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<document id="1F2BA9EE673EA313A34E4B4D54CB37F5" ID-CLB-Dataset="20497" ID-DOI="10.5169/seals-787048" ID-GBIF-Dataset="8c468ca9-0ee3-4200-b5a3-6adfd98caaea" ID-ISSN="2624-9170" ID-Zenodo-Dep="6284316" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637935242462" checkinUser="carolina" docAuthor="Balkenohl, Michael" docDate="2017" docId="2559A84DA87DFFC4D6FFFFC68F9EFB2B" docLanguage="en" docName="ContribNatHist.35.1-66.pdf.imf" docOrigin="Contributions to Natural History 35" docStyle="DocumentStyle:D319E9290ED630111D2A11900DAFC11F.4:ContribNatHist.2011-.journal_article" docStyleId="D319E9290ED630111D2A11900DAFC11F" docStyleName="ContribNatHist.2011-.journal_article" docStyleVersion="4" docTitle="Orictites barclayi Balkenohl, 2017, sp. nov." docType="treatment" docVersion="12" lastPageNumber="49" masterDocId="D960D035A853FFF4D660FF9B8D45FFA1" masterDocTitle="Revision of the genus Orictites ANDREWES, 1931 (Coleoptera, Carabidae, Clivinini)" masterLastPageNumber="68" masterPageNumber="1" pageNumber="47" updateTime="1699470517822" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="3B70C8F7D9840B6EFFA984597B2EFE52">Revision of the genus Orictites ANDREWES, 1931 (Coleoptera, Carabidae, Clivinini)</mods:title>
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<mods:namePart id="DE6C1B2431012A57D52F20EE38B17FCD">Balkenohl, Michael</mods:namePart>
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<mods:title id="27F26957FF2192D5C8D5ABF481AECEA2">Contributions to Natural History</mods:title>
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<mods:date id="65C5CB91D85DAC7B0B3DD53999155C47">2017</mods:date>
<mods:detail id="4C4E9F668C4B35142D19EC7A5DB50E4A" type="pubDate">
<mods:number id="25162B13AE6024C4473DB209A954E16A">2017-07-17</mods:number>
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<mods:number id="9D711FC722D38F2E6595BBDED8B370BA">35</mods:number>
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<mods:classification id="DE0E7BE51456CE75023013AA307D7EE9">journal article</mods:classification>
<mods:identifier id="13E0F9078223A5E5ADE52792C817987D" type="CLB-Dataset">20497</mods:identifier>
<mods:identifier id="269A71619A8C3C2D53E5C67AFD52878B" type="DOI">10.5169/seals-787048</mods:identifier>
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<mods:identifier id="7A56CF51E6D5111E95441757AC28BD0B" type="ISSN">2624-9170</mods:identifier>
<mods:identifier id="DB7C54F068B62CF3BE6DF51933AF8D2D" type="Zenodo-Dep">6284316</mods:identifier>
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<treatment id="2559A84DA87DFFC4D6FFFFC68F9EFB2B" ID-DOI="http://doi.org/10.5281/zenodo.6315585" ID-GBIF-Taxon="193586552" ID-Zenodo-Dep="6315585" LSID="urn:lsid:plazi:treatment:2559A84DA87DFFC4D6FFFFC68F9EFB2B" httpUri="http://treatment.plazi.org/id/2559A84DA87DFFC4D6FFFFC68F9EFB2B" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47">
<subSubSection id="E5EA4AD0A87DFFDAD6FFFFC68CFAFFD4" box="[159,447,93,118]" pageId="46" pageNumber="47" type="nomenclature">
<paragraph id="AD4F195BA87DFFDAD6FFFFC68CFAFFD4" blockId="46.[159,447,93,118]" box="[159,447,93,118]" pageId="46" pageNumber="47">
<heading id="F607AE37A87DFFDAD6FFFFC68CFAFFD4" box="[159,447,93,118]" centered="true" fontSize="12" level="2" pageId="46" pageNumber="47" reason="2">
<taxonomicName id="6AF062D8A87DFFDAD6FFFFC68C24FFD7" ID-CoL="8MG4G" authorityName="Balkenohl" authorityYear="2017" box="[159,353,93,118]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="barclayi" status="sp. nov.">
<emphasis id="9F84C549A87DFFDAD6FFFFC68C24FFD7" bold="true" box="[159,353,93,118]" italics="true" pageId="46" pageNumber="47">Orictites barclayi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84B77832A87DFFDAD708FFC68CFAFFD4" box="[360,447,93,117]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A87DFFDAD6FFFF338CF9FEDB" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="AD4F195BA87DFFDAD6FFFF338CF9FEDB" blockId="46.[159,1044,168,453]" pageId="46" pageNumber="47">
Type material:
<materialsCitation id="1D981306A87DFFDAD734FF338CF9FEDB" ID-GBIF-Occurrence="3497007333" collectingDate="1921-11-15" collectionCode="BMNH" collectorName="H. M. Pendlebury" location="MALAYA SELANGOR" municipality="Gombak Valley" pageId="46" pageNumber="47" specimenCount="1" specimenCount-female="1" stateProvince="MALAYA SELANGOR" typeStatus="holotype">
<typeStatus id="724BA7F9A87DFFDAD734FF338C87FF61" box="[340,450,168,192]" pageId="46" pageNumber="47" type="holotype">Holotype</typeStatus>
:
<specimenCount id="BBF6D2D2A87DFFDAD7B2FF338CA3FF1E" box="[466,486,168,191]" pageId="46" pageNumber="47" type="female">♀</specimenCount>
, labels: beige, black printed and handwritten in black ink: “
<collectingRegion id="6F34D7B9A87DFFDAD75EFF568F6BFF44" box="[318,558,205,229]" pageId="46" pageNumber="47">MALAYA SELANGOR</collectingRegion>
, F.M.S. Gombak Valley Gd.
<date id="D94E3F9BA87DFFDAD510FF568EB4FF44" box="[880,1009,205,229]" pageId="46" pageNumber="47" value="1921-11-15">
<collectingDate id="C90AC673A87DFFDAD510FF568EB4FF44" box="[880,1009,205,229]" pageId="46" pageNumber="47" value="1921-11-15">11 15 1921</collectingDate>
</date>
” / backside printed: “
<collectorName id="00057C8DA87DFFDAD717FF698F77FEAB" box="[375,562,242,266]" pageId="46" pageNumber="47">H.M.Pendlebury</collectorName>
Coll: F.M.S.Museums.” / beige, handwritten in black ink and printed: “
<taxonomicName id="6AF062D8A87DFFDAD794FE838E96FE91" authority="Bates H. E. Andrewes" box="[500,979,280,304]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="costulipennis">Clivina costulipennis Bates H.E.Andrewes</taxonomicName>
det.” / small, beige, black printed: “346” / white, black printed: “Ex F.M.S.Museum: B.M.1955–354.” (
<collectionCode id="CBE1819EA87DFFDAD701FEF98CF4FEDB" box="[353,433,354,378]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="46" pageNumber="47">BMNH</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A87DFFDAD6FFFE138E5CFE64" pageId="46" pageNumber="47" type="discussion">
<paragraph id="AD4F195BA87DFFDAD6FFFE138E5CFE64" blockId="46.[159,1044,168,453]" pageId="46" pageNumber="47">Remark: The specimen was pinned centrally in between the two elytra, so the elytra are opened. It has been mounted on a paper card.</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A87DFFC4D6FFFD868FDBFCC1" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47" type="description">
<paragraph id="AD4F195BA87DFFDAD6FFFD868F66FD94" blockId="46.[159,547,541,566]" box="[159,547,541,566]" pageId="46" pageNumber="47">
<emphasis id="9F84C549A87DFFDAD6FFFD868C67FD97" bold="true" box="[159,290,541,566]" pageId="46" pageNumber="47">Description</emphasis>
(
<figureCitation id="35CB05DEA87DFFDAD74EFD868C37FD94" box="[302,370,541,565]" captionStart="Figs 9–12" captionStartId="9.[121,158,1494,1512]" captionTargetBox="[145,998,148,1407]" captionTargetId="figure-60@9.[567,998,842,1405]" captionTargetPageId="9" captionText="Figs 9–12: Orictites spp., habitus. 9: O. barclayi sp. nov., holotype, female; 10: O. brancuccii sp. nov., holotype, male; 11: O. barclayi minorfrater ssp. nov., holotype, male; 12: O. tubercucollis sp. nov., holotype, male." figureDoi="http://doi.org/10.5281/zenodo.6284330" httpUri="https://zenodo.org/record/6284330/files/figure.png" pageId="46" pageNumber="47">Figs 9</figureCitation>
,
<figureCitation id="35CB05DEA87DFFDAD71FFD868CE5FD94" box="[383,416,541,565]" captionStart="Figs 35–38" captionStartId="35.[121,158,1306,1324]" captionTargetBox="[121,1005,147,1260]" captionTargetId="figure-64@35.[121,552,811,1130]" captionTargetPageId="35" captionText="Figs 35–38: Orictites spp., coxostylus. 35: O. anteriortenuis sp. nov., scale bar 0.1 mm; 36: O. omnipunctatus sp. nov., scale bar 0.1 mm; 37: O. ampliosetosus sp. nov., scale bar 0.1 mm; 38: O. barclayi sp. nov., holotype, scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6284368" httpUri="https://zenodo.org/record/6284368/files/figure.png" pageId="46" pageNumber="47">38</figureCitation>
; pp. 10, 36)
</paragraph>
<paragraph id="AD4F195BA87DFFDAD6FFFDF38FDBFD04" blockId="46.[159,1044,616,1573]" pageId="46" pageNumber="47">
Measurements: Body length
<quantity id="6A08B4BEA87DFFDAD786FDF38F7AFD21" box="[486,575,616,640]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="46" pageNumber="47" unit="mm" value="4.8">4.8 mm</quantity>
, width 1,4 mm; ratio length/width of pronotum 1.03; ratio length/width of elytra 1.93.
</paragraph>
<paragraph id="AD4F195BA87DFFDAD6A4FD298DABFCB4" blockId="46.[159,1044,616,1573]" pageId="46" pageNumber="47">Colour: Shiny. Black. Wings of clypeus and supraantennal plates slightly transparent brown. Labrum black. Antennae and legs dark-brown, palpi red brown.</paragraph>
<paragraph id="AD4F195BA87DFFDBD6A4FCB98E43FD21" blockId="46.[159,1044,616,1573]" lastBlockId="47.[121,1006,93,1573]" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
Head: A fifth narrower than pronotum. Clypeus with small rounded tooth laterally, anterior margin excised, with blunt raised knob in between the two clypeal setae, anteriorly smooth, posterior part of knob covered with big punctures, margin of wings convex, slightly hollowed out, divided from middle part of clypeus by slight notches and carinae, divided from supraantennal plates by distinct obtusely angled notches and ridge, clypeus and wings reflexed margined, supraantennal plates vaulted up to mid-eye level, smooth, with reflexed rounded margin, elongated as supraorbital carina, frons separated from clypeus by flat transverse depression with punctures, with costate, nearly parallel running frontal carinae at each side posteriorly, with broad furrow between frontal carina and supraorbital carina, sharp furrow between frons and frontal carina, frons moderately convex, smooth. Supraorbital setae located in broad furrows between supraorbital carina and frontal carina, the posterior one arising from tubercle. Eyes subglobose, genae indistinct, enclosing eyes posteriorly, separated from eye by small ridge, forming obtuse angle at neck. Grooves for reception of antennae short, length 0.4 of eye diameter. Neck constriction marked by band of big punctures, neck laterally covered with reticulation and densely scattered medium sized punctures. Labrum straight, 7-setose, ciliate laterally, with isodiametric reticulation. Mandibles robust, shorter than head, flattened, stout, broadened at base, acutely curved at apex, carinae of scrobe complete, both mandibles obtusely angled towards base (opened position), both mandibles somewhat hollowed out dorsally. Maxilla distinctly curved, acutely hooked at apex. Apical maxillary palpomere slender, slightly securiform. Apical labial palpomere slender, straight, slightly longer than 2
<superScript id="5A85B413A87CFFDBD74DFF518C01FF7C" attach="left" box="[301,324,202,221]" fontSize="8" pageId="47" pageNumber="48">nd</superScript>
segment, 2
<superScript id="5A85B413A87CFFDBD7ACFF518CA6FF7C" attach="left" box="[460,483,202,221]" fontSize="8" pageId="47" pageNumber="48">nd</superScript>
segment bisetose. Ligula with one long seta; paraglossae slender. Ventral surface of neck with punctures and reticulation. Submentum and mentum distinctly separated; submentum with four, about equally spaced setigerous punctures; mentum at base bilaterally with rounded elevation, with some longitudinal carinae, with fine and complete reflexed margin, lateral lobes projecting, nearly right-angled at tips, with a seta near base of each lobe, median tooth forming broad obtuse triangle, bisetose, not projecting as far as lateral lobes. Antennae of moderate length, reaching up to middle of pronotum, scapus knee-like angled, with one seta dorsoapically, scapus and pedicellus with fine reticulation, segments 5–10 submoniliform (L/
<geoCoordinate id="C8C47F9CA87CFFDBD6F6FDD98DA5FDFB" box="[150,224,578,602]" degrees="0. 9" direction="west" orientation="longitude" pageId="47" pageNumber="48" precision="5555" value="-0.9">W 0.9</geoCoordinate>
), antennomeres somewhat flattened, with shiny areas, segments 4–11 densely pubescent, with a few additional long setae.
</paragraph>
<paragraph id="AD4F195BA87CFFDBD6FEFD168C51FB11" blockId="47.[121,1006,93,1573]" pageId="47" pageNumber="48">Pronotum: Square, slightly longer than wide, sides slightly convex. Anterior margin straight. Reflexed lateral margin smooth. Margin from posterior angle to base running as straight line. Lateral channel conspicuously broad between the two lateral setigerous punctures, with isodiametric reticulation and very few irregular punctures. Anterior setigerous puncture located at the end of anterior quarter adjoining convexity of pronotum, the posterior one located at level of posterior angle, removed from lateral channel by diameter of pore. Anterior angle indistinctly projecting, formed by reflexed lateral margin, posterior angle developed as sharp distinct tooth, projecting laterally. Anterior transverse line narrow, formed by distinct, partly connected punctures. Median line deep and moderately narrow, with some punctures, surpassing anterior transverse line without joining, fine at base, adjoining base. Surface bilaterally with two irregularly longitudinal groups of punctures with distinct basal impression. Basal channel broad, deep. Flange raised keel-like (lateral view).</paragraph>
<paragraph id="AD4F195BA87CFFC4D6FEFB268F17FF3B" blockId="47.[121,1006,93,1573]" lastBlockId="48.[159,1044,90,1162]" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48">
Elytra: Subcylindrical, sides slightly diverging, marginal channel broad, visible from above, with uninterrupted row of big setigerous punctures arising from broad tubercles, with additional row of punctures laterally. Reflexed margin with some scars in anterior third, margin slightly thickened at humerus, bending with channel over rounded humerus up to 5
<superScript id="5A85B413A87CFFDBD56AFAD48E58FAC3" attach="left" box="[778,797,1359,1378]" fontSize="8" pageId="47" pageNumber="48">th</superScript>
stria. No tooth at humerus. Basal tubercle distinct, with setigerous puncture, situated at declivity of 1
<superScript id="5A85B413A87CFFDBD6A7FA018D9DFA0C" attach="left" box="[199,216,1434,1453]" fontSize="8" pageId="47" pageNumber="48">st</superScript>
stria. Striae 1–4 free at base, all striae deep, distinctly punctate, striae 1–2, 3–4 and 5–6 joined apically; striae 6 and 7 shortened at humerus. Striole distinct, interval between striole and sutura raised. All intervals conspicuously convex, thought costate, 3
<superScript id="5A85B413A87CFFDBD78EF9918F44F9BC" attach="left" box="[494,513,1546,1565]" fontSize="8" pageId="47" pageNumber="48">rd</superScript>
broader and distinctly raised at base, with tubercle at base, 8
<superScript id="5A85B413A863FFC4D714FFC18CC2FFCC" attach="left" box="[372,391,90,109]" fontSize="8" pageId="48" pageNumber="49">th</superScript>
carinate. All intervals shiny on disc. Interval 3 with seven setigerous punctures adjoining stria 3.
</paragraph>
<paragraph id="AD4F195BA863FFC4D6A4FF338F4DFF61" blockId="48.[159,1044,90,1162]" box="[196,520,168,192]" pageId="48" pageNumber="49">Hind wings: Fully developed.</paragraph>
<paragraph id="AD4F195BA863FFC4D6A4FF568FF6FE64" blockId="48.[159,1044,90,1162]" pageId="48" pageNumber="49">
Ventral surface: Epipleuron shiny, covered with big scattered punctures and indistinct reticulation, submarginal furrow distinct. Proepisternum and episternum nearly smooth, prosternite smooth, keeled at middle, mesosternum smooth. Sternites with transverse to isodiametric reticulation, each with big punctures at base, denser laterally, 3
<superScript id="5A85B413A863FFC4D414FEC48FC2FED3" attach="left" box="[628,647,351,370]" fontSize="8" pageId="48" pageNumber="49">rd</superScript>
to 5
<superScript id="5A85B413A863FFC4D4DBFEC48F8BFED3" attach="left" box="[699,718,351,370]" fontSize="8" pageId="48" pageNumber="49">th</superScript>
with paralateral ambulatory setae at each side, ventral strigae distinct, sternite 6 smooth at middle, with two widely separated apical setae at each side.
</paragraph>
<paragraph id="AD4F195BA863FFC4D6A4FE498CFBFD21" blockId="48.[159,1044,90,1162]" pageId="48" pageNumber="49">Legs: Anterior tibia with strong digitation, with two big and one small lateral preapical denticles, apical spine curved distinctly towards ventral, no dorsal furrow, reticulated, basal tarsomere elongated, as long as the following four together. Mesotibia with preapical spine, furnished with strong seta. All tarsomeres relatively broad.</paragraph>
<paragraph id="AD4F195BA863FFC4D6A4FD168CA0FD04" blockId="48.[159,1044,90,1162]" box="[196,485,653,677]" pageId="48" pageNumber="49">Male genitalia: Unknown.</paragraph>
<paragraph id="AD4F195BA863FFC4D6A4FD298F5DFC9B" blockId="48.[159,1044,90,1162]" pageId="48" pageNumber="49">
Female genitalia (
<figureCitation id="35CB05DEA863FFC4D7F9FD298CABFD6B" box="[409,494,690,714]" captionStart="Figs 35–38" captionStartId="35.[121,158,1306,1324]" captionTargetBox="[121,1005,147,1260]" captionTargetId="figure-64@35.[121,552,811,1130]" captionTargetPageId="35" captionText="Figs 35–38: Orictites spp., coxostylus. 35: O. anteriortenuis sp. nov., scale bar 0.1 mm; 36: O. omnipunctatus sp. nov., scale bar 0.1 mm; 37: O. ampliosetosus sp. nov., scale bar 0.1 mm; 38: O. barclayi sp. nov., holotype, scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6284368" httpUri="https://zenodo.org/record/6284368/files/figure.png" pageId="48" pageNumber="49">Fig. 38</figureCitation>
): Coxostylus slender, slightly sigmoid, gently curved to apex, with short obtuse knob-like tooth dorsally at base, with two setae apically tight together, with one short robust ensiform seta, with five nematiforme setae. Ramus short.
</paragraph>
<paragraph id="AD4F195BA863FFC4D6A4FCD38FDBFCC1" blockId="48.[159,1044,90,1162]" box="[196,670,840,864]" pageId="48" pageNumber="49">Variation: No bilateral variation observed.</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A863FFC4D6A4FCF68E4DFBBB" pageId="48" pageNumber="49" type="diagnosis">
<paragraph id="AD4F195BA863FFC4D6A4FCF68E4DFBBB" blockId="48.[159,1044,90,1162]" pageId="48" pageNumber="49">Diagnosis: A medium-sized black species with distinct knob on the clypeus and distinct flat transverse depression behind, no humeral tooth, and a tubercle at the base of interval 3 of the elytra. Distinguished from all other species by the presence of seven setigerous punctures on interval 3, and the moderately narrow median line of the pronotum joining base.</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A863FFC4D6A4FBB38FEAFBE1" box="[196,687,1064,1088]" pageId="48" pageNumber="49" type="distribution">
<paragraph id="AD4F195BA863FFC4D6A4FBB38FEAFBE1" blockId="48.[159,1044,90,1162]" box="[196,687,1064,1088]" pageId="48" pageNumber="49">
Distribution: Known from West of
<collectingCountry id="D5E759CBA863FFC4D422FBB38FEEFBE1" box="[578,683,1064,1088]" name="Malaysia" pageId="48" pageNumber="49">Malaysia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A863FFC4D6A4FBD68F9EFB2B" pageId="48" pageNumber="49" type="etymology">
<paragraph id="AD4F195BA863FFC4D6A4FBD68F9EFB2B" blockId="48.[159,1044,90,1162]" pageId="48" pageNumber="49">Etymology. The name is a patronym of Maxwell V. L. Barclay (BMNH) who supported my work over the last time significantly.</paragraph>
</subSubSection>
</treatment>
</document>
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