treatments-xml/data/00/40/1A/00401A42FFBCFFE50693FDD6FBAC6603.xml

<document id="663E8A67B4783096DEEA90DC524F31B0" ID-DOI="10.11646/zootaxa.3972.1.6" ID-GBIF-Dataset="6b920f46-597d-470d-82d4-72d7d4d8428c" ID-ISSN="1175-5326" ID-Zenodo-Dep="288921" ID-ZooBank="7CA5AC46-C2A2-4361-A1A7-C81A0C5A1C1D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461185141162" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2015" docId="00401A42FFBCFFE50693FDD6FBAC6603" docLanguage="en" docName="zt03972p092.pdf" docOrigin="Zootaxa 3972 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Alpheopsis balaeniceps Anker, 2015, sp. nov." docType="treatment" docVersion="6" lastPageNumber="90" masterDocId="FC79623AFFBDFFE30604FFB5FFAE671E" masterDocTitle="Alpheopsis balaeniceps sp. nov., an unusual alpheid shrimp (Decapoda: Caridea) from French Polynesia" masterLastPageNumber="92" masterPageNumber="85" pageNumber="86" updateTime="1698599445152" updateUser="plazi">
<mods:mods id="63969A66BB99B4D74749EF33A83719A7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="22CCCA612EAA50313D111F93B554A06A">
<mods:title id="A843C5199E0623DF04A212E507BADE74">Alpheopsis balaeniceps sp. nov., an unusual alpheid shrimp (Decapoda: Caridea) from French Polynesia</mods:title>
</mods:titleInfo>
<mods:name id="37955A2B962EB9E3A94C7AC505431921" type="personal">
<mods:role id="C1B1C520F334E901D519275AE2DCAA9D">
<mods:roleTerm id="9DAFDCC2F2D9DE99D6CBDE783154E650">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DF48B4632B5615AE2DAFEDDC4B4E865C">Anker, Arthur</mods:namePart>
</mods:name>
<mods:typeOfResource id="51120AEA920FBEE59A01707D092E6EF9">text</mods:typeOfResource>
<mods:relatedItem id="F3F4BDD62AA0216BFFF97D31211B9CC3" type="host">
<mods:titleInfo id="E8745892CC09C8E262BD3B368271E354">
<mods:title id="D1D2C2B7A6EEE8DFDBC4274FCD3FB9AA">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="16120228DD253A9CBA81AA83BF9EB240">
<mods:date id="D95F00268808BDD7E75481BF141564E0">2015</mods:date>
<mods:detail id="367842BD005332033C075BD3D548807D" type="volume">
<mods:number id="010B962A1773D159C371819E9C9B73AE">3972</mods:number>
</mods:detail>
<mods:detail id="522EB4223794D357B8F952568AA59AA1" type="issue">
<mods:number id="3A5011150CDA80BBB011DC4109812987">1</mods:number>
</mods:detail>
<mods:extent id="376819D76F892F2082E3E05492671ADC" unit="page">
<mods:start id="792F564AC40A4EB286C81D81C1164112">85</mods:start>
<mods:end id="90A117ECEB19D0A80C50D6AFF8F43B78">92</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="E32B9570298091775FDE24AAC102766A">journal article</mods:classification>
<mods:identifier id="1D2B96BD242E1DF75728CB7DC4E98F20" type="DOI">10.11646/zootaxa.3972.1.6</mods:identifier>
<mods:identifier id="EAB41EFE2A901EDFF726F9091AD96BA9" type="GBIF-Dataset">6b920f46-597d-470d-82d4-72d7d4d8428c</mods:identifier>
<mods:identifier id="70B7D4968E7EF256E53488CD06F717EA" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="5C054131B61A7F84257A2727F68200B0" type="Zenodo-Dep">288921</mods:identifier>
<mods:identifier id="DECC6639E9731C551FDCF492D60F7625" type="ZooBank">7CA5AC46-C2A2-4361-A1A7-C81A0C5A1C1D</mods:identifier>
</mods:mods>
<treatment id="00401A42FFBCFFE50693FDD6FBAC6603" ID-DOI="http://doi.org/10.5281/zenodo.6110900" ID-GBIF-Taxon="127692026" ID-Zenodo-Dep="6110900" LSID="urn:lsid:plazi:treatment:00401A42FFBCFFE50693FDD6FBAC6603" httpUri="http://treatment.plazi.org/id/00401A42FFBCFFE50693FDD6FBAC6603" lastPageId="6" lastPageNumber="90" pageId="1" pageNumber="86">
<subSubSection id="C0F3F8DFFFBCFFE20693FDD6FEBF6581" pageId="1" pageNumber="86" type="nomenclature">
<paragraph id="8856AB54FFBCFFE20693FDD6FDBA6560" blockId="1.[151,532,611,671]" box="[151,532,611,638]" pageId="1" pageNumber="86">
<heading id="D31E1C38FFBCFFE20693FDD6FDBA6560" bold="true" box="[151,532,611,638]" fontSize="11" level="1" pageId="1" pageNumber="86" reason="1">
<emphasis id="BA9D7746FFBCFFE20693FDD6FDBA6560" bold="true" box="[151,532,611,638]" pageId="1" pageNumber="86">
<taxonomicName id="4FE9D0D7FFBCFFE20693FDD6FE016563" ID-CoL="85LCJ" box="[151,431,611,637]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="86" phylum="Arthropoda" rank="species" species="balaeniceps" status="sp. nov.">
<emphasis id="BA9D7746FFBCFFE20693FDD6FE016563" bold="true" box="[151,431,611,637]" italics="true" pageId="1" pageNumber="86">Alpheopsis balaeniceps</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1AECA3DFFBCFFE207B3FDD1FDBA6560" box="[439,532,612,638]" pageId="1" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8856AB54FFBCFFE20693FD33FEBF6581" blockId="1.[151,532,611,671]" box="[151,273,646,671]" pageId="1" pageNumber="86">
(
<figureCitation id="10D2B7D1FFBCFFE2069BFD33FEA76581" box="[159,265,646,671]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1859,1881]" captionStartId-1="3.[151,250,1176,1198]" captionStartId-2="4.[151,250,1979,2001]" captionTargetBox-0="[207,1369,198,1812]" captionTargetBox-1="[164,1429,201,1146]" captionTargetBox-2="[151,1436,411,1955]" captionTargetId-0="figure@2.[198,1380,195,1829]" captionTargetId-1="figure@3.[151,1436,193,1155]" captionTargetId-2="figure@4.[151,1436,411,1955]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Alpheopsis balaeniceps sp. nov., holotype, male (cl 3.3 mm) from Moorea, French Polynesia (FLMNH UF 23302); A—frontal region, dorsal; B—same, lateral; C—detail of pterygostomial angle, lateral; D—posterior abdominal somites, lateral; E—telson, dorsal; F—epistomial sclerites; G—tooth on ventromesial carina of first article of antennular peduncle, lateral; H—third maxilliped, lateral; I—second pereiopod, lateral; J—third pereiopod, lateral; K—same, detail of distal propodus and dactylus, mesial; L—fifth pereiopod, lateral; M—pleopod, lateral; N—uropod, dorsal. Scale bars as indicated except for C, F, G (drawn without scale)." captionText-1="FIGURE 2. Alpheopsis balaeniceps sp. nov., holotype, male (cl 3.3 mm) from Moorea, French Polynesia (FLMNH UF 23302); A—right (major) cheliped, lateral; B—same, chela with fingers closed, lateral; C—same, chela fingers open, lateral; D—left (minor) cheliped, lateral; E—same, chela, mesial. Scale bars as indicated." captionText-2="FIGURE 3. Alpheopsis balaeniceps sp. nov., holotype, male (cl 3.3 mm) from Moorea, French Polynesia (FLMNH UF 23302), in life; A—dorsal view; B—lateral view. Photographs by the author." httpUri-0="https://zenodo.org/record/288922/files/figure.png" httpUri-1="https://zenodo.org/record/288923/files/figure.png" httpUri-2="https://zenodo.org/record/288924/files/figure.png" pageId="1" pageNumber="86">Figs. 1–3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C0F3F8DFFFBCFFE20693FD7BFDD66431" pageId="1" pageNumber="86" type="materials_examined">
<paragraph id="8856AB54FFBCFFE20693FD7BFDD66431" blockId="1.[151,1437,718,2003]" pageId="1" pageNumber="86">
<emphasis id="BA9D7746FFBCFFE20693FD7BFEEF65F9" bold="true" box="[151,321,718,743]" pageId="1" pageNumber="86">
<typeStatus id="575215F6FFBCFFE20693FD7BFF7C65F9" box="[151,210,718,743]" pageId="1" pageNumber="86">Type</typeStatus>
material
</emphasis>
. 
<typeStatus id="575215F6FFBCFFE20756FD7BFE1165F9" box="[338,447,718,743]" pageId="1" pageNumber="86" type="holotype">Holotype</typeStatus>
: 
<specimenCount id="9EEF60DDFFBCFFE207C8FD7AFDB465F9" box="[460,538,718,743]" pageId="1" pageNumber="86" type="male">1 male</specimenCount>
(cl 
<quantity id="4F1106B1FFBCFFE20448FD7AFD0765F9" box="[588,681,719,743]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="1" pageNumber="86" unit="mm" value="3.3">3.3 mm</quantity>
), 
<collectionCode id="EEF83391FFBCFFE204BBFD7AFC8F65F8" box="[703,801,719,742]" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="1" pageNumber="86">FLMNH</collectionCode>
UF 23302, 
<collectingCountry id="F0FEEBC4FFBCFFE205B4FD7BFBD765F9" box="[944,1145,718,743]" name="French Polynesia" pageId="1" pageNumber="86">French Polynesia</collectingCountry>
, Society Islands, Moorea Island, eastern coast of Cook's Bay, 
<geoCoordinate id="EDDDCD93FFBCFFE20432FD47FD1B6415" box="[566,693,754,779]" direction="south" orientation="latitude" pageId="1" pageNumber="86" precision="15" value="-17.487778">17°29'16&quot;S</geoCoordinate>
149°49'7 W, depth: ~ 
<quantity id="4F1106B1FFBCFFE205B7FD46FC716414" box="[947,991,755,779]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="86" unit="m" value="1.0">1 m</quantity>
, under corals, on fine sand and rubble, coll. A. Anker, 
<date id="FC578D94FFBCFFE20740FCA3FE1A6431" box="[324,436,790,815]" pageId="1" pageNumber="86" value="2009-10-26">26.x.2009</date>
[BMOO-07149].
</paragraph>
</subSubSection>
<subSubSection id="C0F3F8DFFFBCFFE606C3FC8FFBFB665E" lastPageId="5" lastPageNumber="90" pageId="1" pageNumber="86" type="description">
<paragraph id="8856AB54FFBCFFE206C3FC8FFDCD64A1" blockId="1.[151,1437,718,2003]" pageId="1" pageNumber="86">
<emphasis id="BA9D7746FFBCFFE206C3FC8FFEFF644D" bold="true" box="[199,337,826,851]" pageId="1" pageNumber="86">Description</emphasis>
. Small-sized species of 
<taxonomicName id="4FE9D0D7FFBCFFE20475FC8EFD42644D" box="[625,748,827,851]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="86" phylum="Arthropoda" rank="genus">
<emphasis id="BA9D7746FFBCFFE20475FC8EFD42644D" box="[625,748,827,851]" italics="true" pageId="1" pageNumber="86">Alpheopsis</emphasis>
</taxonomicName>
(cl of 
<typeStatus id="575215F6FFBCFFE20546FC8FFC0A644D" box="[834,932,826,851]" pageId="1" pageNumber="86" type="holotype">holotype</typeStatus>
&lt;
<quantity id="4F1106B1FFBCFFE205CEFC8FFB87644D" box="[970,1065,826,851]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="86" unit="mm" value="3.5">3.5 mm</quantity>
). Carapace with well-developed rostrum, much longer than wide, acute distally, tip almost reaching second half of second article of antennular peduncle; orbital hoods broadly rounded anteriorly, without orbital teeth; pterygostomial angle with minute, anteriorly protruding tooth (
<figureCitation id="10D2B7D1FFBCFFE207D5FC13FDB464A1" box="[465,538,934,959]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[207,1369,198,1812]" captionTargetId="figure@2.[198,1380,195,1829]" captionTargetPageId="2" captionText="FIGURE 1. Alpheopsis balaeniceps sp. nov., holotype, male (cl 3.3 mm) from Moorea, French Polynesia (FLMNH UF 23302); A—frontal region, dorsal; B—same, lateral; C—detail of pterygostomial angle, lateral; D—posterior abdominal somites, lateral; E—telson, dorsal; F—epistomial sclerites; G—tooth on ventromesial carina of first article of antennular peduncle, lateral; H—third maxilliped, lateral; I—second pereiopod, lateral; J—third pereiopod, lateral; K—same, detail of distal propodus and dactylus, mesial; L—fifth pereiopod, lateral; M—pleopod, lateral; N—uropod, dorsal. Scale bars as indicated except for C, F, G (drawn without scale)." httpUri="https://zenodo.org/record/288922/files/figure.png" pageId="1" pageNumber="86">Figs. 1</figureCitation>
A–C).
</paragraph>
<paragraph id="8856AB54FFBCFFE206C3FC7FFC1D6389" blockId="1.[151,1437,718,2003]" pageId="1" pageNumber="86">
First four abdominal somites with pleura rounded posteroventrally; fifth pleuron with acutely projecting posteroventral angle; sixth somite with subacute posterior lobe and well-delimited, triangular, articulated plate (
<figureCitation id="10D2B7D1FFBCFFE2069BFBA7FF4F6335" box="[159,225,1042,1067]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[207,1369,198,1812]" captionTargetId="figure@2.[198,1380,195,1829]" captionTargetPageId="2" captionText="FIGURE 1. Alpheopsis balaeniceps sp. nov., holotype, male (cl 3.3 mm) from Moorea, French Polynesia (FLMNH UF 23302); A—frontal region, dorsal; B—same, lateral; C—detail of pterygostomial angle, lateral; D—posterior abdominal somites, lateral; E—telson, dorsal; F—epistomial sclerites; G—tooth on ventromesial carina of first article of antennular peduncle, lateral; H—third maxilliped, lateral; I—second pereiopod, lateral; J—third pereiopod, lateral; K—same, detail of distal propodus and dactylus, mesial; L—fifth pereiopod, lateral; M—pleopod, lateral; N—uropod, dorsal. Scale bars as indicated except for C, F, G (drawn without scale)." httpUri="https://zenodo.org/record/288922/files/figure.png" pageId="1" pageNumber="86">Fig. 1</figureCitation>
D). Telson narrow, distally tapering, about 2.5 as long as anterior width; dorsal surface with two pairs of slender spiniform setae inserted at some distance from lateral margin, at about 0.45 and 0.75 telson length, respectively; posterior margin less than half as long as anterior margin, rounded; each posterolateral angle with one pair of stout spiniform setae, mesial much longer than lateral (
<figureCitation id="10D2B7D1FFBCFFE2054BFBCBFC216389" box="[847,911,1150,1175]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[207,1369,198,1812]" captionTargetId="figure@2.[198,1380,195,1829]" captionTargetPageId="2" captionText="FIGURE 1. Alpheopsis balaeniceps sp. nov., holotype, male (cl 3.3 mm) from Moorea, French Polynesia (FLMNH UF 23302); A—frontal region, dorsal; B—same, lateral; C—detail of pterygostomial angle, lateral; D—posterior abdominal somites, lateral; E—telson, dorsal; F—epistomial sclerites; G—tooth on ventromesial carina of first article of antennular peduncle, lateral; H—third maxilliped, lateral; I—second pereiopod, lateral; J—third pereiopod, lateral; K—same, detail of distal propodus and dactylus, mesial; L—fifth pereiopod, lateral; M—pleopod, lateral; N—uropod, dorsal. Scale bars as indicated except for C, F, G (drawn without scale)." httpUri="https://zenodo.org/record/288922/files/figure.png" pageId="1" pageNumber="86">Fig. 1</figureCitation>
E).
</paragraph>
<paragraph id="8856AB54FFBCFFE206C3FB16FE5E621D" blockId="1.[151,1437,718,2003]" pageId="1" pageNumber="86">
Eyes completely concealed by orbital hoods, not visible in dorsal view, anterior-most portion exposed in lateral view; eyestalk with well-pigmented cornea and small tubercle on anterior margin (
<figureCitation id="10D2B7D1FFBCFFE20236FB73FBDE63C1" box="[1074,1136,1222,1247]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[207,1369,198,1812]" captionTargetId="figure@2.[198,1380,195,1829]" captionTargetPageId="2" captionText="FIGURE 1. Alpheopsis balaeniceps sp. nov., holotype, male (cl 3.3 mm) from Moorea, French Polynesia (FLMNH UF 23302); A—frontal region, dorsal; B—same, lateral; C—detail of pterygostomial angle, lateral; D—posterior abdominal somites, lateral; E—telson, dorsal; F—epistomial sclerites; G—tooth on ventromesial carina of first article of antennular peduncle, lateral; H—third maxilliped, lateral; I—second pereiopod, lateral; J—third pereiopod, lateral; K—same, detail of distal propodus and dactylus, mesial; L—fifth pereiopod, lateral; M—pleopod, lateral; N—uropod, dorsal. Scale bars as indicated except for C, F, G (drawn without scale)." httpUri="https://zenodo.org/record/288922/files/figure.png" pageId="1" pageNumber="86">Fig. 1</figureCitation>
A, B). Epistomial sclerites with sharp processes (
<figureCitation id="10D2B7D1FFBCFFE2078AFB5FFE63621D" box="[398,461,1258,1283]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[207,1369,198,1812]" captionTargetId="figure@2.[198,1380,195,1829]" captionTargetPageId="2" captionText="FIGURE 1. Alpheopsis balaeniceps sp. nov., holotype, male (cl 3.3 mm) from Moorea, French Polynesia (FLMNH UF 23302); A—frontal region, dorsal; B—same, lateral; C—detail of pterygostomial angle, lateral; D—posterior abdominal somites, lateral; E—telson, dorsal; F—epistomial sclerites; G—tooth on ventromesial carina of first article of antennular peduncle, lateral; H—third maxilliped, lateral; I—second pereiopod, lateral; J—third pereiopod, lateral; K—same, detail of distal propodus and dactylus, mesial; L—fifth pereiopod, lateral; M—pleopod, lateral; N—uropod, dorsal. Scale bars as indicated except for C, F, G (drawn without scale)." httpUri="https://zenodo.org/record/288922/files/figure.png" pageId="1" pageNumber="86">Fig. 1</figureCitation>
F).
</paragraph>
<paragraph id="8856AB54FFBCFFE206C3FABBFDA962E1" blockId="1.[151,1437,718,2003]" pageId="1" pageNumber="86">
Antennular peduncles stout; stylocerite slender, with acute tip, latter almost reaching distal margin of second article; ventromesial carina with stout, anteriorly directed, acute tooth; second article stout, slightly wider than long; lateral flagellum biramous; fused portion short, with three joints; accessory ramus long, with three groups of long aesthetascs (
<figureCitation id="10D2B7D1FFBCFFE20759FACFFE32628D" box="[349,412,1402,1427]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[207,1369,198,1812]" captionTargetId="figure@2.[198,1380,195,1829]" captionTargetPageId="2" captionText="FIGURE 1. Alpheopsis balaeniceps sp. nov., holotype, male (cl 3.3 mm) from Moorea, French Polynesia (FLMNH UF 23302); A—frontal region, dorsal; B—same, lateral; C—detail of pterygostomial angle, lateral; D—posterior abdominal somites, lateral; E—telson, dorsal; F—epistomial sclerites; G—tooth on ventromesial carina of first article of antennular peduncle, lateral; H—third maxilliped, lateral; I—second pereiopod, lateral; J—third pereiopod, lateral; K—same, detail of distal propodus and dactylus, mesial; L—fifth pereiopod, lateral; M—pleopod, lateral; N—uropod, dorsal. Scale bars as indicated except for C, F, G (drawn without scale)." httpUri="https://zenodo.org/record/288922/files/figure.png" pageId="1" pageNumber="86">Fig. 1</figureCitation>
A, B, G). Antenna with basicerite ending in sharp tooth distoventrally; scaphocerite ovate, with anteriorly rounded blade and sharp distolateral tooth, latter not reaching beyond anterior margin of blade; anterior margin of scaphocerite not exceeding distal margin of antennular peduncles; carpocerite reaching almost to end of scaphocerite (
<figureCitation id="10D2B7D1FFBCFFE20784FA53FE1062E1" box="[384,446,1510,1535]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[207,1369,198,1812]" captionTargetId="figure@2.[198,1380,195,1829]" captionTargetPageId="2" captionText="FIGURE 1. Alpheopsis balaeniceps sp. nov., holotype, male (cl 3.3 mm) from Moorea, French Polynesia (FLMNH UF 23302); A—frontal region, dorsal; B—same, lateral; C—detail of pterygostomial angle, lateral; D—posterior abdominal somites, lateral; E—telson, dorsal; F—epistomial sclerites; G—tooth on ventromesial carina of first article of antennular peduncle, lateral; H—third maxilliped, lateral; I—second pereiopod, lateral; J—third pereiopod, lateral; K—same, detail of distal propodus and dactylus, mesial; L—fifth pereiopod, lateral; M—pleopod, lateral; N—uropod, dorsal. Scale bars as indicated except for C, F, G (drawn without scale)." httpUri="https://zenodo.org/record/288922/files/figure.png" pageId="1" pageNumber="86">Fig. 1</figureCitation>
A, B).
</paragraph>
<paragraph id="8856AB54FFBCFFE206C3F9BFFD176191" blockId="1.[151,1437,718,2003]" pageId="1" pageNumber="86">
Mouthparts not dissected, appearing typical for family in external view. Third maxilliped slender, pediform; coxa with distally subacute lateral plate above mastigobranch; penultimate article at most three times as long as wide proximally, widening distally; ultimate article tapering to slender corneous tip, with stiff subdistal seta dorsally; arthrobranch well-developed (
<figureCitation id="10D2B7D1FFBCFFE20456F9C3FD3F6191" box="[594,657,1654,1679]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[207,1369,198,1812]" captionTargetId="figure@2.[198,1380,195,1829]" captionTargetPageId="2" captionText="FIGURE 1. Alpheopsis balaeniceps sp. nov., holotype, male (cl 3.3 mm) from Moorea, French Polynesia (FLMNH UF 23302); A—frontal region, dorsal; B—same, lateral; C—detail of pterygostomial angle, lateral; D—posterior abdominal somites, lateral; E—telson, dorsal; F—epistomial sclerites; G—tooth on ventromesial carina of first article of antennular peduncle, lateral; H—third maxilliped, lateral; I—second pereiopod, lateral; J—third pereiopod, lateral; K—same, detail of distal propodus and dactylus, mesial; L—fifth pereiopod, lateral; M—pleopod, lateral; N—uropod, dorsal. Scale bars as indicated except for C, F, G (drawn without scale)." httpUri="https://zenodo.org/record/288922/files/figure.png" pageId="1" pageNumber="86">Fig. 1</figureCitation>
H).
</paragraph>
<paragraph id="8856AB54FFBCFFE006C3F92FFD0262C1" blockId="1.[151,1437,718,2003]" lastBlockId="3.[151,1437,1299,1972]" lastPageId="3" lastPageNumber="88" pageId="1" pageNumber="86">
Chelipeds unequal in size, subsymmetrical in shape, carried extended when not in use (
<figureCitation id="10D2B7D1FFBCFFE202CFF92FFAB061AD" box="[1227,1310,1690,1715]" captionStart="FIGURE 2" captionStartId="3.[151,250,1176,1198]" captionTargetBox="[164,1429,201,1146]" captionTargetId="figure@3.[151,1436,193,1155]" captionTargetPageId="3" captionText="FIGURE 2. Alpheopsis balaeniceps sp. nov., holotype, male (cl 3.3 mm) from Moorea, French Polynesia (FLMNH UF 23302); A—right (major) cheliped, lateral; B—same, chela with fingers closed, lateral; C—same, chela fingers open, lateral; D—left (minor) cheliped, lateral; E—same, chela, mesial. Scale bars as indicated." httpUri="https://zenodo.org/record/288923/files/figure.png" pageId="1" pageNumber="86">Figs. 2</figureCitation>
, 
<figureCitation id="10D2B7D1FFBCFFE2032AF92EFA9061AD" box="[1326,1342,1691,1715]" captionStart="FIGURE 3" captionStartId="4.[151,250,1979,2001]" captionTargetBox="[151,1436,411,1955]" captionTargetId="figure@4.[151,1436,411,1955]" captionTargetPageId="4" captionText="FIGURE 3. Alpheopsis balaeniceps sp. nov., holotype, male (cl 3.3 mm) from Moorea, French Polynesia (FLMNH UF 23302), in life; A—dorsal view; B—lateral view. Photographs by the author." httpUri="https://zenodo.org/record/288924/files/figure.png" pageId="1" pageNumber="86">3</figureCitation>
). Major cheliped moderately robust; ischium with distodorsal spiniform seta; merus slightly more than four times as long as wide, depressed distoventrally, broadening distally, distomesial angle blunt; carpus short, cup-shaped, with dorsal constriction; chela moderately slender, with fingers about 0.9 times as long as palm; palm about 2.5 times as long as high, subcylindrical in cross-section, smooth; fingers balaeniceps-like, not gaping, with distally crossing tips; pollex with row of plumose setae on lateral and mesial surfaces, posterior half of cutting edge with seven irregularly spaced, subtriangular teeth, becoming gradually larger distally; dactylus slightly flattened dorsoventrally, slightly expanded latero-mesially, with row of plumose setae on lateral and mesial surfaces, subdistal portion of cutting edge with few small irregular teeth (
<figureCitation id="10D2B7D1FFBCFFE204FCF80FFC9560CD" box="[760,827,1978,2003]" captionStart="FIGURE 2" captionStartId="3.[151,250,1176,1198]" captionTargetBox="[164,1429,201,1146]" captionTargetId="figure@3.[151,1436,193,1155]" captionTargetPageId="3" captionText="FIGURE 2. Alpheopsis balaeniceps sp. nov., holotype, male (cl 3.3 mm) from Moorea, French Polynesia (FLMNH UF 23302); A—right (major) cheliped, lateral; B—same, chela with fingers closed, lateral; C—same, chela fingers open, lateral; D—left (minor) cheliped, lateral; E—same, chela, mesial. Scale bars as indicated." httpUri="https://zenodo.org/record/288923/files/figure.png" pageId="1" pageNumber="86">Fig. 2</figureCitation>
A–C). Minor cheliped noticeably smaller than major cheliped, generally similar to major cheliped in proportions and armature on fingers, somewhat more slender; chela with fingers subequal to palm in length, balaeniceps-like, not gaping; dactylus and pollex with rows of plumose setae on lateral and mesial surfaces, rows being somewhat shorter compared to those of major cheliped fingers; cutting edge of dactylus with one subtriangular tooth at about 0.7 of dactylar length and one minute subdistal tooth; cutting edge of pollex with at least six small widely spaced subtriangular teeth, becoming gradually larger distally, and one very small subdistal tooth (
<figureCitation id="10D2B7D1FFBEFFE00422FA73FDC762C1" box="[550,617,1478,1503]" captionStart="FIGURE 2" captionStartId="3.[151,250,1176,1198]" captionTargetBox="[164,1429,201,1146]" captionTargetId="figure@3.[151,1436,193,1155]" captionTargetPageId="3" captionText="FIGURE 2. Alpheopsis balaeniceps sp. nov., holotype, male (cl 3.3 mm) from Moorea, French Polynesia (FLMNH UF 23302); A—right (major) cheliped, lateral; B—same, chela with fingers closed, lateral; C—same, chela fingers open, lateral; D—left (minor) cheliped, lateral; E—same, chela, mesial. Scale bars as indicated." httpUri="https://zenodo.org/record/288923/files/figure.png" pageId="3" pageNumber="88">Fig. 2</figureCitation>
D, E).
</paragraph>
<caption id="DC96FBDCFFBFFFE10693F8F6FD3760EC" httpUri="https://zenodo.org/record/288922/files/figure.png" pageId="2" pageNumber="87" targetBox="[207,1369,198,1812]" targetPageId="2">
<paragraph id="8856AB54FFBFFFE10693F8F6FD3760EC" blockId="2.[151,1436,1859,2034]" pageId="2" pageNumber="87">
<emphasis id="BA9D7746FFBFFFE10693F8F6FEBF6046" bold="true" box="[151,273,1859,1881]" pageId="2" pageNumber="87">FIGURE 1</emphasis>
. 
<taxonomicName id="4FE9D0D7FFBFFFE10726F8F6FDBB6047" box="[290,533,1859,1881]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="87" phylum="Arthropoda" rank="species" species="balaeniceps" status="sp. nov.">
<emphasis id="BA9D7746FFBFFFE10726F8F6FDBB6047" box="[290,533,1859,1881]" italics="true" pageId="2" pageNumber="87">Alpheopsis balaeniceps</emphasis>
</taxonomicName>
<emphasis id="BA9D7746FFBFFFE1041BF8F6FDDA6046" bold="true" box="[543,628,1859,1880]" pageId="2" pageNumber="87">
<taxonomicNameLabel id="A1AECA3DFFBFFFE1041BF8F6FDDA6046" box="[543,628,1859,1880]" pageId="2" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, male (cl 3.3 mm) from Moorea, French Polynesia (FLMNH UF 23302); A—frontal region, dorsal; B—same, lateral; C—detail of pterygostomial angle, lateral; D—posterior abdominal somites, lateral; E—telson, dorsal; F—epistomial sclerites; G—tooth on ventromesial carina of first article of antennular peduncle, lateral; H—third maxilliped, lateral; I—second pereiopod, lateral; J—third pereiopod, lateral; K—same, detail of distal propodus and dactylus, mesial; L—fifth pereiopod, lateral; M—pleopod, lateral; N—uropod, dorsal. Scale bars as indicated except for C, F, G (drawn without scale).
</paragraph>
</caption>
<caption id="DC96FBDCFFBEFFE00693FB2DFC7A63F5" httpUri="https://zenodo.org/record/288923/files/figure.png" pageId="3" pageNumber="88" targetBox="[164,1429,201,1146]" targetPageId="3">
<paragraph id="8856AB54FFBEFFE00693FB2DFC7A63F5" blockId="3.[151,1436,1176,1259]" pageId="3" pageNumber="88">
<emphasis id="BA9D7746FFBEFFE00693FB2DFEBF63B3" bold="true" box="[151,273,1176,1198]" pageId="3" pageNumber="88">FIGURE 2</emphasis>
. 
<taxonomicName id="4FE9D0D7FFBEFFE00726FB2DFDBB63B0" box="[290,533,1176,1198]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="88" phylum="Arthropoda" rank="species" species="balaeniceps" status="sp. nov.">
<emphasis id="BA9D7746FFBEFFE00726FB2DFDBB63B0" box="[290,533,1176,1198]" italics="true" pageId="3" pageNumber="88">Alpheopsis balaeniceps</emphasis>
</taxonomicName>
<emphasis id="BA9D7746FFBEFFE0041BFB2DFDDA63B3" bold="true" box="[543,628,1176,1197]" pageId="3" pageNumber="88">
<taxonomicNameLabel id="A1AECA3DFFBEFFE0041BFB2DFDDA63B3" box="[543,628,1176,1197]" pageId="3" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, male (cl 3.3 mm) from Moorea, French Polynesia (FLMNH UF 23302); A—right (major) cheliped, lateral; B—same, chela with fingers closed, lateral; C—same, chela fingers open, lateral; D—left (minor) cheliped, lateral; E—same, chela, mesial. Scale bars as indicated.
</paragraph>
</caption>
<paragraph id="8856AB54FFBEFFE006C3FA5EFAB360AD" blockId="3.[151,1437,1299,1972]" pageId="3" pageNumber="88">
Second pereiopod slender; ischium slightly longer than merus; carpus five-articulated; ratio of carpal articles (from proximal to distal) approximately equal to: 3: 1: 1.3: 1.2: 1.5; chela longer than distal carpal article, with fingers subequal to palm in length, simple (
<figureCitation id="10D2B7D1FFBEFFE00494F986FD7A6155" box="[656,724,1587,1612]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[207,1369,198,1812]" captionTargetId="figure@2.[198,1380,195,1829]" captionTargetPageId="2" captionText="FIGURE 1. Alpheopsis balaeniceps sp. nov., holotype, male (cl 3.3 mm) from Moorea, French Polynesia (FLMNH UF 23302); A—frontal region, dorsal; B—same, lateral; C—detail of pterygostomial angle, lateral; D—posterior abdominal somites, lateral; E—telson, dorsal; F—epistomial sclerites; G—tooth on ventromesial carina of first article of antennular peduncle, lateral; H—third maxilliped, lateral; I—second pereiopod, lateral; J—third pereiopod, lateral; K—same, detail of distal propodus and dactylus, mesial; L—fifth pereiopod, lateral; M—pleopod, lateral; N—uropod, dorsal. Scale bars as indicated except for C, F, G (drawn without scale)." httpUri="https://zenodo.org/record/288922/files/figure.png" pageId="3" pageNumber="88">
Fig. 
<date id="FC578D94FFBEFFE004CCF986FD7A6155" box="[712,724,1587,1611]" pageId="3" pageNumber="88">1</date>
</figureCitation>
I). Third pereiopod slender; ischium with two stout spiniform setae on ventrolateral surface; merus about six times as long as wide, unarmed; carpus about 0.6 length of merus, with small spiniform seta on distoventral margin; propodus with three slender spiniform setae along ventral margin and two distal spiniform setae, one much longer than other, adjacent to dactylus; dactylus about 0.4 length of propodus, slender, simple, subconical, gradually curving distally, with some setae subdistally (
<figureCitation id="10D2B7D1FFBEFFE002C3F976FAA961C5" box="[1223,1287,1731,1756]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[207,1369,198,1812]" captionTargetId="figure@2.[198,1380,195,1829]" captionTargetPageId="2" captionText="FIGURE 1. Alpheopsis balaeniceps sp. nov., holotype, male (cl 3.3 mm) from Moorea, French Polynesia (FLMNH UF 23302); A—frontal region, dorsal; B—same, lateral; C—detail of pterygostomial angle, lateral; D—posterior abdominal somites, lateral; E—telson, dorsal; F—epistomial sclerites; G—tooth on ventromesial carina of first article of antennular peduncle, lateral; H—third maxilliped, lateral; I—second pereiopod, lateral; J—third pereiopod, lateral; K—same, detail of distal propodus and dactylus, mesial; L—fifth pereiopod, lateral; M—pleopod, lateral; N—uropod, dorsal. Scale bars as indicated except for C, F, G (drawn without scale)." httpUri="https://zenodo.org/record/288922/files/figure.png" pageId="3" pageNumber="88">Fig. 1</figureCitation>
J, K). Fourth pereiopod generally similar to third pereiopod in proportions and armature on ischium and propodus. Fifth pereiopod more slender than third and fourth pereiopods; ischium with one spiniform seta on ventrolateral surface; merus at least eight times as long as wide, unarmed; carpus about 0.7 length of merus, with stiff seta on distoventral margin; propodus long and slender, with at least four small spiniform setae along ventral margin (some not visible in lateral view), one long distal spiniform setae adjacent to dactylus, and several rows of serrulate setae on distolateral surface; dactylus about 0.35 length of propodus, similar to that of third pereiopod (
<figureCitation id="10D2B7D1FFBEFFE002BDF82EFB5660AD" box="[1209,1272,1947,1972]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[207,1369,198,1812]" captionTargetId="figure@2.[198,1380,195,1829]" captionTargetPageId="2" captionText="FIGURE 1. Alpheopsis balaeniceps sp. nov., holotype, male (cl 3.3 mm) from Moorea, French Polynesia (FLMNH UF 23302); A—frontal region, dorsal; B—same, lateral; C—detail of pterygostomial angle, lateral; D—posterior abdominal somites, lateral; E—telson, dorsal; F—epistomial sclerites; G—tooth on ventromesial carina of first article of antennular peduncle, lateral; H—third maxilliped, lateral; I—second pereiopod, lateral; J—third pereiopod, lateral; K—same, detail of distal propodus and dactylus, mesial; L—fifth pereiopod, lateral; M—pleopod, lateral; N—uropod, dorsal. Scale bars as indicated except for C, F, G (drawn without scale)." httpUri="https://zenodo.org/record/288922/files/figure.png" pageId="3" pageNumber="88">Fig. 1</figureCitation>
L).
</paragraph>
<paragraph id="8856AB54FFB9FFE706C3FF22FD4E6602" blockId="4.[151,1436,151,392]" pageId="4" pageNumber="89">
Male second pleopod with appendix masculina exceeding appendix interna; apex with several long stiff setae (
<figureCitation id="10D2B7D1FFB9FFE7069BFF09FF4D67CA" box="[159,227,188,213]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[207,1369,198,1812]" captionTargetId="figure@2.[198,1380,195,1829]" captionTargetPageId="2" captionText="FIGURE 1. Alpheopsis balaeniceps sp. nov., holotype, male (cl 3.3 mm) from Moorea, French Polynesia (FLMNH UF 23302); A—frontal region, dorsal; B—same, lateral; C—detail of pterygostomial angle, lateral; D—posterior abdominal somites, lateral; E—telson, dorsal; F—epistomial sclerites; G—tooth on ventromesial carina of first article of antennular peduncle, lateral; H—third maxilliped, lateral; I—second pereiopod, lateral; J—third pereiopod, lateral; K—same, detail of distal propodus and dactylus, mesial; L—fifth pereiopod, lateral; M—pleopod, lateral; N—uropod, dorsal. Scale bars as indicated except for C, F, G (drawn without scale)." httpUri="https://zenodo.org/record/288922/files/figure.png" pageId="4" pageNumber="89">Fig. 1</figureCitation>
M). Uropod with lateral lobe of protopod ending in sharp tooth; endopod and exopod broadly ovate; distolateral tooth strong, sharp; adjacent spiniform seta long, slender; diaresis with subtriangular tooth adjacent to spiniform seta and sinuous mesial portion (
<figureCitation id="10D2B7D1FFB9FFE7047EFEB1FD176602" box="[634,697,260,285]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[207,1369,198,1812]" captionTargetId="figure@2.[198,1380,195,1829]" captionTargetPageId="2" captionText="FIGURE 1. Alpheopsis balaeniceps sp. nov., holotype, male (cl 3.3 mm) from Moorea, French Polynesia (FLMNH UF 23302); A—frontal region, dorsal; B—same, lateral; C—detail of pterygostomial angle, lateral; D—posterior abdominal somites, lateral; E—telson, dorsal; F—epistomial sclerites; G—tooth on ventromesial carina of first article of antennular peduncle, lateral; H—third maxilliped, lateral; I—second pereiopod, lateral; J—third pereiopod, lateral; K—same, detail of distal propodus and dactylus, mesial; L—fifth pereiopod, lateral; M—pleopod, lateral; N—uropod, dorsal. Scale bars as indicated except for C, F, G (drawn without scale)." httpUri="https://zenodo.org/record/288922/files/figure.png" pageId="4" pageNumber="89">Fig. 1</figureCitation>
N).
</paragraph>
<paragraph id="8856AB54FFB9FFE706C3FE92FBEA6696" blockId="4.[151,1436,151,392]" pageId="4" pageNumber="89">Gill-exopod formula: five pleurobranchs above first to fifth pereiopods; five mastigobranchs on coxae of third maxilliped and first to fourth pereiopods; five setobranchs on coxae of first to fifth pereiopods; one arthrobranch at base of third maxilliped; no podobranchs; three exopods on first to third maxilliped.</paragraph>
<caption id="DC96FBDCFFB9FFE70693F80EFC3160EE" httpUri="https://zenodo.org/record/288924/files/figure.png" pageId="4" pageNumber="89" targetBox="[151,1436,411,1955]" targetPageId="4">
<paragraph id="8856AB54FFB9FFE70693F80EFC3160EE" blockId="4.[151,1435,1979,2032]" pageId="4" pageNumber="89">
<emphasis id="BA9D7746FFB9FFE70693F80EFEBF60CE" bold="true" box="[151,273,1979,2001]" pageId="4" pageNumber="89">FIGURE 3</emphasis>
. 
<taxonomicName id="4FE9D0D7FFB9FFE70726F80EFDBB60CF" box="[290,533,1979,2001]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="89" phylum="Arthropoda" rank="species" species="balaeniceps" status="sp. nov.">
<emphasis id="BA9D7746FFB9FFE70726F80EFDBB60CF" box="[290,533,1979,2001]" italics="true" pageId="4" pageNumber="89">Alpheopsis balaeniceps</emphasis>
</taxonomicName>
<emphasis id="BA9D7746FFB9FFE7041BF809FDDA60CF" bold="true" box="[543,628,1980,2001]" pageId="4" pageNumber="89">
<taxonomicNameLabel id="A1AECA3DFFB9FFE7041BF809FDDA60CF" box="[543,628,1980,2001]" pageId="4" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, male (cl 3.3 mm) from Moorea, French Polynesia (FLMNH UF 23302), in life; A—dorsal view; B—lateral view. Photographs by the author.
</paragraph>
</caption>
<paragraph id="8856AB54FFB8FFE606C3FF22FBFB665E" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="90">
<emphasis id="BA9D7746FFB8FFE606C3FF22FED567AE" bold="true" box="[199,379,151,176]" pageId="5" pageNumber="90">Colour pattern</emphasis>
. Carapace semitransparent with red chromatophores forming two diffuse transverse bands in posterior half and with smaller field of red chromatophores in frontal area; abdomen semitransparent with six transverse bands of red chromatophores, one across each somite; antennular and antennal peduncles, uropods and telson almost entirely covered with red chromatophores; chelipeds mostly hyaline-whitish except for some reddish areas with red chromatophores; walking legs and pleopods mostly colourless (
<figureCitation id="10D2B7D1FFB8FFE60207FE92FBE9665E" box="[1027,1095,295,320]" captionStart="FIGURE 3" captionStartId="4.[151,250,1979,2001]" captionTargetBox="[151,1436,411,1955]" captionTargetId="figure@4.[151,1436,411,1955]" captionTargetPageId="4" captionText="FIGURE 3. Alpheopsis balaeniceps sp. nov., holotype, male (cl 3.3 mm) from Moorea, French Polynesia (FLMNH UF 23302), in life; A—dorsal view; B—lateral view. Photographs by the author." httpUri="https://zenodo.org/record/288924/files/figure.png" pageId="5" pageNumber="90">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C0F3F8DFFFB8FFE606C3FEF9FE7566B3" pageId="5" pageNumber="90" type="etymology">
<paragraph id="8856AB54FFB8FFE606C3FEF9FE7566B3" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="90">
<emphasis id="BA9D7746FFB8FFE606C3FEF9FEE8667B" bold="true" box="[199,326,332,357]" pageId="5" pageNumber="90">Etymology</emphasis>
. The new species name refers to the characteristic balaeniceps-like setae on the cheliped fingers (term “ 
<taxonomicName id="4FE9D0D7FFB8FFE606E1FEDAFEC26696" box="[229,364,367,392]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="balaeniceps">balaeniceps</taxonomicName>
” first proposed by 
<bibRefCitation id="EC78D6A5FFB8FFE60440FEDAFD586696" author="Coutiere" box="[580,758,367,392]" pageId="5" pageNumber="90" refString="Coutiere, H. (1899) Les Alpheidae. Morphologie externe et interne, formes larvaires, bionomie. Annales des Sciences Naturelles, Zoologie et Paleontologie, (8), 9, 1 - 559, pls. 1 - 6." type="journal article" year="1899">Coutière (1899)</bibRefCitation>
to describe the minor cheliped in some species of 
<taxonomicName id="4FE9D0D7FFB8FFE60335FEC5FA256696" box="[1329,1419,368,392]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="BA9D7746FFB8FFE60335FEC5FA256696" box="[1329,1419,368,392]" italics="true" pageId="5" pageNumber="90">Alpheus</emphasis>
</taxonomicName>
); used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C0F3F8DFFFB8FFE606C3FE02FD3466CE" box="[199,666,439,464]" pageId="5" pageNumber="90" type="materials_examined">
<paragraph id="8856AB54FFB8FFE606C3FE02FD3466CE" blockId="5.[151,1437,151,2013]" box="[199,666,439,464]" pageId="5" pageNumber="90">
<emphasis id="BA9D7746FFB8FFE606C3FE02FEF366CE" bold="true" box="[199,349,439,464]" pageId="5" pageNumber="90">
<typeStatus id="575215F6FFB8FFE606C3FE02FEAC66CE" box="[199,258,439,464]" pageId="5" pageNumber="90">Type</typeStatus>
locality
</emphasis>
. Moorea, 
<collectingCountry id="F0FEEBC4FFB8FFE607D5FE02FD3866CE" box="[465,662,439,464]" name="French Polynesia" pageId="5" pageNumber="90">French Polynesia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C0F3F8DFFFB8FFE606C3FE69FBEF66EB" box="[199,1089,476,501]" pageId="5" pageNumber="90" type="distribution">
<paragraph id="8856AB54FFB8FFE606C3FE69FBEF66EB" blockId="5.[151,1437,151,2013]" box="[199,1089,476,501]" pageId="5" pageNumber="90">
<emphasis id="BA9D7746FFB8FFE606C3FE69FEF766EB" bold="true" box="[199,345,476,501]" pageId="5" pageNumber="90">Distribution</emphasis>
. Presently known only from the 
<typeStatus id="575215F6FFB8FFE604CDFE68FD5766EB" box="[713,761,477,501]" pageId="5" pageNumber="90">type</typeStatus>
locality in 
<collectingCountry id="F0FEEBC4FFB8FFE6057CFE69FB9066EB" box="[888,1086,476,501]" name="French Polynesia" pageId="5" pageNumber="90">French Polynesia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C0F3F8DFFFB8FFE606C3FE4AFB5B624B" pageId="5" pageNumber="90" type="discussion">
<paragraph id="8856AB54FFB8FFE606C3FE4AFDC8640B" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="90">
<emphasis id="BA9D7746FFB8FFE606C3FE4AFE9A6506" bold="true" box="[199,308,511,536]" pageId="5" pageNumber="90">Remarks</emphasis>
. 
<taxonomicName id="4FE9D0D7FFB8FFE60743FDB5FDFB6506" box="[327,597,512,536]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="balaeniceps" status="sp. nov.">
<emphasis id="BA9D7746FFB8FFE60743FDB5FDFB6506" box="[327,597,512,536]" italics="true" pageId="5" pageNumber="90">Alpheopsis balaeniceps</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1AECA3DFFB8FFE60466FE4AFD6F6506" box="[610,705,511,536]" pageId="5" pageNumber="90" rank="species">
<emphasis id="BA9D7746FFB8FFE60466FE4AFD6F6506" bold="true" box="[610,705,511,536]" pageId="5" pageNumber="90">sp. nov.</emphasis>
</taxonomicNameLabel>
is the only known species in the genus characterised by the presence of balaeniceps-type rows of plumose setae on the cheliped fingers. Two species of 
<taxonomicName id="4FE9D0D7FFB8FFE602A5FD91FAB26522" box="[1185,1308,548,572]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="BA9D7746FFB8FFE602A5FD91FAB26522" box="[1185,1308,548,572]" italics="true" pageId="5" pageNumber="90">Alpheopsis</emphasis>
</taxonomicName>
, namely 
<taxonomicName id="4FE9D0D7FFB8FFE60380FD91FE11657E" authority="Coutiere, 1908" authorityName="Coutiere" authorityYear="1908" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="idiocarpus">
<emphasis id="BA9D7746FFB8FFE60380FD91FEA1657E" italics="true" pageId="5" pageNumber="90">A. idiocarpus</emphasis>
<bibRefCitation id="EC78D6A5FFB8FFE60713FDF2FE11657E" author="Coutiere" box="[279,447,583,608]" pageId="5" pageNumber="90" refString="Coutiere, H. (1908) Sur quelques nouvelles especes d'Alpheidae. Bulletin de la Societe Philomathique de Paris, Serie 9, 10, 191 - 216." type="journal article" year="1908">Coutière, 1908</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4FE9D0D7FFB8FFE607FCFDFDFCC7657E" authority="Banner, 1953" authorityName="Banner" authorityYear="1953" box="[504,873,583,608]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="biunguiculata">
<emphasis id="BA9D7746FFB8FFE607FCFDFDFD1B657E" box="[504,693,584,608]" italics="true" pageId="5" pageNumber="90">A. biunguiculata</emphasis>
(
<bibRefCitation id="EC78D6A5FFB8FFE604C1FDFDFCCF657E" author="Banner" box="[709,865,583,608]" pageId="5" pageNumber="90" refString="Banner, A. H. (1953) The Crangonidae, or snapping shrimp, of Hawaii. Pacific Science, 7, 3 - 147." type="journal article" year="1953">Banner, 1953</bibRefCitation>
)
</taxonomicName>
, were described based on fragmentary specimens lacking both chelipeds (
<bibRefCitation id="EC78D6A5FFB8FFE607A8FDD9FDFA659A" author="Coutiere" box="[428,596,620,645]" pageId="5" pageNumber="90" refString="Coutiere, H. (1908) Sur quelques nouvelles especes d'Alpheidae. Bulletin de la Societe Philomathique de Paris, Serie 9, 10, 191 - 216." type="journal article" year="1908">Coutière 1908</bibRefCitation>
, 
<bibRefCitation id="EC78D6A5FFB8FFE60467FDD9FD30659A" author="Coutiere" box="[611,670,620,644]" pageId="5" pageNumber="90" refString="Coutiere, H. (1921) The Percy Sladen Trust Expedition to the Indian Ocean in 1905, under the leadership of Mr. J. Stanley Gardiner, M. A. No. X. Les especes d'Alpheidae rapportees de l'Ocean Indien par M. J. Stanley Gardiner. Transactions of the Linnean Society of London, Series 2 (Zoology), 17, 413 - 428, pls. 60 - 64." type="journal article" year="1921">1921</bibRefCitation>
; 
<bibRefCitation id="EC78D6A5FFB8FFE604A9FDD8FCEB659B" author="Banner" box="[685,837,620,645]" pageId="5" pageNumber="90" refString="Banner, A. H. (1953) The Crangonidae, or snapping shrimp, of Hawaii. Pacific Science, 7, 3 - 147." type="journal article" year="1953">Banner 1953</bibRefCitation>
). However, 
<taxonomicName id="4FE9D0D7FFB8FFE605D7FDD9FBC3659A" box="[979,1133,620,644]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="idiocarpus">
<emphasis id="BA9D7746FFB8FFE605D7FDD9FBC3659A" box="[979,1133,620,644]" italics="true" pageId="5" pageNumber="90">A. idiocarpus</emphasis>
</taxonomicName>
can be separated from 
<taxonomicName id="4FE9D0D7FFB8FFE60380FDD9FEB365B6" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="balaeniceps" status="sp. nov.">
<emphasis id="BA9D7746FFB8FFE60380FDD9FEB365B6" italics="true" pageId="5" pageNumber="90">A. balaeniceps</emphasis>
</taxonomicName>
<emphasis id="BA9D7746FFB8FFE60720FD3AFED365B6" bold="true" box="[292,381,655,680]" pageId="5" pageNumber="90">
<taxonomicNameLabel id="A1AECA3DFFB8FFE60720FD3AFED365B6" box="[292,381,655,680]" pageId="5" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the second pereiopod carpus subdivided into three articles (
<bibRefCitation id="EC78D6A5FFB8FFE60244FD3AFA9765B6" author="Coutiere" box="[1088,1337,655,680]" pageId="5" pageNumber="90" refString="Coutiere, H. (1921) The Percy Sladen Trust Expedition to the Indian Ocean in 1905, under the leadership of Mr. J. Stanley Gardiner, M. A. No. X. Les especes d'Alpheidae rapportees de l'Ocean Indien par M. J. Stanley Gardiner. Transactions of the Linnean Society of London, Series 2 (Zoology), 17, 413 - 428, pls. 60 - 64." type="journal article" year="1921" yearSuffix="a">Coutière 1921: fig. 2a</bibRefCitation>
) and not five, as in the new species and most other species in the genus, whereas 
<taxonomicName id="4FE9D0D7FFB8FFE6021FFD01FB7365D2" box="[1051,1245,692,716]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="biunguiculata">
<emphasis id="BA9D7746FFB8FFE6021FFD01FB7365D2" box="[1051,1245,692,716]" italics="true" pageId="5" pageNumber="90">A. biunguiculata</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4FE9D0D7FFB8FFE60380FD01FEB365EE" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="balaeniceps" status="sp. nov.">
<emphasis id="BA9D7746FFB8FFE60380FD01FEB365EE" italics="true" pageId="5" pageNumber="90">A. balaeniceps</emphasis>
</taxonomicName>
<emphasis id="BA9D7746FFB8FFE60720FD62FED065EE" bold="true" box="[292,382,727,752]" pageId="5" pageNumber="90">
<taxonomicNameLabel id="A1AECA3DFFB8FFE60720FD62FED065EE" box="[292,382,727,752]" pageId="5" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the shorter descendant rostrum and much stouter biunguiculate dactyli of the third to fifth pereiopods (
<bibRefCitation id="EC78D6A5FFB8FFE60724FD49FE18640B" author="Banner" box="[288,438,764,789]" pageId="5" pageNumber="90" refString="Banner, A. H. (1953) The Crangonidae, or snapping shrimp, of Hawaii. Pacific Science, 7, 3 - 147." type="journal article" year="1953">Banner 1953</bibRefCitation>
: fig. 5a, b, e, f).
</paragraph>
<paragraph id="8856AB54FFB8FFE606C3FCAAFB5B624B" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="90">
The shape of the frontal margin of the carapace, including a relatively long pointed rostrum and the absence of orbital teeth, as well as the general shape of the chelipeds suggest that 
<taxonomicName id="4FE9D0D7FFB8FFE605B1FCF1FB106442" box="[949,1214,836,860]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="balaeniceps" status="sp. nov.">
<emphasis id="BA9D7746FFB8FFE605B1FCF1FB106442" box="[949,1214,836,860]" italics="true" pageId="5" pageNumber="90">Alpheopsis balaeniceps</emphasis>
</taxonomicName>
<emphasis id="BA9D7746FFB8FFE602C1FCF1FA8E6443" bold="true" box="[1221,1312,836,861]" pageId="5" pageNumber="90">
<taxonomicNameLabel id="A1AECA3DFFB8FFE602C1FCF1FA8E6443" box="[1221,1312,836,861]" pageId="5" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
belongs to the 
<taxonomicName id="4FE9D0D7FFB8FFE606C3FCDDFDA6649E" authority="Coutiere, 1897" authorityName="Coutiere" authorityYear="1897" box="[199,520,871,896]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="aequalis">
<emphasis id="BA9D7746FFB8FFE606C3FCDDFEE2649E" box="[199,332,872,896]" italics="true" pageId="5" pageNumber="90">A. aequalis</emphasis>
<bibRefCitation id="EC78D6A5FFB8FFE6075FFCD2FDA6649E" author="Coutiere" box="[347,520,871,896]" pageId="5" pageNumber="90" refString="Coutiere, H. (1897) Note sur quelques genres nouveaux ou peu connus d'Alpheides, formant la sous-famille des Alpheopsides. Bulletin du Museum d'Histoire Naturelle, 2, 380 - 386." type="journal article" year="1897">Coutière, 1897</bibRefCitation>
</taxonomicName>
species complex. This species complex contains, besides the taxonomically problematic 
<taxonomicName id="4FE9D0D7FFB8FFE60720FC39FE0C64BA" box="[292,418,908,932]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="aequalis">
<emphasis id="BA9D7746FFB8FFE60720FC39FE0C64BA" box="[292,418,908,932]" italics="true" pageId="5" pageNumber="90">A. aequalis</emphasis>
</taxonomicName>
(Indo-West Pacific) with two questionable synonyms (
<taxonomicName id="4FE9D0D7FFB8FFE6020FFC39FA3264BA" authority="Coutiere, 1903" authorityName="Coutiere" authorityYear="1903" box="[1035,1436,908,933]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="subSpecies" species="aequalis" subSpecies="truncata">
<emphasis id="BA9D7746FFB8FFE6020FFC39FB4064BA" box="[1035,1262,908,932]" italics="true" pageId="5" pageNumber="90">A. aequalis truncata</emphasis>
<bibRefCitation id="EC78D6A5FFB8FFE602F1FC39FA3264BA" author="Coutiere" box="[1269,1436,908,933]" pageId="5" pageNumber="90" refString="Coutiere, H. (1903) Note sur quelques Alpheidae des Maldives et Laquedives. Bulletin de la Societe Philomathique de Paris, Serie 9, 5, 72 - 90." type="journal article" year="1903">Coutière, 1903</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4FE9D0D7FFB8FFE606C3FC05FD8F64D6" authority="De Man, 1910" authorityName="De Man" authorityYear="1910" box="[199,545,944,968]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="BA9D7746FFB8FFE606C3FC05FEDF64D6" box="[199,369,944,968]" italics="true" pageId="5" pageNumber="90">A. consobrinus</emphasis>
<bibRefCitation id="EC78D6A5FFB8FFE6077DFC05FD8F64D6" author="De" box="[377,545,944,968]" pageId="5" pageNumber="90" refString="De Man, J. G. (1910) Diagnoses of new species of macrurous decapod Crustacea from the &quot; Siboga-Expedition &quot;. Tijdschrift der Nederlandsche Dierkundige Vereeniging, 2 (11), 287 - 319." type="journal article" year="1910">De Man, 1910</bibRefCitation>
</taxonomicName>
), at least four other species: 
<taxonomicName id="4FE9D0D7FFB8FFE6056CFC05FAA964D6" authority="Banner &amp; Banner, 1973" authorityName="Banner &amp; Banner" authorityYear="1973" box="[872,1287,944,968]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="yaldwyni">
<emphasis id="BA9D7746FFB8FFE6056CFC05FC4364D6" box="[872,1005,944,968]" italics="true" pageId="5" pageNumber="90">A. yaldwyni</emphasis>
<bibRefCitation id="EC78D6A5FFB8FFE605F1FC05FAA964D6" author="Banner" box="[1013,1287,944,968]" pageId="5" pageNumber="90" refString="Banner, D. M. &amp; Banner, A. H. (1973) The alpheid shrimp of Australia. Part I: The lower genera. Records of the Australian Museum, 28, 291 - 382. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.28.1973.407" type="journal article" year="1973">Banner &amp; Banner, 1973</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4FE9D0D7FFB8FFE60317FC05FE8064F3" authority="Banner, 1956" authorityName="Banner" authorityYear="1956" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="tetrarthri">
<emphasis id="BA9D7746FFB8FFE60317FC05FA3264D6" box="[1299,1436,944,968]" italics="true" pageId="5" pageNumber="90">A. tetrarthri</emphasis>
<bibRefCitation id="EC78D6A5FFB8FFE60693FC60FE8064F3" author="Banner" box="[151,302,980,1005]" pageId="5" pageNumber="90" refString="Banner, A. H. (1956) Contributions to the knowledge of the alpheid shrimp of the Pacific Ocean. Part I. Collections from the Mariana Archipelago. Pacific Science, 10, 318 - 373." type="journal article" year="1956">Banner, 1956</bibRefCitation>
</taxonomicName>
(both Indo-West Pacific), 
<taxonomicName id="4FE9D0D7FFB8FFE6045BFC61FCE364F2" authority="Chace, 1972" authorityName="Chace" authorityYear="1972" box="[607,845,980,1005]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="labis">
<emphasis id="BA9D7746FFB8FFE6045BFC61FD1864F2" box="[607,694,980,1004]" italics="true" pageId="5" pageNumber="90">A. labis</emphasis>
<bibRefCitation id="EC78D6A5FFB8FFE604BBFC61FCE364F2" author="Chace" box="[703,845,980,1005]" pageId="5" pageNumber="90" refString="Chace, F. A. Jr. (1972) The shrimps of the Smithsonian-Bredin Caribbean expeditions with a summary of the West Indian shallow-water species (Crustacea: Decapoda: Natantia). Smithsonian Contributions to Zoology, 98, 1 - 179. http: // dx. doi. org / 10.5479 / si. 00810282.98" type="journal article" year="1972">Chace, 1972</bibRefCitation>
</taxonomicName>
(western Atlantic), and 
<taxonomicName id="4FE9D0D7FFB8FFE60260FC61FF61630E" authority="Wicksten, 1992" authorityName="Wicksten" authorityYear="1992" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="allanhancocki">
<emphasis id="BA9D7746FFB8FFE60260FC61FA8D64F2" box="[1124,1315,980,1004]" italics="true" pageId="5" pageNumber="90">A. allanhancocki</emphasis>
<bibRefCitation id="EC78D6A5FFB8FFE60328FC61FF61630E" author="Wicksten" pageId="5" pageNumber="90" refString="Wicksten, M. K. (1992) A new species of Alpheopsis (Decapoda, Alpheidae) from the tropical eastern Pacific, with a key to the species of Alpheopsis of the Americas. Crustaceana, 63, 51 - 56. http: // dx. doi. org / 10.1163 / 156854092 X 00271" type="journal article" year="1992">Wicksten, 1992</bibRefCitation>
</taxonomicName>
(eastern Pacific). As already mentioned, none of these species has balaeniceps-type setae on the chelipeds. Examination of 
<collectingCountry id="F0FEEBC4FFB8FFE60748FBA8FE3F632A" box="[332,401,1053,1076]" name="Papua New Guinea" pageId="5" pageNumber="90">Papua</collectingCountry>
New Guinean material identified as 
<taxonomicName id="4FE9D0D7FFB8FFE60529FBA9FC05632A" box="[813,939,1052,1076]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="aequalis">
<emphasis id="BA9D7746FFB8FFE60529FBA9FC05632A" box="[813,939,1052,1076]" italics="true" pageId="5" pageNumber="90">A. aequalis</emphasis>
</taxonomicName>
[
<emphasis id="BA9D7746FFB8FFE605BFFBA8FB82632A" box="[955,1068,1052,1076]" italics="true" pageId="5" pageNumber="90">
<taxonomicName id="4FE9D0D7FFB8FFE605BFFBA8FC57632A" box="[955,1017,1053,1076]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="sensu">sensu</taxonomicName>
lato
</emphasis>
] and 
<taxonomicName id="4FE9D0D7FFB8FFE60268FBA9FB5F632A" box="[1132,1265,1052,1076]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="yaldwyni">
<emphasis id="BA9D7746FFB8FFE60268FBA9FB5F632A" box="[1132,1265,1052,1076]" italics="true" pageId="5" pageNumber="90">A. yaldwyni</emphasis>
</taxonomicName>
confirmed that the cheliped fingers in these specimens only possess simple setae on the lateral and mesial surfaces, in addition to rows of spaced simple setae along the cutting edges, not homologous to the plumose balaeniceps-type setae found in 
<taxonomicName id="4FE9D0D7FFB8FFE606B0FB3DFEF463BE" box="[180,346,1160,1184]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="balaeniceps" status="sp. nov.">
<emphasis id="BA9D7746FFB8FFE606B0FB3DFEF463BE" box="[180,346,1160,1184]" italics="true" pageId="5" pageNumber="90">A. balaeniceps</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1AECA3DFFB8FFE60765FB32FE1563BE" box="[353,443,1159,1184]" pageId="5" pageNumber="90" rank="species">
<emphasis id="BA9D7746FFB8FFE60765FB32FE1A63BE" bold="true" box="[353,436,1159,1184]" pageId="5" pageNumber="90">sp. nov</emphasis>
.
</taxonomicNameLabel>
In addition, 
<taxonomicName id="4FE9D0D7FFB8FFE6044FFB3DFD5F63BE" box="[587,753,1160,1184]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="balaeniceps" status="sp. nov.">
<emphasis id="BA9D7746FFB8FFE6044FFB3DFD5F63BE" box="[587,753,1160,1184]" italics="true" pageId="5" pageNumber="90">A. balaeniceps</emphasis>
</taxonomicName>
<emphasis id="BA9D7746FFB8FFE604FCFB32FCFC63BE" bold="true" box="[760,850,1159,1184]" pageId="5" pageNumber="90">
<taxonomicNameLabel id="A1AECA3DFFB8FFE604FCFB32FCFC63BE" box="[760,850,1159,1184]" pageId="5" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs specifically from 
<taxonomicName id="4FE9D0D7FFB8FFE60274FB3DFB4163BE" box="[1136,1263,1160,1184]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="aequalis">
<emphasis id="BA9D7746FFB8FFE60274FB3DFB4163BE" box="[1136,1263,1160,1184]" italics="true" pageId="5" pageNumber="90">A. aequalis</emphasis>
</taxonomicName>
by the absence of stout spiniform setae on the cheliped merus (present in 
<taxonomicName id="4FE9D0D7FFB8FFE60539FB19FC1063DA" box="[829,958,1196,1220]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="aequalis">
<emphasis id="BA9D7746FFB8FFE60539FB19FC1063DA" box="[829,958,1196,1220]" italics="true" pageId="5" pageNumber="90">A. aequalis</emphasis>
</taxonomicName>
); the presence of a spiniform seta on the ischium of the fifth pereiopod (absent in 
<taxonomicName id="4FE9D0D7FFB8FFE6046AFB65FD4063F6" box="[622,750,1232,1256]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="aequalis">
<emphasis id="BA9D7746FFB8FFE6046AFB65FD4063F6" box="[622,750,1232,1256]" italics="true" pageId="5" pageNumber="90">A. aequalis</emphasis>
</taxonomicName>
); the presence of two spiniform setae on the ischium of the third pereiopod (unarmed in 
<taxonomicName id="4FE9D0D7FFB8FFE607DCFB41FDF86212" box="[472,598,1268,1292]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="aequalis">
<emphasis id="BA9D7746FFB8FFE607DCFB41FDF86212" box="[472,598,1268,1292]" italics="true" pageId="5" pageNumber="90">A. aequalis</emphasis>
</taxonomicName>
); and the red-banded colour pattern (colour described as orange-yellow by 
<bibRefCitation id="EC78D6A5FFB8FFE60693FAA2FEE7622E" author="Coutiere" box="[151,329,1303,1328]" pageId="5" pageNumber="90" refString="Coutiere, H. (1899) Les Alpheidae. Morphologie externe et interne, formes larvaires, bionomie. Annales des Sciences Naturelles, Zoologie et Paleontologie, (8), 9, 1 - 559, pls. 1 - 6." type="journal article" year="1899">Coutière (1899)</bibRefCitation>
for 
<taxonomicName id="4FE9D0D7FFB8FFE6077DFAADFE59622E" box="[377,503,1304,1328]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="aequalis">
<emphasis id="BA9D7746FFB8FFE6077DFAADFE59622E" box="[377,503,1304,1328]" italics="true" pageId="5" pageNumber="90">A. aequalis</emphasis>
</taxonomicName>
) (cf. 
<bibRefCitation id="EC78D6A5FFB8FFE60431FAA2FD77622E" author="Coutiere" box="[565,729,1303,1328]" pageId="5" pageNumber="90" refString="Coutiere, H. (1899) Les Alpheidae. Morphologie externe et interne, formes larvaires, bionomie. Annales des Sciences Naturelles, Zoologie et Paleontologie, (8), 9, 1 - 559, pls. 1 - 6." type="journal article" year="1899">Coutière 1899</bibRefCitation>
; 
<bibRefCitation id="EC78D6A5FFB8FFE604E0FAADFCD6622E" author="Banner" box="[740,888,1303,1328]" pageId="5" pageNumber="90" refString="Banner, A. H. (1953) The Crangonidae, or snapping shrimp, of Hawaii. Pacific Science, 7, 3 - 147." type="journal article" year="1953">Banner 1953</bibRefCitation>
; 
<bibRefCitation id="EC78D6A5FFB8FFE60580FAADFB23622E" author="Banner" box="[900,1165,1304,1328]" pageId="5" pageNumber="90" refString="Banner, D. M. &amp; Banner, A. H. (1973) The alpheid shrimp of Australia. Part I: The lower genera. Records of the Australian Museum, 28, 291 - 382. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.28.1973.407" type="journal article" year="1973">Banner &amp; Banner 1973</bibRefCitation>
; 
<bibRefCitation id="EC78D6A5FFB8FFE6029DFAA2FAB9622E" author="Miya" box="[1177,1303,1303,1328]" pageId="5" pageNumber="90" refString="Miya, Y. (1983) Alpheopsis aequalis Coutiere, 1896, a species new to Japan, with remarks on its mouthparts (Crustacea, Decapoda, Alpheidae). Bulletin of the Faculty of Liberal Arts, Nagasaki University (Natural Science), 23, 17 - 21." type="journal article" year="1983">Miya 1983</bibRefCitation>
). It must be noted that 
<taxonomicName id="4FE9D0D7FFB8FFE60709FA89FE25624A" box="[269,395,1340,1364]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="aequalis">
<emphasis id="BA9D7746FFB8FFE60709FA89FE25624A" box="[269,395,1340,1364]" italics="true" pageId="5" pageNumber="90">A. aequalis</emphasis>
</taxonomicName>
[
<emphasis id="BA9D7746FFB8FFE6079FFA88FDA2624A" box="[411,524,1340,1364]" italics="true" pageId="5" pageNumber="90">
<taxonomicName id="4FE9D0D7FFB8FFE6079FFA88FE77624A" box="[411,473,1341,1364]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="sensu">sensu</taxonomicName>
lato
</emphasis>
] appears to contain several species and needs a thorough revision.
</paragraph>
</subSubSection>
<subSubSection id="C0F3F8DFFFB8FFE506C3FAD5FBAC6603" lastPageId="6" lastPageNumber="91" pageId="5" pageNumber="90" type="reference_group">
<paragraph id="8856AB54FFB8FFE606C3FAD5FC8361FE" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="90">
<taxonomicName id="4FE9D0D7FFB8FFE606C3FAD5FE616266" ID-CoL="85LCJ" box="[199,463,1376,1400]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="balaeniceps" status="sp. nov.">
<emphasis id="BA9D7746FFB8FFE606C3FAD5FE616266" box="[199,463,1376,1400]" italics="true" pageId="5" pageNumber="90">Alpheopsis balaeniceps</emphasis>
</taxonomicName>
<emphasis id="BA9D7746FFB8FFE607D2FAEAFD9E6266" bold="true" box="[470,560,1375,1400]" pageId="5" pageNumber="90">
<taxonomicNameLabel id="A1AECA3DFFB8FFE607D2FAEAFD9E6266" box="[470,560,1375,1400]" pageId="5" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be easily separated from 
<taxonomicName id="4FE9D0D7FFB8FFE6057BFAD5FBAA6266" ID-CoL="85LD8" box="[895,1028,1376,1400]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="yaldwyni">
<emphasis id="BA9D7746FFB8FFE6057BFAD5FBAA6266" box="[895,1028,1376,1400]" italics="true" pageId="5" pageNumber="90">A. yaldwyni</emphasis>
</taxonomicName>
by the more slender chelipeds, with a non-flattened, subcylindrical chelae (noticeably stouter and with a dorso-ventrally flattened chelae in 
<taxonomicName id="4FE9D0D7FFB8FFE60380FA31FF5262DE" ID-CoL="85LD8" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="yaldwyni">
<emphasis id="BA9D7746FFB8FFE60380FA31FF5262DE" italics="true" pageId="5" pageNumber="90">A. yaldwyni</emphasis>
</taxonomicName>
) and the pterygostomial angle bearing a minute tooth visible only at the highest magnification (compared to a conspicuously projecting tooth in 
<taxonomicName id="4FE9D0D7FFB8FFE60446FA79FD6B62FA" ID-CoL="85LD8" box="[578,709,1484,1508]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="yaldwyni">
<emphasis id="BA9D7746FFB8FFE60446FA79FD6B62FA" box="[578,709,1484,1508]" italics="true" pageId="5" pageNumber="90">A. yaldwyni</emphasis>
</taxonomicName>
). The new species also differs from 
<taxonomicName id="4FE9D0D7FFB8FFE6025FFA79FB4C62FA" ID-CoL="85LD5" box="[1115,1250,1484,1508]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="tetrarthri">
<emphasis id="BA9D7746FFB8FFE6025FFA79FB4C62FA" box="[1115,1250,1484,1508]" italics="true" pageId="5" pageNumber="90">A. tetrarthri</emphasis>
</taxonomicName>
, for instance, by the second pereiopod having a five-articulated carpus (four-articulated in 
<taxonomicName id="4FE9D0D7FFB8FFE605CFFA45FBFF6116" ID-CoL="85LD5" box="[971,1105,1520,1544]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="tetrarthri">
<emphasis id="BA9D7746FFB8FFE605CFFA45FBFF6116" box="[971,1105,1520,1544]" italics="true" pageId="5" pageNumber="90">A. tetrarthri</emphasis>
</taxonomicName>
); the much longer, horizontal rostrum (short and descendant in 
<taxonomicName id="4FE9D0D7FFB8FFE60409F9A1FD3A6132" ID-CoL="85LD5" box="[525,660,1556,1580]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="tetrarthri">
<emphasis id="BA9D7746FFB8FFE60409F9A1FD3A6132" box="[525,660,1556,1580]" italics="true" pageId="5" pageNumber="90">A. tetrarthri</emphasis>
</taxonomicName>
); the much longer stylocerite, reaching almost to the distal margin of the second article (not reaching mid-length of the second article in 
<taxonomicName id="4FE9D0D7FFB8FFE605ADF98DFB9D614E" ID-CoL="85LD5" box="[937,1075,1592,1616]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="tetrarthri">
<emphasis id="BA9D7746FFB8FFE605ADF98DFB9D614E" box="[937,1075,1592,1616]" italics="true" pageId="5" pageNumber="90">A. tetrarthri</emphasis>
</taxonomicName>
) (cf. 
<bibRefCitation id="EC78D6A5FFB8FFE60273F98DFAA1614E" author="Banner" box="[1143,1295,1591,1616]" pageId="5" pageNumber="90" refString="Banner, A. H. (1956) Contributions to the knowledge of the alpheid shrimp of the Pacific Ocean. Part I. Collections from the Mariana Archipelago. Pacific Science, 10, 318 - 373." type="journal article" year="1956">Banner 1956</bibRefCitation>
). Finally, 
<taxonomicName id="4FE9D0D7FFB8FFE60380F98DFEB3616A" ID-CoL="85LCJ" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="balaeniceps" status="sp. nov.">
<emphasis id="BA9D7746FFB8FFE60380F98DFEB3616A" italics="true" pageId="5" pageNumber="90">A. balaeniceps</emphasis>
</taxonomicName>
<emphasis id="BA9D7746FFB8FFE60728F9E9FE20616B" bold="true" box="[300,398,1628,1653]" pageId="5" pageNumber="90">
<taxonomicNameLabel id="A1AECA3DFFB8FFE60728F9E9FE20616B" box="[300,398,1628,1653]" pageId="5" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be separated from 
<taxonomicName id="4FE9D0D7FFB8FFE604B9F9E9FCB2616A" ID-CoL="84W69" box="[701,796,1628,1652]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="labis">
<emphasis id="BA9D7746FFB8FFE604B9F9E9FCB2616A" box="[701,796,1628,1652]" italics="true" pageId="5" pageNumber="90">A. labis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4FE9D0D7FFB8FFE60567F9E9FB84616A" ID-CoL="84W64" box="[867,1066,1628,1652]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="allanhancocki">
<emphasis id="BA9D7746FFB8FFE60567F9E9FB84616A" box="[867,1066,1628,1652]" italics="true" pageId="5" pageNumber="90">A. allanhancocki</emphasis>
</taxonomicName>
by the much narrower telson (conspicuously wider in 
<taxonomicName id="4FE9D0D7FFB8FFE607AEF935FE516186" ID-CoL="84W69" box="[426,511,1664,1688]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="labis">
<emphasis id="BA9D7746FFB8FFE607AEF935FE516186" box="[426,511,1664,1688]" italics="true" pageId="5" pageNumber="90">A. labis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4FE9D0D7FFB8FFE60432F935FD5D6186" ID-CoL="84W64" box="[566,755,1664,1688]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="allanhancocki">
<emphasis id="BA9D7746FFB8FFE60432F935FD5D6186" box="[566,755,1664,1688]" italics="true" pageId="5" pageNumber="90">A. allanhancocki</emphasis>
</taxonomicName>
) and from 
<taxonomicName id="4FE9D0D7FFB8FFE60574F935FB836186" ID-CoL="84W64" box="[880,1069,1664,1688]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="allanhancocki">
<emphasis id="BA9D7746FFB8FFE60574F935FB836186" box="[880,1069,1664,1688]" italics="true" pageId="5" pageNumber="90">A. allanhancocki</emphasis>
</taxonomicName>
also by the presence of a minute tooth on the pterygostomial angle (illustrated and described as blunt in 
<taxonomicName id="4FE9D0D7FFB8FFE605CBF911FB2161A2" ID-CoL="84W64" box="[975,1167,1700,1724]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="allanhancocki">
<emphasis id="BA9D7746FFB8FFE605CBF911FB2161A2" box="[975,1167,1700,1724]" italics="true" pageId="5" pageNumber="90">A. allanhancocki</emphasis>
</taxonomicName>
), in addition to several features on the chelipeds (cf. 
<bibRefCitation id="EC78D6A5FFB8FFE607E4F972FDC761FE" author="Chace" box="[480,617,1735,1760]" pageId="5" pageNumber="90" refString="Chace, F. A. Jr. (1972) The shrimps of the Smithsonian-Bredin Caribbean expeditions with a summary of the West Indian shallow-water species (Crustacea: Decapoda: Natantia). Smithsonian Contributions to Zoology, 98, 1 - 179. http: // dx. doi. org / 10.5479 / si. 00810282.98" type="journal article" year="1972">Chace 1972</bibRefCitation>
; 
<bibRefCitation id="EC78D6A5FFB8FFE60470F972FC8E61FE" author="Wicksten" box="[628,800,1735,1760]" pageId="5" pageNumber="90" refString="Wicksten, M. K. (1992) A new species of Alpheopsis (Decapoda, Alpheidae) from the tropical eastern Pacific, with a key to the species of Alpheopsis of the Americas. Crustaceana, 63, 51 - 56. http: // dx. doi. org / 10.1163 / 156854092 X 00271" type="journal article" year="1992">Wicksten 1992</bibRefCitation>
).
</paragraph>
<paragraph id="8856AB54FFB8FFE506C3F958FBAC6603" blockId="5.[151,1437,151,2013]" lastBlockId="6.[151,1437,151,285]" lastPageId="6" lastPageNumber="91" pageId="5" pageNumber="90">
To the author’s best knowledge, 
<taxonomicName id="4FE9D0D7FFB8FFE60432F959FC90601A" ID-CoL="85LCJ" box="[566,830,1772,1796]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="balaeniceps" status="sp. nov.">
<emphasis id="BA9D7746FFB8FFE60432F959FC90601A" box="[566,830,1772,1796]" italics="true" pageId="5" pageNumber="90">Alpheopsis balaeniceps</emphasis>
</taxonomicName>
<emphasis id="BA9D7746FFB8FFE60541F959FC31601B" bold="true" box="[837,927,1772,1797]" pageId="5" pageNumber="90">
<taxonomicNameLabel id="A1AECA3DFFB8FFE60541F959FC31601B" box="[837,927,1772,1797]" pageId="5" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
represents the first species with balaenicepstype setae on the cheliped fingers outside the genus 
<taxonomicName id="4FE9D0D7FFB8FFE604F4F8A5FCE46036" ID-CoL="84LB5" box="[752,842,1808,1832]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="BA9D7746FFB8FFE604F4F8A5FCE46036" box="[752,842,1808,1832]" italics="true" pageId="5" pageNumber="90">Alpheus</emphasis>
</taxonomicName>
. In 
<taxonomicName id="4FE9D0D7FFB8FFE6057FF8A5FC7B6036" ID-CoL="84LB5" box="[891,981,1808,1832]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="BA9D7746FFB8FFE6057FF8A5FC7B6036" box="[891,981,1808,1832]" italics="true" pageId="5" pageNumber="90">Alpheus</emphasis>
</taxonomicName>
, the 
<taxonomicName id="4FE9D0D7FFB8FFE60215F8BAFB3A6036" ID-CoL="85LCJ" box="[1041,1172,1807,1832]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="balaeniceps">balaeniceps</taxonomicName>
setae are present, with various degrees of development, on the minor cheliped of numerous species belonging to several species groups. The 
<taxonomicName id="4FE9D0D7FFB8FFE606CAF8E2FEFF606E" ID-CoL="85LCJ" box="[206,337,1879,1904]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="balaeniceps">balaeniceps</taxonomicName>
condition is typically present only in males or, more rarely, in both sexes, and may help to distinguish closely related species, suggesting that this character is highly flexible, adding to the general plasticity of the alpheid chelipeds. As explained by Banner and Banner (1982), in the most extreme form, the dactylus of the minor chela is conspicously broadened and somewhat excavate, whereas in the least developed form, the dactylus shows lesser degrees of lateral expansion and only bears a low crest befringed with setae. This latter condition is found in 
<taxonomicName id="4FE9D0D7FFBBFFE506F9FF09FE0D67CA" ID-CoL="85LCJ" box="[253,419,188,212]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="balaeniceps" status="sp. nov.">
<emphasis id="BA9D7746FFBBFFE506F9FF09FE0D67CA" box="[253,419,188,212]" italics="true" pageId="6" pageNumber="91">A. balaeniceps</emphasis>
</taxonomicName>
<emphasis id="BA9D7746FFBBFFE507AFFF09FDAB67CB" bold="true" box="[427,517,188,213]" pageId="6" pageNumber="91">
<taxonomicNameLabel id="A1AECA3DFFBBFFE507AFFF09FDAB67CB" box="[427,517,188,213]" pageId="6" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, however, and in contrast to 
<taxonomicName id="4FE9D0D7FFBBFFE5054AFF09FC0667CA" ID-CoL="84LB5" box="[846,936,188,212]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="91" phylum="Arthropoda" rank="genus">
<emphasis id="BA9D7746FFBBFFE5054AFF09FC0667CA" box="[846,936,188,212]" italics="true" pageId="6" pageNumber="91">Alpheus</emphasis>
</taxonomicName>
, on the dactylus of both chelipeds. Whether the female chelipeds of the new species also have 
<taxonomicName id="4FE9D0D7FFBBFFE504E7FF6AFCC867E6" ID-CoL="85LCJ" box="[739,870,223,248]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="balaeniceps">balaeniceps</taxonomicName>
setae is presently unknown. Curiously, the exact function of the 
<taxonomicName id="4FE9D0D7FFBBFFE50747FEB1FE686603" ID-CoL="85LCJ" box="[323,454,260,285]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="balaeniceps">balaeniceps</taxonomicName>
minor chela in alpheid shrimps remains a mystery.
</paragraph>
</subSubSection>
</treatment>
</document>