572659" pageId="0" pageNumber="185">1B</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A overall dorsal view B overall ventral view C, D dorsal view of carapace from different angles." figureDoi="10.3897/zookeys.1053.67326.figure2" httpUri="https://binary.pensoft.net/fig/572660" pageId="0" pageNumber="185">2A, C, D</figureCitation>
) (versus lobes more rounded and distinctly lined with rounded granules) (cf. Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Orientotlos iishibai Sakai, 1980 A holotype female (7.0 by 5.5 mm) (repository unknown), Japan [after Sakai 1980: text-fig. 1] B color in life, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan." figureDoi="10.3897/zookeys.1053.67326.figure1" httpUri="https://binary.pensoft.net/fig/572659" pageId="0" pageNumber="185">1A</figureCitation>
;
<bibRefCitation DOI="https://doi.org/10.18353/rcustacea.10.0_73" author="Sakai, T" journalOrPublisher="Researches on Crustacea" pageId="0" pageNumber="185" pagination="73 - 84" refId="B19" refString="Sakai, T, 1980. On new and rare crabs taken from Japanese and Central Pacific waters. Researches on Crustacea 10: 73 - 84, DOI: https://doi.org/10.18353/rcustacea.10.0_73" title="On new and rare crabs taken from Japanese and Central Pacific waters." url="https://doi.org/10.18353/rcustacea.10.0_73" volume="10" year="1980">Sakai 1980</bibRefCitation>
: text-fig. 1).
</paragraph>
<paragraph pageId="0" pageNumber="185">
Not considering the generic characters, the carapace of
<taxonomicName authorityName="Sakai" authorityYear="1980" class="Malacostraca" family="Leucosiidae" genus="Orientotlos" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orientotlos iishibai" order="Decapoda" pageId="0" pageNumber="185" phylum="Arthropoda" rank="species" species="iishibai">
<emphasis italics="true" pageId="0" pageNumber="185">Orientotlos iishibai</emphasis>
</taxonomicName>
superficially most closely resembles that of
<taxonomicName baseAuthorityName="Ihle" baseAuthorityYear="1918" class="Malacostraca" family="Leucosiidae" genus="Alox" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Alox ornatum" order="Decapoda" pageId="0" pageNumber="185" phylum="Arthropoda" rank="species" species="ornatum">
<emphasis italics="true" pageId="0" pageNumber="185">Alox ornatum</emphasis>
</taxonomicName>
(Ihle, 1918), which also has many rounded granules on its surface. However, in
<taxonomicName lsidName="A. ornatum" pageId="0" pageNumber="185" rank="species" species="ornatum">
<emphasis italics="true" pageId="0" pageNumber="185">A. ornatum</emphasis>
</taxonomicName>
the granules are arranged very differently, being more closely packed and sometimes coalescing (cf.
<bibRefCitation author="Tan, CGS" editor="Richer de Forges, B" journalOrPublisher="2. ORSTOM, 1995" pageId="0" pageNumber="185" pagination="101 - 205" refId="B22" refString="Tan, CGS, Ng, PKL, 1996. A revision of the IndoPacific genus Oreophorus Rueppell, 1830 (Crustacea: Decapoda: Brachyura: Leucosiidae). In: Richer de Forges, B, Ed., Les fonds meubles des lagons de NouvelleCaledonie (Sedimentologie, Benthos), Etudes &amp; Theses Vol. 2. ORSTOM, 1995: 101 - 205" title="A revision of the IndoPacific genus Oreophorus Rueppell, 1830 (Crustacea: Decapoda: Brachyura: Leucosiidae)." volumeTitle="Les fonds meubles des lagons de NouvelleCaledonie (Sedimentologie, Benthos), Etudes &amp; Theses Vol." year="1996">Tan and Ng 1996</bibRefCitation>
: pl. 5A;
<bibRefCitation author="Galil, BS" journalOrPublisher="Raffles Bulletin of Zoology, Supplement" pageId="0" pageNumber="185" pagination="79 - 94" refId="B7" refString="Galil, BS, Ng, PKL, 2007. Leucosiid crabs from Panglao, Philippines, with descriptions of three new species (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16: 79 - 94" title="Leucosiid crabs from Panglao, Philippines, with descriptions of three new species (Crustacea: Decapoda: Brachyura)." volume="16" year="2007">Galil and Ng 2007</bibRefCitation>
: fig. 1C;
<bibRefCitation author="Galil, BS" editor="Corbari, L" journalOrPublisher="Tropical Deep-Sea Benthos, Paris" pageId="0" pageNumber="185" pagination="403 - 444" refId="B8" refString="Galil, BS, Ng, PKL, 2020. New and rare leucosiid crabs (Crustacea: Decapoda: Brachyura) from Papua New Guinea. In: Corbari, L, Ahyong, ST, Chan, T-Y, Eds., Deep-Sea Crustaceans from Papua New Guinea. Tropical Deep-Sea Benthos, Paris 31: 403 - 444" title="New and rare leucosiid crabs (Crustacea: Decapoda: Brachyura) from Papua New Guinea." volume="31" volumeTitle="Deep-Sea Crustaceans from Papua New Guinea." year="2020">Galil and Ng 2020</bibRefCitation>
: fig. 1A).
</paragraph>
<paragraph pageId="0" pageNumber="185">
No males of
<taxonomicName authorityName="Sakai" authorityYear="1980" class="Malacostraca" family="Leucosiidae" genus="Orientotlos" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orientotlos iishibai" order="Decapoda" pageId="0" pageNumber="185" phylum="Arthropoda" rank="species" species="iishibai">
<emphasis italics="true" pageId="0" pageNumber="185">Orientotlos iishibai</emphasis>
</taxonomicName>
have been collected, so the important characters of the male pleon and gonopods are unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="185" type="biology_ecology">
<paragraph pageId="0" pageNumber="185">Biology.</paragraph>
<paragraph pageId="0" pageNumber="185">
Station CP4210 is a hard bottom habitat and the trawl net was seriously damaged, though its cod end was intact. That same haul contained many sponges and crinoids and their associated fauna, including a new species of stenopodid shrimp of the genus
<taxonomicName authorityName="Holthuis" authorityYear="1946" class="Malacostraca" family="Stenopodidae" genus="Odontozona" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Odontozona" order="Decapoda" pageId="0" pageNumber="185" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="185">Odontozona</emphasis>
</taxonomicName>
Holthuis, 1946, often associated with sponges (Chen and Chan, in press). The hard substrate may explain the rarity of
<taxonomicName class="Malacostraca" family="Leucosiidae" genus="Orientotlos" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orientotlos iishibaii" order="Decapoda" pageId="0" pageNumber="185" phylum="Arthropoda" rank="species" species="iishibaii">
<emphasis italics="true" pageId="0" pageNumber="185">Orientotlos iishibaii</emphasis>
</taxonomicName>
in collections, as this habitat is very hard to sample, especially in deeper waters (see
<bibRefCitation DOI="https://doi.org/10.1007/s10531-009-9744-x" author="Mendoza, JCE" journalOrPublisher="Biological Conservation" pageId="0" pageNumber="185" pagination="575 - 586" refId="B13" refString="Mendoza, JCE, Naruse, T, Tan, SH, Chan, T-Y, Richer De Forges, B, Ng, PKL, 2010. Case studies on decapod crustaceans from the Philippines reveal deep, steep underwater slopes as prime habitats for 'rare' species. Biological Conservation 19: 575 - 586, DOI: https://doi.org/10.1007/s10531-009-9744-x" title="Case studies on decapod crustaceans from the Philippines reveal deep, steep underwater slopes as prime habitats for ' rare' species." url="https://doi.org/10.1007/s10531-009-9744-x" volume="19" year="2010">Mendoza et al. 2010</bibRefCitation>
). In fact, the rare deep-water western Pacific leucosiid
<taxonomicName authorityName="P.K.L.Ng &amp; Richer de Forges" authorityYear="2007" class="Malacostraca" family="Leucosiidae" genus="Galilia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Galilia" order="Decapoda" pageId="0" pageNumber="185" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="185">Galilia</emphasis>
</taxonomicName>
Ng &amp; Richer de Forges, 2007 originates from a similar habitat (see
<bibRefCitation author="Ng, PKL" journalOrPublisher="Zootaxa" pageId="0" pageNumber="185" pagination="15 - 24" refId="B16" refString="Ng, PKL, Richer de Forges, B, 2007. A new genus and new species of leucosiid crab from New Caledonia, with a note on the validity of Tanaoa serenei (Richer de Forges, 1983) (Crustacea: Decapoda: Brachyura). Zootaxa 1662: 15 - 24" title="A new genus and new species of leucosiid crab from New Caledonia, with a note on the validity of Tanaoa serenei (Richer de Forges, 1983) (Crustacea: Decapoda: Brachyura)." volume="1662" year="2007">Ng and Richer de Forges 2007</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3764.3.3" author="Komai, T" journalOrPublisher="Zootaxa" pageId="0" pageNumber="185" pagination="279 - 316" refId="B12" refString="Komai, T, Tsuchida, S, 2014. Deep-sea decapod crustaceans (Caridea, Polychelida, Anomura and Brachyura) collected from the Nikko Seamounts, Mariana Arc, using a remotely operated vehicle &quot;Hyper-Dolphin&quot;. Zootaxa 3764: 279 - 316, DOI: https://doi.org/10.11646/zootaxa.3764.3.3" title="Deep-sea decapod crustaceans (Caridea, Polychelida, Anomura and Brachyura) collected from the Nikko Seamounts, Mariana Arc, using a remotely operated vehicle &quot; Hyper-Dolphin &quot;." url="https://doi.org/10.11646/zootaxa.3764.3.3" volume="3764" year="2014">Komai and Tsuchida 2014</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4052.1.7" author="Shih, Y-J" journalOrPublisher="Zootaxa" pageId="0" pageNumber="185" pagination="127 - 134" refId="B21" refString="Shih, Y-J, Ho, P-H, Chan, T-Y, 2015. Leucosiid crabs (Crustacea: Decapoda: Brachyura) from Taiwan, with three new records. Zootaxa 4052: 127 - 134, DOI: https://doi.org/10.11646/zootaxa.4052.1.7" title="Leucosiid crabs (Crustacea: Decapoda: Brachyura) from Taiwan, with three new records." url="https://doi.org/10.11646/zootaxa.4052.1.7" volume="4052" year="2015">Shih et al. 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>