218 lines
19 KiB
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218 lines
19 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.57.5339" ID-PMC="PMC4698512" ID-Pensoft-Pub="1314-2003-57-1" ID-Pensoft-UUID="FFACFFC1FFEEFFA5C525FFE19915867A" ID-PubMed="26751914" ID-Zenodo-Dep="576307" ModsDocID="1314-2003-57-1" checkinTime="1451251344257" checkinUser="pensoft" docAuthor="Lorence, David H., Dietzsch, Anke C. & Kelly, Daniel L." docDate="2015" docId="6401EA2ACC71770CDB353C29AFA89FB1" docLanguage="en" docName="PhytoKeys 57: 1-9" docOrigin="PhytoKeys 57" docPubDate="2015-10-13" docSource="http://dx.doi.org/10.3897/phytokeys.57.5339" docTitle="Sommera cusucoana Lorence, D. Kelly & A. Dietzsch 2015, sp. nov." docType="treatment" docVersion="6" id="FFACFFC1FFEEFFA5C525FFE19915867A" lastPageNumber="3" masterDocId="FFACFFC1FFEEFFA5C525FFE19915867A" masterDocTitle="Sommera cusucoana, a new species of Rubiaceae from Honduras" masterLastPageNumber="9" masterPageNumber="1" pageNumber="1" updateTime="1668141479705" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Sommera cusucoana, a new species of Rubiaceae from Honduras</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lorence, David H.</mods:namePart>
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<mods:affiliation>National Tropical Botanical Garden, 3530 Papalina Road, Kalaheo, HI 96741 USA</mods:affiliation>
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<mods:nameIdentifier type="email">lorence@ntbg.org</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Dietzsch, Anke C.</mods:namePart>
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<mods:affiliation>Department of Botany, School of Natural Sciences, Trinity College, the University of Dublin, Dublin 2, Ireland</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kelly, Daniel L.</mods:namePart>
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<mods:affiliation>Department of Botany, School of Natural Sciences, Trinity College, the University of Dublin, Dublin 2, Ireland</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2015-10-13</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>57</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>9</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.57.5339</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.57.5339</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-57-1</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FFACFFC1FFEEFFA5C525FFE19915867A</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576307</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152026308" LSID="urn:lsid:plazi:treatment:6401EA2ACC71770CDB353C29AFA89FB1" httpUri="http://treatment.plazi.org/id/6401EA2ACC71770CDB353C29AFA89FB1" lastPageId="2" lastPageNumber="3" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="6401EA2A-CC71-770C-DB35-3C29AFA89FB1" authority="Lorence, D. Kelly & A. Dietzsch" authorityName="Lorence, D. Kelly & A. Dietzsch" authorityYear="2015" class="Magnoliopsida" family="Rubiaceae" genus="Sommera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sommera cusucoana" order="Gentianales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="cusucoana" status="sp. nov.">Sommera cusucoana Lorence, D. Kelly & A. Dietzsch</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Scan of Holotype specimen of Sommera cusucoana (to be deposited at TEFH)." figureDoi="10.3897/phytokeys.57.5339.figure2" httpUri="https://binary.pensoft.net/fig/60129" pageId="0" pageNumber="1">Figures 2</figureCitation>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Sommera cusucoana. A Tip of shoot with infructescence and leaf pair emerging between pair of stipules. Note red color of stipules, petioles, and infructescences B Freshly cut branch with inflorescence and infructescences. Photos by A. C. Dietzsch, 5 July 2013." figureDoi="10.3897/phytokeys.57.5339.figure3" httpUri="https://binary.pensoft.net/fig/60130" pageId="0" pageNumber="1">, 3</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Sommera cusucoana. A Tip of shoot with flower, developing fruits, and leaf pair emerging between pair of stipules B Flower at anthesis and developing fruit (appearing dark because in shadow). Photos by A. C. Dietzsch, 5 July 2013." figureDoi="10.3897/phytokeys.57.5339.figure4" httpUri="https://binary.pensoft.net/fig/60131" pageId="0" pageNumber="1">, 4</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type">
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<paragraph pageId="0" pageNumber="1">Type.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Honduras. Prov. Cortes, W of San Pedro Sula. El Cortecito campsite (near left bank of river), Parque Nacional El Cusuco, Sierra de
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<normalizedToken originalValue="Merendón">Merendon</normalizedToken>
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, UTM 361834 1716534 + 11 m, 1333 m alt., 5 July 2013, D. L. Kelly, A. C. Dietzsch & W. Lopez 15079 (Holotype TEFH!, Isotypes MO and TCD).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">Differs from its congeners by the combination of large, obovate leaves with long red petioles, glabrous or glabrate intervenal areas, red stipules, inflorescences 2-4-flowered, sparsely pubescent, with red axes, flowers with red hypanthium and calyx, and mature fruits dark red with long pedicels.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="description">
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<paragraph pageId="1" pageNumber="2">
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<pageBreakToken pageId="1" pageNumber="2" start="start">Description</pageBreakToken>
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.
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</paragraph>
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<paragraph lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
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Tree 10 m high, branchlets 5-6 mm in diam., glabrous, finely ribbed when dry, with sparse, pale +/- ellipsoidal lenticels.
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<emphasis bold="true" pageId="1" pageNumber="2">Leaves</emphasis>
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opposite, blades 21.5-30
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<normalizedToken originalValue="×">x</normalizedToken>
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10.2-15 cm, obovate, acuminate, finely pointed, cuneate and +/- asymmetrical at base, drying membranaceous, glabrous above, strigillose beneath on costa and 2°-4° veins, intervenal areas glabrous, 2° veins 13-14 pairs, eucamptodromous, venation prominulous; petioles 4.5-9 cm long, sparsely strigillose, red when fresh;
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<emphasis bold="true" pageId="1" pageNumber="2">stipules narrowly</emphasis>
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deltate to lanceolate, 3-3.8 cm long, when fresh red with thin white margins, glabrous, caducous.
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<emphasis bold="true" pageId="1" pageNumber="2">Inflorescences</emphasis>
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5.5-6.5
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<normalizedToken originalValue="×">x</normalizedToken>
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3-7 cm, dichasial, 2-3-4-flowered, axes sparsely strigillose, red when fresh; peduncle 2.5-3.8 cm long, bracts oblong-elliptic, c. 1 mm long, caducous; pedicels 1-2.8 cm long, with bract scars medially.
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<emphasis bold="true" pageId="1" pageNumber="2">Flowers</emphasis>
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with hypanthium 5
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<normalizedToken originalValue="×">x</normalizedToken>
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3-4 mm, turbinate to obovoid, glabrous;
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<pageBreakToken pageId="2" pageNumber="3" start="start">calyx</pageBreakToken>
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limb red, cupuliform, 2-3 mm long with the tube 0.6-1 mm long, externally glabrous, lobes 5, 1-1.2
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<normalizedToken originalValue="×">x</normalizedToken>
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2 mm, broadly triangular, obtuse, equal, margins densely ciliolate; corolla cylindrical-funnelform, yellowish-cream, fleshy, tube 10-11
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<normalizedToken originalValue="×">x</normalizedToken>
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2.8-3 mm, externally densely hirtellous-tomentose, lobes 5, 1.5
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<normalizedToken originalValue="×">x</normalizedToken>
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3 mm, triangular-ovate, densely papillose-puberulent internally; anthers, style and stigmas not seen.
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<emphasis bold="true" pageId="2" pageNumber="3">Fruits</emphasis>
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12-15
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<normalizedToken originalValue="×">x</normalizedToken>
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9-13 mm, subglobose to broadly ellipsoid, dark red at maturitiy, glabrous.
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<emphasis bold="true" pageId="2" pageNumber="3">Seeds</emphasis>
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numerous, 1-1.8 mm long, irregularly polygonal-angulate, testa dark brown, reticulate. (Figures
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Sommera cusucoana. A Tip of shoot with infructescence and leaf pair emerging between pair of stipules. Note red color of stipules, petioles, and infructescences B Freshly cut branch with inflorescence and infructescences. Photos by A. C. Dietzsch, 5 July 2013." figureDoi="10.3897/phytokeys.57.5339.figure3" httpUri="https://binary.pensoft.net/fig/60130" pageId="2" pageNumber="3">3</figureCitation>
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,
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Sommera cusucoana. A Tip of shoot with flower, developing fruits, and leaf pair emerging between pair of stipules B Flower at anthesis and developing fruit (appearing dark because in shadow). Photos by A. C. Dietzsch, 5 July 2013." figureDoi="10.3897/phytokeys.57.5339.figure4" httpUri="https://binary.pensoft.net/fig/60131" pageId="2" pageNumber="3">4</figureCitation>
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).
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</paragraph>
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<caption ID-Zenodo-Dep="966308" doi="10.3897/phytokeys.57.5339.figure2" httpUri="https://binary.pensoft.net/fig/60129" pageId="2" pageNumber="3" start="Figure 2" startId="F2">
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<paragraph pageId="2" pageNumber="3">
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<emphasis bold="true" pageId="2" pageNumber="3">Figure 2.</emphasis>
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Scan of Holotype specimen of
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<taxonomicName authorityName="Lorence, D. Kelly & A. Dietzsch" authorityYear="2015" class="Magnoliopsida" family="Rubiaceae" genus="Sommera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sommera cusucoana" order="Gentianales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="cusucoana">
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<emphasis italics="true" pageId="2" pageNumber="3">Sommera cusucoana</emphasis>
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</taxonomicName>
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(to be deposited at TEFH).
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</paragraph>
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</caption>
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<caption ID-Zenodo-Dep="966310" doi="10.3897/phytokeys.57.5339.figure3" httpUri="https://binary.pensoft.net/fig/60130" pageId="2" pageNumber="3" start="Figure 3" startId="F3">
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<paragraph pageId="2" pageNumber="3">
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<emphasis bold="true" pageId="2" pageNumber="3">Figure 3.</emphasis>
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<taxonomicName authorityName="Lorence, D. Kelly & A. Dietzsch" authorityYear="2015" class="Magnoliopsida" family="Rubiaceae" genus="Sommera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sommera cusucoana" order="Gentianales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="cusucoana">
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<emphasis italics="true" pageId="2" pageNumber="3">Sommera cusucoana</emphasis>
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</taxonomicName>
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.
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<emphasis bold="true" pageId="2" pageNumber="3">A</emphasis>
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Tip of shoot with infructescence and leaf pair emerging between pair of stipules. Note red color of stipules, petioles, and infructescences
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<emphasis bold="true" pageId="2" pageNumber="3">B</emphasis>
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Freshly cut branch with inflorescence and infructescences. Photos by A.C. Dietzsch, 5 July 2013.
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</paragraph>
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</caption>
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<caption ID-Zenodo-Dep="966312" doi="10.3897/phytokeys.57.5339.figure4" httpUri="https://binary.pensoft.net/fig/60131" pageId="2" pageNumber="3" start="Figure 4" startId="F4">
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<paragraph pageId="2" pageNumber="3">
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<emphasis bold="true" pageId="2" pageNumber="3">Figure 4.</emphasis>
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<taxonomicName authorityName="Lorence, D. Kelly & A. Dietzsch" authorityYear="2015" class="Magnoliopsida" family="Rubiaceae" genus="Sommera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sommera cusucoana" order="Gentianales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="cusucoana">
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<emphasis italics="true" pageId="2" pageNumber="3">Sommera cusucoana</emphasis>
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</taxonomicName>
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.
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<emphasis bold="true" pageId="2" pageNumber="3">A</emphasis>
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Tip of shoot with flower, developing fruits, and leaf pair emerging between pair of stipules
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<emphasis bold="true" pageId="2" pageNumber="3">B</emphasis>
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Flower at anthesis and developing fruit (appearing dark because in shadow). Photos by A.C. Dietzsch, 5 July 2013.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="3" type="habitat and ecology">
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<paragraph pageId="2" pageNumber="3">Habitat and ecology.</paragraph>
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<paragraph pageId="2" pageNumber="3">
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Only two individual trees were located, about the same size and within a few meters of each other. The site is within Cusuco National Park, in the upper slopes of the Sierra del
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<normalizedToken originalValue="Merendón">Merendon</normalizedToken>
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. These upper slopes (highest point 2242 m) are largely covered by montane rain forest vegetation. The bedrock is composed of a mixture of gneiss and schist (
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<bibRefCitation author="Lorence, DH" journalOrPublisher="Flora Mesoamericana" pageId="4" pageNumber="5" refId="B10" refString="The Nature Conservancy, Fundacion Ecologista "Hector Rodrigo Pastor Fasquelle" & Paca 1994. Evaluacion Ecologica Rapida (EER) Parque Nacional "El Cusuco" y Cordillera del Merendon. Unpublished report." year="2012">The Nature Conservancy et al. 1994</bibRefCitation>
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); slopes are steep and soils are strongly acidic (D.L. Kelly & A.C. Dietzsch, unpublished data).
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</paragraph>
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<paragraph pageId="2" pageNumber="3">
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The type locality, at 1333 m, is at the bottom of a deep, narrow valley, about 25-50 m from the bank of a small river. The site is riparian rain forest, dominated by tall trees, mainly
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<taxonomicName class="Dicotyledoneae" family="Hamamelidaceae" genus="Liquidambar" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Liquidambar styraciflua" order="Rosales" pageId="2" pageNumber="3" phylum="Angiospermae" rank="species" species="styraciflua">
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<emphasis italics="true" pageId="2" pageNumber="3">Liquidambar styraciflua</emphasis>
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</taxonomicName>
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L. and
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<taxonomicName class="Dicotyledoneae" family="Meliaceae" genus="Cedrela" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Cedrela odorata" order="Rutales" pageId="2" pageNumber="3" phylum="Angiospermae" rank="species" species="odorata">
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<emphasis italics="true" pageId="2" pageNumber="3">Cedrela odorata</emphasis>
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</taxonomicName>
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L. The microclimate is moist and the vegetation lush and species-rich. The vicinity shows minor levels of disturbance: human disturbance due to the proximity of a seasonal camp-site with radiating trails, and natural disturbance due to wind-throw, and land-slips on the steeper slopes.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="3" type="etymology">
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<paragraph pageId="2" pageNumber="3">Etymology.</paragraph>
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<paragraph pageId="2" pageNumber="3">The name honors the Cusuco National Park in which it was found.</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="3" type="discussion">
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<paragraph pageId="2" pageNumber="3">Discussion.</paragraph>
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<paragraph pageId="2" pageNumber="3">
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<taxonomicName authorityName="Lorence, D. Kelly & A. Dietzsch" authorityYear="2015" class="Magnoliopsida" family="Rubiaceae" genus="Sommera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sommera cusucoana" order="Gentianales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="cusucoana">
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<emphasis italics="true" pageId="2" pageNumber="3">Sommera cusucoana</emphasis>
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</taxonomicName>
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differs from its Mesoamerican congeners by the combination of large, obovate leaves with long red petioles, glabrous or glabrate intervenal areas, red stipules, lax, sparsely pubescent 2-4-flowered inflorescences with red axes, flowers with red hypanthium and calyx, and long flowering and fruiting pedicels. Floral hypanthia and fruits of
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<taxonomicName authorityName="Lorence, D. Kelly & A. Dietzsch" authorityYear="2015" class="Magnoliopsida" family="Rubiaceae" genus="Sommera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sommera cusucoana" order="Gentianales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="cusucoana">
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<emphasis italics="true" pageId="2" pageNumber="3">Sommera cusucoana</emphasis>
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</taxonomicName>
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are bright red at all developmental stages. Herbarium label notes for other
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<taxonomicName authorityName="Schlechtendal" authorityYear="1835" class="Magnoliopsida" family="Rubiaceae" genus="Sommera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sommera" order="Gentianales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="2" pageNumber="3">Sommera</emphasis>
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</taxonomicName>
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species indicate fruits are green when immature and ripen red, at least in
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<taxonomicName authorityName="Brandegee" authorityYear="1915" class="Magnoliopsida" family="Rubiaceae" genus="Sommera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sommera chiapensis" order="Gentianales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="chiapensis">
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<emphasis italics="true" pageId="2" pageNumber="3">Sommera chiapensis</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Standl" authorityYear="1914" class="Magnoliopsida" family="Rubiaceae" genus="Sommera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sommera donnell-smithii" order="Gentianales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="donnell-smithii">
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<emphasis italics="true" pageId="2" pageNumber="3">Sommera donnell-smithii</emphasis>
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</taxonomicName>
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and
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<taxonomicName authorityName="L. O. Williams" authorityYear="1973" class="Magnoliopsida" family="Rubiaceae" genus="Sommera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sommera montana" order="Gentianales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="montana">
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<emphasis italics="true" pageId="2" pageNumber="3">Sommera montana</emphasis>
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</taxonomicName>
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, and possibly white in
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<taxonomicName class="Magnoliopsida" family="Rubiaceae" genus="Sommera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sommera grandis" order="Gentianales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="grandis">
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<emphasis italics="true" pageId="2" pageNumber="3">Sommera grandis</emphasis>
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</taxonomicName>
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(Mexico). Only
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<taxonomicName authorityName="Brandegee" authorityYear="1915" class="Magnoliopsida" family="Rubiaceae" genus="Sommera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sommera chiapensis" order="Gentianales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="chiapensis">
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<emphasis italics="true" pageId="2" pageNumber="3">Sommera chiapensis</emphasis>
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</taxonomicName>
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Brandegee (Chiapas, Guatemala, Honduras) has flowers with similarly short, broadly triangular calyx lobes 1-2 mm long, but it differs in having shorter petioles 2-5 cm long, cymes with (4-)6-12 flowers, shorter corollas with tube 4-8 mm long, and smaller fruits 5-9 mm in diameter.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |