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<document id="F3240CBF1AD2839449D0EC563F9FEA8D" ID-CLB-Dataset="7893" ID-DOI="10.11646/zootaxa.4938.3.2" ID-GBIF-Dataset="640aac64-3963-4435-8e04-7af89edd428b" ID-ISSN="1175-5326" ID-Zenodo-Dep="4569653" ID-ZooBank="DF5541CF-E4BE-43DF-BE55-615F4D4CEFBD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614597281385" checkinUser="plazi" docAuthor="Uiblein, Franz &amp; Motomura, Hiroyuki" docDate="2021" docId="039187F5F2592B65FF78FD9FCC6FDB07" docLanguage="en" docName="zootaxa.4938.3.2.pdf" docOrigin="Zootaxa 4938 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Upeneus willwhite Uiblein &amp; Motomura 2021, n. sp." docType="treatment" docUuid="DF1D9A1D-97C4-44E5-8B54-8C621DDAE887" docUuidSource="ZooBank" docVersion="7" lastPageNumber="314" masterDocId="FFA8FF8DF2562B75FFEFFFF4C976DF7A" masterDocTitle="Three new goatfishes of the genus Upeneus from the Eastern Indian Ocean and Western Pacific, with an updated taxonomic account for U. itoui (Mullidae: japonicus-species group)" masterLastPageNumber="324" masterPageNumber="298" pageNumber="313" updateTime="1698931649437" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="ACCDB769FE2A66D36F56250115820E15">Three new goatfishes of the genus Upeneus from the Eastern Indian Ocean and Western Pacific, with an updated taxonomic account for U. itoui (Mullidae: japonicus-species group)</mods:title>
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<mods:namePart id="9357626939E770A1D775C6D27185F4D0">Uiblein, Franz</mods:namePart>
<mods:affiliation id="755E4E5A99CCF3007A6DDA650F34AA80">Institute of Marine Research, P. O. Box 1870 Nordnes, N- 5817 Bergen, Norway &amp; National Research Foundation - South African Institute for Aquatic Biodiversity, Makhanda, South Africa &amp; Natural History Museum, University of Oslo, Norway &amp; Vietnam National Museum of Nature, Vietnam Academy of Science and Technology, Ha Noi, Vietnam</mods:affiliation>
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<mods:namePart id="1ED0FA41688C277AB81E0C2427A13E7C">Motomura, Hiroyuki</mods:namePart>
<mods:affiliation id="DDC8EA7807C71A231ACC6560F92BA1F5">The Kagoshima University Museum, Korimoto, Kagoshima, Japan.</mods:affiliation>
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<mods:title id="C8CA0AD9201DC942E6DA1400677D916E">Zootaxa</mods:title>
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<mods:date id="C70F42F543683D70D2567B69D011ABA9">2021</mods:date>
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<treatment id="039187F5F2592B65FF78FD9FCC6FDB07" ID-DOI="http://doi.org/10.5281/zenodo.4582353" ID-GBIF-Taxon="179989605" ID-Zenodo-Dep="4582353" LSID="urn:lsid:zoobank.org:act:DF1D9A1D-97C4-44E5-8B54-8C621DDAE887" httpUri="http://treatment.plazi.org/id/039187F5F2592B65FF78FD9FCC6FDB07" lastPageId="16" lastPageNumber="314" pageId="15" pageNumber="313">
<subSubSection id="C3226568F2592B7AFF78FD9FC823DDD3" pageId="15" pageNumber="313" type="nomenclature">
<paragraph id="8B8736E3F2592B7AFF78FD9FC8C9DDFF" blockId="15.[151,480,618,717]" box="[151,447,618,645]" pageId="15" pageNumber="313">
<heading id="D0CF818FF2592B7AFF78FD9FC8C9DDFF" bold="true" box="[151,447,618,645]" fontSize="11" level="1" pageId="15" pageNumber="313" reason="1">
<emphasis id="B94CEAF1F2592B7AFF78FD9FC8C9DDFF" bold="true" box="[151,447,618,645]" pageId="15" pageNumber="313">
<taxonomicName id="4C384D60F2592B7AFF78FD9FC803DDFF" authority="Uiblein &amp; Motomura, 2021" authorityName="Uiblein &amp; Motomura" authorityYear="2021" box="[151,373,618,645]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="313" phylum="Chordata" rank="species" species="willwhite" status="sp. nov.">
<emphasis id="B94CEAF1F2592B7AFF78FD9FC803DDFF" bold="true" box="[151,373,618,645]" italics="true" pageId="15" pageNumber="313">Upeneus willwhite</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27F578AF2592B7AFE92FD9FC8C9DDFF" box="[381,447,619,645]" pageId="15" pageNumber="313" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8736E3F2592B7AFF78FD7BC823DDD3" blockId="15.[151,480,618,717]" box="[151,341,655,681]" pageId="15" pageNumber="313">Whites Goatfish</paragraph>
</subSubSection>
<subSubSection id="C3226568F2592B7AFF78FD47CAD4DC75" pageId="15" pageNumber="313" type="description">
<paragraph id="8B8736E3F2592B7AFF78FD47C896DDB7" blockId="15.[151,480,618,717]" box="[151,480,691,717]" pageId="15" pageNumber="313">
(
<figureCitation id="13032A66F2592B7AFF70FD47C86FDDB7" box="[159,281,691,717]" captionStart="FIGURE 1" captionStartId="5.[151,250,1713,1737]" captionTargetBox="[151,1435,381,1689]" captionTargetId="figure-125@5.[151,1436,381,1690]" captionTargetPageId="5" captionText="FIGURE 1. A: Upeneus dimipavlov n. sp., VNMN-I 2056, HT, 120 mm SL, N of Hon Tre Island, Nha Trang, S-central Vietnam (D.A. Pavlov); B, C: Upeneus elongatus n. sp., KAUM-I. 58746, HT, 89 mm SL, Tanega-shima Island, S Japan (H. Motomura); D, E: Upeneus itoui, Kagoshima Prefecture, S Japan; D: KAUM-I. 13591, PT, 104 mm SL; E: KAUM-I. 13595, PT, 118 mm SL; F: U. willwhite n. sp., CSIRO 7217-07, HT, 90 mm SL, Lombok, S Indonesia (W.T. White); GJ: U. cf. elongatus, Central Philippines, Visayas, adults, unvouchered; G: Panay; H: Leyte; I, J: Negros (P. &amp; G. Poppe https://poppe-images.com)." figureDoi="http://doi.org/10.5281/zenodo.4569657" httpUri="https://zenodo.org/record/4569657/files/figure.png" pageId="15" pageNumber="313">Figures 1F</figureCitation>
,
<figureCitation id="13032A66F2592B7AFECAFD47C824DDB7" box="[293,338,691,717]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStartId-0="7.[151,250,1865,1889]" captionStartId-1="8.[151,250,1239,1263]" captionStartId-2="9.[151,250,1939,1963]" captionStartId-3="10.[151,250,1649,1673]" captionStartId-4="11.[151,250,1787,1811]" captionTargetBox-0="[157,1424,422,1835]" captionTargetBox-1="[155,1429,188,1213]" captionTargetBox-2="[159,1429,608,1908]" captionTargetBox-3="[158,1432,193,1612]" captionTargetBox-4="[206,1383,188,1761]" captionTargetId-0="figure-142@7.[151,1436,417,1842]" captionTargetId-1="figure-17@8.[151,1436,181,1215]" captionTargetId-2="figure-242@9.[151,1436,597,1916]" captionTargetId-3="figure-17@10.[151,1436,181,1626]" captionTargetId-4="figure-29@11.[189,1398,181,1764]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="10" captionTargetPageId-4="11" captionText-0="FIGURE 2. Relationship among two meristic and five morphometric characters in the four featured species of Upeneus and U. asymmetricus and U. pori." captionText-1="FIGURE 3. Relationship among four morphometric characters in three featured species of Upeneus and U. guttatus." captionText-2="FIGURE 4. Relationship among six morphometric characters in three featured species of Upeneus and U. australiae." captionText-3="FIGURE 5. Relationship among one meristic and five morphometric characters in three featured species of Upeneus and U. floros." captionText-4="FIGURE 6. Relationship among seven morphometric characters in three featured species of Upeneus and U. heterospinus." figureDoi-0="http://doi.org/10.5281/zenodo.4569659" figureDoi-1="http://doi.org/10.5281/zenodo.4569661" figureDoi-2="http://doi.org/10.5281/zenodo.4569665" figureDoi-3="http://doi.org/10.5281/zenodo.4569667" figureDoi-4="http://doi.org/10.5281/zenodo.4569671" httpUri-0="https://zenodo.org/record/4569659/files/figure.png" httpUri-1="https://zenodo.org/record/4569661/files/figure.png" httpUri-2="https://zenodo.org/record/4569665/files/figure.png" httpUri-3="https://zenodo.org/record/4569667/files/figure.png" httpUri-4="https://zenodo.org/record/4569671/files/figure.png" pageId="15" pageNumber="313">26</figureCitation>
;
<tableCitation id="C6BA0358F2592B7AFEB2FD47C8CBDDB7" box="[349,445,691,717]" captionStart="TABLE 2" captionStartId="3.[150,216,469,488]" captionText="TABLE 2. (A) Number of specimens, SL, meristic and colour characters and (B) morphometric characters as ratios of SL for the four features species, all other japonicus-group species and U. heterospinus (margarethae-species group)." pageId="15" pageNumber="313">Tables 2</tableCitation>
,
<tableCitation id="C6BA0358F2592B7AFE26FD47C8A1DDB7" box="[457,471,691,717]" captionStart="TABLE 3" captionStartId="13.[150,234,479,504]" captionText="TABLE 3. Morphometric and meristic characters for three new species of Upeneus and Upeneus itoui." pageId="15" pageNumber="313">3</tableCitation>
)
</paragraph>
<paragraph id="8B8736E3F2592B7AFF78FD0CCAD4DC75" blockId="15.[151,930,759,784]" box="[151,930,759,784]" pageId="15" pageNumber="313">
<taxonomicName id="4C384D60F2592B7AFF78FD0CC8CFDC75" authority="Day, 1868" authorityName="Day" authorityYear="1868" box="[151,441,759,784]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="313" phylum="Chordata" rank="species" species="guttatus">
<emphasis id="B94CEAF1F2592B7AFF78FD0CC83EDC75" box="[151,328,760,783]" italics="true" pageId="15" pageNumber="313">Upeneus guttatus</emphasis>
Day, 1868
</taxonomicName>
: White
<emphasis id="B94CEAF1F2592B7AFDE8FD0CCB4CDC75" box="[519,570,759,783]" italics="true" pageId="15" pageNumber="313">et al.</emphasis>
2013;
<bibRefCitation id="EFA94B12F2592B7AFD6FFD03CA3CDC75" author="Uiblein, F. &amp; Gledhill, D. C. &amp; Peristiwady T." box="[640,842,759,783]" pageId="15" pageNumber="313" pagination="295 - 311" refId="ref19927" refString="Uiblein, F., Gledhill, D. C. &amp; Peristiwady T. (2017 a) Two new goatfishes of the genus Upeneus (Mullidae) from Australia and Indonesia. Zootaxa, 4318 (2), 295 - 311. https: // doi. org / 10.11646 / zootaxa. 4318.2.4" type="journal article" year="2017">
Uiblein
<emphasis id="B94CEAF1F2592B7AFD3CFD0CCA70DC75" box="[723,774,759,783]" italics="true" pageId="15" pageNumber="313">et al.</emphasis>
2017a
</bibRefCitation>
(in part)
</paragraph>
</subSubSection>
<subSubSection id="C3226568F2592B7AFF78FCC9CAB2DCE5" pageId="15" pageNumber="313" type="materials_examined">
<paragraph id="8B8736E3F2592B7AFF78FCC9CBEFDC01" blockId="15.[151,1437,829,2008]" pageId="15" pageNumber="313">
<emphasis id="B94CEAF1F2592B7AFF78FCC9C80EDC2D" bold="true" box="[151,376,829,855]" pageId="15" pageNumber="313">Material examined</emphasis>
.
<materialsCitation id="3B503CBEF2592B7AFE69FCC9CBEFDC01" ID-GBIF-Occurrence="3045898314" collectionCode="EIO" country="Indonesia" county="William White" location="Lombok" municipality="Tanjung Luar" pageId="15" pageNumber="313" specimenCode="CSIRO 7217-07" specimenCount="1" typeStatus="holotype">
<emphasis id="B94CEAF1F2592B7AFE69FCC9C89BDC2D" box="[390,493,829,855]" italics="true" pageId="15" pageNumber="313">
<typeStatus id="54838841F2592B7AFE69FCC9C89BDC2D" box="[390,493,829,855]" pageId="15" pageNumber="313" type="holotype">Holotype</typeStatus>
</emphasis>
.
<specimenCode id="DB9E9E98F2592B7AFE16FCC9CBC4DC2D" box="[505,690,829,855]" collectionCode="CSIRO" country="Australia" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="15" pageNumber="313" type="Museum">CSIRO 7217-07</specimenCode>
,
<quantity id="4CC09B06F2592B7AFD52FCC9CA7DDC22" box="[701,779,829,856]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="15" pageNumber="313" unit="mm" value="90.0">90 mm</quantity>
SL,
<collectionCode id="ED29AE26F2592B7AFCD1FCC9CA19DC2D" box="[830,879,829,855]" pageId="15" pageNumber="313">EIO</collectionCode>
, S
<collectingCountry id="F32F7673F2592B7AFC7FFCC9CD76DC2D" box="[912,1024,829,855]" name="Indonesia" pageId="15" pageNumber="313">Indonesia</collectingCountry>
,
<location id="8EE76038F2592B7AFBE5FCC9CD1ADC2D" LSID="urn:lsid:plazi:treatment:039187F5F2592B65FF78FD9FCC6FDB07:8EE76038F2592B7AFBE5FCC9CD1ADC2D" box="[1034,1132,829,855]" country="Indonesia" county="William White" municipality="Tanjung Luar" name="Lombok" pageId="15" pageNumber="313">Lombok</location>
,
<collectingMunicipality id="6BE3AC99F2592B7AFB9AFCC9CC78DC22" box="[1141,1294,829,856]" pageId="15" pageNumber="313">Tanjung Luar</collectingMunicipality>
, fish market,
<collectingCounty id="62E64E6FF2592B7AFF78FC95C84ADC01" box="[151,316,865,891]" pageId="15" pageNumber="313">William White</collectingCounty>
(photo of fresh HT,
<figureCitation id="13032A66F2592B7AFDCDFC95CBE6DC01" box="[546,656,865,891]" captionStart="FIGURE 1" captionStartId="5.[151,250,1713,1737]" captionTargetBox="[151,1435,381,1689]" captionTargetId="figure-125@5.[151,1436,381,1690]" captionTargetPageId="5" captionText="FIGURE 1. A: Upeneus dimipavlov n. sp., VNMN-I 2056, HT, 120 mm SL, N of Hon Tre Island, Nha Trang, S-central Vietnam (D.A. Pavlov); B, C: Upeneus elongatus n. sp., KAUM-I. 58746, HT, 89 mm SL, Tanega-shima Island, S Japan (H. Motomura); D, E: Upeneus itoui, Kagoshima Prefecture, S Japan; D: KAUM-I. 13591, PT, 104 mm SL; E: KAUM-I. 13595, PT, 118 mm SL; F: U. willwhite n. sp., CSIRO 7217-07, HT, 90 mm SL, Lombok, S Indonesia (W.T. White); GJ: U. cf. elongatus, Central Philippines, Visayas, adults, unvouchered; G: Panay; H: Leyte; I, J: Negros (P. &amp; G. Poppe https://poppe-images.com)." figureDoi="http://doi.org/10.5281/zenodo.4569657" httpUri="https://zenodo.org/record/4569657/files/figure.png" pageId="15" pageNumber="313">Figure 1F</figureCitation>
)
</materialsCitation>
</paragraph>
<paragraph id="8B8736E3F2592B7AFF28FC72CAB2DCE5" blockId="15.[151,1437,829,2008]" box="[199,964,901,928]" pageId="15" pageNumber="313">
<materialsCitation id="3B503CBEF2592B7AFF28FC72CAB6DCE5" ID-GBIF-Occurrence="3045898321" accessionNumber="MZB 23014" box="[199,960,901,928]" collectionCode="MZB" country="Indonesia" county="William White" location="Lombok" municipality="Tanjung Luar" pageId="15" pageNumber="313" specimenCode="92" specimenCount="1" typeStatus="paratype">
<emphasis id="B94CEAF1F2592B7AFF28FC72C858DCE5" box="[199,302,902,927]" italics="true" pageId="15" pageNumber="313">
<typeStatus id="54838841F2592B7AFF28FC72C858DCE5" box="[199,302,902,927]" pageId="15" pageNumber="313" type="paratype">Paratype</typeStatus>
</emphasis>
.
<collectionCode id="ED29AE26F2592B7AFED3FC72C80FDCDA" box="[316,377,902,928]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="15" pageNumber="313" type="Museum">MZB</collectionCode>
23014,
<specimenCode id="DB9E9E98F2592B7AFE3CFC71C899DCE5" box="[467,495,901,927]" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="15" pageNumber="313" type="Museum">92</specimenCode>
mm SL, same sampling data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3226568F2592B7AFF28FC5DC839DAED" pageId="15" pageNumber="313" type="diagnosis">
<paragraph id="8B8736E3F2592B7AFF28FC5DC839DAED" blockId="15.[151,1437,829,2008]" pageId="15" pageNumber="313">
<emphasis id="B94CEAF1F2592B7AFF28FC5DC834DCB9" bold="true" box="[199,322,937,963]" pageId="15" pageNumber="313">Diagnosis.</emphasis>
Dorsal fins VII + 9; pectoral fins 14; gill rakers 6 + 1517 = 2123; body moderately elongate and laterally slightly compressed; measurements as % SL: body depth at first dorsal-fin origin 2324; body depth at anal-fin origin 2021; caudal-peduncle depth 9.69.9, caudal-peduncle width 4.24.3; maximum head depth 2021; head depth through eye 16; head length 29; snout length 1011; orbit length 7.17.5; upper-jaw length 12; barbel length 20; caudal-fin length 2830; anal-fin height 1617; pelvic-fin length 2021; pectoral-fin length 1920; first dorsal-fin height 20, dorsal-fin spines proportionally decreasing in height; second dorsal-fin height 17; total oblique bars on caudal fin 11
<emphasis id="B94CEAF1F2592B7AFE92FB75C8FDDBE1" bold="true" box="[381,395,1153,1179]" pageId="15" pageNumber="313"></emphasis>
14, upper caudal-fin lobe with 56 brown mostly straight bars; lower caudal-fin lobe with 68 short brown bars ventrally, connecting dorsally to a brown stripe of about orbit diameter in its maximum width, and with about 3 short red, brownish-ochre bars along dorsal margin that do not correspond with bars on ventral-lobe margin; caudal-fin stripe and bars retained after preservation; dorsal fins in fresh fish with 34 thin brown stripes, retained in preservative; barbels reddish white when fresh; body colour in freshly deceased fish dark brown dorsally and pale brown ventrally, ventral margin white; mid-lateral body stripe dark brown, rather inconspicuous on snout and posterior half of body; head entirely grey brown; body and head dark brown dorsally and pale brown ventrally when preserved.
</paragraph>
</subSubSection>
<subSubSection id="C3226568F2592B65FF28FA55CBE1DCFB" lastPageId="16" lastPageNumber="314" pageId="15" pageNumber="313" type="description">
<paragraph id="8B8736E3F2592B7AFF28FA55C8A5D9C2" blockId="15.[151,1437,829,2008]" pageId="15" pageNumber="313">
<emphasis id="B94CEAF1F2592B7AFF28FA55C82FDAC1" bold="true" box="[199,345,1441,1467]" pageId="15" pageNumber="313">Description.</emphasis>
Measurements in % SL and counts are given in
<tableCitation id="C6BA0358F2592B7AFC67FA55CAA8DAC1" box="[904,990,1441,1467]" captionStart="TABLE 3" captionStartId="13.[150,234,479,504]" captionText="TABLE 3. Morphometric and meristic characters for three new species of Upeneus and Upeneus itoui." pageId="15" pageNumber="313">Table 3</tableCitation>
; morphometric data as ratios of SL for
<typeStatus id="54838841F2592B7AFF78FA31C98ADAA5" box="[151,252,1477,1503]" pageId="15" pageNumber="313" type="holotype">holotype</typeStatus>
, data for
<typeStatus id="54838841F2592B7AFE89FA32C8B0DA9A" box="[358,454,1478,1504]" pageId="15" pageNumber="313" type="paratype">paratype</typeStatus>
in brackets: body moderately elongate and laterally slightly compressed, body depth at first dorsal-fin origin 4.3 [4.3]; body depth at anal-fin origin 4.9 [4.9]; head depth through eye 5.1 [4.8]; head length 3.4 [3.5], larger than maximum body depth and subequal to caudal-fin length (3.6 [3.4]); first dorsal-fin height 5.1 [4.9], subequal to barbel length (5.1 [5.1]) and larger than second dorsal-fin height (6.0 [5.8]); pelvic-fin length 5.0 [4.7], slightly larger than pectoral-fin length (5.2 [4.9]) and subequal to body depth at anal-fin origin; caudal-peduncle depth 10 [10], distinctly larger than orbit length (14 [13]); and caudal-peduncle width 23 [24], subequal to pectoral-fin width (22 [23]).
</paragraph>
<paragraph id="8B8736E3F2592B65FF28F935C89CDD8B" blockId="15.[151,1437,829,2008]" lastBlockId="16.[151,1437,151,1149]" lastPageId="16" lastPageNumber="314" pageId="15" pageNumber="313">
<emphasis id="B94CEAF1F2592B7AFF28F935C856D9A1" bold="true" box="[199,288,1729,1755]" pageId="15" pageNumber="313">Colour.</emphasis>
<emphasis id="B94CEAF1F2592B7AFEC7F935C8E3D9A1" box="[296,405,1729,1755]" italics="true" pageId="15" pageNumber="313">Fresh HT</emphasis>
. Dorsal part of head and body grey brown from snout to above eye and above lateral line; dorsal body colour appearing marbled due to grey scales with dark grey-brown edges; dark grey-brown band of about pupil diameter in width along entire dorsal body margin, becoming more conspicuous behind first dorsal fin and narrows behind second dorsal fin to a dark grey, partly blackish stripe; a similarly coloured dark grey, partly blackish mid-lateral body stripe of pupil diameter in width from behind eye to caudal-fin base crossing the faint white-greyish lateral line below second dorsal fin; below of mid-lateral body stripe scales pale grey or white with brown-greenish edges forming four oblique bands that become weaker and more ochre-coloured distally, before ending on lower part of caudal-fin base or above posterioventral half of body; a fifth faint-yellowish band from below pectoral-fin base to above anal-fin origin; head below eye from behind snout pale grey with grey-brown dots or streaks and two faint-reddish blotches on ventral margin below and behind eye; iris red to rose whitish; ventral part of head and body white creamy; caudal fin with five oblique brown-ochre bars of about half pupil width on upper lobe, the most anterior bar at lobe base bent, the remaining bars more straight; white-hyaline interspaces between bars increasing in width from being narrower than bars proximally to double-bar width distally; tip of upper caudal-fin lobe white hyaline; lower caudal-fin lobe with eight short brown bars, along ventral margin of lobe brown streaks on ventral-most fin rays; white-hyaline interspaces between bars increasing distally to more than double-bar width; dark brown stripe along middle of lower caudal-fin lobe of about orbit diameter in its maximum width, stripe originating from a large, vertically extended dark-brown patch on caudal-fin base, connecting there with mid-lateral body stripe; lower caudal-fin lobe stripe changes from dark brown to brown red distally, ending as a dark-brown lobe tip; dorsal part of lower caudal-fin lobe with three brownish-ochre bars of about twice the width of bars on ventral side of lobe and with white-hyaline interspaces as wide as these bars; first dorsal fin hyaline with rather irregularly arranged pale-brown pigmentation spots, streaks or small blotches on or between individual spines giving a weak indication of a horizontal stripe close to fin base and three more oblique stripes further above; first dorsal-fin tip unpigmented; second dorsal fin hyaline with pale-brown spots arranged into three horizontal stripes, fin tip pale-brown; pectoral, pelvic and anal fins hyaline, weakly pale reddish-brown along several fin rays; barbels rose whitish, pale red on base.
</paragraph>
<paragraph id="8B8736E3F2462B65FF28FD0FCBE1DCFB" blockId="16.[151,1437,151,1149]" pageId="16" pageNumber="314">
<emphasis id="B94CEAF1F2462B65FF28FD0FC814DC6E" box="[199,354,763,789]" italics="true" pageId="16" pageNumber="314">Preserved HT</emphasis>
. Dark brown dorsally at level of snout to termination of lateral line at caudal-fin base; pale brown to beige ventrally below eyes and level of pectoral-fin base; behind pectoral-fin base to second dorsal-fin base two weakly indicated brown bands intermingled with beige; caudal-fin bars and stripe and dorsal-fin pigmentation well retained; other fins pale hyaline; barbels pale.
</paragraph>
</subSubSection>
<subSubSection id="C3226568F2462B65FF28FC7FCA2ADB6B" pageId="16" pageNumber="314" type="etymology">
<paragraph id="8B8736E3F2462B65FF28FC7FCA2ADB6B" blockId="16.[151,1437,151,1149]" pageId="16" pageNumber="314">
<emphasis id="B94CEAF1F2462B65FF28FC7FC838DCDF" bold="true" box="[199,334,907,933]" pageId="16" pageNumber="314">Etymology.</emphasis>
The name “
<taxonomicName id="4C384D60F2462B65FE35FC7FCB37DCDF" authorityName="Uiblein &amp; Motomura" authorityYear="2021" box="[474,577,907,933]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="314" phylum="Chordata" rank="species" species="willwhite">
<emphasis id="B94CEAF1F2462B65FE35FC7FCB37DCDF" box="[474,577,907,933]" italics="true" pageId="16" pageNumber="314">willwhite</emphasis>
</taxonomicName>
” is used as a composed noun in apposition. It honours the ichthyologist Dr. William T. White, Australian National Fish Collection, CSIRO National Research Collections
<collectingCountry id="F32F7673F2462B65FB3FFC5BCC4CDCB3" box="[1232,1338,943,969]" name="Australia" pageId="16" pageNumber="314">Australia</collectingCountry>
, Hobart,
<collectingCountry id="F32F7673F2462B65FF78FC27C877DC97" box="[151,257,979,1005]" name="Australia" pageId="16" pageNumber="314">Australia</collectingCountry>
, for collecting and photographing mullid specimens from S
<collectingCountry id="F32F7673F2462B65FC2EFC27CD58DC97" box="[961,1070,979,1005]" name="Indonesia" pageId="16" pageNumber="314">Indonesia</collectingCountry>
and making them available for taxonomic research (including the
<typeStatus id="54838841F2462B65FDF3FC03CB21DB6B" box="[540,599,1015,1041]" pageId="16" pageNumber="314">types</typeStatus>
of
<taxonomicName id="4C384D60F2462B65FD93FC03CA3ADB6B" authorityName="Uiblein &amp; Motomura" authorityYear="2021" box="[636,844,1015,1041]" class="Actinopterygii" family="Mullidae" genus="Upeneus" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="314" phylum="Chordata" rank="species" species="willwhite">
<emphasis id="B94CEAF1F2462B65FD93FC03CA3ADB6B" box="[636,844,1015,1041]" italics="true" pageId="16" pageNumber="314">Upeneus willwhite</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3226568F2462B65FF28FBEECB55DB23" pageId="16" pageNumber="314" type="distribution">
<paragraph id="8B8736E3F2462B65FF28FBEECB55DB23" blockId="16.[151,1437,151,1149]" pageId="16" pageNumber="314">
<emphasis id="B94CEAF1F2462B65FF28FBEECB76DB4E" bold="true" box="[199,512,1050,1076]" pageId="16" pageNumber="314">Distribution, habitat, size.</emphasis>
Only known from Lombok, S
<collectingCountry id="F32F7673F2462B65FC8FFBEFCAA7DB4F" box="[864,977,1051,1077]" name="Indonesia" pageId="16" pageNumber="314">Indonesia</collectingCountry>
; occurs most probably on shallow sandy bottoms; attains at least
<quantity id="4CC09B06F2462B65FE49FBCBC883DB23" box="[422,501,1087,1113]" metricMagnitude="-2" metricUnit="m" metricValue="9.2" pageId="16" pageNumber="314" unit="mm" value="92.0">92 mm</quantity>
SL.
</paragraph>
</subSubSection>
<subSubSection id="C3226568F2462B65FF28FB97CC6FDB07" box="[199,1305,1123,1149]" pageId="16" pageNumber="314" type="discussion">
<paragraph id="8B8736E3F2462B65FF28FB97CC6FDB07" blockId="16.[151,1437,151,1149]" box="[199,1305,1123,1149]" pageId="16" pageNumber="314">
<emphasis id="B94CEAF1F2462B65FF28FB97C84DDB07" bold="true" box="[199,315,1123,1149]" pageId="16" pageNumber="314">Remarks.</emphasis>
This species was obtained at a fish market and hence may have local commercial value.
</paragraph>
</subSubSection>
</treatment>
</document>