treatments-xml/data/1D/42/F2/1D42F25BFF85FF9781D0FF565C6CDA63.xml

<document id="1311F8343C28D76043E79A2D3183C934" ID-DOI="10.11646/zootaxa.4007.3.9" ID-GBIF-Dataset="6d1f4ddf-caa4-4e3d-9e95-47b5c572b706" ID-ISSN="1175-5326" ID-Zenodo-Dep="237728" ID-ZooBank="43A658BB-8864-4C85-B8E5-B7801A07A6F3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461219940471" checkinUser="plazi" docAuthor="Houart, Roland" docDate="2015" docId="1D42F25BFF85FF9781D0FF565C6CDA63" docLanguage="en" docName="zt04007p432.pdf" docOrigin="Zootaxa 4007 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Typhinellus jacolombi Houart, 2015, n. sp." docType="treatment" docVersion="7" lastPageNumber="431" masterDocId="E17B8A23FF84FF938147FFCE5F07DC3C" masterDocTitle="Description of a new species of Typhinellus (Gastropoda: Muricidae: Typhinae) from the Western Atlantic" masterLastPageNumber="432" masterPageNumber="427" pageNumber="428" updateTime="1698603848174" updateUser="plazi">
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<mods:title id="7A0E2165968C0AD9F655069525829FCF">Description of a new species of Typhinellus (Gastropoda: Muricidae: Typhinae) from the Western Atlantic</mods:title>
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<mods:namePart id="D6BA3682730E95AF8B8AFC3D8FC89B23">Houart, Roland</mods:namePart>
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<mods:date id="A8C140307A11788D1860A14478489EFE">2015</mods:date>
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<treatment id="1D42F25BFF85FF9781D0FF565C6CDA63" ID-DOI="http://doi.org/10.5281/zenodo.6108033" ID-GBIF-Taxon="119631457" ID-Zenodo-Dep="6108033" LSID="urn:lsid:plazi:treatment:1D42F25BFF85FF9781D0FF565C6CDA63" httpUri="http://treatment.plazi.org/id/1D42F25BFF85FF9781D0FF565C6CDA63" lastPageId="4" lastPageNumber="431" pageId="1" pageNumber="428">
<subSubSection id="DDF110C6FF85FF9281D0FF565E59DCED" pageId="1" pageNumber="428" type="nomenclature">
<paragraph id="9554434DFF85FF9281D0FF565EEADC8E" blockId="1.[151,493,152,209]" box="[151,493,152,178]" pageId="1" pageNumber="428">
<heading id="CE1CF421FF85FF9281D0FF565EEADC8E" bold="true" box="[151,493,152,178]" fontSize="11" level="1" pageId="1" pageNumber="428" reason="1">
<emphasis id="A79F9F5FFF85FF9281D0FF565EEADC8E" bold="true" box="[151,493,152,178]" pageId="1" pageNumber="428">
<taxonomicName id="52EB38CEFF85FF9281D0FF565EA4DC8E" ID-CoL="7DB6H" box="[151,419,152,178]" class="Gastropoda" family="Muricidae" genus="Typhinellus" kingdom="Animalia" order="Neogastropoda" pageId="1" pageNumber="428" phylum="Mollusca" rank="species" species="jacolombi" status="sp. nov.">
<emphasis id="A79F9F5FFF85FF9281D0FF565EA4DC8E" bold="true" box="[151,419,152,178]" italics="true" pageId="1" pageNumber="428">Typhinellus jacolombi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCAC2224FF85FF9280ECFF565EEADC8E" box="[427,493,152,178]" pageId="1" pageNumber="428" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="9554434DFF85FF9281D0FF745E59DCED" blockId="1.[151,493,152,209]" box="[151,350,186,209]" pageId="1" pageNumber="428">
(
<figureCitation id="0DD05FC8FF85FF9281D9FF745FE6DCED" box="[158,225,186,209]" captionStart="FIGURE 1" captionStartId="1.[151,244,1898,1920]" captionTargetBox="[226,1359,453,1882]" captionTargetId="figure@1.[226,1360,447,1883]" captionTargetPageId="1" captionText="FIGURE 1. Spiral cord morphology. a – c. Typhinellus jacolombi n. sp. a. Panama, Portobelo Bay, 37 – 74 m, holotype MNHN IM- 2000 - 30465, length: 19.2 mm; b. Panama, Portobelo Bay, 60 m, muddy substrate, coll. 1989, paratype RH, length: 28.7 mm; c. Panama, Portobelo, 19 – 37 m, paratype JC, length: 19.3 mm; d. Typhinellus labiatus (Cristofori &amp; Jan, 1832), Kerkennah, Tunisia, 0.5 m, RH, length: 20.1 mm; e. Typhinellus lamyi Garrigues &amp; Merle, 2014, Guadeloupe, Port Louis, 66 m, holotype MNHN-IM- 2013 - 7776, length: 13.1 mm." httpUri="https://zenodo.org/record/237729/files/figure.png" pageId="1" pageNumber="428">Figs 1</figureCitation>
a–c; 2 a–k)
</paragraph>
</subSubSection>
<subSubSection id="DDF110C6FF85FF9181D0FECE5C70DBA2" lastPageId="2" lastPageNumber="429" pageId="1" pageNumber="428" type="materials_examined">
<paragraph id="9554434DFF85FF9281D0FECE5BA4DD60" blockId="1.[151,1435,256,417]" pageId="1" pageNumber="428">
<emphasis id="A79F9F5FFF85FF9281D0FECE5E3BDD2B" bold="true" box="[151,316,256,279]" pageId="1" pageNumber="428">
<typeStatus id="4A50FDEFFF85FF9281D0FECE5FC8DD2B" box="[151,207,256,279]" pageId="1" pageNumber="428">Type</typeStatus>
material.
</emphasis>
<typeStatus id="4A50FDEFFF85FF928004FECE5EA1DD2B" box="[323,422,256,279]" pageId="1" pageNumber="428" type="holotype">Holotype</typeStatus>
(
<collectionCode id="F3FADB88FF85FF9280F3FECE5D00DD2B" LSID="urn:lsid:biocol.org:col:34988" box="[436,519,256,279]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="428">MNHN</collectionCode>
<collectionCode id="F3FADB88FF85FF928349FECE5D3FDD2B" box="[526,568,256,279]" pageId="1" pageNumber="428">IM-</collectionCode>
2000-30465: 
<figureCitation id="0DD05FC8FF85FF928381FECE5C02DD2B" box="[710,773,256,279]" captionStart="FIGURE 1" captionStartId="1.[151,244,1898,1920]" captionTargetBox="[226,1359,453,1882]" captionTargetId="figure@1.[226,1360,447,1883]" captionTargetPageId="1" captionText="FIGURE 1. Spiral cord morphology. a – c. Typhinellus jacolombi n. sp. a. Panama, Portobelo Bay, 37 – 74 m, holotype MNHN IM- 2000 - 30465, length: 19.2 mm; b. Panama, Portobelo Bay, 60 m, muddy substrate, coll. 1989, paratype RH, length: 28.7 mm; c. Panama, Portobelo, 19 – 37 m, paratype JC, length: 19.3 mm; d. Typhinellus labiatus (Cristofori &amp; Jan, 1832), Kerkennah, Tunisia, 0.5 m, RH, length: 20.1 mm; e. Typhinellus lamyi Garrigues &amp; Merle, 2014, Guadeloupe, Port Louis, 66 m, holotype MNHN-IM- 2013 - 7776, length: 13.1 mm." httpUri="https://zenodo.org/record/237729/files/figure.png" pageId="1" pageNumber="428">Figs 1</figureCitation>
a; 2a–c), Caribbean Sea, 
<collectingCountry id="EDFC03DDFF85FF928555FECE5B6EDD2B" box="[1042,1129,256,279]" name="Panama" pageId="1" pageNumber="428">Panama</collectingCountry>
, Portobelo Bay, 
<quantity id="5213EEA8FF85FF92845EFECE5A7DDD2B" box="[1305,1402,256,279]" metricMagnitude="1" metricUnit="m" metricValue="5.55" metricValueMax="7.4" metricValueMin="3.7" pageId="1" pageNumber="428" unit="m" value="55.5" valueMax="74.0" valueMin="37.0">
37– 
<specimenCount id="83ED88C4FF85FF928407FECE5A7DDD2B" box="[1344,1402,256,279]" pageId="1" pageNumber="428" type="generic">74 m</specimenCount>
</quantity>
. 
<typeStatus id="4A50FDEFFF85FF928180FEEC5E34DD05" box="[199,307,290,313]" pageId="1" pageNumber="428" type="paratype">Paratypes</typeStatus>
: Caribbean Sea, 
<collectingCountry id="EDFC03DDFF85FF9280A3FEEC5D3CDD05" box="[484,571,290,313]" name="Panama" pageId="1" pageNumber="428">Panama</collectingCountry>
, Portobelo Bay, east of Canal Zone, 
<quantity id="5213EEA8FF85FF928287FEEC5CFFDD05" box="[960,1016,290,313]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="428" unit="m" value="60.0">60 m</quantity>
, on muddy sand substrate, 1989, 2 RH; Caribbean Sea, 
<collectingCountry id="EDFC03DDFF85FF928078FE8B5E91DD60" box="[319,406,325,348]" name="Panama" pageId="1" pageNumber="428">Panama</collectingCountry>
, Portobelo, 
<quantity id="5213EEA8FF85FF928352FE8B5D71DD60" box="[533,630,325,348]" metricMagnitude="1" metricUnit="m" metricValue="2.8" metricValueMax="3.7" metricValueMin="1.9" pageId="1" pageNumber="428" unit="m" value="28.0" valueMax="37.0" valueMin="19.0">19–37 m</quantity>
, 1 JC; Caribbean Sea, 
<collectingCountry id="EDFC03DDFF85FF928220FE8B5CBCDD60" box="[871,955,325,348]" name="Panama" pageId="1" pageNumber="428">Panama</collectingCountry>
(no other data), 1 JC.
</paragraph>
<paragraph id="9554434DFF85FF928180FEA65C51DD9D" blockId="1.[151,1435,256,417]" pageId="1" pageNumber="428">
<emphasis id="A79F9F5FFF85FF928180FEA65EE8DD43" bold="true" box="[199,495,360,383]" pageId="1" pageNumber="428">Other material examined.</emphasis>
Caribbean Sea, 
<collectingCountry id="EDFC03DDFF85FF9283E6FEA65DF2DD43" box="[673,757,360,383]" name="Panama" pageId="1" pageNumber="428">Panama</collectingCountry>
(no other data) 
<quantity id="5213EEA8FF85FF9282E3FEA65CDADD43" box="[932,989,360,383]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="428" unit="m" value="25.0">25 m</quantity>
, on coralline algae, 1 JC; Caribbean Sea, 
<collectingCountry id="EDFC03DDFF85FF9281D0FE445FE9DD9D" box="[151,238,394,417]" name="Panama" pageId="1" pageNumber="428">Panama</collectingCountry>
, off San Blas Islands, dredged at 
<quantity id="5213EEA8FF85FF928316FE445D90DD9D" box="[593,663,394,417]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="428" unit="m" value="120.0">120 m</quantity>
, 
<date id="E155658DFF85FF9283E5FE445C2CDD9D" box="[674,811,394,417]" pageId="1" pageNumber="428" value="2000-11-01">Nov. 2000, 1</date>
JC.
</paragraph>
<caption id="C19413C5FF85FF9281D0F8A45E7FDBC9" httpUri="https://zenodo.org/record/237729/files/figure.png" pageId="1" pageNumber="428" targetBox="[226,1359,453,1882]" targetPageId="1">
<paragraph id="9554434DFF85FF9281D0F8A45E7FDBC9" blockId="1.[151,1436,1898,2037]" pageId="1" pageNumber="428">
<emphasis id="A79F9F5FFF85FF9281D0F8A45E08DBBC" bold="true" box="[151,271,1898,1920]" pageId="1" pageNumber="428">FIGURE 1.</emphasis>
Spiral cord morphology. a–c. 
<taxonomicName id="52EB38CEFF85FF92837CF8A45C16DB43" box="[571,785,1898,1919]" class="Gastropoda" family="Muricidae" genus="Typhinellus" kingdom="Animalia" order="Neogastropoda" pageId="1" pageNumber="428" phylum="Mollusca" rank="species" species="jacolombi" status="sp. nov.">
<emphasis id="A79F9F5FFF85FF92837CF8A45C16DB43" box="[571,785,1898,1919]" italics="true" pageId="1" pageNumber="428">Typhinellus jacolombi</emphasis>
</taxonomicName>
<emphasis id="A79F9F5FFF85FF92825EF8A45C55DBBC" bold="true" box="[793,850,1898,1920]" pageId="1" pageNumber="428">
<taxonomicNameLabel id="BCAC2224FF85FF92825EF8A45C55DBBC" box="[793,850,1898,1920]" pageId="1" pageNumber="428" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
a. Panama, Portobelo Bay, 37–74 m, holotype MNHN IM- 2000-30465, length: 19.2 mm; b. Panama, Portobelo Bay, 60 m, muddy substrate, coll. 1989, paratype RH, length: 28.7 mm; c. Panama, Portobelo, 19–37 m, paratype JC, length: 19.3 mm; d. 
<taxonomicName id="52EB38CEFF85FF9283BDF86B5BACDB86" authority="Cristofori &amp; Jan, 1832" authorityName="Cristofori &amp; Jan" authorityYear="1832" box="[762,1195,1957,1978]" class="Gastropoda" family="Muricidae" genus="Typhinellus" kingdom="Animalia" order="Neogastropoda" pageId="1" pageNumber="428" phylum="Mollusca" rank="species" species="labiatus">
<emphasis id="A79F9F5FFF85FF9283BDF86B5CBCDB86" box="[762,955,1957,1978]" italics="true" pageId="1" pageNumber="428">Typhinellus labiatus</emphasis>
(Cristofori &amp; Jan, 1832)
</taxonomicName>
, Kerkennah, Tunisia, 0.5 m, RH, length: 20.1 mm; e. 
<taxonomicName id="52EB38CEFF85FF9280ECF80C5C53DBEB" authority="Garrigues &amp; Merle, 2014" authorityName="Garrigues &amp; Merle" authorityYear="2014" box="[427,852,1986,2007]" class="Gastropoda" family="Muricidae" genus="Typhinellus" kingdom="Animalia" order="Neogastropoda" pageId="1" pageNumber="428" phylum="Mollusca" rank="species" species="lamyi">
<emphasis id="A79F9F5FFF85FF9280ECF80C5D52DBEB" box="[427,597,1986,2007]" italics="true" pageId="1" pageNumber="428">Typhinellus lamyi</emphasis>
Garrigues &amp; Merle, 2014
</taxonomicName>
, Guadeloupe, Port Louis, 66 m, holotype MNHN-IM-2013- 7776, length: 13.1 mm.
</paragraph>
</caption>
<caption id="C19413C5FF86FF9181D0F9195ED2DB5D" httpUri="https://zenodo.org/record/237730/files/figure.png" pageId="2" pageNumber="429" targetBox="[191,1396,191,1717]" targetPageId="2">
<paragraph id="9554434DFF86FF9181D0F9195ED2DB5D" blockId="2.[151,1436,1751,1889]" pageId="2" pageNumber="429">
<emphasis id="A79F9F5FFF86FF9181D0F9195E0ADAD1" bold="true" box="[151,269,1751,1773]" pageId="2" pageNumber="429">FIGURE 2.</emphasis>
a–k. 
<taxonomicName id="52EB38CEFF86FF918004F9195D10DAD0" box="[323,535,1751,1772]" class="Gastropoda" family="Muricidae" genus="Typhinellus" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="429" phylum="Mollusca" rank="species" species="jacolombi" status="sp. nov.">
<emphasis id="A79F9F5FFF86FF918004F9195D10DAD0" box="[323,535,1751,1772]" italics="true" pageId="2" pageNumber="429">Typhinellus jacolombi</emphasis>
</taxonomicName>
<emphasis id="A79F9F5FFF86FF918359F9195D52DAD1" bold="true" box="[542,597,1751,1773]" pageId="2" pageNumber="429">
<taxonomicNameLabel id="BCAC2224FF86FF918359F9195D52DAD1" box="[542,597,1751,1773]" pageId="2" pageNumber="429" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
a–c. Panama, Portobelo Bay, 37–74 m, holotype MNHN IM-2000-30465, length: 19.2 mm; d–f. Panama, Portobelo Bay, 60 m, muddy substrate, coll. 1989, paratype RH, length: 28.7 mm; g–h. Panama, Portobelo, 19–37 m, paratype JC, length: 19.3 mm; i–j. Panama, Caribbean (no other data), paratype JC, length: 21.7 mm; k. Protoconch of specimen figured in i–j (scale bar 500 µm). l–n. 
<taxonomicName id="52EB38CEFF86FF91834BF8E15CB3DB78" authority="Garrigues &amp; Merle, 2014" authorityName="Garrigues &amp; Merle" authorityYear="2014" box="[524,948,1839,1860]" class="Gastropoda" family="Muricidae" genus="Typhinellus" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="429" phylum="Mollusca" rank="species" species="lamyi">
<emphasis id="A79F9F5FFF86FF91834BF8E15DB1DB78" box="[524,694,1839,1860]" italics="true" pageId="2" pageNumber="429">Typhinellus lamyi</emphasis>
Garrigues &amp; Merle, 2014
</taxonomicName>
, Guadeloupe, Port Louis, 66 m, holotype MNHN- IM-2013-7776, length: 13.1 mm.
</paragraph>
</caption>
<paragraph id="9554434DFF86FF918180F8495C70DBA2" blockId="2.[151,1436,1927,2019]" box="[199,887,1927,1950]" pageId="2" pageNumber="429">
<emphasis id="A79F9F5FFF86FF918180F8495E59DBA2" bold="true" box="[199,350,1927,1950]" pageId="2" pageNumber="429">
<typeStatus id="4A50FDEFFF86FF918180F8495FF8DBA2" box="[199,255,1927,1950]" pageId="2" pageNumber="429">Type</typeStatus>
locality.
</emphasis>
Portobelo Bay, Caribbean Sea, 
<collectingCountry id="EDFC03DDFF86FF9183F6F8495C0FDBA2" box="[689,776,1927,1950]" name="Panama" pageId="2" pageNumber="429">Panama</collectingCountry>
, 
<quantity id="5213EEA8FF86FF918255F8495C74DBA2" box="[786,883,1927,1950]" metricMagnitude="1" metricUnit="m" metricValue="5.55" metricValueMax="7.4" metricValueMin="3.7" pageId="2" pageNumber="429" unit="m" value="55.5" valueMax="74.0" valueMin="37.0">
37– 
<specimenCount id="83ED88C4FF86FF91827DF8495C74DBA2" box="[826,883,1927,1950]" pageId="2" pageNumber="429" type="generic">74 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="DDF110C6FF86FF918180F8645D14DBDF" pageId="2" pageNumber="429" type="etymology">
<paragraph id="9554434DFF86FF918180F8645D14DBDF" blockId="2.[151,1436,1927,2019]" pageId="2" pageNumber="429">
<emphasis id="A79F9F5FFF86FF918180F8645E40DBFD" bold="true" box="[199,327,1962,1985]" pageId="2" pageNumber="429">Etymology.</emphasis>
The species is named for Jacques Colomb (Marseille, 
<collectingCountry id="EDFC03DDFF86FF9182E4F8645CF7DBFD" box="[931,1008,1962,1985]" name="France" pageId="2" pageNumber="429">France</collectingCountry>
), who kindly donated the 
<typeStatus id="4A50FDEFFF86FF918449F8645A6CDBFD" box="[1294,1387,1962,1985]" pageId="2" pageNumber="429" type="holotype">holotype</typeStatus>
and loaned material from his collection.
</paragraph>
</subSubSection>
<caption id="C19413C5FF87FF9081D0F9BF5EB2DAE2" httpUri="https://zenodo.org/record/237731/files/figure.png" pageId="3" pageNumber="430" targetBox="[226,1359,191,1627]" targetPageId="3">
<paragraph id="9554434DFF87FF9081D0F9BF5EB2DAE2" blockId="3.[151,1436,1649,1758]" pageId="3" pageNumber="430">
<emphasis id="A79F9F5FFF87FF9081D0F9BF5E09DABB" bold="true" box="[151,270,1649,1671]" pageId="3" pageNumber="430">FIGURE 3.</emphasis>
<taxonomicName id="52EB38CEFF87FF908051F9BF5DD5DABA" authority="Cristofori &amp; Jan, 1832" authorityName="Cristofori &amp; Jan" authorityYear="1832" box="[278,722,1649,1670]" class="Gastropoda" family="Muricidae" genus="Typhinellus" kingdom="Animalia" order="Neogastropoda" pageId="3" pageNumber="430" phylum="Mollusca" rank="species" species="labiatus">
<emphasis id="A79F9F5FFF87FF908051F9BF5EDEDABA" box="[278,473,1649,1670]" italics="true" pageId="3" pageNumber="430">Typhinellus labiatus</emphasis>
(Cristofori &amp; Jan, 1832)
</taxonomicName>
. a–c. Kerkennah, Tunisia, 0.5 m, RH, length: 20.1 mm; d–f. Kerkennah, Tunisia, 0.5 m, RH, length: 19.7 mm; g–h. Kerkennah, Tunisia, 0.5 m, RH, length: 18.1 mm; i. Mallorca, Porto Colom, 1978, RH, length: 16.6 mm; j. Italy, Livorno, 26 m, 1978, RH, length: 13.6 mm; k–m. Sardinia, RH, 14.4 mm; n. Protoconch, Sicily, Brucoli, 1990, RH (scale bar 500 µm).
</paragraph>
</caption>
<subSubSection id="DDF110C6FF87FF978180F8C75E83DE35" lastPageId="4" lastPageNumber="431" pageId="3" pageNumber="430" type="description">
<paragraph id="9554434DFF87FF978180F8C75BF4DD60" blockId="3.[151,1436,1801,2032]" lastBlockId="4.[151,1436,152,1631]" lastPageId="4" lastPageNumber="431" pageId="3" pageNumber="430">
<emphasis id="A79F9F5FFF87FF908180F8C75D0CDB1C" bold="true" box="[199,523,1801,1824]" pageId="3" pageNumber="430">
Description of the 
<typeStatus id="4A50FDEFFF87FF9080E5F8C75D01DB1C" box="[418,518,1801,1824]" pageId="3" pageNumber="430" type="holotype">holotype</typeStatus>
.
</emphasis>
Shell large for the genus, 
<quantity id="5213EEA8FF87FF908272F8C75C9FDB1C" box="[821,920,1801,1824]" metricMagnitude="-2" metricUnit="m" metricValue="1.92" pageId="3" pageNumber="430" unit="mm" value="19.2">19.2 mm</quantity>
in length at maturity. Length/width ratio 1.75, broad, nearly smooth, lightly built. Subsutural ramp broad, gently sloping, weakly concave. Light tan with lighter colored siphonal canal, dark brown subsutural area and 3 or 4 light brown spots on abapical part of outer apertural lip. Aperture white within. Spire high with 5 broad, convex, angulate, strongly shouldered, weakly spinose teleoconch whorls. Suture of whorls impressed. Protoconch eroded in the 
<typeStatus id="4A50FDEFFF87FF908274F85A5C93DB97" box="[819,916,1940,1963]" pageId="3" pageNumber="430" type="holotype">holotype</typeStatus>
. Axial sculpture of teleoconch whorls consisting of 4 high, thin, sharp lamellate varices per whorl, each with a short, open, inwardly curved shoulder spine. Apertural varix very broad, flange-like, lightly reflected dorsally, with broadly open, short spinelets at outer edge. Variceal flange extended to nearly the tip of siphonal canal. Shoulder spine of apertural lamellae strongly dorsally curved dorsally, connected to last teleoconch whorl by a broad, high laminar buttress (partition). Spiral sculpture of low, rounded, broad, smooth primary and secondary cords, and narrow tertiary cords: P1 (anal tube), P2 (shoulder spine), t, s2, t, P3, s3, P4, s4, P5, P6, ADP, MP and ABP. P1 with long, tapering, broad, ventrally sealed anal tube, forming an angle of approximately 85° with axis of shell. Only last tube completely hollow and functional, older tubes broken off and closed. Other spiral cords, from P2 to P6 corresponding to short, broadly open, blunt spinelets at apertural varix.
</paragraph>
<paragraph id="9554434DFF80FF978180FEA65FCEDDF8" blockId="4.[151,1436,152,1631]" pageId="4" pageNumber="431">
Aperture moderately large, roundly ovate, forming a continuous peristome. Columellar lip narrow, smooth within. Siphonal canal short, broad, tip partly broken in 
<typeStatus id="4A50FDEFFF80FF9783E2FE445C02DD9D" box="[677,773,394,417]" pageId="4" pageNumber="431" type="holotype">holotype</typeStatus>
, ventrally sealed. Left side of canal weakly overlapping right side.
</paragraph>
<paragraph id="9554434DFF80FF978180FE1E5C41DDDB" blockId="4.[151,1436,152,1631]" box="[199,838,464,487]" pageId="4" pageNumber="431">Operculum light brown, thin, ovate, with subapical nucleus.</paragraph>
<paragraph id="9554434DFF80FF978180FE3C5E83DE35" blockId="4.[151,1436,152,1631]" box="[199,388,498,521]" pageId="4" pageNumber="431">Radula unknown.</paragraph>
</subSubSection>
<subSubSection id="DDF110C6FF80FF978180FDDB5CD6DEC0" pageId="4" pageNumber="431" type="distribution">
<paragraph id="9554434DFF80FF978180FDDB5E22DE73" blockId="4.[151,1436,152,1631]" pageId="4" pageNumber="431">
<emphasis id="A79F9F5FFF80FF978180FDDB5E5EDE10" bold="true" box="[199,345,533,556]" pageId="4" pageNumber="431">Distribution.</emphasis>
Caribbean Sea, off the coast of northern 
<collectingCountry id="EDFC03DDFF80FF978252FDDB5C6BDE10" box="[789,876,533,556]" name="Panama" pageId="4" pageNumber="431">Panama</collectingCountry>
, from the San Blas Islands to the Gulf of Portobelo, in 
<quantity id="5213EEA8FF80FF9781F5FDF65E26DE73" box="[178,289,568,591]" metricMagnitude="1" metricUnit="m" metricValue="7.25" metricValueMax="12.0" metricValueMin="2.5" pageId="4" pageNumber="431" unit="m" value="72.5" valueMax="120.0" valueMin="25.0">
25– 
<specimenCount id="83ED88C4FF80FF97819CFDF65E26DE73" box="[219,289,568,591]" pageId="4" pageNumber="431" type="generic">120 m</specimenCount>
</quantity>
.
</paragraph>
<paragraph id="9554434DFF80FF978180FD945CD6DEC0" blockId="4.[151,1436,152,1631]" pageId="4" pageNumber="431">
<bibRefCitation id="F17A3EBCFF80FF978180FD945EAFDE4D" author="Gertman" box="[199,424,602,625]" pageId="4" pageNumber="431" refString="Gertman, R. L. (1969) Cenozoic Typhinae (Mollusca: Gastropoda) of the Western Atlantic region. Tulane Studies in Geology and Paleontology, 7 (4), 143 - 191." type="journal article" year="1969">Gertman (1969: 156)</bibRefCitation>
also mentioned Portobelo for the species he identified as 
<taxonomicName id="52EB38CEFF80FF97855FFD945A55DE4D" box="[1048,1362,602,625]" class="Gastropoda" family="Muricidae" genus="Typhis" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="431" phylum="Mollusca" rank="species" species="sowerbii" subGenus="Typhinellus">
<emphasis id="A79F9F5FFF80FF97855FFD945B5ADE4D" box="[1048,1117,602,625]" italics="true" pageId="4" pageNumber="431">Typhis</emphasis>
(
<emphasis id="A79F9F5FFF80FF97852AFD945BE0DE4D" box="[1133,1255,602,625]" italics="true" pageId="4" pageNumber="431">Typhinellus</emphasis>
) 
<emphasis id="A79F9F5FFF80FF9785B0FD945A55DE4D" box="[1271,1362,602,625]" italics="true" pageId="4" pageNumber="431">sowerbii</emphasis>
</taxonomicName>
but he also added Egmont Key, Florida, at 41 fms (
<quantity id="5213EEA8FF80FF9783C3FDB35DC6DEA8" box="[644,705,637,660]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="4" pageNumber="431" unit="m" value="75.0">75 m</quantity>
), the Texas coast at 28 fms (
<quantity id="5213EEA8FF80FF978544FDB35B39DEA8" box="[1027,1086,637,660]" metricMagnitude="1" metricUnit="m" metricValue="5.1" pageId="4" pageNumber="431" unit="m" value="51.0">51 m</quantity>
) and the Leeward Islands (near 
<collectingCountry id="EDFC03DDFF80FF9781D0FD6E5FDADE8B" box="[151,221,672,695]" name="Saint Kitts and Nevis" pageId="4" pageNumber="431">Nevis)</collectingCountry>
at 50–60 fms (
<quantity id="5213EEA8FF80FF9780C6FD6E5EF6DE8B" box="[385,497,672,695]" metricMagnitude="2" metricUnit="m" metricValue="1.005" metricValueMax="1.1" metricValueMin="0.9099999999999999" pageId="4" pageNumber="431" unit="m" value="100.5" valueMax="110.0" valueMin="91.0">91–110 m</quantity>
). The specimen illustrated by 
<bibRefCitation id="F17A3EBCFF80FF978272FD6E5CDBDE8B" author="Gertman" box="[821,988,672,695]" pageId="4" pageNumber="431" refString="Gertman, R. L. (1969) Cenozoic Typhinae (Mollusca: Gastropoda) of the Western Atlantic region. Tulane Studies in Geology and Paleontology, 7 (4), 143 - 191." type="journal article" year="1969">Gertman (1969</bibRefCitation>
: 157, pl. 1, figs 5a–5c) from the Leeward Islands is close to 
<taxonomicName id="52EB38CEFF80FF978019FD0C5EB4DEE5" box="[350,435,706,729]" class="Gastropoda" family="Muricidae" genus="Typhinellus" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="431" phylum="Mollusca" rank="species" species="lamyi">
<emphasis id="A79F9F5FFF80FF978019FD0C5EB4DEE5" box="[350,435,706,729]" italics="true" pageId="4" pageNumber="431">T. lamyi</emphasis>
</taxonomicName>
and is here considered conspecific with that species. Having not seen the other specimens, I will at this time not enlarge the geographical range to the other two localities.
</paragraph>
</subSubSection>
<subSubSection id="DDF110C6FF80FF978180FCC65C6CDA63" pageId="4" pageNumber="431" type="discussion">
<paragraph id="9554434DFF80FF978180FCC65E3EDFF0" blockId="4.[151,1436,152,1631]" pageId="4" pageNumber="431">
<emphasis id="A79F9F5FFF80FF978180FCC65E32DF23" bold="true" box="[199,309,776,799]" pageId="4" pageNumber="431">Remarks.</emphasis>
The presence of Indo-West Pacific species in the Caribbean is highly improbable and their shell morphology differs in many ways from 
<taxonomicName id="52EB38CEFF80FF978325FCE45C54DF7D" box="[610,851,810,833]" class="Gastropoda" family="Muricidae" genus="Typhinellus" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="431" phylum="Mollusca" rank="species" species="jacolombi" status="sp. nov.">
<emphasis id="A79F9F5FFF80FF978325FCE45C54DF7D" box="[610,851,810,833]" italics="true" pageId="4" pageNumber="431">Typhinellus jacolombi</emphasis>
</taxonomicName>
<emphasis id="A79F9F5FFF80FF978227FCE45CA6DF7D" bold="true" box="[864,929,810,833]" pageId="4" pageNumber="431">
<taxonomicNameLabel id="BCAC2224FF80FF978227FCE45CA6DF7D" box="[864,929,810,833]" pageId="4" pageNumber="431" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
<taxonomicName id="52EB38CEFF80FF9782E9FCE45A4DDF7D" authority="Garrard, 1963" authorityName="Garrard" authorityYear="1963" box="[942,1354,810,833]" class="Gastropoda" family="Muricidae" genus="Typhinellus" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="431" phylum="Mollusca" rank="species" species="occlusus">
<emphasis id="A79F9F5FFF80FF9782E9FCE45B88DF7D" box="[942,1167,810,833]" italics="true" pageId="4" pageNumber="431">Typhinellus occlusus</emphasis>
(
<bibRefCitation id="F17A3EBCFF80FF9785E3FCE45A46DF7D" author="Garrard" box="[1188,1345,810,833]" pageId="4" pageNumber="431" refString="Garrard, T. A. (1963) New species of Mollusca from eastern Australia. Journal of the Malacological Society of Australia, 1 (7), 42 - 46." type="journal article" year="1963">Garrard, 1963</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="52EB38CEFF80FF9784CFFCE45E9DDF58" authority="Houart, 1991" authorityName="Houart" authorityYear="1991" class="Gastropoda" family="Muricidae" genus="Typhinellus" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="431" phylum="Mollusca" rank="species" species="insolitus">
<emphasis id="A79F9F5FFF80FF9784CFFCE45FF6DF58" italics="true" pageId="4" pageNumber="431">T. insolitus</emphasis>
(
<bibRefCitation id="F17A3EBCFF80FF978046FC835E95DF58" author="Houart" box="[257,402,845,868]" pageId="4" pageNumber="431" refString="Houart, R. (1991) Description of thirteen new species of Muricidae (Gastropoda) from Australia and the New Caledonian region, with range extensions to South Africa. Journal of the Malacological Society of Australia, 12, 35 - 55." type="journal article" year="1991">Houart, 1991</bibRefCitation>
)
</taxonomicName>
have a narrower shell with fewer spiral cords and a more strongly constricted last teleoconch whorl. 
<taxonomicName id="52EB38CEFF80FF9781A0FCBE5EBEDFBB" box="[231,441,880,903]" class="Gastropoda" family="Muricidae" genus="Typhinellus" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="431" phylum="Mollusca" rank="species" species="bicolor">
<emphasis id="A79F9F5FFF80FF9781A0FCBE5EBEDFBB" box="[231,441,880,903]" italics="true" pageId="4" pageNumber="431">Typhinellus bicolor</emphasis>
</taxonomicName>
and 
<taxonomicName id="52EB38CEFF80FF9780B1FCBE5D93DFBB" box="[502,660,880,903]" class="Gastropoda" family="Muricidae" genus="Typhinellus" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="431" phylum="Mollusca" rank="species" species="androyensis">
<emphasis id="A79F9F5FFF80FF9780B1FCBE5D93DFBB" box="[502,660,880,903]" italics="true" pageId="4" pageNumber="431">T. androyensis</emphasis>
</taxonomicName>
also have less spiral cords and a comparatively smaller shell and 
<taxonomicName id="52EB38CEFF80FF9784CFFCBE5FF1DF95" class="Gastropoda" family="Muricidae" genus="Typhinellus" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="431" phylum="Mollusca" rank="species" species="amoenus">
<emphasis id="A79F9F5FFF80FF9784CFFCBE5FF1DF95" italics="true" pageId="4" pageNumber="431">T. amoenus</emphasis>
</taxonomicName>
differs in having a narrower shell with a more strongly constricted last teleoconch whorl and less expanded variceal wings.
</paragraph>
<paragraph id="9554434DFF80FF978180FC165E8BD8A0" blockId="4.[151,1436,152,1631]" pageId="4" pageNumber="431">
Of the two species reported from the western Atlantic, one, 
<taxonomicName id="52EB38CEFF80FF97822CFC165A22DFD3" authority="Gertman, 1969" authorityName="Gertman" authorityYear="1969" box="[875,1317,984,1007]" class="Gastropoda" family="Muricidae" genus="Typhinellus" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="431" phylum="Mollusca" rank="species" species="chipolanus">
<emphasis id="A79F9F5FFF80FF97822CFC165B62DFD3" box="[875,1125,984,1007]" italics="true" pageId="4" pageNumber="431">Typhinellus chipolanus</emphasis>
(
<bibRefCitation id="F17A3EBCFF80FF978530FC165A19DFD3" author="Gertman" box="[1143,1310,984,1007]" pageId="4" pageNumber="431" refString="Gertman, R. L. (1969) Cenozoic Typhinae (Mollusca: Gastropoda) of the Western Atlantic region. Tulane Studies in Geology and Paleontology, 7 (4), 143 - 191." type="journal article" year="1969">Gertman, 1969</bibRefCitation>
)
</taxonomicName>
is a fossil species from the late lower Miocene (Burdigalian) of the Chipola Formation in Florida. It is different from the Recent species in having a smaller and slenderer shell, reaching 
<quantity id="5213EEA8FF80FF978242FBD35C61D808" box="[773,870,1053,1076]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="4" pageNumber="431" unit="mm" value="12.5">12.5 mm</quantity>
in the 
<typeStatus id="4A50FDEFFF80FF9782F2FBD35B12D808" box="[949,1045,1053,1076]" pageId="4" pageNumber="431" type="holotype">holotype</typeStatus>
, and in having fewer and shallower spiral cords. The other species, 
<taxonomicName id="52EB38CEFF80FF9780A1FB8E5D3ED86B" box="[486,569,1088,1111]" class="Gastropoda" family="Muricidae" genus="Typhinellus" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="431" phylum="Mollusca" rank="species" species="lamyi">
<emphasis id="A79F9F5FFF80FF9780A1FB8E5D3ED86B" box="[486,569,1088,1111]" italics="true" pageId="4" pageNumber="431">T. lamyi</emphasis>
</taxonomicName>
(
<figureCitation id="0DD05FC8FF80FF97830EFB8E5D8FD86B" box="[585,648,1088,1111]" captionStart="FIGURE 2. a – k" captionStartId="2.[151,244,1751,1773]" captionTargetBox="[191,1396,191,1717]" captionTargetId="figure@2.[191,1396,191,1717]" captionTargetPageId="2" captionText="FIGURE 2. a – k. Typhinellus jacolombi n. sp. a – c. Panama, Portobelo Bay, 37 – 74 m, holotype MNHN IM- 2000 - 30465, length: 19.2 mm; d – f. Panama, Portobelo Bay, 60 m, muddy substrate, coll. 1989, paratype RH, length: 28.7 mm; g – h. Panama, Portobelo, 19 – 37 m, paratype JC, length: 19.3 mm; i – j. Panama, Caribbean (no other data), paratype JC, length: 21.7 mm; k. Protoconch of specimen figured in i – j (scale bar 500 µm). l – n. Typhinellus lamyi Garrigues &amp; Merle, 2014, Guadeloupe, Port Louis, 66 m, holotype MNHN- IM- 2013 - 7776, length: 13.1 mm." httpUri="https://zenodo.org/record/237730/files/figure.png" pageId="4" pageNumber="431">Fig. 2</figureCitation>
l–n) (
<bibRefCitation id="F17A3EBCFF80FF978381FB8E5B0CD86B" author="Merle" box="[710,1035,1088,1111]" pageId="4" pageNumber="431" refString="Merle, D. &amp; Garrigues, B. (2014) Nine new species of Muricidae Rafinesque, 1815 (Mollusca, Gastropoda) from the French Antilles. Zoosystema, 36 (4), 841 - 864." type="journal article" year="2014">Merle &amp; Garrigues, 2014: 844</bibRefCitation>
) from 
<collectingCountry id="EDFC03DDFF80FF978514FB8E5BD3D86B" box="[1107,1236,1088,1111]" name="Guadeloupe" pageId="4" pageNumber="431">Guadeloupe</collectingCountry>
differs in having a smaller shell relative to the number of teleoconch whorls, in having a less expanded variceal flange and in lacking any secondary spiral cords.
</paragraph>
<paragraph id="9554434DFF80FF978180FB665D7CD9CB" blockId="4.[151,1436,152,1631]" pageId="4" pageNumber="431">
The Mediterranean 
<taxonomicName id="52EB38CEFF80FF9780D9FB665D08D883" box="[414,527,1192,1215]" class="Gastropoda" family="Muricidae" genus="Typhinellus" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="431" phylum="Mollusca" rank="species" species="labiatus">
<emphasis id="A79F9F5FFF80FF9780D9FB665D08D883" box="[414,527,1192,1215]" italics="true" pageId="4" pageNumber="431">T. labiatus</emphasis>
</taxonomicName>
(
<figureCitation id="0DD05FC8FF80FF978366FB665D65D883" box="[545,610,1192,1215]" captionStart="FIGURE 3" captionStartId="3.[151,244,1649,1671]" captionTargetBox="[226,1359,191,1627]" captionTargetId="figure@3.[226,1360,191,1627]" captionTargetPageId="3" captionText="FIGURE 3. Typhinellus labiatus (Cristofori &amp; Jan, 1832). a – c. Kerkennah, Tunisia, 0.5 m, RH, length: 20.1 mm; d – f. Kerkennah, Tunisia, 0.5 m, RH, length: 19.7 mm; g – h. Kerkennah, Tunisia, 0.5 m, RH, length: 18.1 mm; i. Mallorca, Porto Colom, 1978, RH, length: 16.6 mm; j. Italy, Livorno, 26 m, 1978, RH, length: 13.6 mm; k – m. Sardinia, RH, 14.4 mm; n. Protoconch, Sicily, Brucoli, 1990, RH (scale bar 500 µm)." httpUri="https://zenodo.org/record/237731/files/figure.png" pageId="4" pageNumber="431">Fig. 3</figureCitation>
a–n) (
<bibRefCitation id="F17A3EBCFF80FF9783ECFB665CCED883" author="Cristofori" box="[683,969,1192,1215]" pageId="4" pageNumber="431" refString="Cristofori, J. de &amp; Jan, G. (1832) Catalogus in IV. sectiones divisus rerum naturalium in museo exstantium Josephi de Cristofori et Georgii Jan plurium Acad. Scient. et Societ. Nat. Cur. Sodalium complectens adumbrationem oryctognosiae et geognosiae atque prodromum fauna et florae Italiae superioris. Sectio II. Pars I. Fasc. I. Coi Tipi Di Giovanni Pirotta, Milano, 16 pp. http: // dx. doi. org / 10.5962 / bhl. title. 59812" type="book" year="1832">
Cristofori &amp; 
<date id="E155658DFF80FF97827BFB665CA5D883" box="[828,930,1192,1215]" pageId="4" pageNumber="431" value="1832-01">Jan 1832</date>
: 11
</bibRefCitation>
) also occurs in a small area in the eastern Atlantic and has been confused with the new Panamanian species (see above) but it consistently differs in having a generally smaller shell, although one of the 
<typeStatus id="4A50FDEFFF80FF978329FB235DCCD938" box="[622,715,1261,1284]" pageId="4" pageNumber="431" type="syntype">syntypes</typeStatus>
of 
<taxonomicName id="52EB38CEFF80FF9783B6FB235C9AD938" box="[753,925,1261,1284]" class="Gastropoda" family="Muricidae" genus="Typhis" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="431" phylum="Mollusca" rank="species" species="sowerbyi">
<emphasis id="A79F9F5FFF80FF9783B6FB235C9AD938" box="[753,925,1261,1284]" italics="true" pageId="4" pageNumber="431">Typhis sowerbyi</emphasis>
</taxonomicName>
, a synonym, in NHMUK measures 
<quantity id="5213EEA8FF80FF978466FB235A87D938" box="[1313,1408,1261,1284]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" pageId="4" pageNumber="431" unit="mm" value="24.5">24.5 mm</quantity>
in length (
<bibRefCitation id="F17A3EBCFF80FF9781A0FADE5D09D91B" author="Houart" box="[231,526,1296,1319]" pageId="4" pageNumber="431" refString="Houart, R. (2001) A review of the Recent Mediterranean and Northeastern Atlantic species of Muricidae, 1 - 227. [Evolver]" type="book chapter" year="2001">Houart, 2001: 122, fig. 129</bibRefCitation>
). However, that species also differs in having a narrower shell compared to its length, with a length/width ratio of 1.68–1.77 compared to 
<quantity id="5213EEA8FF80FF978391FAFC5C67D975" box="[726,864,1330,1353]" metricMagnitude="-2" metricUnit="m" metricValue="4.0893999999999995" metricValueMax="4.3688" metricValueMin="3.81" pageId="4" pageNumber="431" unit="in" value="1.6099999999999999" valueMax="1.72" valueMin="1.5">1.50–1.72 in</quantity>
<taxonomicName id="52EB38CEFF80FF97822DFAFC5CF6D975" box="[874,1009,1330,1353]" class="Gastropoda" family="Muricidae" genus="Typhinellus" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="431" phylum="Mollusca" rank="species" species="jacolombi" status="sp. nov.">
<emphasis id="A79F9F5FFF80FF97822DFAFC5CF6D975" box="[874,1009,1330,1353]" italics="true" pageId="4" pageNumber="431">T. jacolombi</emphasis>
</taxonomicName>
<emphasis id="A79F9F5FFF80FF9782BBFAFC5B3CD975" bold="true" box="[1020,1083,1330,1353]" pageId="4" pageNumber="431">
<taxonomicNameLabel id="BCAC2224FF80FF9782BBFAFC5B3CD975" box="[1020,1083,1330,1353]" pageId="4" pageNumber="431" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and in having a different spiral sculpture morphology (
<figureCitation id="0DD05FC8FF80FF9780D3FA9B5ED3D950" box="[404,468,1365,1388]" captionStart="FIGURE 1" captionStartId="1.[151,244,1898,1920]" captionTargetBox="[226,1359,453,1882]" captionTargetId="figure@1.[226,1360,447,1883]" captionTargetPageId="1" captionText="FIGURE 1. Spiral cord morphology. a – c. Typhinellus jacolombi n. sp. a. Panama, Portobelo Bay, 37 – 74 m, holotype MNHN IM- 2000 - 30465, length: 19.2 mm; b. Panama, Portobelo Bay, 60 m, muddy substrate, coll. 1989, paratype RH, length: 28.7 mm; c. Panama, Portobelo, 19 – 37 m, paratype JC, length: 19.3 mm; d. Typhinellus labiatus (Cristofori &amp; Jan, 1832), Kerkennah, Tunisia, 0.5 m, RH, length: 20.1 mm; e. Typhinellus lamyi Garrigues &amp; Merle, 2014, Guadeloupe, Port Louis, 66 m, holotype MNHN-IM- 2013 - 7776, length: 13.1 mm." httpUri="https://zenodo.org/record/237729/files/figure.png" pageId="4" pageNumber="431">Fig. 1</figureCitation>
d), 
<taxonomicName id="52EB38CEFF80FF978345FA9B5D73D950" box="[514,628,1365,1388]" class="Gastropoda" family="Muricidae" genus="Typhinellus" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="431" phylum="Mollusca" rank="species" species="labiatus">
<emphasis id="A79F9F5FFF80FF978345FA9B5D73D950" box="[514,628,1365,1388]" italics="true" pageId="4" pageNumber="431">T. labiatus</emphasis>
</taxonomicName>
has the primary cords IP to MP with s 
<quantity id="5213EEA8FF80FF978559FA9B5B4DD950" box="[1054,1098,1365,1388]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="4" pageNumber="431" unit="in" value="2.0">2 in</quantity>
all the examined shells and an occasional s 
<date id="E155658DFF80FF97805FFAB65E3CD9B3" box="[280,315,1400,1423]" pageId="4" pageNumber="431">4. I</date>
never observed any other secondary cords in 
<taxonomicName id="52EB38CEFF80FF978258FAB65C89D9B3" box="[799,910,1400,1423]" class="Gastropoda" family="Muricidae" genus="Typhinellus" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="431" phylum="Mollusca" rank="species" species="labiatus">
<emphasis id="A79F9F5FFF80FF978258FAB65C89D9B3" box="[799,910,1400,1423]" italics="true" pageId="4" pageNumber="431">T. labiatus</emphasis>
</taxonomicName>
contrary to 
<taxonomicName id="52EB38CEFF80FF978549FAB65B96D9B3" box="[1038,1169,1400,1423]" class="Gastropoda" family="Muricidae" genus="Typhinellus" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="431" phylum="Mollusca" rank="species" species="jacolombi" status="sp. nov.">
<emphasis id="A79F9F5FFF80FF978549FAB65B96D9B3" box="[1038,1169,1400,1423]" italics="true" pageId="4" pageNumber="431">T. jacolombi</emphasis>
</taxonomicName>
<emphasis id="A79F9F5FFF80FF9785DEFAB65BD3D9B3" bold="true" box="[1177,1236,1400,1423]" pageId="4" pageNumber="431">
<taxonomicNameLabel id="BCAC2224FF80FF9785DEFAB65BD3D9B3" box="[1177,1236,1400,1423]" pageId="4" pageNumber="431" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, which always has a s2 spiral cord, often s3, less often s4 and rarely s5 and abs (
<figureCitation id="0DD05FC8FF80FF978260FA545C61D98D" box="[807,870,1434,1457]" captionStart="FIGURE 1" captionStartId="1.[151,244,1898,1920]" captionTargetBox="[226,1359,453,1882]" captionTargetId="figure@1.[226,1360,447,1883]" captionTargetPageId="1" captionText="FIGURE 1. Spiral cord morphology. a – c. Typhinellus jacolombi n. sp. a. Panama, Portobelo Bay, 37 – 74 m, holotype MNHN IM- 2000 - 30465, length: 19.2 mm; b. Panama, Portobelo Bay, 60 m, muddy substrate, coll. 1989, paratype RH, length: 28.7 mm; c. Panama, Portobelo, 19 – 37 m, paratype JC, length: 19.3 mm; d. Typhinellus labiatus (Cristofori &amp; Jan, 1832), Kerkennah, Tunisia, 0.5 m, RH, length: 20.1 mm; e. Typhinellus lamyi Garrigues &amp; Merle, 2014, Guadeloupe, Port Louis, 66 m, holotype MNHN-IM- 2013 - 7776, length: 13.1 mm." httpUri="https://zenodo.org/record/237729/files/figure.png" pageId="4" pageNumber="431">Fig. 1</figureCitation>
a–c). 
<taxonomicName id="52EB38CEFF80FF9782EDFA545B29D98D" box="[938,1070,1434,1457]" class="Gastropoda" family="Muricidae" genus="Typhinellus" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="431" phylum="Mollusca" rank="species" species="jacolombi">
<emphasis id="A79F9F5FFF80FF9782EDFA545B29D98D" box="[938,1070,1434,1457]" italics="true" pageId="4" pageNumber="431">T. jacolombi</emphasis>
</taxonomicName>
occasionally has a smoother shell than observed in the 
<typeStatus id="4A50FDEFFF80FF97803CFA735EDFD9E8" box="[379,472,1469,1492]" pageId="4" pageNumber="431" type="holotype">holotype</typeStatus>
(
<figureCitation id="0DD05FC8FF80FF9780AFFA735D52D9E8" box="[488,597,1469,1492]" captionStart="FIGURE 2. a – k" captionStartId="2.[151,244,1751,1773]" captionTargetBox="[191,1396,191,1717]" captionTargetId="figure@2.[191,1396,191,1717]" captionTargetPageId="2" captionText="FIGURE 2. a – k. Typhinellus jacolombi n. sp. a – c. Panama, Portobelo Bay, 37 – 74 m, holotype MNHN IM- 2000 - 30465, length: 19.2 mm; d – f. Panama, Portobelo Bay, 60 m, muddy substrate, coll. 1989, paratype RH, length: 28.7 mm; g – h. Panama, Portobelo, 19 – 37 m, paratype JC, length: 19.3 mm; i – j. Panama, Caribbean (no other data), paratype JC, length: 21.7 mm; k. Protoconch of specimen figured in i – j (scale bar 500 µm). l – n. Typhinellus lamyi Garrigues &amp; Merle, 2014, Guadeloupe, Port Louis, 66 m, holotype MNHN- IM- 2013 - 7776, length: 13.1 mm." httpUri="https://zenodo.org/record/237730/files/figure.png" pageId="4" pageNumber="431">
Fig. 
<quantity id="5213EEA8FF80FF97835CFA735D39D9E8" box="[539,574,1469,1492]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0" pageId="4" pageNumber="431" unit="g" value="2.0">2 g</quantity>
–j
</figureCitation>
) but 
<taxonomicName id="52EB38CEFF80FF9783C9FA735C07D9E8" box="[654,768,1469,1492]" class="Gastropoda" family="Muricidae" genus="Typhinellus" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="431" phylum="Mollusca" rank="species" species="labiatus">
<emphasis id="A79F9F5FFF80FF9783C9FA735C07D9E8" box="[654,768,1469,1492]" italics="true" pageId="4" pageNumber="431">T. labiatus</emphasis>
</taxonomicName>
has a less broad and less expanded apertural wing abapically and a more constricted last teleoconch whorl.
</paragraph>
<paragraph id="9554434DFF80FF978180F9CC5C6CDA63" blockId="4.[151,1436,152,1631]" pageId="4" pageNumber="431">
<taxonomicName id="52EB38CEFF80FF978180F9CC5EB1DA25" box="[199,438,1538,1561]" class="Gastropoda" family="Muricidae" genus="Typhinellus" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="431" phylum="Mollusca" rank="species" species="jacolombi">
<emphasis id="A79F9F5FFF80FF978180F9CC5EB1DA25" box="[199,438,1538,1561]" italics="true" pageId="4" pageNumber="431">Typhinellus jacolombi</emphasis>
</taxonomicName>
varies in shell colour, ranging from completely white with darker brown subsutural area (
<typeStatus id="4A50FDEFFF80FF9781D9F9EB5FFBDA00" box="[158,252,1573,1596]" pageId="4" pageNumber="431" type="paratype">paratype</typeStatus>
RH) to completely light brown (
<typeStatus id="4A50FDEFFF80FF97831FF9EB5DB2DA00" box="[600,693,1573,1596]" pageId="4" pageNumber="431" type="paratype">paratype</typeStatus>
RH) (
<figureCitation id="0DD05FC8FF80FF9783BEF9EB5C60DA00" box="[761,871,1573,1596]" captionStart="FIGURE 2. a – k" captionStartId="2.[151,244,1751,1773]" captionTargetBox="[191,1396,191,1717]" captionTargetId="figure@2.[191,1396,191,1717]" captionTargetPageId="2" captionText="FIGURE 2. a – k. Typhinellus jacolombi n. sp. a – c. Panama, Portobelo Bay, 37 – 74 m, holotype MNHN IM- 2000 - 30465, length: 19.2 mm; d – f. Panama, Portobelo Bay, 60 m, muddy substrate, coll. 1989, paratype RH, length: 28.7 mm; g – h. Panama, Portobelo, 19 – 37 m, paratype JC, length: 19.3 mm; i – j. Panama, Caribbean (no other data), paratype JC, length: 21.7 mm; k. Protoconch of specimen figured in i – j (scale bar 500 µm). l – n. Typhinellus lamyi Garrigues &amp; Merle, 2014, Guadeloupe, Port Louis, 66 m, holotype MNHN- IM- 2013 - 7776, length: 13.1 mm." httpUri="https://zenodo.org/record/237730/files/figure.png" pageId="4" pageNumber="431">Fig. 2 d–f</figureCitation>
). The protoconch (
<typeStatus id="4A50FDEFFF80FF978569F9EB5B8CDA00" box="[1070,1163,1573,1596]" pageId="4" pageNumber="431" type="paratype">paratype</typeStatus>
JC) is small and rounded (
<figureCitation id="0DD05FC8FF80FF9781D8F9865FF4DA63" box="[159,243,1608,1631]" captionStart="FIGURE 2. a – k" captionStartId="2.[151,244,1751,1773]" captionTargetBox="[191,1396,191,1717]" captionTargetId="figure@2.[191,1396,191,1717]" captionTargetPageId="2" captionText="FIGURE 2. a – k. Typhinellus jacolombi n. sp. a – c. Panama, Portobelo Bay, 37 – 74 m, holotype MNHN IM- 2000 - 30465, length: 19.2 mm; d – f. Panama, Portobelo Bay, 60 m, muddy substrate, coll. 1989, paratype RH, length: 28.7 mm; g – h. Panama, Portobelo, 19 – 37 m, paratype JC, length: 19.3 mm; i – j. Panama, Caribbean (no other data), paratype JC, length: 21.7 mm; k. Protoconch of specimen figured in i – j (scale bar 500 µm). l – n. Typhinellus lamyi Garrigues &amp; Merle, 2014, Guadeloupe, Port Louis, 66 m, holotype MNHN- IM- 2013 - 7776, length: 13.1 mm." httpUri="https://zenodo.org/record/237730/files/figure.png" pageId="4" pageNumber="431">Fig. 2 k</figureCitation>
) and the total shell length may attain 
<quantity id="5213EEA8FF80FF9783C3F9865DE3DA63" box="[644,740,1608,1631]" metricMagnitude="-2" metricUnit="m" metricValue="2.87" pageId="4" pageNumber="431" unit="mm" value="28.7">28.7 mm</quantity>
(
<figureCitation id="0DD05FC8FF80FF9783B5F9865C67DA63" box="[754,864,1608,1631]" captionStart="FIGURE 2. a – k" captionStartId="2.[151,244,1751,1773]" captionTargetBox="[191,1396,191,1717]" captionTargetId="figure@2.[191,1396,191,1717]" captionTargetPageId="2" captionText="FIGURE 2. a – k. Typhinellus jacolombi n. sp. a – c. Panama, Portobelo Bay, 37 – 74 m, holotype MNHN IM- 2000 - 30465, length: 19.2 mm; d – f. Panama, Portobelo Bay, 60 m, muddy substrate, coll. 1989, paratype RH, length: 28.7 mm; g – h. Panama, Portobelo, 19 – 37 m, paratype JC, length: 19.3 mm; i – j. Panama, Caribbean (no other data), paratype JC, length: 21.7 mm; k. Protoconch of specimen figured in i – j (scale bar 500 µm). l – n. Typhinellus lamyi Garrigues &amp; Merle, 2014, Guadeloupe, Port Louis, 66 m, holotype MNHN- IM- 2013 - 7776, length: 13.1 mm." httpUri="https://zenodo.org/record/237730/files/figure.png" pageId="4" pageNumber="431">Fig. 2 d–f</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>