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<document id="D3728D1254AE0516915184D8A9573A43" ID-CLB-Dataset="29001" ID-DOI="10.11646/zootaXa.4460.1.1" ID-GBIF-Dataset="84973bd6-9d66-4172-8f50-2a482edccf2a" ID-ISSN="1175-5326" ID-Zenodo-Dep="1459506" ID-ZooBank="DB841017-698F-4D44-A633-461D350DC984" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539326878349" checkinUser="plazi" docAuthor="Choi, Jinyeong, Soysouvanh, Pheophanh, Lee, Seunghwan &amp; Hong, Ki-Jeong" docDate="2018" docId="0974884CB674FFD6FF6CFE870386FCD1" docLanguage="en" docName="zootaxa.4460.1.1.pdf.imf" docOrigin="Zootaxa 4460 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Drepanococcus Williams &amp; WatsOn 1990" docType="treatment" docVersion="7" lastPageNumber="8" masterDocId="F54DF034B673FFD1FFFBFFA5024BFFD6" masterDocTitle="Review of the family Coccidae (Hemiptera: Coccomorpha) in Laos" masterLastPageNumber="62" masterPageNumber="1" pageNumber="8" updateTime="1698579179695" updateUser="ExternalLinkService">
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<mods:title id="44576B386864CD099BBB40FDD5479EF0">Review of the family Coccidae (Hemiptera: Coccomorpha) in Laos</mods:title>
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<mods:namePart id="1FCF1F5FF6134FF66C8D8F9238AF04A2">Choi, Jinyeong</mods:namePart>
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<mods:namePart id="979FB4E782A4D5103EF20F6911B1C365">Soysouvanh, Pheophanh</mods:namePart>
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<mods:namePart id="8F0E994F0791A2156B3579C16CF6D41D">Lee, Seunghwan</mods:namePart>
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<mods:namePart id="803D9634884F6D578423D5AB0933E27B">Hong, Ki-Jeong</mods:namePart>
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<mods:date id="784003AF4F2D80AB927E9C86BD110A4A">2018</mods:date>
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<treatment id="0974884CB674FFD6FF6CFE870386FCD1" ID-DOI="http://doi.org/10.5281/zenodo.5966414" ID-GBIF-Taxon="148695339" ID-Zenodo-Dep="5966414" LSID="urn:lsid:plazi:treatment:0974884CB674FFD6FF6CFE870386FCD1" httpUri="http://treatment.plazi.org/id/0974884CB674FFD6FF6CFE870386FCD1" lastPageNumber="8" pageId="7" pageNumber="8">
<subSubSection id="C9C76AD1B674FFD6FF6CFE8700A4FEEA" box="[151,751,290,316]" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="8162395AB674FFD6FF6CFE8700A4FEEA" blockId="7.[151,751,290,316]" box="[151,751,290,316]" pageId="7" pageNumber="8">
<heading id="DA2A8E36B674FFD6FF6CFE8700A4FEEA" bold="true" box="[151,751,290,316]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
<emphasis id="B3A9E548B674FFD6FF6CFE8700A4FEEA" bold="true" box="[151,751,290,316]" pageId="7" pageNumber="8">
Genus
<taxonomicName id="46DD42D9B674FFD6FF15FE8700A4FEEA" ID-CoL="488K" ID-ENA="1290250" authority="Williams &amp; Watson, 1990" authorityName="Williams &amp; WatsOn" authorityYear="1990" box="[238,751,290,316]" class="Insecta" family="Coccidae" genus="Drepanococcus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B3A9E548B674FFD6FF15FE8703E2FEEA" bold="true" box="[238,425,290,316]" italics="true" pageId="7" pageNumber="8">Drepanococcus</emphasis>
<bibRefCitation id="E54C44ABB674FFD6FE4BFE8700A4FEEA" box="[432,751,290,316]" pageId="7" pageNumber="8" refId="ref29081" refString="Williams, D. J. &amp; Watson, G. W. (1990) The scale insects of the tropical South Pacific Region. Pt. 3: The soft scales (Coccidae) and other families. CAB International, Wallingford, Oxon, 267 pp." type="book">Williams &amp; Watson, 1990</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C9C76AD1B674FFD6FF6CFECC013DFEA8" box="[151,886,360,382]" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="8162395AB674FFD6FF6CFECC013DFEA8" blockId="7.[151,886,360,382]" box="[151,886,360,382]" pageId="7" pageNumber="8">
<emphasis id="B3A9E548B674FFD6FF6CFECC0355FEA8" box="[151,286,361,382]" italics="true" pageId="7" pageNumber="8">Type species:</emphasis>
<taxonomicName id="46DD42D9B674FFD6FEDFFECD0030FEA8" authority="Maskell, 1891 a" authorityName="Maskell" authorityYear="1891" box="[292,635,360,382]" class="Magnoliopsida" family="Amaranthaceae" genus="Eriochiton" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="cajani">
<emphasis id="B3A9E548B674FFD6FEDFFECD039DFEA8" box="[292,470,360,382]" italics="true" pageId="7" pageNumber="8">Eriochiton cajani</emphasis>
<bibRefCitation id="E54C44ABB674FFD6FE27FECD0030FEA8" box="[476,635,360,382]" pageId="7" pageNumber="8" refId="ref26880" refString="Maskell, W. M. (1891 a) Descriptions of new Coccidae. Indian Museum Notes, 2, 59 - 62." type="journal article">Maskell, 1891a</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
<subSubSection id="C9C76AD1B674FFD6FF6CFE0F03A6FDA1" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="8162395AB674FFD6FF6CFE0F03A6FDA1" blockId="7.[151,1436,426,775]" pageId="7" pageNumber="8">
<emphasis id="B3A9E548B674FFD6FF6CFE0F035AFE15" bold="true" box="[151,273,426,451]" pageId="7" pageNumber="8">Diagnosis.</emphasis>
Anal cleft short, about 1/9 of body length. Dorsum with anal plates each with conical inner margin setae and apical setae (
<figureCitation id="19E625DFB674FFD6FEA2FE6B03F8FE31" box="[345,435,462,487]" captionStart="FIGURE 3" captionStartId="8.[151,250,1947,1969]" captionTargetBox="[166,1361,222,1887]" captionTargetId="figure@8.[154,1398,209,1909]" captionTargetPageId="8" captionText="FIGURE 3. Drepanococcus cajani (Maskell, 1891), adult female, from Hodgson (1994). A, stigmatic spine; B1-2, marginal setae; C1-3, dorsal microducts; D, preopercular pore; E, anal plates; F, leg; G, multilocular disc-pore; H1-2, ventral tubular ducts; I, ventral microduct; J, spiracular disc-pore." httpUri="https://zenodo.org/record/1459512/files/figure.png" pageId="7" pageNumber="8">Figs 3E</figureCitation>
,
<figureCitation id="19E625DFB674FFD6FE3BFE6B03AEFE31" box="[448,485,462,487]" captionStart="FIGURE 5" captionStartId="10.[151,250,1605,1627]" captionTargetBox="[314,1279,199,1546]" captionTargetId="figure@10.[302,1285,193,1583]" captionTargetPageId="10" captionText="FIGURE 5. Drepanococcus chiton (Green, 1909), adult female, from Williams &amp; Watson (1990). A, dorsal microduct; B, simple pore; C, preopercular pores; D, anal plates; E. multilocular disc-pore; F. anterior and lateral marginal setae on anogenital fold; G, ventral tubular duct; H, spiracular disc-pore; I, stigmatic spine (broken) and marginal setae; J, ventral microduct." httpUri="https://zenodo.org/record/1459516/files/figure.png" pageId="7" pageNumber="8">5D</figureCitation>
); dorsal setae absent; and tubular ducts absent. Marginal setae conical (
<figureCitation id="19E625DFB674FFD6FAF0FE6B072CFE31" box="[1291,1383,462,487]" captionStart="FIGURE 3" captionStartId="8.[151,250,1947,1969]" captionTargetBox="[166,1361,222,1887]" captionTargetId="figure@8.[154,1398,209,1909]" captionTargetPageId="8" captionText="FIGURE 3. Drepanococcus cajani (Maskell, 1891), adult female, from Hodgson (1994). A, stigmatic spine; B1-2, marginal setae; C1-3, dorsal microducts; D, preopercular pore; E, anal plates; F, leg; G, multilocular disc-pore; H1-2, ventral tubular ducts; I, ventral microduct; J, spiracular disc-pore." httpUri="https://zenodo.org/record/1459512/files/figure.png" pageId="7" pageNumber="8">Figs 3B</figureCitation>
,
<figureCitation id="19E625DFB674FFD6FA8FFE6B07C5FE31" box="[1396,1422,462,487]" captionStart="FIGURE 5" captionStartId="10.[151,250,1605,1627]" captionTargetBox="[314,1279,199,1546]" captionTargetId="figure@10.[302,1285,193,1583]" captionTargetPageId="10" captionText="FIGURE 5. Drepanococcus chiton (Green, 1909), adult female, from Williams &amp; Watson (1990). A, dorsal microduct; B, simple pore; C, preopercular pores; D, anal plates; E. multilocular disc-pore; F. anterior and lateral marginal setae on anogenital fold; G, ventral tubular duct; H, spiracular disc-pore; I, stigmatic spine (broken) and marginal setae; J, ventral microduct." httpUri="https://zenodo.org/record/1459516/files/figure.png" pageId="7" pageNumber="8">
<date id="F5631F9AB674FFD6FA8FFE6B07C5FE31" box="[1396,1422,462,487]" pageId="7" pageNumber="8">5I</date>
</figureCitation>
); stigmatic clefts absent, each with a single stigmatic spine (
<figureCitation id="19E625DFB674FFD6FCC4FE5701D4FDDD" box="[831,927,498,523]" captionStart="FIGURE 2" captionStartId="7.[151,250,1576,1598]" captionTargetBox="[159,1444,1037,1550]" captionTargetId="figure@7.[159,1444,1037,1550]" captionTargetPageId="7" captionText="FIGURE 2. Drepanococcus cajani (Maskell, 1891). A, population of females in life; B, slide-mounted adult female; C, antenna; D, stigmatic spine. Scale lines for B = 0.5 mm; C = 100 µm; D = 50 µm." httpUri="https://zenodo.org/record/1459510/files/figure.png" pageId="7" pageNumber="8">Figs 2D</figureCitation>
,
<figureCitation id="19E625DFB674FFD6FC56FE56019AFDDD" box="[941,977,499,523]" captionStart="FIGURE 3" captionStartId="8.[151,250,1947,1969]" captionTargetBox="[166,1361,222,1887]" captionTargetId="figure@8.[154,1398,209,1909]" captionTargetPageId="8" captionText="FIGURE 3. Drepanococcus cajani (Maskell, 1891), adult female, from Hodgson (1994). A, stigmatic spine; B1-2, marginal setae; C1-3, dorsal microducts; D, preopercular pore; E, anal plates; F, leg; G, multilocular disc-pore; H1-2, ventral tubular ducts; I, ventral microduct; J, spiracular disc-pore." httpUri="https://zenodo.org/record/1459512/files/figure.png" pageId="7" pageNumber="8">3A</figureCitation>
,
<figureCitation id="19E625DFB674FFD6FC1BFE56064EFDDD" box="[992,1029,499,523]" captionStart="FIGURE 4" captionStartId="9.[151,250,1676,1698]" captionTargetBox="[340,1247,1270,1636]" captionTargetId="figure@9.[340,1247,1270,1636]" captionTargetPageId="9" captionText="FIGURE 4. Drepanococcus chiton (Green, 1909). A, adult female in life; B, slide-mounted adult female; C, antenna; D, stigmatic spine. Scale lines for B = 0.5 mm; C = 100 µm; D = 50 µm." httpUri="https://zenodo.org/record/1459514/files/figure.png" pageId="7" pageNumber="8">4D</figureCitation>
,
<figureCitation id="19E625DFB674FFD6FBE8FE570666FDDD" box="[1043,1069,498,523]" captionStart="FIGURE 5" captionStartId="10.[151,250,1605,1627]" captionTargetBox="[314,1279,199,1546]" captionTargetId="figure@10.[302,1285,193,1583]" captionTargetPageId="10" captionText="FIGURE 5. Drepanococcus chiton (Green, 1909), adult female, from Williams &amp; Watson (1990). A, dorsal microduct; B, simple pore; C, preopercular pores; D, anal plates; E. multilocular disc-pore; F. anterior and lateral marginal setae on anogenital fold; G, ventral tubular duct; H, spiracular disc-pore; I, stigmatic spine (broken) and marginal setae; J, ventral microduct." httpUri="https://zenodo.org/record/1459516/files/figure.png" pageId="7" pageNumber="8">
<date id="F5631F9AB674FFD6FBE8FE570666FDDD" box="[1043,1069,498,523]" pageId="7" pageNumber="8">5I</date>
</figureCitation>
). Venter with multilocular discpores each with 10 loculi (
<figureCitation id="19E625DFB674FFD6FE31FDB3006DFDF9" box="[458,550,534,559]" captionStart="FIGURE 3" captionStartId="8.[151,250,1947,1969]" captionTargetBox="[166,1361,222,1887]" captionTargetId="figure@8.[154,1398,209,1909]" captionTargetPageId="8" captionText="FIGURE 3. Drepanococcus cajani (Maskell, 1891), adult female, from Hodgson (1994). A, stigmatic spine; B1-2, marginal setae; C1-3, dorsal microducts; D, preopercular pore; E, anal plates; F, leg; G, multilocular disc-pore; H1-2, ventral tubular ducts; I, ventral microduct; J, spiracular disc-pore." httpUri="https://zenodo.org/record/1459512/files/figure.png" pageId="7" pageNumber="8">Figs 3G</figureCitation>
,
<figureCitation id="19E625DFB674FFD6FDC8FDB3001EFDF9" box="[563,597,534,559]" captionStart="FIGURE 5" captionStartId="10.[151,250,1605,1627]" captionTargetBox="[314,1279,199,1546]" captionTargetId="figure@10.[302,1285,193,1583]" captionTargetPageId="10" captionText="FIGURE 5. Drepanococcus chiton (Green, 1909), adult female, from Williams &amp; Watson (1990). A, dorsal microduct; B, simple pore; C, preopercular pores; D, anal plates; E. multilocular disc-pore; F. anterior and lateral marginal setae on anogenital fold; G, ventral tubular duct; H, spiracular disc-pore; I, stigmatic spine (broken) and marginal setae; J, ventral microduct." httpUri="https://zenodo.org/record/1459516/files/figure.png" pageId="7" pageNumber="8">5E</figureCitation>
); tubular ducts of 1 or 2 types present over entire surface (
<figureCitation id="19E625DFB674FFD6FB07FDB30710FDF9" box="[1276,1371,534,559]" captionStart="FIGURE 3" captionStartId="8.[151,250,1947,1969]" captionTargetBox="[166,1361,222,1887]" captionTargetId="figure@8.[154,1398,209,1909]" captionTargetPageId="8" captionText="FIGURE 3. Drepanococcus cajani (Maskell, 1891), adult female, from Hodgson (1994). A, stigmatic spine; B1-2, marginal setae; C1-3, dorsal microducts; D, preopercular pore; E, anal plates; F, leg; G, multilocular disc-pore; H1-2, ventral tubular ducts; I, ventral microduct; J, spiracular disc-pore." httpUri="https://zenodo.org/record/1459512/files/figure.png" pageId="7" pageNumber="8">Figs 3H</figureCitation>
,
<figureCitation id="19E625DFB674FFD6FA92FDB307C5FDF9" box="[1385,1422,534,559]" captionStart="FIGURE 5" captionStartId="10.[151,250,1605,1627]" captionTargetBox="[314,1279,199,1546]" captionTargetId="figure@10.[302,1285,193,1583]" captionTargetPageId="10" captionText="FIGURE 5. Drepanococcus chiton (Green, 1909), adult female, from Williams &amp; Watson (1990). A, dorsal microduct; B, simple pore; C, preopercular pores; D, anal plates; E. multilocular disc-pore; F. anterior and lateral marginal setae on anogenital fold; G, ventral tubular duct; H, spiracular disc-pore; I, stigmatic spine (broken) and marginal setae; J, ventral microduct." httpUri="https://zenodo.org/record/1459516/files/figure.png" pageId="7" pageNumber="8">5G</figureCitation>
); antenna 7 or 8 segmented (
<figureCitation id="19E625DFB674FFD6FE2CFD9F007DFD85" box="[471,566,570,595]" captionStart="FIGURE 2" captionStartId="7.[151,250,1576,1598]" captionTargetBox="[159,1444,1037,1550]" captionTargetId="figure@7.[159,1444,1037,1550]" captionTargetPageId="7" captionText="FIGURE 2. Drepanococcus cajani (Maskell, 1891). A, population of females in life; B, slide-mounted adult female; C, antenna; D, stigmatic spine. Scale lines for B = 0.5 mm; C = 100 µm; D = 50 µm." httpUri="https://zenodo.org/record/1459510/files/figure.png" pageId="7" pageNumber="8">Figs 2C</figureCitation>
,
<figureCitation id="19E625DFB674FFD6FDBDFD9E0021FD85" box="[582,618,571,595]" captionStart="FIGURE 4" captionStartId="9.[151,250,1676,1698]" captionTargetBox="[340,1247,1270,1636]" captionTargetId="figure@9.[340,1247,1270,1636]" captionTargetPageId="9" captionText="FIGURE 4. Drepanococcus chiton (Green, 1909). A, adult female in life; B, slide-mounted adult female; C, antenna; D, stigmatic spine. Scale lines for B = 0.5 mm; C = 100 µm; D = 50 µm." httpUri="https://zenodo.org/record/1459514/files/figure.png" pageId="7" pageNumber="8">4C</figureCitation>
); legs without tibio-tarsal articulatory scleroses (
<figureCitation id="19E625DFB674FFD6FB58FD9F06B1FD85" box="[1187,1274,570,595]" captionStart="FIGURE 3" captionStartId="8.[151,250,1947,1969]" captionTargetBox="[166,1361,222,1887]" captionTargetId="figure@8.[154,1398,209,1909]" captionTargetPageId="8" captionText="FIGURE 3. Drepanococcus cajani (Maskell, 1891), adult female, from Hodgson (1994). A, stigmatic spine; B1-2, marginal setae; C1-3, dorsal microducts; D, preopercular pore; E, anal plates; F, leg; G, multilocular disc-pore; H1-2, ventral tubular ducts; I, ventral microduct; J, spiracular disc-pore." httpUri="https://zenodo.org/record/1459512/files/figure.png" pageId="7" pageNumber="8">Fig. 3F</figureCitation>
) (
<bibRefCitation id="E54C44ABB674FFD6FAEFFD9F0366FDA1" pageId="7" pageNumber="8" refId="ref29081" refString="Williams, D. J. &amp; Watson, G. W. (1990) The scale insects of the tropical South Pacific Region. Pt. 3: The soft scales (Coccidae) and other families. CAB International, Wallingford, Oxon, 267 pp." type="book">Williams &amp; Watson 1990</bibRefCitation>
;
<bibRefCitation id="E54C44ABB674FFD6FEC3FDFB03ABFDA1" box="[312,480,606,631]" pageId="7" pageNumber="8" refId="ref25777" refString="Hodgson, C. J. (1994) The scale insect family Coccidae: an identification manual to genera. CAB International, Wallingford, Oxon, 639 pp." type="book">Hodgson 1994</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C9C76AD1B674FFD6FF3CFD270386FCD1" pageId="7" pageNumber="8" type="discussion">
<paragraph id="8162395AB674FFD6FF3CFD270386FCD1" blockId="7.[151,1436,426,775]" pageId="7" pageNumber="8">
<emphasis id="B3A9E548B674FFD6FF3CFD270370FD4D" bold="true" box="[199,315,642,667]" pageId="7" pageNumber="8">Remarks.</emphasis>
<taxonomicName id="46DD42D9B674FFD6FEBFFD2603B8FD4C" authorityName="Williams &amp; WatsOn" authorityYear="1990" box="[324,499,643,666]" class="Insecta" family="Coccidae" genus="Drepanococcus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B3A9E548B674FFD6FEBFFD2603B8FD4C" box="[324,499,643,666]" italics="true" pageId="7" pageNumber="8">Drepanococcus</emphasis>
</taxonomicName>
contains 4 species, mostly distributed on South-East Asia (
<bibRefCitation id="E54C44ABB674FFD6FB64FD26029FFD69" pageId="7" pageNumber="8" refId="ref24883" refString="Garcia Morales, M., Denno, B. D., Miller, D. R., Miller, G. L., Ben-Dov, Y. &amp; Hardy, N. B. (2016) ScaleNet: a literature-based model of scale insect biology and systematics. Database. Available from: http: // scalenet. info (accessed 3 July 2018) https: // doi. org / 10.1093 / database / bav 118" type="url">
García Morales
<emphasis id="B3A9E548B674FFD6FAA0FD2607C5FD4D" box="[1371,1422,643,667]" italics="true" pageId="7" pageNumber="8">et al</emphasis>
., 2016
</bibRefCitation>
). Adult females of the genus are completely covered with a test of granular glassy wax in life. In slidemounted material, this genus is easily recognized by the single and long stigmatic spine opposite each spiracle (
<bibRefCitation id="E54C44ABB674FFD6FF65FD4B038BFCD1" box="[158,448,750,775]" pageId="7" pageNumber="8" refId="ref29081" refString="Williams, D. J. &amp; Watson, G. W. (1990) The scale insects of the tropical South Pacific Region. Pt. 3: The soft scales (Coccidae) and other families. CAB International, Wallingford, Oxon, 267 pp." type="book">Williams &amp; Watson 1990</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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