503 lines
54 KiB
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503 lines
54 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.884.39322" ID-GBIF-Dataset="8bef7f29-d28a-4701-87a4-465512797381" ID-PMC="PMC6834722" ID-Pensoft-Pub="1313-2970-884-69" ID-Pensoft-UUID="594C3C225AA7588F8CEA3ECD7803687A" ID-PubMed="31723327" ModsDocID="1313-2970-884-69" checkinTime="1572509033109" checkinUser="pensoft" docAuthor="Silva, Fernando A. B. & Genier, Francois" docDate="2019" docId="656E1DFA1C075C7BA80D56082C7957BE" docLanguage="en" docName="ZooKeys 884: 69-80" docOrigin="ZooKeys 884" docSource="http://dx.doi.org/10.3897/zookeys.884.39322" docTitle="Scybalocanthon ashei Silva & Genier, 2019, sp. nov." docType="treatment" docVersion="5" id="594C3C225AA7588F8CEA3ECD7803687A" lastPageNumber="69" masterDocId="594C3C225AA7588F8CEA3ECD7803687A" masterDocTitle="A new Peruvian species of Scybalocanthon Martinez, 1948 (Coleoptera, Scarabaeidae, Scarabaeinae, Deltochilini) and some remarkable intrapopulational variation in the endophallus of S. pinopterus (Kirsch, 1873)" masterLastPageNumber="80" masterPageNumber="69" pageNumber="69" updateTime="1668167995861" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>A new Peruvian species of Scybalocanthon Martinez, 1948 (Coleoptera, Scarabaeidae, Scarabaeinae, Deltochilini) and some remarkable intrapopulational variation in the endophallus of S. pinopterus (Kirsch, 1873)</mods:title>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Silva, Fernando A. B.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Genier, Francois</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>884</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>69</mods:start>
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<mods:end>80</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.884.39322</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.884.39322</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-884-69</mods:identifier>
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<mods:identifier type="Pensoft-UUID">594C3C225AA7588F8CEA3ECD7803687A</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="159932893" LSID="urn:lsid:zoobank.org:act:9E58A29A-B2F0-4893-8AF4-5EAA15821193" httpUri="http://treatment.plazi.org/id/656E1DFA1C075C7BA80D56082C7957BE" lastPageNumber="69" pageId="0" pageNumber="69">
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<subSubSection pageId="0" pageNumber="69" type="nomenclature">
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<paragraph pageId="0" pageNumber="69">
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<taxonomicName LSID="656E1DFA-1C07-5C7B-A80D-56082C7957BE" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scybalocanthon ashei" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="ashei">Scybalocanthon ashei</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="69">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Scybalocanthon ashei sp. nov. A Holotype (CMNC), dorsal view B holotype (CMNC), ventral view C aedeagus (right side) D aedeagus (left side) E set of bristles F fronto-lateral peripheral (FLP) sclerite (left), and Complex of axial and subaxial (A + SA) sclerites (right) G Superior right peripheral (SRP) sclerite." figureDoi="10.3897/zookeys.884.39322.figure1" httpUri="https://binary.pensoft.net/fig/351073" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure1">
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Figs 1
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<normalizedToken originalValue="A–G">A-G</normalizedToken>
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</figureCitation>
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,
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Known distributions of species of Scybalocanthon. A S. adisi and S. aereus B S. acrianus, S. arnaudi, S. chamorroi, S. kaestneri and S. pinopterus C S. ashei sp. nov., S. federicoescobari, S. martinezi, S. papaxibe, S. pygidialis, and S. uniplagiatus." figureDoi="10.3897/zookeys.884.39322.figure5" httpUri="https://binary.pensoft.net/fig/351077" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure5">5C</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="69" type="materials_examined">
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<paragraph pageId="0" pageNumber="69">Material studied.</paragraph>
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<paragraph pageId="0" pageNumber="69">
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<emphasis bold="true" pageId="0" pageNumber="69">
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<emphasis italics="true" pageId="0" pageNumber="69">Holotype</emphasis>
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.
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</emphasis>
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PERU: MADRE DE DIOS, Reserva Cuzco Amazonica, 15 km NE Puerto Maldonado,
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<geoCoordinate degrees="69" direction="west" minutes="03" orientation="longitude" precision="925" value="-69.05">69°03'W</geoCoordinate>
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,
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<geoCoordinate degrees="12" direction="south" minutes="33" orientation="latitude" precision="925" value="-12.55">12°33'S</geoCoordinate>
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, 200m, 4.VII.1989, Ashe and Leschen legs (1♂ CMNC).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="69">Paratypes</emphasis>
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[5♂ and 7♀]. PERU: MADRE DE DIOS, Reserva Cuzco Amazonica, 15 km NE Puerto Maldonado,
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<geoCoordinate degrees="69" direction="west" minutes="03" orientation="longitude" precision="925" value="-69.05">69°03'W</geoCoordinate>
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,
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<geoCoordinate degrees="12" direction="south" minutes="33" orientation="latitude" precision="925" value="-12.55">12°33'S</geoCoordinate>
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, 200m, 7.VII.1989, Ashe and Leschen legs (1♀ CMNC); Reserva Cuzco Amazonica, 15 km NE Puerto Maldonado,
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<geoCoordinate degrees="69" direction="west" minutes="03" orientation="longitude" precision="925" value="-69.05">69°03'W</geoCoordinate>
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,
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<geoCoordinate degrees="12" direction="south" minutes="33" orientation="latitude" precision="925" value="-12.55">12°33'S</geoCoordinate>
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, 200m, 13.VII.1989, Ashe and Leschen legs (1♀ CMNC); Reserva Cuzco Amazonica, 15 km NE Puerto Maldonado,
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<geoCoordinate degrees="69" direction="west" minutes="03" orientation="longitude" precision="925" value="-69.05">69°03'W</geoCoordinate>
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,
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<geoCoordinate degrees="12" direction="south" minutes="33" orientation="latitude" precision="925" value="-12.55">12°33'S</geoCoordinate>
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, 200m, 17.VI.1989, Ashe and Leschen legs (1♀ CMNC); Reserva Cuzco Amazonica, 15 km NE Puerto Maldonado,
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<geoCoordinate degrees="69" direction="west" minutes="03" orientation="longitude" precision="925" value="-69.05">69°03'W</geoCoordinate>
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,
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<geoCoordinate degrees="12" direction="south" minutes="33" orientation="latitude" precision="925" value="-12.55">12°33'S</geoCoordinate>
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, 200m, 24.VI.1989, Ashe and Leschen legs (1♀ CMNC); Reserva Cuzco Amazonica, 15 km NE Puerto Maldonado,
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<geoCoordinate degrees="69" direction="west" minutes="03" orientation="longitude" precision="925" value="-69.05">69°03'W</geoCoordinate>
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,
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<geoCoordinate degrees="12" direction="south" minutes="33" orientation="latitude" precision="925" value="-12.55">12°33'S</geoCoordinate>
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, 200m, 26.VI.1989, Ashe and Leschen legs (1♂ CMNC, 1♂ MZUFPA, 1♂ CEMT); Reserva Cuzco Amazonica, 15 km NE Puerto Maldonado,
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<geoCoordinate degrees="69" direction="west" minutes="03" orientation="longitude" precision="925" value="-69.05">69°03'W</geoCoordinate>
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,
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<geoCoordinate degrees="12" direction="south" minutes="33" orientation="latitude" precision="925" value="-12.55">12°33'S</geoCoordinate>
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, 200m, 20.VI.1989, Ashe and Leschen legs (1♂ CMNC); Reserva Cuzco Amazonica, 15 km NE Puerto Maldonado,
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<geoCoordinate degrees="69" direction="west" minutes="03" orientation="longitude" precision="925" value="-69.05">69°03'W</geoCoordinate>
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,
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<geoCoordinate degrees="12" direction="south" minutes="33" orientation="latitude" precision="925" value="-12.55">12°33'S</geoCoordinate>
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, 200m, 30.VI.1989, Ashe and Leschen legs (1♀ CMNC); Parque Nacional del
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<normalizedToken originalValue="Manú">Manu</normalizedToken>
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, 15-30.VIII.1986, A. Forsyth leg. (1♂ and 1♀ CMNC, 1♀ MZUFPA).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="69" type="diagnosis">
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<paragraph pageId="0" pageNumber="69">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="69">
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Specimens of
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<taxonomicName class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scybalocanthon ashei" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="ashei">
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<emphasis italics="true" pageId="0" pageNumber="69">Scybalocanthon ashei</emphasis>
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</taxonomicName>
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sp. nov. (
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Scybalocanthon ashei sp. nov. A Holotype (CMNC), dorsal view B holotype (CMNC), ventral view C aedeagus (right side) D aedeagus (left side) E set of bristles F fronto-lateral peripheral (FLP) sclerite (left), and Complex of axial and subaxial (A + SA) sclerites (right) G Superior right peripheral (SRP) sclerite." figureDoi="10.3897/zookeys.884.39322.figure1" httpUri="https://binary.pensoft.net/fig/351073" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure1">Fig. 1A</figureCitation>
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) are similar to those of
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<taxonomicName lsidName="S. arnaudi" pageId="0" pageNumber="69" rank="species" species="arnaudi">
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<emphasis italics="true" pageId="0" pageNumber="69">S. arnaudi</emphasis>
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</taxonomicName>
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(
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Species of Scybalocanthon, dorsal views. A S. acrianus B S. adisi C S. aereus D S. arnaudi E S. chamorroi F S. federicoescobari G S. kaestneri (specimen from Pastaza, Ecuador) H S. kaestneri (specimen from Napo, Ecuador) I S. martinezi J S. papaxibe K S. pinopterus (specimen from Madre de Dios, Peru) L S. pinopterus (specimen from Tingo Maria, Huanuco, Peru) M S. pinopterus (specimen from Orellana, Ecuador) N S. pygidialis O S. uniplagiatus, Scale bars: 2 mm." figureDoi="10.3897/zookeys.884.39322.figure2" httpUri="https://binary.pensoft.net/fig/351074" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure2">Fig. 2D</figureCitation>
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),
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<taxonomicName lsidName="S. federicoescobari" pageId="0" pageNumber="69" rank="species" species="federicoescobari">
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<emphasis italics="true" pageId="0" pageNumber="69">S. federicoescobari</emphasis>
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</taxonomicName>
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(
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Species of Scybalocanthon, dorsal views. A S. acrianus B S. adisi C S. aereus D S. arnaudi E S. chamorroi F S. federicoescobari G S. kaestneri (specimen from Pastaza, Ecuador) H S. kaestneri (specimen from Napo, Ecuador) I S. martinezi J S. papaxibe K S. pinopterus (specimen from Madre de Dios, Peru) L S. pinopterus (specimen from Tingo Maria, Huanuco, Peru) M S. pinopterus (specimen from Orellana, Ecuador) N S. pygidialis O S. uniplagiatus, Scale bars: 2 mm." figureDoi="10.3897/zookeys.884.39322.figure2" httpUri="https://binary.pensoft.net/fig/351074" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure2">Fig. 2F</figureCitation>
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),
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<taxonomicName lsidName="S. martinezi" pageId="0" pageNumber="69" rank="species" species="martinezi">
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<emphasis italics="true" pageId="0" pageNumber="69">S. martinezi</emphasis>
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</taxonomicName>
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(
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Species of Scybalocanthon, dorsal views. A S. acrianus B S. adisi C S. aereus D S. arnaudi E S. chamorroi F S. federicoescobari G S. kaestneri (specimen from Pastaza, Ecuador) H S. kaestneri (specimen from Napo, Ecuador) I S. martinezi J S. papaxibe K S. pinopterus (specimen from Madre de Dios, Peru) L S. pinopterus (specimen from Tingo Maria, Huanuco, Peru) M S. pinopterus (specimen from Orellana, Ecuador) N S. pygidialis O S. uniplagiatus, Scale bars: 2 mm." figureDoi="10.3897/zookeys.884.39322.figure2" httpUri="https://binary.pensoft.net/fig/351074" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure2">Fig. 2I</figureCitation>
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),
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<taxonomicName lsidName="S. papaxibe" pageId="0" pageNumber="69" rank="species" species="papaxibe">
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<emphasis italics="true" pageId="0" pageNumber="69">S. papaxibe</emphasis>
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</taxonomicName>
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(
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Species of Scybalocanthon, dorsal views. A S. acrianus B S. adisi C S. aereus D S. arnaudi E S. chamorroi F S. federicoescobari G S. kaestneri (specimen from Pastaza, Ecuador) H S. kaestneri (specimen from Napo, Ecuador) I S. martinezi J S. papaxibe K S. pinopterus (specimen from Madre de Dios, Peru) L S. pinopterus (specimen from Tingo Maria, Huanuco, Peru) M S. pinopterus (specimen from Orellana, Ecuador) N S. pygidialis O S. uniplagiatus, Scale bars: 2 mm." figureDoi="10.3897/zookeys.884.39322.figure2" httpUri="https://binary.pensoft.net/fig/351074" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure2">Fig. 2J</figureCitation>
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),
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<taxonomicName lsidName="S. pinopterus" pageId="0" pageNumber="69" rank="species" species="pinopterus">
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<emphasis italics="true" pageId="0" pageNumber="69">S. pinopterus</emphasis>
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</taxonomicName>
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(
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Species of Scybalocanthon, dorsal views. A S. acrianus B S. adisi C S. aereus D S. arnaudi E S. chamorroi F S. federicoescobari G S. kaestneri (specimen from Pastaza, Ecuador) H S. kaestneri (specimen from Napo, Ecuador) I S. martinezi J S. papaxibe K S. pinopterus (specimen from Madre de Dios, Peru) L S. pinopterus (specimen from Tingo Maria, Huanuco, Peru) M S. pinopterus (specimen from Orellana, Ecuador) N S. pygidialis O S. uniplagiatus, Scale bars: 2 mm." figureDoi="10.3897/zookeys.884.39322.figure2" httpUri="https://binary.pensoft.net/fig/351074" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure2">
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Fig. 2
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<normalizedToken originalValue="K–M">K-M</normalizedToken>
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</figureCitation>
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), and
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<taxonomicName lsidName="S. uniplagiatus" pageId="0" pageNumber="69" rank="species" species="uniplagiatus">
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<emphasis italics="true" pageId="0" pageNumber="69">S. uniplagiatus</emphasis>
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</taxonomicName>
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(
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Species of Scybalocanthon, dorsal views. A S. acrianus B S. adisi C S. aereus D S. arnaudi E S. chamorroi F S. federicoescobari G S. kaestneri (specimen from Pastaza, Ecuador) H S. kaestneri (specimen from Napo, Ecuador) I S. martinezi J S. papaxibe K S. pinopterus (specimen from Madre de Dios, Peru) L S. pinopterus (specimen from Tingo Maria, Huanuco, Peru) M S. pinopterus (specimen from Orellana, Ecuador) N S. pygidialis O S. uniplagiatus, Scale bars: 2 mm." figureDoi="10.3897/zookeys.884.39322.figure2" httpUri="https://binary.pensoft.net/fig/351074" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure2">Fig. 2O</figureCitation>
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) in having the pronotum uniformly colored, or with one elliptical spot on the central portion; femora almost completely yellow or brown, with black spots only on the tips (
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Scybalocanthon ashei sp. nov. A Holotype (CMNC), dorsal view B holotype (CMNC), ventral view C aedeagus (right side) D aedeagus (left side) E set of bristles F fronto-lateral peripheral (FLP) sclerite (left), and Complex of axial and subaxial (A + SA) sclerites (right) G Superior right peripheral (SRP) sclerite." figureDoi="10.3897/zookeys.884.39322.figure1" httpUri="https://binary.pensoft.net/fig/351073" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure1">Fig. 1B</figureCitation>
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); eighth elytral stria with thin carina anteriorly (see
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<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4629.3.1" author="Silva, FAB" journalOrPublisher="Zootaxa" pageId="0" pageNumber="69" pagination="301 - 341" refId="B3" refString="Silva, FAB, Valois, M, 2019. A taxonomic revision of the genus Scybalocanthon Martinez, 1948 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Zootaxa 4629: 301 - 341" title="A taxonomic revision of the genus Scybalocanthon Martinez, 1948 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini)." url="https://doi.org/10.11646/zootaxa.4629.3.1" volume="4629" year="2019">Silva and Valois 2019</bibRefCitation>
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, fig. 3D); endophallus with bristles right beside the FLP sclerite (
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Scybalocanthon ashei sp. nov. A Holotype (CMNC), dorsal view B holotype (CMNC), ventral view C aedeagus (right side) D aedeagus (left side) E set of bristles F fronto-lateral peripheral (FLP) sclerite (left), and Complex of axial and subaxial (A + SA) sclerites (right) G Superior right peripheral (SRP) sclerite." figureDoi="10.3897/zookeys.884.39322.figure1" httpUri="https://binary.pensoft.net/fig/351073" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure1">Fig. 1E</figureCitation>
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); and additional sclerite (AS) absent. They can be distinguished from those of
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<taxonomicName lsidName="S. arnaudi" pageId="0" pageNumber="69" rank="species" species="arnaudi">
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<emphasis italics="true" pageId="0" pageNumber="69">S. arnaudi</emphasis>
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</taxonomicName>
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and
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<taxonomicName lsidName="S. pinopterus" pageId="0" pageNumber="69" rank="species" species="pinopterus">
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<emphasis italics="true" pageId="0" pageNumber="69">S. pinopterus</emphasis>
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</taxonomicName>
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, however, by the strongly asymmetrical parameres; left paramere with acute projection on dorsal margin and bilobate excavation on ventral margin (
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Scybalocanthon ashei sp. nov. A Holotype (CMNC), dorsal view B holotype (CMNC), ventral view C aedeagus (right side) D aedeagus (left side) E set of bristles F fronto-lateral peripheral (FLP) sclerite (left), and Complex of axial and subaxial (A + SA) sclerites (right) G Superior right peripheral (SRP) sclerite." figureDoi="10.3897/zookeys.884.39322.figure1" httpUri="https://binary.pensoft.net/fig/351073" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure1">Fig. 1C, D</figureCitation>
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) (other species with slightly asymmetrical parameres, lacking acute projection and bilobate excavation on dorsal and ventral margins (
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Aedeagus, detail of parameres in left and right view. A S. acrianus B S. adisi C S. aereus D S. arnaudi E S. chamorroi F S. federicoescobari G S. kaestneri (specimen from Pastaza, Ecuador) H S. kaestneri (specimen from Napo, Ecuador) I S. martinezi J S. papaxibe K S. pinopterus (specimen from Madre de Dios, Peru) L S. pinopterus (specimen from Tingo Maria, Huanuco, Peru) M S. pinopterus (specimen from Tingo Maria, Huanuco, Peru) N S. pinopterus (specimen from Orellana, Ecuador) O S. pygidialis P S. uniplagiatus, Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.884.39322.figure3" httpUri="https://binary.pensoft.net/fig/351075" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure3">
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Fig. 3D,
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<normalizedToken originalValue="K–N">K-N</normalizedToken>
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</figureCitation>
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); from those of
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<taxonomicName lsidName="S. federicoescobari" pageId="0" pageNumber="69" rank="species" species="federicoescobari">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. federicoescobari</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="S. martinezi" pageId="0" pageNumber="69" rank="species" species="martinezi">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. martinezi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="S. papaxibe" pageId="0" pageNumber="69" rank="species" species="papaxibe">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. papaxibe</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName lsidName="S. uniplagiatus" pageId="0" pageNumber="69" rank="species" species="uniplagiatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. uniplagiatus</emphasis>
|
||
</taxonomicName>
|
||
by the bilobate excavation of the ventral margin of the left paramere wider, extending along two-third of the paramere margin (
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Scybalocanthon ashei sp. nov. A Holotype (CMNC), dorsal view B holotype (CMNC), ventral view C aedeagus (right side) D aedeagus (left side) E set of bristles F fronto-lateral peripheral (FLP) sclerite (left), and Complex of axial and subaxial (A + SA) sclerites (right) G Superior right peripheral (SRP) sclerite." figureDoi="10.3897/zookeys.884.39322.figure1" httpUri="https://binary.pensoft.net/fig/351073" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure1">Fig. 1D</figureCitation>
|
||
) (in
|
||
<taxonomicName lsidName="S. federicoescobari" pageId="0" pageNumber="69" rank="species" species="federicoescobari">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. federicoescobari</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Aedeagus, detail of parameres in left and right view. A S. acrianus B S. adisi C S. aereus D S. arnaudi E S. chamorroi F S. federicoescobari G S. kaestneri (specimen from Pastaza, Ecuador) H S. kaestneri (specimen from Napo, Ecuador) I S. martinezi J S. papaxibe K S. pinopterus (specimen from Madre de Dios, Peru) L S. pinopterus (specimen from Tingo Maria, Huanuco, Peru) M S. pinopterus (specimen from Tingo Maria, Huanuco, Peru) N S. pinopterus (specimen from Orellana, Ecuador) O S. pygidialis P S. uniplagiatus, Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.884.39322.figure3" httpUri="https://binary.pensoft.net/fig/351075" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure3">Fig. 3F</figureCitation>
|
||
) and
|
||
<taxonomicName lsidName="S. papaxibe" pageId="0" pageNumber="69" rank="species" species="papaxibe">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. papaxibe</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Aedeagus, detail of parameres in left and right view. A S. acrianus B S. adisi C S. aereus D S. arnaudi E S. chamorroi F S. federicoescobari G S. kaestneri (specimen from Pastaza, Ecuador) H S. kaestneri (specimen from Napo, Ecuador) I S. martinezi J S. papaxibe K S. pinopterus (specimen from Madre de Dios, Peru) L S. pinopterus (specimen from Tingo Maria, Huanuco, Peru) M S. pinopterus (specimen from Tingo Maria, Huanuco, Peru) N S. pinopterus (specimen from Orellana, Ecuador) O S. pygidialis P S. uniplagiatus, Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.884.39322.figure3" httpUri="https://binary.pensoft.net/fig/351075" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure3">Fig. 3J</figureCitation>
|
||
) the excavation extending along one-fourth of the paramere margin; in
|
||
<taxonomicName lsidName="S. uniplagiatus" pageId="0" pageNumber="69" rank="species" species="uniplagiatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. uniplagiatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Aedeagus, detail of parameres in left and right view. A S. acrianus B S. adisi C S. aereus D S. arnaudi E S. chamorroi F S. federicoescobari G S. kaestneri (specimen from Pastaza, Ecuador) H S. kaestneri (specimen from Napo, Ecuador) I S. martinezi J S. papaxibe K S. pinopterus (specimen from Madre de Dios, Peru) L S. pinopterus (specimen from Tingo Maria, Huanuco, Peru) M S. pinopterus (specimen from Tingo Maria, Huanuco, Peru) N S. pinopterus (specimen from Orellana, Ecuador) O S. pygidialis P S. uniplagiatus, Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.884.39322.figure3" httpUri="https://binary.pensoft.net/fig/351075" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure3">Fig. 3P</figureCitation>
|
||
) the excavation extending along one-third of the paramere margin; in
|
||
<taxonomicName lsidName="S. martinezi" pageId="0" pageNumber="69" rank="species" species="martinezi">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. martinezi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Aedeagus, detail of parameres in left and right view. A S. acrianus B S. adisi C S. aereus D S. arnaudi E S. chamorroi F S. federicoescobari G S. kaestneri (specimen from Pastaza, Ecuador) H S. kaestneri (specimen from Napo, Ecuador) I S. martinezi J S. papaxibe K S. pinopterus (specimen from Madre de Dios, Peru) L S. pinopterus (specimen from Tingo Maria, Huanuco, Peru) M S. pinopterus (specimen from Tingo Maria, Huanuco, Peru) N S. pinopterus (specimen from Orellana, Ecuador) O S. pygidialis P S. uniplagiatus, Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.884.39322.figure3" httpUri="https://binary.pensoft.net/fig/351075" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure3">Fig. 3I</figureCitation>
|
||
) the excavation is deeper, extending about one-half of the paramere margin).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.884.39322.figure1" httpUri="https://binary.pensoft.net/fig/351073" pageId="0" pageNumber="69" start="Figure 1" startId="F1">
|
||
<paragraph pageId="0" pageNumber="69">
|
||
<emphasis bold="true" pageId="0" pageNumber="69">Figure 1.</emphasis>
|
||
<taxonomicName class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scybalocanthon ashei" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="ashei">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Scybalocanthon ashei</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="69">A</emphasis>
|
||
Holotype (CMNC), dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="69">B</emphasis>
|
||
holotype (CMNC), ventral view
|
||
<emphasis bold="true" pageId="0" pageNumber="69">C</emphasis>
|
||
aedeagus (right side)
|
||
<emphasis bold="true" pageId="0" pageNumber="69">D</emphasis>
|
||
aedeagus (left side)
|
||
<emphasis bold="true" pageId="0" pageNumber="69">E</emphasis>
|
||
set of bristles
|
||
<emphasis bold="true" pageId="0" pageNumber="69">F</emphasis>
|
||
fronto-lateral peripheral (FLP) sclerite (left), and Complex of axial and subaxial (A+SA) sclerites (right)
|
||
<emphasis bold="true" pageId="0" pageNumber="69">G</emphasis>
|
||
Superior right peripheral (SRP) sclerite.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.884.39322.figure2" httpUri="https://binary.pensoft.net/fig/351074" pageId="0" pageNumber="69" start="Figure 2" startId="F2">
|
||
<paragraph pageId="0" pageNumber="69">
|
||
<emphasis bold="true" pageId="0" pageNumber="69">Figure 2.</emphasis>
|
||
Species of
|
||
<taxonomicName class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scybalocanthon" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">Scybalocanthon</emphasis>
|
||
</taxonomicName>
|
||
, dorsal views.
|
||
<emphasis bold="true" pageId="0" pageNumber="69">A</emphasis>
|
||
<taxonomicName lsidName="S. acrianus" pageId="0" pageNumber="69" rank="species" species="acrianus">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. acrianus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="69">B</emphasis>
|
||
<taxonomicName lsidName="S. adisi" pageId="0" pageNumber="69" rank="species" species="adisi">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. adisi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="69">C</emphasis>
|
||
<taxonomicName lsidName="S. aereus" pageId="0" pageNumber="69" rank="species" species="aereus">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. aereus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="69">D</emphasis>
|
||
<taxonomicName lsidName="S. arnaudi" pageId="0" pageNumber="69" rank="species" species="arnaudi">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. arnaudi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="69">E</emphasis>
|
||
<taxonomicName lsidName="S. chamorroi" pageId="0" pageNumber="69" rank="species" species="chamorroi">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. chamorroi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="69">F</emphasis>
|
||
<taxonomicName lsidName="S. federicoescobari" pageId="0" pageNumber="69" rank="species" species="federicoescobari">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. federicoescobari</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="69">G</emphasis>
|
||
<taxonomicName lsidName="S. kaestneri" pageId="0" pageNumber="69" rank="species" species="kaestneri">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. kaestneri</emphasis>
|
||
</taxonomicName>
|
||
(specimen from Pastaza, Ecuador)
|
||
<emphasis bold="true" pageId="0" pageNumber="69">H</emphasis>
|
||
<taxonomicName lsidName="S. kaestneri" pageId="0" pageNumber="69" rank="species" species="kaestneri">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. kaestneri</emphasis>
|
||
</taxonomicName>
|
||
(specimen from Napo, Ecuador)
|
||
<emphasis bold="true" pageId="0" pageNumber="69">I</emphasis>
|
||
<taxonomicName lsidName="S. martinezi" pageId="0" pageNumber="69" rank="species" species="martinezi">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. martinezi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="69">J</emphasis>
|
||
<taxonomicName lsidName="S. papaxibe" pageId="0" pageNumber="69" rank="species" species="papaxibe">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. papaxibe</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="69">K</emphasis>
|
||
<taxonomicName lsidName="S. pinopterus" pageId="0" pageNumber="69" rank="species" species="pinopterus">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. pinopterus</emphasis>
|
||
</taxonomicName>
|
||
(specimen from Madre de Dios, Peru)
|
||
<emphasis bold="true" pageId="0" pageNumber="69">L</emphasis>
|
||
<taxonomicName lsidName="S. pinopterus" pageId="0" pageNumber="69" rank="species" species="pinopterus">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. pinopterus</emphasis>
|
||
</taxonomicName>
|
||
(specimen from Tingo Maria, Huanuco, Peru)
|
||
<emphasis bold="true" pageId="0" pageNumber="69">M</emphasis>
|
||
<taxonomicName lsidName="S. pinopterus" pageId="0" pageNumber="69" rank="species" species="pinopterus">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. pinopterus</emphasis>
|
||
</taxonomicName>
|
||
(specimen from Orellana, Ecuador)
|
||
<emphasis bold="true" pageId="0" pageNumber="69">N</emphasis>
|
||
<taxonomicName lsidName="S. pygidialis" pageId="0" pageNumber="69" rank="species" species="pygidialis">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. pygidialis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="69">O</emphasis>
|
||
<taxonomicName lsidName="S. uniplagiatus" pageId="0" pageNumber="69" rank="species" species="uniplagiatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. uniplagiatus</emphasis>
|
||
</taxonomicName>
|
||
, Scale bars: 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.884.39322.figure3" httpUri="https://binary.pensoft.net/fig/351075" pageId="0" pageNumber="69" start="Figure 3" startId="F3">
|
||
<paragraph pageId="0" pageNumber="69">
|
||
<emphasis bold="true" pageId="0" pageNumber="69">Figure 3.</emphasis>
|
||
Aedeagus, detail of parameres in left and right view.
|
||
<emphasis bold="true" pageId="0" pageNumber="69">A</emphasis>
|
||
<taxonomicName lsidName="S. acrianus" pageId="0" pageNumber="69" rank="species" species="acrianus">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. acrianus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="69">B</emphasis>
|
||
<taxonomicName lsidName="S. adisi" pageId="0" pageNumber="69" rank="species" species="adisi">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. adisi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="69">C</emphasis>
|
||
<taxonomicName lsidName="S. aereus" pageId="0" pageNumber="69" rank="species" species="aereus">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. aereus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="69">D</emphasis>
|
||
<taxonomicName lsidName="S. arnaudi" pageId="0" pageNumber="69" rank="species" species="arnaudi">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. arnaudi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="69">E</emphasis>
|
||
<taxonomicName lsidName="S. chamorroi" pageId="0" pageNumber="69" rank="species" species="chamorroi">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. chamorroi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="69">F</emphasis>
|
||
<taxonomicName lsidName="S. federicoescobari" pageId="0" pageNumber="69" rank="species" species="federicoescobari">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. federicoescobari</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="69">G</emphasis>
|
||
<taxonomicName lsidName="S. kaestneri" pageId="0" pageNumber="69" rank="species" species="kaestneri">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. kaestneri</emphasis>
|
||
</taxonomicName>
|
||
(specimen from Pastaza, Ecuador)
|
||
<emphasis bold="true" pageId="0" pageNumber="69">H</emphasis>
|
||
<taxonomicName lsidName="S. kaestneri" pageId="0" pageNumber="69" rank="species" species="kaestneri">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. kaestneri</emphasis>
|
||
</taxonomicName>
|
||
(specimen from Napo, Ecuador)
|
||
<emphasis bold="true" pageId="0" pageNumber="69">I</emphasis>
|
||
<taxonomicName lsidName="S. martinezi" pageId="0" pageNumber="69" rank="species" species="martinezi">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. martinezi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="69">J</emphasis>
|
||
<taxonomicName lsidName="S. papaxibe" pageId="0" pageNumber="69" rank="species" species="papaxibe">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. papaxibe</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="69">K</emphasis>
|
||
<taxonomicName lsidName="S. pinopterus" pageId="0" pageNumber="69" rank="species" species="pinopterus">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. pinopterus</emphasis>
|
||
</taxonomicName>
|
||
(specimen from Madre de Dios, Peru)
|
||
<emphasis bold="true" pageId="0" pageNumber="69">L</emphasis>
|
||
<taxonomicName lsidName="S. pinopterus" pageId="0" pageNumber="69" rank="species" species="pinopterus">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. pinopterus</emphasis>
|
||
</taxonomicName>
|
||
(specimen from Tingo Maria, Huanuco, Peru)
|
||
<emphasis bold="true" pageId="0" pageNumber="69">M</emphasis>
|
||
<taxonomicName lsidName="S. pinopterus" pageId="0" pageNumber="69" rank="species" species="pinopterus">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. pinopterus</emphasis>
|
||
</taxonomicName>
|
||
(specimen from Tingo Maria, Huanuco, Peru)
|
||
<emphasis bold="true" pageId="0" pageNumber="69">N</emphasis>
|
||
<taxonomicName lsidName="S. pinopterus" pageId="0" pageNumber="69" rank="species" species="pinopterus">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. pinopterus</emphasis>
|
||
</taxonomicName>
|
||
(specimen from Orellana, Ecuador)
|
||
<emphasis bold="true" pageId="0" pageNumber="69">O</emphasis>
|
||
<taxonomicName lsidName="S. pygidialis" pageId="0" pageNumber="69" rank="species" species="pygidialis">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. pygidialis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="69">P</emphasis>
|
||
<taxonomicName lsidName="S. uniplagiatus" pageId="0" pageNumber="69" rank="species" species="uniplagiatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. uniplagiatus</emphasis>
|
||
</taxonomicName>
|
||
, Scale bars: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="69" type="description">
|
||
<paragraph pageId="0" pageNumber="69">Description.</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="69">Body</emphasis>
|
||
: Oval-elongated (
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Scybalocanthon ashei sp. nov. A Holotype (CMNC), dorsal view B holotype (CMNC), ventral view C aedeagus (right side) D aedeagus (left side) E set of bristles F fronto-lateral peripheral (FLP) sclerite (left), and Complex of axial and subaxial (A + SA) sclerites (right) G Superior right peripheral (SRP) sclerite." figureDoi="10.3897/zookeys.884.39322.figure1" httpUri="https://binary.pensoft.net/fig/351073" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure1">Fig. 1A</figureCitation>
|
||
). General surface opaque, completely microgranulated. Pronotum with silky sheen.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="69">Color</emphasis>
|
||
: Most of pronotum, hypomera (except internal and posterior margins), metaventrite, metepisterna, abdominal ventrites, pygidium, and middle of femora yellow or light brown. Head, internal and posterior margins of hypomera, prosternum, mesoventrite, mesepisterna, elytra, trochanters, proximal and distal portions of femora, tibiae and tarsomeres dark brown.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="69">Length</emphasis>
|
||
: 8.8-11.5 mm.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="69">Head</emphasis>
|
||
: Dorsal surface with fine microgranulation. Clypeal margin with two small, triangular central teeth. Eye comma-shaped in dorsal view.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="69">Thorax</emphasis>
|
||
: Pronotum twice as wide as long, anterior angles acute, directed forward. Pronotum usually with one brown rounded spot at the anterocentral portion; in some specimens, spot absent. Anterior angles of pronotum approximately 80°. Lateral margin strongly curved outward.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="69">Elytra</emphasis>
|
||
: Striae thin and shiny, punctures conspicuous. Eighth stria with a thin carina anteriorly.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="69">Legs</emphasis>
|
||
: Protibiae with three lateral teeth. Anterior and posterior edge of meso- and metafemora not margined. Mesotibiae smoothly arched toward body. Metatibiae almost straight. First meso- and metatarsomeres short, external margin half the length of second tarsomere, and obliquely truncated apically. Lateral margins of tarsomeres parallel, forming even margin along length of tarsus. Overall shape of meso- and metatarsomeres 2-4 varying from quadrate to rectangular. Dorsal (internal) surface of mesotibiae with dense setae, randomly distributed.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="69">Secondary sexual characters</emphasis>
|
||
: Females can be distinguished from males by the sixth abdominal ventrite longer than in males, and the anterocentral portion of sixth abdominal ventrite more swollen than the posterocentral portion in lateral view (males have, in general, the posterocentral portion more swollen).
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="69">Genital capsule</emphasis>
|
||
: Parameres strongly asymmetrical (
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Scybalocanthon ashei sp. nov. A Holotype (CMNC), dorsal view B holotype (CMNC), ventral view C aedeagus (right side) D aedeagus (left side) E set of bristles F fronto-lateral peripheral (FLP) sclerite (left), and Complex of axial and subaxial (A + SA) sclerites (right) G Superior right peripheral (SRP) sclerite." figureDoi="10.3897/zookeys.884.39322.figure1" httpUri="https://binary.pensoft.net/fig/351073" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure1">Fig. 1C, D</figureCitation>
|
||
). Dorsal margin of right paramere curved inward, apex obliquely truncated. Ventral margin of right paramere with a rounded excavation at the basal portion (
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Scybalocanthon ashei sp. nov. A Holotype (CMNC), dorsal view B holotype (CMNC), ventral view C aedeagus (right side) D aedeagus (left side) E set of bristles F fronto-lateral peripheral (FLP) sclerite (left), and Complex of axial and subaxial (A + SA) sclerites (right) G Superior right peripheral (SRP) sclerite." figureDoi="10.3897/zookeys.884.39322.figure1" httpUri="https://binary.pensoft.net/fig/351073" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure1">Fig. 1C</figureCitation>
|
||
). Dorsal margin of left paramere curved inward from basal to medial portions, medial portion with a short and pointed projection, apex obliquely truncated (
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Scybalocanthon ashei sp. nov. A Holotype (CMNC), dorsal view B holotype (CMNC), ventral view C aedeagus (right side) D aedeagus (left side) E set of bristles F fronto-lateral peripheral (FLP) sclerite (left), and Complex of axial and subaxial (A + SA) sclerites (right) G Superior right peripheral (SRP) sclerite." figureDoi="10.3897/zookeys.884.39322.figure1" httpUri="https://binary.pensoft.net/fig/351073" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure1">Fig. 1D</figureCitation>
|
||
). Ventral margin of left paramere with a bilobate excavation extending along two-third of the paramere margin; apical third obliquely truncated (
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Scybalocanthon ashei sp. nov. A Holotype (CMNC), dorsal view B holotype (CMNC), ventral view C aedeagus (right side) D aedeagus (left side) E set of bristles F fronto-lateral peripheral (FLP) sclerite (left), and Complex of axial and subaxial (A + SA) sclerites (right) G Superior right peripheral (SRP) sclerite." figureDoi="10.3897/zookeys.884.39322.figure1" httpUri="https://binary.pensoft.net/fig/351073" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure1">Fig. 1D</figureCitation>
|
||
).
|
||
<taxonomicName class="Agaricomycetes" family="Phallaceae" genus="Endophallus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Endophallus" order="Phallales" pageId="0" pageNumber="69" phylum="Basidiomycota" rank="genus">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="69">Endophallus</emphasis>
|
||
</taxonomicName>
|
||
: SRP circular, with rounded handle-shaped extension (
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Scybalocanthon ashei sp. nov. A Holotype (CMNC), dorsal view B holotype (CMNC), ventral view C aedeagus (right side) D aedeagus (left side) E set of bristles F fronto-lateral peripheral (FLP) sclerite (left), and Complex of axial and subaxial (A + SA) sclerites (right) G Superior right peripheral (SRP) sclerite." figureDoi="10.3897/zookeys.884.39322.figure1" httpUri="https://binary.pensoft.net/fig/351073" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure1">Fig. 1G</figureCitation>
|
||
). FLP short (
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Scybalocanthon ashei sp. nov. A Holotype (CMNC), dorsal view B holotype (CMNC), ventral view C aedeagus (right side) D aedeagus (left side) E set of bristles F fronto-lateral peripheral (FLP) sclerite (left), and Complex of axial and subaxial (A + SA) sclerites (right) G Superior right peripheral (SRP) sclerite." figureDoi="10.3897/zookeys.884.39322.figure1" httpUri="https://binary.pensoft.net/fig/351073" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure1">Fig. 1F</figureCitation>
|
||
, left), comma-shaped, with three sets of bristles (
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Scybalocanthon ashei sp. nov. A Holotype (CMNC), dorsal view B holotype (CMNC), ventral view C aedeagus (right side) D aedeagus (left side) E set of bristles F fronto-lateral peripheral (FLP) sclerite (left), and Complex of axial and subaxial (A + SA) sclerites (right) G Superior right peripheral (SRP) sclerite." figureDoi="10.3897/zookeys.884.39322.figure1" httpUri="https://binary.pensoft.net/fig/351073" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure1">Fig. 1E</figureCitation>
|
||
) right beside it. A+SA with two superposed and elongate sclerites (
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Scybalocanthon ashei sp. nov. A Holotype (CMNC), dorsal view B holotype (CMNC), ventral view C aedeagus (right side) D aedeagus (left side) E set of bristles F fronto-lateral peripheral (FLP) sclerite (left), and Complex of axial and subaxial (A + SA) sclerites (right) G Superior right peripheral (SRP) sclerite." figureDoi="10.3897/zookeys.884.39322.figure1" httpUri="https://binary.pensoft.net/fig/351073" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure1">Fig. 1F</figureCitation>
|
||
, right).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="69" type="etymology">
|
||
<paragraph pageId="0" pageNumber="69">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">Named in honor of the late James S. Ashe, collector of most of the known specimens.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="69" type="habitat">
|
||
<paragraph pageId="0" pageNumber="69">Habitat.</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">
|
||
Amazon rainforest. Known from Peru (
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Known distributions of species of Scybalocanthon. A S. adisi and S. aereus B S. acrianus, S. arnaudi, S. chamorroi, S. kaestneri and S. pinopterus C S. ashei sp. nov., S. federicoescobari, S. martinezi, S. papaxibe, S. pygidialis, and S. uniplagiatus." figureDoi="10.3897/zookeys.884.39322.figure5" httpUri="https://binary.pensoft.net/fig/351077" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure5">Fig. 5C</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="69">Endemism areas</emphasis>
|
||
:
|
||
<emphasis bold="true" pageId="0" pageNumber="69">Brazilian sub-region</emphasis>
|
||
: South Brazilian dominion:
|
||
<normalizedToken originalValue="Rondônia">Rondonia</normalizedToken>
|
||
province (see
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3782.1.1" author="Morrone, JJ" journalOrPublisher="Zootaxa" pageId="0" pageNumber="69" pagination="1 - 110" refId="B1" refString="Morrone, JJ, 2014. Biogeographical regionalisation of the Neotropical region. Zootaxa 3782: 1 - 110" title="Biogeographical regionalisation of the Neotropical region." url="https://doi.org/10.11646/zootaxa.3782.1.1" volume="3782" year="2014">Morrone 2014</bibRefCitation>
|
||
; fig. 12).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="69" type="remarks">
|
||
<paragraph pageId="0" pageNumber="69">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="69">
|
||
According to aedeagus characters,
|
||
<taxonomicName lsidName="S. ashei" pageId="0" pageNumber="69" rank="species" species="ashei">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. ashei</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Scybalocanthon ashei sp. nov. A Holotype (CMNC), dorsal view B holotype (CMNC), ventral view C aedeagus (right side) D aedeagus (left side) E set of bristles F fronto-lateral peripheral (FLP) sclerite (left), and Complex of axial and subaxial (A + SA) sclerites (right) G Superior right peripheral (SRP) sclerite." figureDoi="10.3897/zookeys.884.39322.figure1" httpUri="https://binary.pensoft.net/fig/351073" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure1">Fig. 1C, D</figureCitation>
|
||
) is closely related to
|
||
<taxonomicName lsidName="S. federicoescobari" pageId="0" pageNumber="69" rank="species" species="federicoescobari">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. federicoescobari</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Aedeagus, detail of parameres in left and right view. A S. acrianus B S. adisi C S. aereus D S. arnaudi E S. chamorroi F S. federicoescobari G S. kaestneri (specimen from Pastaza, Ecuador) H S. kaestneri (specimen from Napo, Ecuador) I S. martinezi J S. papaxibe K S. pinopterus (specimen from Madre de Dios, Peru) L S. pinopterus (specimen from Tingo Maria, Huanuco, Peru) M S. pinopterus (specimen from Tingo Maria, Huanuco, Peru) N S. pinopterus (specimen from Orellana, Ecuador) O S. pygidialis P S. uniplagiatus, Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.884.39322.figure3" httpUri="https://binary.pensoft.net/fig/351075" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure3">Fig. 3F</figureCitation>
|
||
),
|
||
<taxonomicName lsidName="S. martinezi" pageId="0" pageNumber="69" rank="species" species="martinezi">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. martinezi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Aedeagus, detail of parameres in left and right view. A S. acrianus B S. adisi C S. aereus D S. arnaudi E S. chamorroi F S. federicoescobari G S. kaestneri (specimen from Pastaza, Ecuador) H S. kaestneri (specimen from Napo, Ecuador) I S. martinezi J S. papaxibe K S. pinopterus (specimen from Madre de Dios, Peru) L S. pinopterus (specimen from Tingo Maria, Huanuco, Peru) M S. pinopterus (specimen from Tingo Maria, Huanuco, Peru) N S. pinopterus (specimen from Orellana, Ecuador) O S. pygidialis P S. uniplagiatus, Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.884.39322.figure3" httpUri="https://binary.pensoft.net/fig/351075" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure3">Fig. 3I</figureCitation>
|
||
),
|
||
<taxonomicName lsidName="S. papaxibe" pageId="0" pageNumber="69" rank="species" species="papaxibe">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. papaxibe</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Aedeagus, detail of parameres in left and right view. A S. acrianus B S. adisi C S. aereus D S. arnaudi E S. chamorroi F S. federicoescobari G S. kaestneri (specimen from Pastaza, Ecuador) H S. kaestneri (specimen from Napo, Ecuador) I S. martinezi J S. papaxibe K S. pinopterus (specimen from Madre de Dios, Peru) L S. pinopterus (specimen from Tingo Maria, Huanuco, Peru) M S. pinopterus (specimen from Tingo Maria, Huanuco, Peru) N S. pinopterus (specimen from Orellana, Ecuador) O S. pygidialis P S. uniplagiatus, Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.884.39322.figure3" httpUri="https://binary.pensoft.net/fig/351075" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure3">Fig. 3J</figureCitation>
|
||
),
|
||
<taxonomicName lsidName="S. pygidialis" pageId="0" pageNumber="69" rank="species" species="pygidialis">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. pygidialis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Aedeagus, detail of parameres in left and right view. A S. acrianus B S. adisi C S. aereus D S. arnaudi E S. chamorroi F S. federicoescobari G S. kaestneri (specimen from Pastaza, Ecuador) H S. kaestneri (specimen from Napo, Ecuador) I S. martinezi J S. papaxibe K S. pinopterus (specimen from Madre de Dios, Peru) L S. pinopterus (specimen from Tingo Maria, Huanuco, Peru) M S. pinopterus (specimen from Tingo Maria, Huanuco, Peru) N S. pinopterus (specimen from Orellana, Ecuador) O S. pygidialis P S. uniplagiatus, Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.884.39322.figure3" httpUri="https://binary.pensoft.net/fig/351075" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure3">Fig. 3O</figureCitation>
|
||
), and
|
||
<taxonomicName lsidName="S. uniplagiatus" pageId="0" pageNumber="69" rank="species" species="uniplagiatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="69">S. uniplagiatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Aedeagus, detail of parameres in left and right view. A S. acrianus B S. adisi C S. aereus D S. arnaudi E S. chamorroi F S. federicoescobari G S. kaestneri (specimen from Pastaza, Ecuador) H S. kaestneri (specimen from Napo, Ecuador) I S. martinezi J S. papaxibe K S. pinopterus (specimen from Madre de Dios, Peru) L S. pinopterus (specimen from Tingo Maria, Huanuco, Peru) M S. pinopterus (specimen from Tingo Maria, Huanuco, Peru) N S. pinopterus (specimen from Orellana, Ecuador) O S. pygidialis P S. uniplagiatus, Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.884.39322.figure3" httpUri="https://binary.pensoft.net/fig/351075" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure3">Fig. 3P</figureCitation>
|
||
) by having the parameres strongly asymmetric, with different shape and length (dorsal margin of left paramere with projection; ventral margin of left paramere with bilobate excavation; ventral margin of right paramere with rounded excavation), and FLP sclerite short (
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Scybalocanthon ashei sp. nov. A Holotype (CMNC), dorsal view B holotype (CMNC), ventral view C aedeagus (right side) D aedeagus (left side) E set of bristles F fronto-lateral peripheral (FLP) sclerite (left), and Complex of axial and subaxial (A + SA) sclerites (right) G Superior right peripheral (SRP) sclerite." figureDoi="10.3897/zookeys.884.39322.figure1" httpUri="https://binary.pensoft.net/fig/351073" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure1">Fig. 1F</figureCitation>
|
||
, left), with three sets of bristles right beside it (
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Scybalocanthon ashei sp. nov. A Holotype (CMNC), dorsal view B holotype (CMNC), ventral view C aedeagus (right side) D aedeagus (left side) E set of bristles F fronto-lateral peripheral (FLP) sclerite (left), and Complex of axial and subaxial (A + SA) sclerites (right) G Superior right peripheral (SRP) sclerite." figureDoi="10.3897/zookeys.884.39322.figure1" httpUri="https://binary.pensoft.net/fig/351073" pageId="0" pageNumber="69" tableDoi="10.3897/zookeys.884.39322.figure1">Fig. 1E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |