142 lines
13 KiB
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142 lines
13 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.271.4062" ID-GBIF-Dataset="c93c3a93-d2e1-4c85-8ece-bf3aa1a5109d" ID-PMC="PMC3652427" ID-Pensoft-Pub="1313-2970-271-1" ID-PubMed="23717185" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-271-1" ModsDocOrigin="ZooKeys 271" ModsDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" checkinTime="1451247655320" checkinUser="pensoft" docAuthor="Caterino, Michael S. & Tishechkin, Alexey K." docDate="2013" docId="A4E3348149EA24CF2C2F5EB00682F8D5" docLanguage="en" docName="ZooKeys 271: 1-401" docOrigin="ZooKeys 271" docSource="http://dx.doi.org/10.3897/zookeys.271.4062" docTitle="Operclipygus ashei Caterino & Tishechkin, 2013, sp. n." docType="treatment" docVersion="4" lastPageNumber="315" masterDocId="4403FF80FFE7FF95FFDA5D0E29259B2F" masterDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" masterLastPageNumber="401" masterPageNumber="1" pageNumber="314" updateTime="1668155382655" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Caterino, Michael S.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tishechkin, Alexey K.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>271</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>401</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.271.4062</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.271.4062</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-271-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152040887" LSID="urn:lsid:zoobank.org:act:77891E8F-F464-495F-AFB1-9510FBCA8E93" httpUri="http://treatment.plazi.org/id/A4E3348149EA24CF2C2F5EB00682F8D5" lastPageId="314" lastPageNumber="315" pageId="313" pageNumber="314">
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<subSubSection pageId="313" pageNumber="314" type="nomenclature">
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<paragraph pageId="313" pageNumber="314">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:77891E8F-F464-495F-AFB1-9510FBCA8E93" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus ashei" order="Coleoptera" pageId="313" pageNumber="314" phylum="Arthropoda" rank="species" species="ashei">Operclipygus ashei</taxonomicName>
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<taxonomicNameLabel pageId="313" pageNumber="314">sp. n.</taxonomicNameLabel>
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Figs 83G84GMap 30
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</paragraph>
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</subSubSection>
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<subSubSection pageId="313" pageNumber="314" type="type locality">
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<paragraph pageId="313" pageNumber="314">Type locality.</paragraph>
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<paragraph pageId="313" pageNumber="314">
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COSTA RICA: Puntarenas: Las Cruces Biological Station [
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="8.783334">8°47'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-82.95">82°57'W</geoCoordinate>
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].
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</paragraph>
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</subSubSection>
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<subSubSection pageId="313" pageNumber="314" type="type material">
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<paragraph pageId="313" pageNumber="314">Type material.</paragraph>
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<paragraph pageId="313" pageNumber="314">
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Holotype male: "COSTA RICA: Puntarenas Prov. Las Cruces Biol. Sta. 1330m,
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<geoCoordinate direction="north" orientation="latitude" precision="9" value="8.785666">08°47.14'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="9" value="-82.95966">82°57.58'W</geoCoordinate>
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, 28-31-V-2004. J.S. Ashe, Z.Falin, I.Hinojosa. Ex: flight intercept trap. CR1AFH04 060"/ "SM0621562 KUNHM-ENT" (INBIO). Paratype (1): COSTA RICA: Puntarenas: Coto Brus, Est. Biol. Las Cruces,
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="8.283334">8°17'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-82.95">82°57'W</geoCoordinate>
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, 1100m, 8-20.vi.2005, FIT, M. Ferro (LSAM).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="314" lastPageNumber="315" pageId="313" pageNumber="314" type="description">
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<paragraph pageId="313" pageNumber="314">Diagnostic description.</paragraph>
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<paragraph lastPageId="314" lastPageNumber="315" pageId="313" pageNumber="314">
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This species is very similar to
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus selvorum" order="Coleoptera" pageId="313" pageNumber="314" phylum="Arthropoda" rank="species" species="selvorum">Operclipygus selvorum</taxonomicName>
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, differing mainly in the following characters: length: 2.31-2.50 mm, width: 1.97-2.06 mm; body slightly broader, subquadrate; frons and epistoma depressed at middle; frontal stria carinate at sides, continued onto epistoma as lateral carinae, but also complete across the anterior margin of the frons as a simple stria; all dorsal elytral striae fine but distinct, striae 1-3 complete, 4th barely abbreviated at base, 5th present in apical half,
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<pageBreakToken pageId="314" pageNumber="315" start="start">sutural</pageBreakToken>
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present in apical two-thirds; prosternal lobe larger, more broadly rounded, with fine marginal stria which is obsolete at sides; propygidium with a few secondary punctures along basal margin. Male genitalia: segments 8-10 very similar to those of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus selvorum" order="Coleoptera" pageId="314" pageNumber="315" phylum="Arthropoda" rank="species" species="selvorum">Operclipygus selvorum</taxonomicName>
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, and indistinguishable from that of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus dentatus" order="Coleoptera" pageId="314" pageNumber="315" phylum="Arthropoda" rank="species" species="dentatus">Operclipygus dentatus</taxonomicName>
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(see Figs 83F,
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<normalizedToken originalValue="H–K">H-K</normalizedToken>
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and preceding description); tegmen (Fig. 83G) longer, with slightly more arcuate sides.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="314" pageNumber="315" type="remarks">
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<paragraph pageId="314" pageNumber="315">Remarks.</paragraph>
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<paragraph pageId="314" pageNumber="315">
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This species falls right between the preceding two morphologically, having the strong frontal carina of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus selvorum" order="Coleoptera" pageId="314" pageNumber="315" phylum="Arthropoda" rank="species" species="selvorum">Operclipygus selvorum</taxonomicName>
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(Fig. 84G), but the fine, more or less complete elytral striae of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus dentatus" order="Coleoptera" pageId="314" pageNumber="315" phylum="Arthropoda" rank="species" species="dentatus">Operclipygus dentatus</taxonomicName>
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(see Fig. 84E). The combination of frontal carinae that descend onto the epistoma along with a fine frontal stria on the frons is unique in this species.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="314" pageNumber="315" type="etymology">
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<paragraph pageId="314" pageNumber="315">Etymology.</paragraph>
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<paragraph pageId="314" pageNumber="315">
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We name this species in honor of the late James
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<normalizedToken originalValue="ʽsteve’">ʽsteve'</normalizedToken>
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Ashe (1947-2005), formerly of the University of Kansas, in recognition of his exceptionally productive fieldwork in Central America.
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</paragraph>
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<subSection pageId="314" pageNumber="315" type="operclipygus, incertae sedis">
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<paragraph pageId="314" pageNumber="315">
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus" order="Coleoptera" pageId="314" pageNumber="315" phylum="Arthropoda" rank="genus">Operclipygus</taxonomicName>
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, incertae sedis
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</paragraph>
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<paragraph pageId="314" pageNumber="315">
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The following 27 species are not assigned to any species group. Most are rather generalized, but lack obvious similarities to species in other recognized groups. The majority of these have a detached anterior submarginal pronotal stria, and median pronotal gland openings only slightly displaced posterad the anterior pronotal margin. These characters might appear to ally them with species in the
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus panamensis" order="Coleoptera" pageId="314" pageNumber="315" phylum="Arthropoda" rank="species" species="panamensis">Operclipygus panamensis</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus kerga" order="Coleoptera" pageId="314" pageNumber="315" phylum="Arthropoda" rank="species" species="kerga">Operclipygus kerga</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus sejunctus" order="Coleoptera" pageId="314" pageNumber="315" phylum="Arthropoda" rank="species" species="sejunctus">Operclipygus sejunctus</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus impuncticollis" order="Coleoptera" pageId="314" pageNumber="315" phylum="Arthropoda" rank="species" species="impuncticollis">Operclipygus impuncticollis</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus hospes" order="Coleoptera" pageId="314" pageNumber="315" phylum="Arthropoda" rank="species" species="hospes">Operclipygus hospes</taxonomicName>
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, or
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus dubius" order="Coleoptera" pageId="314" pageNumber="315" phylum="Arthropoda" rank="species" species="dubius">Operclipygus dubius</taxonomicName>
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groups. But this condition is likely to be a symplesiomorphy shared by all of these, and it seems more likely that these incertae sedis species form a grade out of which some of the more distinctive groups have arisen. More detailed phylogenetic analyses focusing on the species of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus" order="Coleoptera" pageId="314" pageNumber="315" phylum="Arthropoda" rank="genus">Operclipygus</taxonomicName>
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should be able to resolve some of these relationships more satisfactorily.
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</paragraph>
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<paragraph pageId="314" pageNumber="315">
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The last two species treated in this section,
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus angustisternus" order="Coleoptera" pageId="314" pageNumber="315" phylum="Arthropoda" rank="species" species="angustisternus">Operclipygus angustisternus</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus shorti" order="Coleoptera" pageId="314" pageNumber="315" phylum="Arthropoda" rank="species" species="shorti">Operclipygus shorti</taxonomicName>
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, do not fit this picture. In fact, based on external characters they might have been assigned to the
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus fossipygus" order="Coleoptera" pageId="314" pageNumber="315" phylum="Arthropoda" rank="species" species="fossipygus">Operclipygus fossipygus</taxonomicName>
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group, as they are both relatively large, convex species with well developed pygidial sulci and strongly displaced median pronotal gland openings. However, both lack the highly specialized and distinctive male genitalia of members of the
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus fossipygus" order="Coleoptera" pageId="314" pageNumber="315" phylum="Arthropoda" rank="species" species="fossipygus">Operclipygus fossipygus</taxonomicName>
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group. It is possible that one or both are its sister group, but that requires much further exploration.
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</paragraph>
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</subSection>
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</subSubSection>
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</treatment>
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</document> |