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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.107.26548" ID-GBIF-Dataset="b09b328e-de27-4071-8af6-eec29dc3b1df" ID-PMC="PMC6160804" ID-Pensoft-Pub="1314-2003-107-1" ID-Pensoft-UUID="686CFF85FFADFFEAC033443CF54BFFFC" ID-PubMed="30271249" ID-Zenodo-Dep="1346433" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1314-2003-107-1" ModsDocOrigin="PhytoKeys 107" ModsDocTitle="A revision of Dissochaeta (Melastomataceae, Dissochaeteae)" checkinTime="1555330651054" checkinUser="pensoft" docAuthor="Kartonegoro, Abdulrokhman, Veldkamp, Jan Frits, Hovenkamp, Peter &amp; Welzen, Peter van" docDate="2018" docId="4AF7104912EF0CCE37CEED54B92369F3" docLanguage="en" docName="PhytoKeys 107: 1-178" docOrigin="PhytoKeys 107" docSource="http://dx.doi.org/10.3897/phytokeys.107.26548" docTitle="Dissochaeta biligulata Korth. in Temminck, Verh. Nat. Gesch. Ned. Bezitt., Bot.: 240. 1844" docType="treatment" docVersion="6" id="686CFF85FFADFFEAC033443CF54BFFFC" lastPageNumber="1" masterDocId="686CFF85FFADFFEAC033443CF54BFFFC" masterDocTitle="A revision of Dissochaeta (Melastomataceae, Dissochaeteae)" masterLastPageNumber="178" masterPageNumber="1" pageNumber="1" updateTime="1668139007331" updateUser="ExternalLinkService">
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<mods:title>A revision of Dissochaeta (Melastomataceae, Dissochaeteae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Kartonegoro, Abdulrokhman</mods:namePart>
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<mods:name type="personal">
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<mods:namePart>Veldkamp, Jan Frits</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Hovenkamp, Peter</mods:namePart>
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<mods:namePart>Welzen, Peter van</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>PhytoKeys</mods:title>
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<mods:date>2018</mods:date>
<mods:detail type="volume">
<mods:number>107</mods:number>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.107.26548</mods:identifier>
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<treatment ID-GBIF-Taxon="156201507" LSID="urn:lsid:plazi:treatment:4AF7104912EF0CCE37CEED54B92369F3" httpUri="http://treatment.plazi.org/id/4AF7104912EF0CCE37CEED54B92369F3" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
10.
<taxonomicName LSID="3AD36EDD-7D52-5D85-B51B-B1F9A41F097B" authority="Korth. in Temminck, Verh. Nat. Gesch. Ned. Bezitt., Bot.: 240. 1844." class="Magnoliopsida" family="Melastomataceae" genus="Dissochaeta" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dissochaeta biligulata" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="biligulata">Dissochaeta biligulata Korth. in Temminck, Verh. Nat. Gesch. Ned. Bezitt., Bot.: 240. 1844.</taxonomicName>
<figureCitation captionStart="Figure 7" captionStartId="F12" captionText="Figure 7. Dissochaeta biligulata a habit b branchlet c hypanthium d flower e fruits. Photographs by D. Penneys; vouchers: Penneys 2488 (WNC) &amp; Penneys 2509 (WNC)." figureDoi="10.3897/phytokeys.107.26548.figure7" httpUri="https://binary.pensoft.net/fig/222106" pageId="0" pageNumber="1">Fig. 7</figureCitation>
<figureCitation captionStart="Map 6" captionStartId="F13" captionText="Map 6. Distribution of D. biligulata (●)." figureDoi="10.3897/phytokeys.107.26548.map6" httpUri="https://binary.pensoft.net/fig/222107" pageId="0" pageNumber="1">, Map 6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Naud" authorityYear="1851" class="Magnoliopsida" family="Melastomataceae" genus="Dissochaeta" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dissochaeta microcarpa" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="microcarpa">Dissochaeta microcarpa</taxonomicName>
Naudin, Ann. Sci. Nat., Bot.
<normalizedToken originalValue="sér">ser</normalizedToken>
. 3, 15: 72. 1851. Type: Singapore. Singapore Island, Feb 1837, C.
<normalizedToken originalValue="Gaudichaud-Beaupré">Gaudichaud-Beaupre</normalizedToken>
80 (holotype: P [P02274814, image seen]!).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Dissochaeta" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dissochaeta bancana" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="bancana">Dissochaeta bancana</taxonomicName>
Miq., Fl. Ned. Ind. 1(1): 529. 1855. Type: Indonesia. Bangka, T. Horsfield 19 (lectotype, designated here: K [K000859500]!; isolectotype: K [K000859501]!).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Anplectrum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Anplectrum biligulatum" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="biligulatum">Anplectrum biligulatum</taxonomicName>
(Korth.) Triana, Trans. Linn. Soc. London 28: 85. 1872.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Dissochaeta" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dissochaeta celebica" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="celebica">Dissochaeta celebica</taxonomicName>
auct. non Blume: Triana, Trans. Linn. Soc. London 28: 83. 1872; C.B.Clarke in Hook.
<emphasis italics="true" pageId="0" pageNumber="1">f.</emphasis>
, Fl. Brit. India 2: 544. 1879; Cogn. in Boerl., Handl. Fl. Ned. Ind. 2: 533. 1890; in A.DC. &amp; C.DC., Monogr. Phan. 7: 561. 1891; in H.J.P.Winkl., Bot. Jahrb. Syst. 48: 108. 1913; King, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 69(2): 54. 1900; Ridl., Fl. Malay Penins. 1: 798. 1922; Merr., Univ. Calif. Publ. Bot. 15: 224. 1929; Craib, Fl. Siam. Enum. 1, 4: 697. 1931; Bygrave &amp; A.P.Davis, Checkl. Fl. Pl. Gymnosperms Brunei Darussalam: 186. 1996; Cellin. in J.H.Beaman &amp; C.E.Anderson, Pl. Mt. Kinabalu 5: 99. 2004.
<emphasis italics="true" pageId="0" pageNumber="1">p.p.</emphasis>
, excl. type.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Dissochaeta" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dissochaeta intermedia" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="intermedia">Dissochaeta intermedia</taxonomicName>
auct. non Blume: C.B.Clarke in Hook.
<emphasis italics="true" pageId="0" pageNumber="1">f.</emphasis>
, Fl. Brit. India 2: 544. 1879.
<emphasis italics="true" pageId="0" pageNumber="1">p.p.</emphasis>
, excl. type.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Kuntze" authorityYear="1891" baseAuthorityName="Korth." class="Magnoliopsida" family="Melastomataceae" genus="Diplectria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Diplectria biligulata" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="biligulata">Diplectria biligulata</taxonomicName>
(Korth.) Kuntze, Revis. Gen. Pl. 1: 246. 1891.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authority="Blume" class="Magnoliopsida" family="Melastomataceae" genus="Dissochaeta" higherTaxonomySource="CoL" infraspecific-rank="var." kingdom="Plantae" lsidName="Dissochaeta celebica var. contracta" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="variety" species="celebica" variety="contracta">Dissochaeta celebica Blume var. contracta</taxonomicName>
King, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 69(2): 54. 1900. Type: Malaysia. Perak: Larut, Apr 1882,
<normalizedToken originalValue="Kings">King's</normalizedToken>
collector 2911 (lectotype, designated here: K [K000859535]!; isolectotypes: BM!, CAL
<emphasis italics="true" pageId="0" pageNumber="1">n.v.</emphasis>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="King" authorityYear="1900" class="Magnoliopsida" family="Melastomataceae" genus="Dissochaeta" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dissochaeta scortechinii" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="scortechinii">Dissochaeta scortechinii</taxonomicName>
King, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 69(2): 55. 1900. Type: Malaysia. Perak: B. Scortechini 23 (lectotype, designated here: K [K000859536]!; isolectotype: SING
<emphasis italics="true" pageId="0" pageNumber="1">n.v.</emphasis>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Bakh fil" authorityYear="1943" baseAuthorityName="Korth." class="Magnoliopsida" family="Melastomataceae" genus="Neodissochaeta" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Neodissochaeta biligulata" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="biligulata">Neodissochaeta biligulata</taxonomicName>
(Korth.) Bakh.
<emphasis italics="true" pageId="0" pageNumber="1">f.</emphasis>
, Contr. Melastom.: 140. 1943.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Bakh fil" authorityYear="1943" baseAuthorityName="Bl." class="Magnoliopsida" family="Melastomataceae" genus="Neodissochaeta" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Neodissochaeta celebica" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="celebica">Neodissochaeta celebica</taxonomicName>
(Blume) Bakh.
<emphasis italics="true" pageId="0" pageNumber="1">f.</emphasis>
, Contr. Melastom.: 141. 1943.
<emphasis italics="true" pageId="0" pageNumber="1">p.p.</emphasis>
, excl. type.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Dissochaeta" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dissochaeta monticola" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="monticola">Dissochaeta monticola</taxonomicName>
auct. non Blume: Clausing in S.S.Renner
<emphasis italics="true" pageId="0" pageNumber="1">et al</emphasis>
., Fl. Thailand 7(3): 428. 2001.
<emphasis italics="true" pageId="0" pageNumber="1">p.p.</emphasis>
, excl. type.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Dissochaeta" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dissochaeta intermedia" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="intermedia">Dissochaeta intermedia</taxonomicName>
Blume var.
<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Dissochaeta" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="sagittata" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="sagittata">sagittata</taxonomicName>
auct. non. J.F.Maxwell: Cellin. in J.H.Beaman &amp; C.E. Anderson, Pl. Mt. Kinabalu 5: 99. 2004.
<emphasis italics="true" pageId="0" pageNumber="1">p.p.</emphasis>
, excl. type.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Type.</paragraph>
<paragraph pageId="0" pageNumber="1">Indonesia. West Sumatra: Gunung Paauw, P.W. Korthals s.n. (lectotype, designated here: L [L0537285]!; isolectotype: L [L0537284]!).</paragraph>
<caption ID-Zenodo-Dep="1346487" doi="10.3897/phytokeys.107.26548.figure7" httpUri="https://binary.pensoft.net/fig/222106" pageId="0" pageNumber="1" start="Figure 7" startId="F12">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 7.</emphasis>
<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Dissochaeta" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dissochaeta biligulata" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="biligulata">
<emphasis italics="true" pageId="0" pageNumber="1">Dissochaeta biligulata</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
habit
<emphasis bold="true" pageId="0" pageNumber="1">b</emphasis>
branchlet
<emphasis bold="true" pageId="0" pageNumber="1">c</emphasis>
hypanthium
<emphasis bold="true" pageId="0" pageNumber="1">d</emphasis>
flower
<emphasis bold="true" pageId="0" pageNumber="1">e</emphasis>
fruits. Photographs by D. Penneys; vouchers: Penneys 2488 (WNC) &amp; Penneys 2509 (WNC).
</paragraph>
</caption>
<caption ID-Zenodo-Dep="1346543" doi="10.3897/phytokeys.107.26548.map6" httpUri="https://binary.pensoft.net/fig/222107" pageId="0" pageNumber="1" start="Map 6" startId="F13">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Map 6.</emphasis>
Distribution of
<taxonomicName class="Magnoliopsida" family="Melastomataceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="D. biligulata" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="biligulata">
<emphasis italics="true" pageId="0" pageNumber="1">D. biligulata</emphasis>
</taxonomicName>
(●).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
Climbing up to 5 m height. Branchlets terete, 3-4 mm in diameter, densely brown stellate-furfuraceous; nodes swollen, with interpetiolar line; internodes 4-6.5 cm long. Leaves: petioles terete, 6-8 mm long, brown stellate-furfuraceous; blades ovate-oblong or elliptic-oblong, 7-15
<normalizedToken originalValue="×">x</normalizedToken>
2.5-5.4 cm, membranous, base rounded, margin entire, apex acuminate, tip ca. 1 cm long; nervation with 1 pair of lateral nerves and 1 pair of intramarginal nerves; adaxially glabrous, yellowish-green, abaxially densely covered with minute brown stellate-furfuraceous hairs. Inflorescences terminal, 10-16 cm long, many-flowered; main axis angular, brown stellate-furfuraceous; primary axes up to 15 cm long with 6-8 nodes, secondary axes 2-4 cm long with 2 or 3 nodes, tertiary axes 0.5-1 cm long with 1 node; bracts linear, ca. 3 mm long, brown stellate-furfuraceous; bracteoles linear, 2-2.5 mm long, brown stellate-furfuraceous; pedicel densely stellate-furfuraceous, 1-2 mm long in central flowers, ca. 0.5 mm long or absent in lateral flowers. Hypanthium suburceolate, 3-4
<normalizedToken originalValue="×">x</normalizedToken>
2-2.5 mm, brown stellate-furfuraceous; calyx lobes truncate with 4 undulate tips, ca. 1 mm long; petal buds conical, 2-3 mm long, glabrous; mature petals ovate to elliptic, 5-6
<normalizedToken originalValue="×">x</normalizedToken>
3-3.5 mm, reflexed, base clawed, apex obtuse, glabrous, white or pinkish-white, sometimes transparent. Stamens 4, equal, alternipetalous, filaments curved sideways, ca. 5 mm long, white, pinkish at base, anthers oblong-lanceolate, thecae 3.5-5 mm long, straight, pink, pedoconnective ca. 1 mm long, basal crest triangular or erose with irregular margin, yellow, ca. 0.5 mm long, lateral appendages paired, filiform, 2-3 mm long. Ovary
<normalizedToken originalValue="¾">3/4</normalizedToken>
of hypanthium in length, apex villous; style glabrous, ca. 9 mm long, pointing in direction opposite to filaments, curved at tip; stigma minute; extra-ovarial chambers 4, shallow, reaching to upper third of ovary. Fruits globose, ca. 6
<normalizedToken originalValue="×">x</normalizedToken>
4-5 mm, glabrous, bright green, slightly 8 vertical lines present; calyx lobes remnant persistent. Seeds ca. 0.5 mm long.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">India (Nicobar Islands), Myanmar, Indochina (Cambodia and Vietnam), southwards through Thailand to western Malesia (Malay Peninsula, Sumatra and Borneo) and Philippines (Palawan).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="ecology">
<paragraph pageId="0" pageNumber="1">Ecology and habitat.</paragraph>
<paragraph pageId="0" pageNumber="1">Secondary lowland forest, opened primary forest or disturbed forest, dipterocarp forest or montane forest from sea level to 1500 m elevation.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="vernacular names">
<paragraph pageId="0" pageNumber="1">Vernacular names.</paragraph>
<paragraph pageId="0" pageNumber="1">
Sumatra:
<emphasis italics="true" pageId="0" pageNumber="1">akar kemunting</emphasis>
(Bintan). Borneo:
<emphasis italics="true" pageId="0" pageNumber="1">kemanti Omang</emphasis>
(Iban);
<emphasis italics="true" pageId="0" pageNumber="1">luka kalapat</emphasis>
(Penan).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="notes">
<paragraph pageId="0" pageNumber="1">Note.</paragraph>
<paragraph pageId="0" pageNumber="1">
Generally recognised as
<taxonomicName class="Magnoliopsida" family="Melastomataceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="D. celebica" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="celebica">
<emphasis italics="true" pageId="0" pageNumber="1">D. celebica</emphasis>
</taxonomicName>
by most authors because of the similarity in the small flowers with only four fertile stamens.
<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Dissochaeta" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dissochaeta biligulata" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="biligulata">
<emphasis italics="true" pageId="0" pageNumber="1">Dissochaeta biligulata</emphasis>
</taxonomicName>
differs from
<taxonomicName class="Magnoliopsida" family="Melastomataceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="D. celebica" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="celebica">
<emphasis italics="true" pageId="0" pageNumber="1">D. celebica</emphasis>
</taxonomicName>
by its dark brown furfuraceous indumentum on the leaves below, a truncate calyx with lobes with undulate tips, a white or pinkish-white corolla and globose fruits with distinct lines. In
<taxonomicName class="Magnoliopsida" family="Melastomataceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="D. celebica" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="celebica">
<emphasis italics="true" pageId="0" pageNumber="1">D. celebica</emphasis>
</taxonomicName>
, the indumentum underneath the leaf blades is less dense, the calyx lobes are glabrous, purple and have a triangular apex, the corolla is dark purple and the fruit ovoid without distinct lines. This species is very common in the open dipterocarp forest in western Malesia.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Selected specimens examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">INDIA. Nicobar Islands</emphasis>
: East West Road, 20 Jul 1976, M.S. Balakrishnan 3899 (L).
<emphasis bold="true" pageId="0" pageNumber="1">MYANMAR. Tenasserim</emphasis>
: Domel Island, 14 Jan 1829, J.W. Helfer 2286 (K).
<emphasis bold="true" pageId="0" pageNumber="1">CAMBODIA. Kah Kong</emphasis>
: Lhuom Rovaal, 600 m, 27 Feb 1966, Anon 758 (P); Mount Rodam Meong Daeum, 650 m, 23 Feb 1966, Martin 348 (P).
<emphasis bold="true" pageId="0" pageNumber="1">VIETNAM. Ha Tay</emphasis>
: Mount Bavi, Tonkin River, Jun 1908, C.
<normalizedToken originalValue="dÁlleizette">dAlleizette</normalizedToken>
2420 (L).
<emphasis bold="true" pageId="0" pageNumber="1">THAILAND. Narathiwat</emphasis>
: Sungei Kolok, 4 Mar 1974. K. Larsen &amp; S.S. Larsen 32923 (K, L).
<emphasis bold="true" pageId="0" pageNumber="1">Pattalung</emphasis>
: Tamote Falls, 225 m, 9 Aug 1986, J.F. Maxwell 86-552 (L).
<emphasis bold="true" pageId="0" pageNumber="1">Pattani</emphasis>
: Bukit, 25 Jan 1931, P. Put 3635 (K).
<emphasis bold="true" pageId="0" pageNumber="1">Surat Thani</emphasis>
: Ko Panghan, 3 Jun 1927, P. Put 778 (BM, K).
<emphasis bold="true" pageId="0" pageNumber="1">Trang</emphasis>
: Khao Pap Pa, 13 Mar 1974, K. Larsen &amp; S.S. Larsen 33279 (P).
<emphasis bold="true" pageId="0" pageNumber="1">Yala</emphasis>
: Nikhom Kulong, 500 m, 31 Dec 1972, T. Santisuk &amp; B. Nimanong BKF 54194 (K, L).
<emphasis bold="true" pageId="0" pageNumber="1">MALAYSIA. Johor</emphasis>
: Kluang, 24 Nov 1967, Alphonso, Sanusi &amp; Sidek S.200 (K, L); Gunung Pulai, 550 m, 29 Jan 1978, J.F. Maxwell 78-24 (L).
<emphasis bold="true" pageId="0" pageNumber="1">Malacca</emphasis>
: W. Griffith KD 2288 (BO, K).
<emphasis bold="true" pageId="0" pageNumber="1">Negri Sembilan</emphasis>
: Gunung Angsi, 730 m, 25 Nov 1923, M. Nur SFN 11708 (BO, K); Nilai, near Seremban, Jinderam Estate, 90 m, 22 Sep 1957, M. Shah 129 (BO, K, L).
<emphasis bold="true" pageId="0" pageNumber="1">Pahang</emphasis>
: Sungai Tahan, 27 Jul 1936, Kiah SFN 31912 (BO, K, PNH);
<normalizedToken originalValue="Frasers">Fraser's</normalizedToken>
Hill, Girdle Road, 1200 m, 26 Aug 1959, M. Shah &amp; Kadim 680A (BO, K, L).
<emphasis bold="true" pageId="0" pageNumber="1">Penang</emphasis>
: N. Wallich 4050 (K, P); Government Hill, 20 Jan 1903, C. Curtis 3806 (BM, BO).
<emphasis bold="true" pageId="0" pageNumber="1">Perak</emphasis>
: B. Scortechini 23 (K); Larut,
<normalizedToken originalValue="Kings">King's</normalizedToken>
collector 2911 (BM, K); Trolak, 22 Mar 1967, S. Chelliah KEP 104685 (K, L); Ulu Bubong, 120-180 m, Jun 1886,
<normalizedToken originalValue="Kings">King's</normalizedToken>
collector 10290 (BM, K).
<emphasis bold="true" pageId="0" pageNumber="1">Selangor</emphasis>
: Ulu Gombak, 21 Aug 1968, Y. Teo &amp; J.W. Purseglove 165 (K, L); Kuala Lumpur, 5 Mar 1915, H.N. Ridley s.n. (BM, K, L).
<emphasis bold="true" pageId="0" pageNumber="1">Terengganu</emphasis>
: Kemaman, Sungai Nipah, 24 Jun 1932, E.J.H. Corner SFN 25848 (K).
<emphasis bold="true" pageId="0" pageNumber="1">Sabah</emphasis>
: Beaufort, Klas FR, 23 Jul 1973, Dewol &amp; Abdul Karim SAN 77821 (K, L); Keningau, Kitau, 8 Sep 1982, Amin SAN 95456 (K, L); Kalabakan, Seranum, 21 Sep 1983, Fidilis &amp; Suali SAN 101240 (K, L); Lahad Datu, Danum Valley, 238 m, 6 Jul 2006, Rosalia et al. SAN 145986 (K); Lamag, Sungai Tangkulap, 30 Jun 1983, Amin SAN 97451 (L); Nabawan, Jalan Nabawan-Pandewan, 12 Feb 1990, Sumbing SAN 128120 (K); Pensiangan, Pensiangan Kayu FR., 28 Jan 1994, Fidilis SAN 136931 (K); Tawau, Oct 1922-Mar 1923, A.D.E. Elmer 21840 (BM, BO, K, L, P, PNH, U); Ranau, Kampung Kiau, 30 Nov 1915, M.S. Clemens 10122 (PNH);
<emphasis italics="true" pageId="0" pageNumber="1">Ibid</emphasis>
., Kampung Merungin, 300 m, 18 Nov 1975, Leopold &amp; Saikeh SAN 82620 (K, L); Mt. Kinabalu, 1060 m, 14 Jun 1961, W.L. Chew, E.J.H. Corner &amp; A. Stainton RSNB 68 (K, L);
<emphasis italics="true" pageId="0" pageNumber="1">Ibid.</emphasis>
, Kelawat, near Tamparuli, 900 m, 2 Mar 1954, S. Darnton 230 (BM, L); Sipitang, Mesapol FR, 30 m, 25 Nov 1968, K. Ogata 11671 (L).
<emphasis bold="true" pageId="0" pageNumber="1">Sarawak</emphasis>
: Baram, Kelabit Highland, 1066 m, 6 Nov 1974, P.K. Chai S.35335 (K, L); Belaga, Sungai Murum, 600 m, 13 May 1994, S.H. Lai et al. S.68539 (K); Kuching, Oct 1892, G.D. Haviland 1901 (BM, K);
<emphasis italics="true" pageId="0" pageNumber="1">Ibid.</emphasis>
, Selang FR., 30 m, 25 Jul 1957, Ilias Paie S.8460 (BO, K, L); Limbang, Bukit Pagon, Sungai Sipayan, 540 m, 3 Aug 1984, D. Awa &amp; B. Lee S.47645 (L); Marudi, Bario, 1100 m, 12 Apr 1995, T.E. Beaman &amp; R. Repin 164 (K); Miri, Lambir National Park, 5 Apr 1966, Sibat ak Luang S.25075 (BO, K, L);
<emphasis italics="true" pageId="0" pageNumber="1">Ibid.</emphasis>
, Riam Road, 30 m, 3 Dec 1962, Joseph Au S.17254 (BO, K, L); Betong, Bukit Sadok, 15 Oct 1982, Banyeng &amp; Ilias Paie S.45085 (K, L); Dataran Tinggi Merurong, Sungai Jelalong, 330 m, 9 Oct 1984, Othman &amp; P.C. Yii S.48814 (K, L); Baram, Ulu Koyan, Mount Dulit, 900 m, 15 Sep 1932, P.W. Richards 1818 (K, L).
<emphasis bold="true" pageId="0" pageNumber="1">SINGAPORE.</emphasis>
Feb 1837, C.
<normalizedToken originalValue="Gaudichaud-Beaupré">Gaudichaud-Beaupre</normalizedToken>
80 (P); Oct 1861, T. Anderson 67 (BM, K, P); Seletar, 27 Mar 1889, H.N. Ridley 2025 (BM); MacRitchie Reservoir 10 m, 15 Jul 1982, J.F. Maxwell 82-194 (L); Pierce Reservoir, 60 m, 20 Oct 1957, H.M. Burkill HMB 1227 (K, L, PNH).
<emphasis bold="true" pageId="0" pageNumber="1">BRUNEI. Belait</emphasis>
: Jalan Merangking-Buau, 10 Aug 1991, N. Nangkat 265 (K, L); Bukit Teraja, 160 m, 27 Sep 1957, P.S. Ashton BRUN 671 (BO, K, L).
<emphasis bold="true" pageId="0" pageNumber="1">Seria</emphasis>
: Andulau FR, 25 m, 31 Jul 1963, H.P. Fuchs &amp; S. Muller 21150 (K, L).
<emphasis bold="true" pageId="0" pageNumber="1">Temburong</emphasis>
: Batu Apoi, 350 m, 29 Oct 1991, D.A. Simpson &amp; S. Marsh 2502 (BO, K, L).
<emphasis bold="true" pageId="0" pageNumber="1">INDONESIA. Aceh</emphasis>
: Mt. Leuser, Gunung Bandahara, 1350 m, 20 Feb 1980, S. Prawiroatmodjo 2386 (BO, K);
<emphasis italics="true" pageId="0" pageNumber="1">Ibid.</emphasis>
, Klut Nature Reserve, Pucuk Lembang, 40 m, 8 Jul 1985, W.J.J.O. de Wilde &amp; B.E.E. de Wilde-Duyfjes 19788 (BO).
<emphasis bold="true" pageId="0" pageNumber="1">Bangka-Belitung</emphasis>
: Bangka, T. Horsfield 19 (K);
<emphasis italics="true" pageId="0" pageNumber="1">Ibid.</emphasis>
, Menumbing, J.E. Teijsmann s.n. (BM, BO, K, U).
<emphasis bold="true" pageId="0" pageNumber="1">Bengkulu</emphasis>
: Lebong Tandai, Apr 1922, C.J. Brooks 6675 (K).
<emphasis bold="true" pageId="0" pageNumber="1">Jambi</emphasis>
: Bangko, 160 m, 10 Aug 1925, O. Posthumus 687 (BO, L); Harapan Rain forest, 31 Mar 2013, Wardi et al. BOHK 390 (BO, K).
<emphasis bold="true" pageId="0" pageNumber="1">North Sumatra</emphasis>
: Asahan, Dolok Tomuan, 1000 m, 20 Aug 1936, Rahmat Si Boeea 9964 (L); Padang Sidempuan, Padang Lawas, Huta Imbaru, 20-21 Jun 1933, Rahmat Si Toroes 4634 (K, L); South Tapanuli, Batang Toru, 640 m, 6 Jun 2003, W. Takeuchi &amp; E.N. Sambas 18228 (BO); Pematang Siantar, 6 Apr 1954, A.H.G. Alston 15233 (BM, L); Besitang, Sikundur, 100 m, 15 Aug 1971, K. Iwatsuki et al. S-401 (BO, L).
<emphasis bold="true" pageId="0" pageNumber="1">Riau</emphasis>
: Indragiri Hulu, Berapit, 13 Apr 1939, P. Buwalda 6535 (BO, K, L, PNH); Rengat, Bukit Tigapuluh, 100 m, 14 Nov 1988, J.S. Burley &amp; Tukirin 1472 (BO, K, L).
<emphasis bold="true" pageId="0" pageNumber="1">Riau Archipelago</emphasis>
: Bintan Island, Gunung Bintan, 40 m, 12 Jun 1919, H.A.B.
<normalizedToken originalValue="Bünnemeijer">Buennemeijer</normalizedToken>
6114 (BO, L).
<emphasis bold="true" pageId="0" pageNumber="1">West Sumatra</emphasis>
: Tanah Datar, Lembah Anai, 600 m, 24 Dec 1983, M. Rahayu &amp; Maskuri 473 (BO, K); Batu Sangkar, Mt. Sago, 1200 m, 10 Mar 1989, H. Nagamasu 3773 (BO, L); Lima Puluh Kota, Gunung Malintang, 1100 m, 18 Jul 1918, H.A.B.
<normalizedToken originalValue="Bünnemeijer">Buennemeijer</normalizedToken>
3587 (BO, K, L, P);
<emphasis italics="true" pageId="0" pageNumber="1">Ibid.</emphasis>
, Harau Valley, Sarasah Bonta, 700 m, 12 Sep 2017, A. Kartonegoro 1088 (BO, L); Lubuk Sikaping, Mt. Gadang, 700 m, 15 Jun 1953, J. van Borssum-Waalkes 1888 (BO); Paauw, P.W. Korthals s.n. (L).
<emphasis bold="true" pageId="0" pageNumber="1">Central Kalimantan</emphasis>
: Barito Ulu, 25 Jun 1990, C.E. Ridsdale PBU 692 (BO, L); Bukit Raya, Tumbang Tapi, 100 m, 18 Jan 1983, J.F. Veldkamp 8297 (BO, L).
<emphasis bold="true" pageId="0" pageNumber="1">East Kalimantan</emphasis>
: West Kutai, Besilau Tuwa, 20 m, 22 Jun 1925, F.H. Endert 1593 (BO, L); Sangkulirang, Sampayau, 25 m, 9 Jun 1937, Aet 669 (BO, K, L); East Kutai, Sungai Menubar Region, 5 m, 5 Jun 1951, A.J.G.H. Kostermans 4960 (BO, K, L); Wanariset Research Area, Semoi Road, 20 m, 10 Mar 1992, Ambriansyah &amp; Z. Arifin AA 468 (K, L); Between Papadi &amp; Pamilau, 700-800 m, 9 Aug 1981, R. Geesink 9295 (L, P); Long Iram, Maruwai, Lampunut, 310 m, 19 Mar 1999, P.J.A. Kessler et al. 2657 (L); Wain River, 24 Jun 1910, L.M.R. Rutten 170 (U).
<emphasis bold="true" pageId="0" pageNumber="1">North Kalimantan</emphasis>
: Tarakan, 25 Oct 1953, W. Meijer 1866 (BO, K, L).
<emphasis bold="true" pageId="0" pageNumber="1">South Kalimantan</emphasis>
: Hayup, 28 Jun 1908, HJ.P. Winkler 2627 (BM, BO, K, L, P).
<emphasis bold="true" pageId="0" pageNumber="1">West Kalimantan</emphasis>
: Sungai
<normalizedToken originalValue="Bluu">Blu'u</normalizedToken>
, 1896, Jaheri 704 (BO); Ketapang, Gunung Palung National Park, Cabang Panti, 20 m, 24 Oct 1996, T.G. Laman et al. 110 (BO, K, L).
<emphasis bold="true" pageId="0" pageNumber="1">PHILIPPINES. Palawan</emphasis>
: Taytay, Ibangley, Pagdanan Range, 50-70 m, 31 Jan 1991, D.D. Soejarto &amp; E. Fernando, 7458 (L);
<emphasis italics="true" pageId="0" pageNumber="1">Ibid.</emphasis>
, B.C. Stone et al. PPI 368 (L).
</paragraph>
</subSubSection>
</treatment>
</document>