263 lines
32 KiB
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263 lines
32 KiB
XML
<document id="2CED4FC3CC465DB992E4F4A7097AE7D5" ID-DOI="10.11646/zootaxa.5315.4.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="8142274" ID-ZooBank="29F4CB0A-16C0-4BAE-88F1-958762C99938" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1689235878063" checkinUser="plazi" docAuthor="Provenzano-Rizzi, Francisco" docDate="2023" docId="094C87FDF6599001FDB943BC15C2FF28" docLanguage="en" docName="zootaxa.5315.4.2.pdf" docOrigin="Zootaxa 5315 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5315.4.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cheilonimata Provenzano-Rizzi 2023, new genus" docType="treatment" docVersion="3" lastPageNumber="318" masterDocId="F575FF85F65B9002FD2E41321709FFD1" masterDocTitle="A new genus and species of whiptail armored catfish (Siluriformes: Loricariinae) from southern Venezuela" masterLastPageNumber="328" masterPageNumber="315" pageNumber="317" updateTime="1689278290879" updateUser="julia" zenodo-license-document="CLOSED">
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<mods:title id="79839654FD17ABFDD2DB6F54BE8E12F0">A new genus and species of whiptail armored catfish (Siluriformes: Loricariinae) from southern Venezuela</mods:title>
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<mods:namePart id="70380A9102D53CB5A1D41982B053FC3D">Provenzano-Rizzi, Francisco</mods:namePart>
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<mods:date id="6EF1C386D9C8A733916824837071B6BB">2023</mods:date>
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<mods:number id="2C1006A761311DDEA5E87069995BD22D">2023-07-11</mods:number>
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<treatment id="094C87FDF6599001FDB943BC15C2FF28" LSID="urn:lsid:plazi:treatment:094C87FDF6599001FDB943BC15C2FF28" httpUri="http://treatment.plazi.org/id/094C87FDF6599001FDB943BC15C2FF28" lastPageId="3" lastPageNumber="318" pageId="2" pageNumber="317">
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<subSubSection id="C9FF6560F6599000FDB943BC16CCFD78" box="[151,453,654,681]" pageId="2" pageNumber="317" type="nomenclature">
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<paragraph id="815A36EBF6599000FDB943BC16CCFD78" blockId="2.[151,453,654,681]" box="[151,453,654,681]" pageId="2" pageNumber="317">
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<heading id="DA128187F6599000FDB943BC16CCFD78" bold="true" box="[151,453,654,681]" fontSize="11" level="1" pageId="2" pageNumber="317" reason="1">
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<emphasis id="B391EAF9F6599000FDB943BC16CCFD78" bold="true" box="[151,453,654,681]" pageId="2" pageNumber="317">
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<taxonomicName id="46E54D68F6599000FDB943BC1635FD78" authority="Provenzano-Rizzi, 2023" authorityName="Provenzano-Rizzi" authorityYear="2023" box="[151,316,654,681]" family="Loricariidae" genus="Cheilonimata" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="genus" status="new genus">
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<emphasis id="B391EAF9F6599000FDB943BC1635FD78" bold="true" box="[151,316,654,681]" italics="true" pageId="2" pageNumber="317">Cheilonimata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A8A25782F6599000FC6D43BD16CCFD78" box="[323,453,655,681]" pageId="2" pageNumber="317" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C9FF6560F6599000FDB943E214CEFD3B" box="[151,967,720,747]" pageId="2" pageNumber="317" type="type_taxon">
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<paragraph id="815A36EBF6599000FDB943E214CEFD3B" blockId="2.[151,1437,720,2043]" box="[151,967,720,747]" pageId="2" pageNumber="317">
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<emphasis id="B391EAF9F6599000FDB943E2163CFD3B" bold="true" box="[151,309,720,746]" pageId="2" pageNumber="317">Type-species.</emphasis>
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<taxonomicName id="46E54D68F6599000FC1243E21520FD3B" authority="Provenzano-Rizzi, 2023" authorityName="Provenzano-Rizzi" authorityYear="2023" box="[316,553,720,746]" family="Loricariidae" genus="Cheilonimata" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="species" species="minuta" status="new species">
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<emphasis id="B391EAF9F6599000FC1243E21520FD3B" box="[316,553,720,746]" italics="true" pageId="2" pageNumber="317">Cheilonimata minuta</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A8A25782F6599000FF1E43E315BFFD3B" box="[560,694,720,747]" pageId="2" pageNumber="317" rank="species">new species</taxonomicNameLabel>
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by original designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="C9FF6560F6599000FDE943C6144BFC87" pageId="2" pageNumber="317" type="description">
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<paragraph id="815A36EBF6599000FDE943C6144BFC87" blockId="2.[151,1437,720,2043]" pageId="2" pageNumber="317">
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<emphasis id="B391EAF9F6599000FDE943C6163AFCDF" bold="true" box="[199,307,756,782]" pageId="2" pageNumber="317">Includes.</emphasis>
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<taxonomicName id="46E54D68F6599000FC1143C61538FCDF" authority="Provenzano-Rizzi, 2023" authorityName="Provenzano-Rizzi" authorityYear="2023" box="[319,561,756,782]" family="Loricariidae" genus="Cheilonimata" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="species" species="minuta" status="new species">
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<emphasis id="B391EAF9F6599000FC1143C61538FCDF" box="[319,561,756,782]" italics="true" pageId="2" pageNumber="317">Cheilonimata minuta</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A8A25782F6599000FF1243C715C3FCDF" box="[572,714,756,783]" pageId="2" pageNumber="317" rank="species">new species</taxonomicNameLabel>
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.
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<taxonomicName id="46E54D68F6599000FFF443C615BEFCE3" authority="Crispim-Rodrigues, Bernt, Waltz, Silva, Benine, Oliveira, Covain, and Roxo 2023" authorityName="Crispim-Rodrigues, Bernt, Waltz, Silva, Benine, Oliveira, Covain, and Roxo" authorityYear="2023" family="Loricariidae" genus="Rhadinoloricaria" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="species" species="papillosa">
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<emphasis id="B391EAF9F6599000FFF443C61319FCDF" box="[730,1040,756,782]" italics="true" pageId="2" pageNumber="317">Rhadinoloricaria papillosa</emphasis>
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<bibRefCitation id="E5744B1AF6599000F93543C615BEFCE3" author="Crispim-Rodrigues, J. L. & Bernt, M. J. & Waltz, B. T. & Silva, G. S. C. & Benine, R. C. & Oliveira, C. & Covain, R. & Roxo, F. F." pageId="2" pageNumber="317" pagination="1 - 10" refId="ref7651" refString="Crispim-Rodrigues, J. L., Bernt, M. J., Waltz, B. T., Silva, G. S. C., Benine, R. C., Oliveira, C., Covain, R. & Roxo, F. F. (2023) A new species of mailed catfish of genus Rhadinoloricaria (Siluriformes: Loricariidae: Loricariinae) from Rio Negro basin, Brazil. Journal of Fish Biology, 1 - 10. [published online] https: // doi. org / 10.1111 / jfb. 15402" type="book chapter" year="2023">Crispim-Rodrigues, Bernt, Waltz, Silva, Benine, Oliveira, Covain, and Roxo 2023</bibRefCitation>
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</taxonomicName>
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.
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<taxonomicName id="46E54D68F6599000FFEC422A157BFC87" authority="(Crispim-Rodrigues, Bernt, Waltz, Silva, Benine, Oliveira, Covain, and Roxo 2023)" authorityName="Provenzano-Rizzi" authorityYear="2023" baseAuthorityName="Crispim-Rodrigues, Bernt, Waltz, Silva, Benine, Oliveira, Covain, and Roxo" baseAuthorityYear="2023" family="Loricariidae" genus="Cheilonimata" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="species" species="papillosa" status="new combination">
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<emphasis id="B391EAF9F6599000FFEC422A14C3FCE3" box="[706,970,792,818]" italics="true" pageId="2" pageNumber="317">Cheilonimata papillosa</emphasis>
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(
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<bibRefCitation id="E5744B1AF6599000FEF7422A1563FC87" author="Crispim-Rodrigues, J. L. & Bernt, M. J. & Waltz, B. T. & Silva, G. S. C. & Benine, R. C. & Oliveira, C. & Covain, R. & Roxo, F. F." pageId="2" pageNumber="317" pagination="1 - 10" refId="ref7651" refString="Crispim-Rodrigues, J. L., Bernt, M. J., Waltz, B. T., Silva, G. S. C., Benine, R. C., Oliveira, C., Covain, R. & Roxo, F. F. (2023) A new species of mailed catfish of genus Rhadinoloricaria (Siluriformes: Loricariidae: Loricariinae) from Rio Negro basin, Brazil. Journal of Fish Biology, 1 - 10. [published online] https: // doi. org / 10.1111 / jfb. 15402" type="book chapter" year="2023">Crispim-Rodrigues, Bernt, Waltz, Silva, Benine, Oliveira, Covain, and Roxo 2023</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="A8A25782F6599000FF57420F1437FC87" box="[633,830,828,855]" pageId="2" pageNumber="317" rank="species">new combination</taxonomicNameLabel>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C9FF6560F6599000FDE942521305F82B" pageId="2" pageNumber="317" type="diagnosis">
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<paragraph id="815A36EBF6599000FDE9425213F5FB83" blockId="2.[151,1437,720,2043]" pageId="2" pageNumber="317">
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<emphasis id="B391EAF9F6599000FDE94252164BFCAB" bold="true" box="[199,322,864,890]" pageId="2" pageNumber="317">Diagnosis.</emphasis>
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The new genus
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<taxonomicName id="46E54D68F6599000FF0E425215B1FCAB" authorityName="Provenzano-Rizzi" authorityYear="2023" box="[544,696,864,890]" family="Loricariidae" genus="Cheilonimata" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="genus">
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<emphasis id="B391EAF9F6599000FF0E425215B1FCAB" box="[544,696,864,890]" italics="true" pageId="2" pageNumber="317">Cheilonimata</emphasis>
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</taxonomicName>
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is included in the subfamily
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<taxonomicName id="46E54D68F6599000F917425213CCFCAB" box="[1081,1221,864,890]" family="Loricariidae" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="subFamily" subFamily="Loricariinae">Loricariinae</taxonomicName>
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, tribe
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<taxonomicName id="46E54D68F6599000F83342521291FCAB" box="[1309,1432,864,890]" family="Loricariidae" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="tribe" tribe="Loricariini">Loricariini</taxonomicName>
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,
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<taxonomicName id="46E54D68F6599000FDB942B61651FC4F" box="[151,344,900,926]" family="Loricariidae" genus="Pseudohemiodon" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="genus">
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<emphasis id="B391EAF9F6599000FDB942B61651FC4F" box="[151,344,900,926]" italics="true" pageId="2" pageNumber="317">Pseudohemiodon</emphasis>
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</taxonomicName>
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group (
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<bibRefCitation id="E5744B1AF6599000FC9B42B6159BFC4F" author="Covain, R. & Fisch & Muller, S. & Oliveira, C. & Mol, J. H. & Montoya-Burgos, J. I. & Dray, S." box="[437,658,900,927]" pageId="2" pageNumber="317" pagination="492 - 517" refId="ref7568" refString="Covain, R., Fisch - Muller, S., Oliveira, C., Mol, J. H., Montoya-Burgos, J. I. & Dray, S. (2016) Molecular phylogeny of the highly diversified catfish subfamily Loricariinae (Siluriformes, Loricariidae) reveals incongruence with morphological classification. Molecular Phylogenetics and Evolution, 94, 492 - 517. https: // doi. org / 10.1016 / j. ympev. 2015.10.018" type="journal article" year="2016">
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Covain
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<emphasis id="B391EAF9F6599000FF3F42B7154CFC4F" box="[529,581,900,926]" italics="true" pageId="2" pageNumber="317">et al</emphasis>
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. 2016
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</bibRefCitation>
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;
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<bibRefCitation id="E5744B1AF6599000FFB142B7146CFC4F" author="Roxo, F. F. & Ochoa, L. E. & Sabaj, M. H. & Lujan, N. K. & Covain, R. & Silva, G. S. C. & Melo, B. F. & Albert, J. S. & Chang, J. & Foresti, F. & Alfaro, M. E. & Oliveira, C." box="[671,869,900,927]" pageId="2" pageNumber="317" pagination="148 - 165" refId="ref10700" refString="Roxo, F. F., Ochoa, L. E., Sabaj, M. H., Lujan, N. K., Covain, R., Silva, G. S. C., Melo, B. F., Albert, J. S., Chang, J., Foresti, F., Alfaro, M. E. & Oliveira, C. (2019) Phylogenomic reappraisal of the Neotropical catfish family Loricariidae (Teleostei, Siluriformes) using ultraconserved elements. Molecular Phylogenetics and Evolution, 135, 148 - 165. https: // doi. org / 10.1016 / j. ympev. 2019.02.017" type="journal article" year="2019">
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Roxo
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<emphasis id="B391EAF9F6599000FFC842B71413FC4F" box="[742,794,900,926]" italics="true" pageId="2" pageNumber="317">et al</emphasis>
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. 2019
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</bibRefCitation>
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;
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<bibRefCitation id="E5744B1AF6599000FE5A42B613EAFC4F" author="Londono-Burbano, A. & Reis, R. E." box="[884,1251,900,926]" pageId="2" pageNumber="317" pagination="0247747" refId="ref8944" refString="Londono-Burbano, A. & Reis, R. E. (2021) A combined molecular and morphological phylogeny of the Loricariinae (Siluriformes, Loricariidae), with emphasis on the Harttiini and Farlowellini. PLoS ONE, 16 (3), e 0247747. https: // doi. org / 10.1371 / journal. pone. 0247747" type="journal article" year="2021">Londoño-Burbano & Reis 2021</bibRefCitation>
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). Species of the
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<taxonomicName id="46E54D68F6599000FDB9429A1651FC13" box="[151,344,936,962]" family="Loricariidae" genus="Pseudohemiodon" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="genus">
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<emphasis id="B391EAF9F6599000FDB9429A1651FC13" box="[151,344,936,962]" italics="true" pageId="2" pageNumber="317">Pseudohemiodon</emphasis>
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</taxonomicName>
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group share the following external characters: head and body very depressed, especially the caudal peduncle; caudal-fin with ten branched rays; maxillary barbel usually reaches or surpasses gill opening or pectoral-fin origin; 16 or fewer teeth in each mandible, teeth small or very small, in some cases an edentulous upper jaw; lip surfaces with papillae and/or barblets; lip borders with barblets branched or not; oral cavity with barblets, mostly on palate; post-orbital notch small and abdomen surface from totally plated to partially plated or naked.
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</paragraph>
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<paragraph id="815A36EBF6599000FDE9456E1621FA47" blockId="2.[151,1437,720,2043]" pageId="2" pageNumber="317">
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<taxonomicName id="46E54D68F6599000FDE9456E1656FBA7" authority="Provenzano-Rizzi, 2023" authorityName="Provenzano-Rizzi" authorityYear="2023" box="[199,351,1116,1142]" family="Loricariidae" genus="Cheilonimata" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="genus" status="n. gen.">
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<emphasis id="B391EAF9F6599000FDE9456E1656FBA7" box="[199,351,1116,1142]" italics="true" pageId="2" pageNumber="317">Cheilonimata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A8A25782F6599000FC42456F16B4FBA6" box="[364,445,1117,1143]" pageId="2" pageNumber="317" rank="genus">n. gen.</taxonomicNameLabel>
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can be recognized among all genera of the subfamily
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<taxonomicName id="46E54D68F6599000F961456E13D2FBA7" box="[1103,1243,1116,1142]" family="Loricariidae" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="subFamily" subFamily="Loricariinae">Loricariinae</taxonomicName>
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, including those belonging to the
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<taxonomicName id="46E54D68F6599000FC4F45B316C5FB4B" authorityName="J. V. F. Lamouroux in Bory de St. -Vincent" authorityYear="1825" box="[353,460,1153,1178]" family="Loricariidae" genus="Loricaria" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="genus">
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<emphasis id="B391EAF9F6599000FC4F45B316C5FB4B" box="[353,460,1153,1178]" italics="true" pageId="2" pageNumber="317">Loricaria</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="46E54D68F6599000FF2545B215C5FB4B" box="[523,716,1152,1178]" family="Loricariidae" genus="Pseudohemiodon" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="genus">
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<emphasis id="B391EAF9F6599000FF2545B215C5FB4B" box="[523,716,1152,1178]" italics="true" pageId="2" pageNumber="317">Pseudohemiodon</emphasis>
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</taxonomicName>
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groups by the presence of two unique and distinctive buccal characters: 1) Presence of translucent membrane that contain and interconnects the unbranched barblets found on upper lip border and on maxillary barbel, leaving only the tips of the barblets free
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<emphasis id="B391EAF9F6599000F96C45FB1369FB33" box="[1090,1120,1225,1250]" italics="true" pageId="2" pageNumber="317">vs.</emphasis>
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barblets found on upper lip border and on maxillary barbel free, without the interconnecting translucent membrane or in some cases, a very narrow translucent membrane at base of the barblets, never as described for
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<taxonomicName id="46E54D68F6599000F927442213A8FAFB" authorityName="Provenzano-Rizzi" authorityYear="2023" box="[1033,1185,1296,1322]" family="Loricariidae" genus="Cheilonimata" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="genus">
|
|
<emphasis id="B391EAF9F6599000F927442213A8FAFB" box="[1033,1185,1296,1322]" italics="true" pageId="2" pageNumber="317">Cheilonimata</emphasis>
|
|
</taxonomicName>
|
|
. 2) Lower lip surface with cylindrical, lengthened and unbranched barblets, some of them surpassing the posterior lower lip margin
|
|
<emphasis id="B391EAF9F6599000F85044071295FA9F" box="[1406,1436,1333,1358]" italics="true" pageId="2" pageNumber="317">vs.</emphasis>
|
|
lower lip surface with papillae or with papillae and barblets, some species may have lower lip with a thick pad of musculature.
|
|
</paragraph>
|
|
<paragraph id="815A36EBF6599000FDE944921305F82B" blockId="2.[151,1437,720,2043]" pageId="2" pageNumber="317">
|
|
Further distinguished from all species of the
|
|
<taxonomicName id="46E54D68F6599000FFEC4492148AFA6B" box="[706,899,1440,1466]" family="Loricariidae" genus="Pseudohemiodon" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="genus">
|
|
<emphasis id="B391EAF9F6599000FFEC4492148AFA6B" box="[706,899,1440,1466]" italics="true" pageId="2" pageNumber="317">Pseudohemiodon</emphasis>
|
|
</taxonomicName>
|
|
group, except
|
|
<taxonomicName id="46E54D68F6599000F905449213BCFA6B" authorityName="Provenzano-Rizzi" authorityYear="2023" baseAuthorityName="Crispim-Rodrigues, Bernt, Waltz, Silva, Benine, Oliveira, Covain, and Roxo" baseAuthorityYear="2023" box="[1067,1205,1440,1466]" family="Loricariidae" genus="Cheilonimata" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="species" species="papillosa">
|
|
<emphasis id="B391EAF9F6599000F905449213BCFA6B" box="[1067,1205,1440,1466]" italics="true" pageId="2" pageNumber="317">C. papillosa</emphasis>
|
|
</taxonomicName>
|
|
by lower lip margin with unbranched barblets and without protuberances
|
|
<emphasis id="B391EAF9F6599000FFD044F71415FA0F" box="[766,796,1477,1502]" italics="true" pageId="2" pageNumber="317">vs.</emphasis>
|
|
lower lip margin with branched barblets or with small protuberances, in some cases. From all species of the
|
|
<taxonomicName id="46E54D68F6599000FFD044DA14B6F9D3" box="[766,959,1512,1538]" family="Loricariidae" genus="Pseudohemiodon" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="genus">
|
|
<emphasis id="B391EAF9F6599000FFD044DA14B6F9D3" box="[766,959,1512,1538]" italics="true" pageId="2" pageNumber="317">Pseudohemiodon</emphasis>
|
|
</taxonomicName>
|
|
group except species of
|
|
<taxonomicName id="46E54D68F6599000F9F544DA1295F9D3" box="[1243,1436,1512,1538]" family="Loricariidae" genus="Pseudohemiodon" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="genus">
|
|
<emphasis id="B391EAF9F6599000F9F544DA1295F9D3" box="[1243,1436,1512,1538]" italics="true" pageId="2" pageNumber="317">Pseudohemiodon</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="46E54D68F6599000FDE9473E1658F9F7" authorityName="Provenzano-Rizzi" authorityYear="2023" baseAuthorityName="Crispim-Rodrigues, Bernt, Waltz, Silva, Benine, Oliveira, Covain, and Roxo" baseAuthorityYear="2023" box="[199,337,1548,1574]" family="Loricariidae" genus="Cheilonimata" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="species" species="papillosa">
|
|
<emphasis id="B391EAF9F6599000FDE9473E1658F9F7" box="[199,337,1548,1574]" italics="true" pageId="2" pageNumber="317">C. papillosa</emphasis>
|
|
</taxonomicName>
|
|
(except the
|
|
<typeStatus id="5E5E8849F6599000FCF2473E1549F9F7" box="[476,576,1548,1574]" pageId="2" pageNumber="317">holotype</typeStatus>
|
|
) by palate behind premaxillaries at midline, with one elongated, cylindrical, unbranched, fleshy barblet
|
|
<emphasis id="B391EAF9F6599000FCE4470316E1F99B" box="[458,488,1585,1610]" italics="true" pageId="2" pageNumber="317">vs.</emphasis>
|
|
two (one just anterior the other) elongated, cylindrical, unbranched, fleshy barblets, or with one thick barblet, ramified into three or four conical arms, similarly or variable sized, or with six or seven elongated, branched and unbranched barblets. From
|
|
<taxonomicName id="46E54D68F6599000FFCC474A14E9F943" box="[738,992,1656,1682]" family="Loricariidae" genus="Apistoloricaria" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="species" species="condei">
|
|
<emphasis id="B391EAF9F6599000FFCC474A14E9F943" box="[738,992,1656,1682]" italics="true" pageId="2" pageNumber="317">Apistoloricaria condei</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<emphasis id="B391EAF9F6599000FEC3474A13D5F943" box="[1005,1244,1656,1682]" italics="true" pageId="2" pageNumber="317">
|
|
<taxonomicName id="46E54D68F6599000FEC3474A1341F943" authorityName="Nijssen & Isbrucker" authorityYear="1988" box="[1005,1096,1656,1682]" family="Loricariidae" genus="Apistoloricaria" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="species" species="laani">A. laani</taxonomicName>
|
|
,
|
|
<taxonomicName id="46E54D68F6599000F97A474A13D5F943" authorityName="Provenzano-Rizzi" authorityYear="2023" baseAuthorityName="Crispim-Rodrigues, Bernt, Waltz, Silva, Benine, Oliveira, Covain, and Roxo" baseAuthorityYear="2023" box="[1108,1244,1656,1682]" family="Loricariidae" genus="Cheilonimata" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="species" species="papillosa">C. papillosa</taxonomicName>
|
|
</emphasis>
|
|
,
|
|
<taxonomicName id="46E54D68F6599000F9C4474A17FAF967" family="Loricariidae" genus="Crossoloricaria" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="species" species="bahuaja">
|
|
<emphasis id="B391EAF9F6599000F9C4474A17FAF967" italics="true" pageId="2" pageNumber="317">Crossoloricaria bahuaja</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<emphasis id="B391EAF9F6599000FC2E47AE13C5F967" box="[256,1228,1692,1718]" italics="true" pageId="2" pageNumber="317">
|
|
<taxonomicName id="46E54D68F6599000FC2E47AE1512F967" baseAuthorityName="Isbrucker" baseAuthorityYear="1971" box="[256,539,1692,1718]" family="Loricariidae" genus="Planiloricaria" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="species" species="cryptodon">Planiloricaria cryptodon</taxonomicName>
|
|
,
|
|
<taxonomicName id="46E54D68F6599000FF0847AE147AF967" baseAuthorityName="Nijssen & Isbrücker" baseAuthorityYear="1988" box="[550,883,1692,1718]" family="Loricariidae" genus="Rhadinoloricaria" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="species" species="listrorhinos">Rhadinoloricaria listrorhinos</taxonomicName>
|
|
,
|
|
<taxonomicName id="46E54D68F6599000FE5347AF133AF967" baseAuthorityName="Gunther" baseAuthorityYear="1869" box="[893,1075,1693,1718]" family="Loricariidae" genus="Rhadinoloricaria" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="species" species="macromystax">R. macromystax</taxonomicName>
|
|
,
|
|
<taxonomicName id="46E54D68F6599000F91347AF13C5F967" box="[1085,1228,1693,1718]" family="Loricariidae" genus="Rhadinoloricaria" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="species" species="ommation">R. ommation</taxonomicName>
|
|
</emphasis>
|
|
and
|
|
<taxonomicName id="46E54D68F6599000F82F47AF1273F967" authorityName="Provenzano & Barriga Salazar" authorityYear="2020" box="[1281,1402,1693,1718]" family="Loricariidae" genus="Rhadinoloricaria" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="species" species="stewarti">
|
|
<emphasis id="B391EAF9F6599000F82F47AF1273F967" box="[1281,1402,1693,1718]" italics="true" pageId="2" pageNumber="317">R. stewarti</emphasis>
|
|
</taxonomicName>
|
|
by maxillary barbel only reaches pectoral-fin origin
|
|
<emphasis id="B391EAF9F6599000FFED47F315EBF90B" box="[707,738,1729,1754]" italics="true" pageId="2" pageNumber="317">vs.</emphasis>
|
|
maxillary barbel surpasses pectoral-fin base. From
|
|
<taxonomicName id="46E54D68F6599000F80247F31621F92F" authorityName="Martin Salazar, Isbrucker & Nijssen" authorityYear="1982" family="Loricariidae" genus="Dentectus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="species" species="barbarmatus">
|
|
<emphasis id="B391EAF9F6599000F80247F31621F92F" italics="true" pageId="2" pageNumber="317">Dentectus barbarmatus</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="46E54D68F6599000FC4D47D6156FF92F" authorityName="Isbrucker & Nijssen" authorityYear="1984" box="[355,614,1764,1790]" family="Loricariidae" genus="Pyxiloricaria" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="species" species="menezesi">
|
|
<emphasis id="B391EAF9F6599000FC4D47D6156FF92F" box="[355,614,1764,1790]" italics="true" pageId="2" pageNumber="317">Pyxiloricaria menezesi</emphasis>
|
|
</taxonomicName>
|
|
by maxillary barbel reaches pectoral-fin origin
|
|
<emphasis id="B391EAF9F6599000F9A347D713A5F92F" box="[1165,1196,1765,1790]" italics="true" pageId="2" pageNumber="317">vs.</emphasis>
|
|
maxillary barbel not surpassing gill opening. Also distinguishable from
|
|
<taxonomicName id="46E54D68F6599000FFC6463B1496F8F3" authorityName="Martin Salazar, Isbrucker & Nijssen" authorityYear="1982" box="[744,927,1800,1826]" family="Loricariidae" genus="Dentectus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="species" species="barbarmatus">
|
|
<emphasis id="B391EAF9F6599000FFC6463B1496F8F3" box="[744,927,1800,1826]" italics="true" pageId="2" pageNumber="317">D. barbarmatus</emphasis>
|
|
</taxonomicName>
|
|
by maxillary barbel surface without plates, barblets on upper lip margin not covering the mouth opening and upper jaw teeth visible to naked eye
|
|
<emphasis id="B391EAF9F6599000F822461F1223F897" box="[1292,1322,1837,1862]" italics="true" pageId="2" pageNumber="317">vs.</emphasis>
|
|
maxillary barbel surface with plates, barblets on upper lip margin lengthened to cover the mouth opening and upper jaw teeth very small, hard to observe to naked eyes. From
|
|
<taxonomicName id="46E54D68F6599000FF984647144AF85F" baseAuthorityName="Isbrucker" baseAuthorityYear="1971" box="[694,835,1908,1934]" family="Loricariidae" genus="Planiloricaria" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="species" species="cryptodon">
|
|
<emphasis id="B391EAF9F6599000FF984647144AF85F" box="[694,835,1908,1934]" italics="true" pageId="2" pageNumber="317">P. cryptodon</emphasis>
|
|
</taxonomicName>
|
|
by upper jaw teeth present
|
|
<emphasis id="B391EAF9F6599000F9584647139DF85F" box="[1142,1172,1909,1934]" italics="true" pageId="2" pageNumber="317">vs.</emphasis>
|
|
upper jaw teeth absent. From
|
|
<taxonomicName id="46E54D68F6599000FDF046AB166BF863" authorityName="Isbrucker & Nijssen" authorityYear="1984" box="[222,354,1945,1970]" family="Loricariidae" genus="Pyxiloricaria" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="species" species="menezesi">
|
|
<emphasis id="B391EAF9F6599000FDF046AB166BF863" box="[222,354,1945,1970]" italics="true" pageId="2" pageNumber="317">P. menezesi</emphasis>
|
|
</taxonomicName>
|
|
by head contour not triangular and without fleshy flap on branchiostegal membrane
|
|
<emphasis id="B391EAF9F6599000F81146AB1254F863" box="[1343,1373,1945,1970]" italics="true" pageId="2" pageNumber="317">vs.</emphasis>
|
|
head contour triangular and with fleshy flap covering branchiostegal membrane. From
|
|
<taxonomicName id="46E54D68F6599000F91A468F13E0F807" baseAuthorityName="Gunther" baseAuthorityYear="1869" box="[1076,1257,1981,2006]" family="Loricariidae" genus="Rhadinoloricaria" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="species" species="macromystax">
|
|
<emphasis id="B391EAF9F6599000F91A468F13E0F807" box="[1076,1257,1981,2006]" italics="true" pageId="2" pageNumber="317">R. macromystax</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="46E54D68F6599000F80C468F1295F807" authorityName="Provenzano & Barriga Salazar" authorityYear="2020" box="[1314,1436,1981,2006]" family="Loricariidae" genus="Rhadinoloricaria" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="317" phylum="Chordata" rank="species" species="stewarti">
|
|
<emphasis id="B391EAF9F6599000F80C468F1295F807" box="[1314,1436,1981,2006]" italics="true" pageId="2" pageNumber="317">R. stewarti</emphasis>
|
|
</taxonomicName>
|
|
by snout not projected and spatula-shape
|
|
<emphasis id="B391EAF9F6599000FF4946D3158CF82B" box="[615,645,2017,2042]" italics="true" pageId="2" pageNumber="317">vs.</emphasis>
|
|
snout projected and spatula shape.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C9FF6560F6589001FDE941A515C2FF28" pageId="3" pageNumber="318" type="etymology">
|
|
<paragraph id="815A36EBF6589001FDE941A515C2FF28" blockId="3.[151,1437,151,249]" pageId="3" pageNumber="318">
|
|
<emphasis id="B391EAF9F6589001FDE941A51647FF60" bold="true" box="[199,334,151,177]" pageId="3" pageNumber="318">Etymology.</emphasis>
|
|
<taxonomicName id="46E54D68F6589001FC7941A516E6FF60" authorityName="Provenzano-Rizzi" authorityYear="2023" box="[343,495,151,177]" family="Loricariidae" genus="Cheilonimata" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="318" phylum="Chordata" rank="genus">
|
|
<emphasis id="B391EAF9F6589001FC7941A516E6FF60" box="[343,495,151,177]" italics="true" pageId="3" pageNumber="318">Cheilonimata</emphasis>
|
|
</taxonomicName>
|
|
is derived from the Greek, “Χείλος
|
|
<emphasis id="B391EAF9F6589001FEA341A514F5FF60" box="[909,1020,151,177]" italics="true" pageId="3" pageNumber="318">” Cheilos</emphasis>
|
|
meaning lip and “νημάτια
|
|
<emphasis id="B391EAF9F6589001F80041A51294FF60" box="[1326,1437,151,177]" italics="true" pageId="3" pageNumber="318">” nimátia</emphasis>
|
|
or
|
|
<emphasis id="B391EAF9F6589001FD974189160EFF05" box="[185,263,187,212]" italics="true" pageId="3" pageNumber="318">nimata</emphasis>
|
|
meaning threads or filaments, alluding to the cylindrical, elongated, unbranched and fleshy barblets observed over lower lip surface. Gender feminine.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |