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<document id="0630D6977FA5AA14B3EF4758F0E1E8DD" ID-CLB-Dataset="28052" ID-DOI="10.11646/zootaxa.4508.2.2" ID-GBIF-Dataset="cf4afb1e-da33-46d0-a3de-3e5092b4de01" ID-ISSN="1175-5326" ID-Zenodo-Dep="2606976" ID-ZooBank="28A35297-48D9-4CEE-97A7-B294C2D271E5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553585755780" checkinUser="plazi" docAuthor="Ho, Hsuan-Ching &amp; Last, Peter R." docDate="2018" docId="9309EE74C5726F41FF10FF18B95673B9" docLanguage="en" docName="zootaxa.4508.2.2.pdf" docOrigin="Zootaxa 4508 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Halicmetus reticulatus Smith &amp; Radcliffe in Radcliffe 1912" docType="treatment" docVersion="9" lastPageNumber="194" masterDocId="6F30960CC57C6F4EFF87FF8FBC5F736C" masterDocTitle="Review of the batfish genus Halicmetus Alcock, 1891 from Australian waters, with descriptions of two new species (Lophiiformes: Ogcocephalidae)" masterLastPageNumber="196" masterPageNumber="179" pageNumber="193" updateTime="1698712994816" updateUser="ExternalLinkService">
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<mods:title id="C2218C45B479B1DD1D1E169452E3FF32">Review of the batfish genus Halicmetus Alcock, 1891 from Australian waters, with descriptions of two new species (Lophiiformes: Ogcocephalidae)</mods:title>
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<mods:namePart id="8569B8BB77CF6B5A91CCE222931EAF03">Ho, Hsuan-Ching</mods:namePart>
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<treatment id="9309EE74C5726F41FF10FF18B95673B9" ID-DOI="http://doi.org/10.5281/zenodo.5296344" ID-GBIF-Taxon="154495746" ID-Zenodo-Dep="5296344" LSID="urn:lsid:plazi:treatment:9309EE74C5726F41FF10FF18B95673B9" httpUri="http://treatment.plazi.org/id/9309EE74C5726F41FF10FF18B95673B9" lastPageId="15" lastPageNumber="194" pageId="14" pageNumber="193">
<subSubSection id="53BA0CE9C5726F40FF10FF18BEF673BF" pageId="14" pageNumber="193" type="nomenclature">
<paragraph id="1B1F5F62C5726F40FF10FF18BE8173DE" blockId="14.[151,734,151,247]" box="[151,734,151,178]" pageId="14" pageNumber="193">
<heading id="4057E80EC5726F40FF10FF18BE8173DE" bold="true" box="[151,734,151,178]" fontSize="11" level="1" pageId="14" pageNumber="193" reason="1">
<taxonomicName id="DCA024E1C5726F40FF10FF18BE8173DE" ID-CoL="6KXK9" authority="Smith &amp; Radcliffe, 1912" authorityName="Smith &amp; Radcliffe in Radcliffe" authorityYear="1912" box="[151,734,151,178]" class="Actinopterygii" family="Ogcocephalidae" genus="Halicmetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lophiiformes" pageId="14" pageNumber="193" phylum="Chordata" rank="species" species="reticulatus">
<emphasis id="29D48370C5726F40FF10FF18BE8173DE" bold="true" box="[151,734,151,178]" pageId="14" pageNumber="193">
<emphasis id="29D48370C5726F40FF10FF18BDF673DD" bold="true" box="[151,425,151,177]" italics="true" pageId="14" pageNumber="193">Halicmetus reticulatus</emphasis>
Smith &amp;
<bibRefCitation id="7F312293C5726F40FDA5FF17BE8173DE" author="Radcliffe, L." box="[546,734,152,178]" pageId="14" pageNumber="193" pagination="199 - 214" refId="ref11499" refString="Radcliffe, L. (1912) New pediculate fishes from the Philippine Islands and contiguous waters. [Scientific results of the Philippine cruise of the Fisheries steamer &quot; Albatross, &quot; 1907 - 1910. No. 16]. Proceedings of the United States National Museum, 42 (1896), 199 - 214, pls. 16 - 27." type="journal article" year="1912">Radcliffe, 1912</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="1B1F5F62C5726F40FF10FF35BEF673BF" blockId="14.[151,734,151,247]" box="[151,681,186,211]" pageId="14" pageNumber="193">Vernacular name: Reticulated Shortnose Seabst</paragraph>
</subSubSection>
<subSubSection id="53BA0CE9C5726F40FF10FF51BD8E739B" box="[151,465,222,247]" pageId="14" pageNumber="193" type="description">
<paragraph id="1B1F5F62C5726F40FF10FF51BD8E739B" blockId="14.[151,734,151,247]" box="[151,465,222,247]" pageId="14" pageNumber="193">
<figureCitation id="839B43E7C5726F40FF10FF51BD4C739B" box="[151,275,222,247]" captionStart="FIGURE 2" captionStartId="7.[151,250,1782,1804]" captionTargetBox="[150,1436,481,1761]" captionTargetId="figure@7.[150,1436,481,1761]" captionTargetPageId="7" captionText="FIGURE 2. Front view of five Halicmetus species. A. H. marmoratus, CSIRO H 954-03, 57.8 mm SL. B. Halicmetus westraliensis sp. nov., CSIRO CA3647, holotype. C. H. drypus sp. nov., holotype. D. H. ruber, WAM P.31798-003, 65.9 mm SL. E. H. reticulatus, CSIRO H 4070-48, 74.7 mm. All preserved specimens, not to scale." httpUri="https://zenodo.org/record/2606982/files/figure.png" pageId="14" pageNumber="193">Figures 2E</figureCitation>
,
<figureCitation id="839B43E7C5726F40FE98FF50BD70739A" box="[287,303,223,246]" captionStart="FIGURE 7" captionStartId="14.[151,250,1723,1745]" captionTargetBox="[156,1431,849,1648]" captionTargetId="figure@14.[151,1436,793,1702]" captionTargetPageId="14" captionText="FIGURE 7. Specimen collection sites of Halicmetus ruber (red triangles), Halicmetus reticulatus (green diamond), and Halicmetus niger (blue circles). Specimens from Australia only." httpUri="https://zenodo.org/record/2606992/files/figure.png" pageId="14" pageNumber="193">7</figureCitation>
,
<figureCitation id="839B43E7C5726F40FEBCFF50BD14739B" box="[315,331,223,247]" captionStart="FIGURE 8" captionStartId="16.[151,250,993,1015]" captionTargetBox="[150,1436,194,972]" captionTargetId="figure@16.[150,1436,194,972]" captionTargetPageId="16" captionText="FIGURE 8. Halicmetus reticulatus Smith and Radcliffe, 1912. CSIRO H 4070-48, 74.7 mm SL." httpUri="https://zenodo.org/record/2606994/files/figure.png" pageId="14" pageNumber="193">8</figureCitation>
;
<tableCitation id="56226AD9C5726F40FEDFFF51BD8E739B" box="[344,465,222,247]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStartId-0="4.[147,231,649,671]" captionStartId-1="8.[151,239,908,931]" captionText-0="TABLE 1. Morphometric values of Halicmetus species studied in present work." captionText-1="TABLE 2. Distribution of dorsal-fin and pectoral-fin ray counts for five Australian species of Halicmetus. Note that H. ruber with 3–4 dorsal-fin rays are juveniles." pageId="14" pageNumber="193">Tables 1–2</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="53BA0CE9C5726F40FF10FEA9B8B67237" pageId="14" pageNumber="193" type="reference_group">
<paragraph id="1B1F5F62C5726F40FF10FEA9B8B67237" blockId="14.[151,1436,294,347]" pageId="14" pageNumber="193">
<treatmentCitationGroup id="3BB0784CC5726F40FF10FEA9B8B67237" pageId="14" pageNumber="193">
<taxonomicName id="DCA024E1C5726F40FF10FEA9BF667250" ID-CoL="6KXK9" authority="Smith &amp; Radcliffe in Radcliffe, 1912: 208" authorityName="Smith &amp; Radcliffe in Radcliffe" authorityPageNumber="208" authorityYear="1912" box="[151,825,294,316]" class="Actinopterygii" family="Ogcocephalidae" genus="Halicmetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lophiiformes" pageId="14" pageNumber="193" phylum="Chordata" rank="species" species="reticulatus">
<emphasis id="29D48370C5726F40FF10FEA9BD207250" box="[151,383,294,316]" italics="true" pageId="14" pageNumber="193">Halicmetus reticulatus</emphasis>
Smith &amp; Radcliffe in
<treatmentCitation id="9A017973C5726F40FDEDFEA9BF667250" author="Radcliffe, L." box="[618,825,294,316]" page="208" pageId="14" pageNumber="193" year="1912">
<bibRefCitation id="7F312293C5726F40FDEDFEA9BF527250" author="Radcliffe, L." box="[618,781,294,316]" pageId="14" pageNumber="193" pagination="199 - 214" refId="ref11499" refString="Radcliffe, L. (1912) New pediculate fishes from the Philippine Islands and contiguous waters. [Scientific results of the Philippine cruise of the Fisheries steamer &quot; Albatross, &quot; 1907 - 1910. No. 16]. Proceedings of the United States National Museum, 42 (1896), 199 - 214, pls. 16 - 27." type="journal article" year="1912">Radcliffe, 1912</bibRefCitation>
:208
</treatmentCitation>
</taxonomicName>
, pls. 20, fig. 2, 21 fig. (
<typeStatus id="C41BE1C0C5726F40FBB3FEA9B83E7250" box="[1076,1121,294,316]" pageId="14" pageNumber="193">type</typeStatus>
locality: off Sombrero Island, southern Luzon Island,
<collectingCountry id="63B71FF2C5726F40FE31FECABE747237" box="[438,555,325,347]" name="Philippines" pageId="14" pageNumber="193">Philippines</collectingCountry>
,
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).
<treatmentCitation id="9A017973C5726F40FD0EFECABF087237" author="Bradbury, M. G." box="[649,855,325,347]" page="415" pageId="14" pageNumber="193" year="1967">
<bibRefCitation id="7F312293C5726F40FD0EFECABF757237" author="Bradbury, M. G." box="[649,810,325,347]" pageId="14" pageNumber="193" pagination="399 - 422" refId="ref10862" refString="Bradbury, M. G. (1967) The genera of batfishes. Copeia, 1967 (2), 399 - 422. https: // doi. org / 10.2307 / 1442130" type="journal article" year="1967">Bradbury, 1967</bibRefCitation>
:415
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.
<treatmentCitation id="9A017973C5726F40FCE7FECAB84B7237" author="Bradbury, M. G." box="[864,1044,325,347]" page="5" pageId="14" pageNumber="193" year="2003">
<bibRefCitation id="7F312293C5726F40FCE7FECAB85A7237" author="Bradbury, M. G." box="[864,1029,325,347]" pageId="14" pageNumber="193" pagination="1 - 17" refId="ref10949" refString="Bradbury, M. G. (2003) Family Ogcocephalidae Jordan 1895 - batfishes. California Academy of Sciences Annotated Checklists of Fishes, 17, 1 - 17." type="journal article" year="2003">Bradbury, 2003</bibRefCitation>
:5
</treatmentCitation>
.
<treatmentCitation id="9A017973C5726F40FB99FEC9B8BA7237" author="Ho, H. - C. &amp; Endo, H. &amp; Sakamaki, K." box="[1054,1253,325,347]" page="772" pageId="14" pageNumber="193" year="2008">
<bibRefCitation id="7F312293C5726F40FB99FEC9B8E67237" author="Ho, H. - C. &amp; Endo, H. &amp; Sakamaki, K." box="[1054,1209,325,347]" pageId="14" pageNumber="193" pagination="767 - 773" refId="ref11109" refString="Ho, H. - C., Endo, H. &amp; Sakamaki, K. (2008) A new species of Halicmetus (Lophiiformes: Ogcocephalidae) from the western Pacific, with comments on congeners. Zoological Studies, 47 (6), 767 - 773." type="journal article" year="2008">
Ho
<emphasis id="29D48370C5726F40FBC4FEC9B8297237" box="[1091,1142,325,347]" italics="true" pageId="14" pageNumber="193">et al.</emphasis>
, 2008
</bibRefCitation>
:772
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="53BA0CE9C5726F40FF10FE08BC8972A9" pageId="14" pageNumber="193" type="materials_examined">
<paragraph id="1B1F5F62C5726F40FF10FE08BC8972A9" blockId="14.[151,1436,391,741]" pageId="14" pageNumber="193">
<emphasis id="29D48370C5726F40FF10FE08BDDD72CC" bold="true" box="[151,386,391,416]" pageId="14" pageNumber="193">Material examined.</emphasis>
<materialsCitation id="ABC8553FC5726F40FE0CFE07BC8D72A9" ID-GBIF-Occurrence="2028138788" collectingDate="1995-09-08" collectionCode="CSIRO" elevation="268" latitude="-18.2" location="Port Hedland" longLatPrecision="1276" longitude="118.23333" pageId="14" pageNumber="193" specimenCode="H 4070-48" specimenCount="1">
<collectionCode id="7DB1C7A7C5726F40FE0CFE07BD8172CC" box="[395,478,392,416]" country="Australia" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="14" pageNumber="193" type="Museum">CSIRO</collectionCode>
<specimenCode id="4B06F719C5726F40FE60FE07BE3D72CC" box="[487,610,392,416]" collectionCode="CSIRO" country="Australia" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="14" pageNumber="193" type="Museum">H 4070-48</specimenCode>
(1, 74.7),
<geoCoordinate id="7E9439A5C5726F40FD5AFE07BF6572CC" box="[733,826,392,416]" direction="south" orientation="latitude" pageId="14" pageNumber="193" precision="925" value="-18.2">18°12’S</geoCoordinate>
,
<geoCoordinate id="7E9439A5C5726F40FCCEFE07BFE972CC" box="[841,950,392,416]" direction="east" orientation="longitude" pageId="14" pageNumber="193" precision="925" value="118.23333">118°14’E</geoCoordinate>
, NW of
<location id="1E7F09B9C5726F40FB9BFE07B8E972CC" LSID="urn:lsid:plazi:treatment:9309EE74C5726F41FF10FF18B95673B9:1E7F09B9C5726F40FB9BFE07B8E972CC" box="[1052,1206,391,416]" latitude="-18.2" longLatPrecision="1276" longitude="118.23333" name="Port Hedland" pageId="14" pageNumber="193">Port Hedland</location>
,
<quantity id="DC58F287C5726F40FB44FE08B91972F3" box="[1219,1350,391,416]" metricMagnitude="2" metricUnit="m" metricValue="2.675" metricValueMax="2.69" metricValueMin="2.66" pageId="14" pageNumber="193" unit="m" value="267.5" valueMax="269.0" valueMin="266.0">
<elevation id="908DB851C5726F40FB44FE08B91972F3" box="[1219,1350,391,416]" metricMagnitude="2" metricUnit="m" metricValue="2.675" metricValueMax="2.69" metricValueMin="2.66" pageId="14" pageNumber="193" unit="m" value="267.5" valueMax="269.0" valueMin="266.0">266–269 m</elevation>
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,
<date id="6F1E79A2C5726F40FAD3FE07BC8D72A9" pageId="14" pageNumber="193" value="1995-09-08">
<collectingDate id="7F5A804AC5726F40FAD3FE07BC8D72A9" pageId="14" pageNumber="193" value="1995-09-08">8 Sep. 1995</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="53BA0CE9C5726F40FF40FE40BD6B7189" pageId="14" pageNumber="193" type="diagnosis">
<paragraph id="1B1F5F62C5726F40FF40FE40BD6B7189" blockId="14.[151,1436,391,741]" pageId="14" pageNumber="193">
<emphasis id="29D48370C5726F40FF40FE40BD1E7284" bold="true" box="[199,321,463,488]" pageId="14" pageNumber="193">Diagnosis.</emphasis>
A species of
<taxonomicName id="DCA024E1C5726F40FE5FFE40BE087284" authorityName="Alcock" authorityYear="1891" box="[472,599,463,488]" class="Actinopterygii" family="Ogcocephalidae" genus="Halicmetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lophiiformes" pageId="14" pageNumber="193" phylum="Chordata" rank="genus">
<emphasis id="29D48370C5726F40FE5FFE40BE087284" box="[472,599,463,488]" italics="true" pageId="14" pageNumber="193">Halicmetus</emphasis>
</taxonomicName>
distinguished by the following combination of characters: disk moderately broad, width ~68% SL; eye small, orbit diameter 7.5–8.3% SL; interorbit broad, 6.0–8.1% SL; illicial trough opening high; mid-dorsal disk covered with very short, simple and multicuspid spinules and dense coverage of minute, spiny tubercles (barely visible without magnification); belly covered with very short, widely spaced, simple and multifid spinules and very short multicuspid tubercles; dorsal-fin rays usually present; pectoral-fin rays 12–14 (modally 12); dorsal surface pale yellowish pink, overlain with dense greenish grey reticulations, pattern persistent when preserved; peritoneal membrane pale with sparse melanophores; distal parts of pectoral, pelvic and anal fins pale.
</paragraph>
</subSubSection>
<caption id="4FDF0FEAC5726F40FF10F934BF40759C" httpUri="https://zenodo.org/record/2606992/files/figure.png" pageId="14" pageNumber="193" startId="14.[151,250,1723,1745]" targetBox="[156,1431,849,1648]" targetPageId="14">
<paragraph id="1B1F5F62C5726F40FF10F934BF40759C" blockId="14.[151,1435,1723,1776]" pageId="14" pageNumber="193">
<emphasis id="29D48370C5726F40FF10F934BD4775BC" bold="true" box="[151,280,1723,1745]" pageId="14" pageNumber="193">FIGURE 7.</emphasis>
Specimen collection sites of
<taxonomicName id="DCA024E1C5726F40FDDCF934BF4E75BD" authorityName="Alcock" authorityYear="1891" box="[603,785,1723,1745]" class="Actinopterygii" family="Ogcocephalidae" genus="Halicmetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lophiiformes" pageId="14" pageNumber="193" phylum="Chordata" rank="species" species="ruber">
<emphasis id="29D48370C5726F40FDDCF934BF4E75BD" box="[603,785,1723,1745]" italics="true" pageId="14" pageNumber="193">Halicmetus ruber</emphasis>
</taxonomicName>
(red triangles),
<taxonomicName id="DCA024E1C5726F40FC44F934B8F275BD" authorityName="Smith &amp; Radcliffe in Radcliffe" authorityYear="1912" box="[963,1197,1723,1745]" class="Actinopterygii" family="Ogcocephalidae" genus="Halicmetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lophiiformes" pageId="14" pageNumber="193" phylum="Chordata" rank="species" species="reticulatus">
<emphasis id="29D48370C5726F40FC44F934B8F275BD" box="[963,1197,1723,1745]" italics="true" pageId="14" pageNumber="193">Halicmetus reticulatus</emphasis>
</taxonomicName>
(green diamond), and
<taxonomicName id="DCA024E1C5726F40FF10F955BD1A759C" authorityName="Ho, Endo &amp; Sakamaki" authorityYear="2008" box="[151,325,1754,1776]" class="Actinopterygii" family="Ogcocephalidae" genus="Halicmetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lophiiformes" pageId="14" pageNumber="193" phylum="Chordata" rank="species" species="niger">
<emphasis id="29D48370C5726F40FF10F955BD1A759C" box="[151,325,1754,1776]" italics="true" pageId="14" pageNumber="193">Halicmetus niger</emphasis>
</taxonomicName>
(blue circles). Specimens from Australia only.
</paragraph>
</caption>
<subSubSection id="53BA0CE9C5726F41FF40F893B95673B9" lastPageId="15" lastPageNumber="194" pageId="14" pageNumber="193" type="discussion">
<paragraph id="1B1F5F62C5726F40FF40F893BF9F7485" blockId="14.[151,1436,1820,2025]" pageId="14" pageNumber="193">
<emphasis id="29D48370C5726F40FF40F893BD647459" bold="true" box="[199,315,1820,1845]" pageId="14" pageNumber="193">Remarks.</emphasis>
Australian material and the diagnosis given above is consistent with the description of this species provided by
<bibRefCitation id="7F312293C5726F40FEA3F8CEBD877435" author="Ho, H. - C. &amp; Endo, H. &amp; Sakamaki, K." box="[292,472,1856,1881]" pageId="14" pageNumber="193" pagination="767 - 773" refId="ref11109" refString="Ho, H. - C., Endo, H. &amp; Sakamaki, K. (2008) A new species of Halicmetus (Lophiiformes: Ogcocephalidae) from the western Pacific, with comments on congeners. Zoological Studies, 47 (6), 767 - 773." type="journal article" year="2008">
Ho
<emphasis id="29D48370C5726F40FECBF8CDBDDB7435" box="[332,388,1856,1881]" italics="true" pageId="14" pageNumber="193">et al.</emphasis>
(2008)
</bibRefCitation>
. The CSIRO specimen is the only individual known from the Australian region. When fresh it had a white reticulate colour pattern, small eye and pale peritoneal membrane with scattered pepper dots, typical of populations of
<taxonomicName id="DCA024E1C5726F40FE3BF807BEE374CD" authorityName="Smith &amp; Radcliffe in Radcliffe" authorityYear="1912" box="[444,700,1928,1953]" class="Actinopterygii" family="Ogcocephalidae" genus="Halicmetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lophiiformes" pageId="14" pageNumber="193" phylum="Chordata" rank="species" species="reticulatus">
<emphasis id="29D48370C5726F40FE3BF807BEE374CD" box="[444,700,1928,1953]" italics="true" pageId="14" pageNumber="193">Halicmetus reticulatus</emphasis>
</taxonomicName>
from other parts of the Pacific. However, it is notable that this specimen has many more bifurcated tubercles on the dorsal surface and these are slightly different in shape to those of western North Pacific populations, which have only simple tubercles.
</paragraph>
<paragraph id="1B1F5F62C5736F41FF40FF17B95673B9" blockId="15.[151,1436,151,213]" pageId="15" pageNumber="194">
Paxton
<emphasis id="29D48370C5736F41FE9BFF16BD1273DC" box="[284,333,151,176]" italics="true" pageId="15" pageNumber="194">et al</emphasis>
. (2006) recorded this species from off southern Queensland to southern New South
<collectingCountry id="63B71FF2C5736F41FA81FF18B91273DC" box="[1286,1357,151,176]" name="United Kingdom" pageId="15" pageNumber="194">Wales</collectingCountry>
. These records are most likely misidentifications of
<taxonomicName id="DCA024E1C5736F41FD0BFF32BF6873B8" authorityName="Weber" authorityYear="1913" box="[652,823,189,212]" class="Actinopterygii" family="Ogcocephalidae" genus="Halicmetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lophiiformes" pageId="15" pageNumber="194" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="29D48370C5736F41FD0BFF32BF6873B8" box="[652,823,189,212]" italics="true" pageId="15" pageNumber="194">H. marmoratus</emphasis>
</taxonomicName>
based on the distribution range provided.
</paragraph>
</subSubSection>
</treatment>
</document>
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