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<document id="D6804DFBC8A78D0E58FABBDED3C11295" ID-CLB-Dataset="8744" ID-DOI="10.11646/zootaxa.4838.2.1" ID-GBIF-Dataset="761fcec7-c79d-4e80-bb2b-23cc9e78e0b6" ID-ISSN="1175-5326" ID-Zenodo-Dep="4403783" ID-ZooBank="0AD43D95-7508-466C-980F-CBEF38CA1198" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609352386978" checkinUser="plazi" docAuthor="Gompel, Nicolas" docDate="2020" docId="039187D8FFC1BB16FF188676FBBFFA01" docLanguage="en" docName="zootaxa.4838.2.1.pdf" docOrigin="Zootaxa 4838 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zarcosia gerstmeieri Gompel 2020, n. sp." docType="treatment" docUuid="BB4E56EC-90C9-4561-B3A6-FD6428719AD9" docUuidSource="ZooBank" docVersion="8" lastPageNumber="158" masterDocId="FFA8FFA0FFC6BB11FF8F844AFF9DFFFF" masterDocTitle="Synopsis of the Asian species in the genus Zarcosia Collado &amp; Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species" masterLastPageNumber="178" masterPageNumber="151" pageNumber="158" updateTime="1698850145988" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A96813CC4384EFC90AD7032C5B2B775C">Synopsis of the Asian species in the genus Zarcosia Collado &amp; Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species</mods:title>
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<mods:namePart id="AA4F4EFF50814B1E7B04F56520F0C666">Gompel, Nicolas</mods:namePart>
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<mods:date id="3B8B351F37131C8D3334D09430A1EEE6">2020</mods:date>
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<mods:identifier id="7CF5E0926F638D6EA9D30ADB622615BA" type="DOI">10.11646/zootaxa.4838.2.1</mods:identifier>
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<treatment id="039187D8FFC1BB16FF188676FBBFFA01" ID-DOI="http://doi.org/10.5281/zenodo.4404251" ID-GBIF-Taxon="177012537" ID-Zenodo-Dep="4404251" LSID="urn:lsid:zoobank.org:act:BB4E56EC-90C9-4561-B3A6-FD6428719AD9" httpUri="http://treatment.plazi.org/id/039187D8FFC1BB16FF188676FBBFFA01" lastPageNumber="158" pageId="7" pageNumber="158">
<subSubSection id="C3226545FFC1BB16FF188676FE44FDA8" box="[151,473,572,599]" pageId="7" pageNumber="158" type="nomenclature">
<paragraph id="8B8736CEFFC1BB16FF188676FE44FDA8" blockId="7.[151,473,572,635]" box="[151,473,572,599]" pageId="7" pageNumber="158">
<heading id="D0CF81A2FFC1BB16FF188676FE44FDA8" bold="true" box="[151,473,572,599]" fontSize="11" level="1" pageId="7" pageNumber="158" reason="1">
<emphasis id="B94CEADCFFC1BB16FF188676FE44FDA8" bold="true" box="[151,473,572,599]" pageId="7" pageNumber="158">
<taxonomicName id="4C384D4DFFC1BB16FF188676FE12FDA9" authority="Gompel, 2020" authorityName="Gompel" authorityYear="2020" box="[151,399,572,598]" class="Insecta" family="Aderidae" genus="Zarcosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="gerstmeieri" status="sp. nov.">
<emphasis id="B94CEADCFFC1BB16FF188676FE12FDA9" bold="true" box="[151,399,572,598]" italics="true" pageId="7" pageNumber="158">Zarcosia gerstmeieri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27F57A7FFC1BB16FE188677FE44FDA8" box="[407,473,573,599]" pageId="7" pageNumber="158" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3226545FFC1BB16FF18862AFE5AFD84" box="[151,455,608,635]" pageId="7" pageNumber="158" type="description">
<paragraph id="8B8736CEFFC1BB16FF18862AFE5AFD84" blockId="7.[151,473,572,635]" box="[151,455,608,635]" pageId="7" pageNumber="158">
(
<figureCitation id="13032A4BFFC1BB16FF10862AFE96FD84" box="[159,267,608,635]" captionStart="FIGURE 1" captionStartId="3.[151,250,1474,1498]" captionTargetBox="[158,1432,565,1446]" captionTargetId="figure-172@3.[151,1437,561,1451]" captionTargetPageId="3" captionText="FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line." figureDoi="http://doi.org/10.5281/zenodo.4403787" httpUri="https://zenodo.org/record/4403787/files/figure.png" pageId="7" pageNumber="158">Figures 1</figureCitation>
;
<figureCitation id="13032A4BFFC1BB16FE97862AFEAAFD85" box="[280,311,608,634]" captionStart="FIGURE 3" captionStartId="20.[151,250,188,212]" captionTargetId="figure-15@19.[244,1343,179,2016]" captionTargetPageId="19" captionText="FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4453388" httpUri="https://zenodo.org/record/4453388/files/figure.png" pageId="7" pageNumber="158">3d</figureCitation>
;
<figureCitation id="13032A4BFFC1BB16FECC862AFEC3FD85" box="[323,350,608,634]" captionStart="FIGURE 4" captionStartId="22.[151,250,186,210]" captionTargetId="figure-15@21.[275,1312,181,1998]" captionTargetPageId="21" captionText="FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4453392" httpUri="https://zenodo.org/record/4453392/files/figure.png" pageId="7" pageNumber="158">4f</figureCitation>
;
<figureCitation id="13032A4BFFC1BB16FEE6862AFE0BFD84" box="[361,406,608,635]" captionStart="FIGURE 5" captionStartId="25.[151,250,1602,1626]" captionTargetBox="[170,1408,188,1563]" captionTargetId="figure-15@25.[169,1419,181,1578]" captionTargetPageId="25" captionText="FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4403791" httpUri="https://zenodo.org/record/4403791/files/figure.png" pageId="7" pageNumber="158">5j,v</figureCitation>
;
<figureCitation id="13032A4BFFC1BB16FE2C862AFE23FD84" box="[419,446,608,635]" captionStart="FIGURE 6" captionStartId="26.[151,250,1631,1655]" captionTargetBox="[158,1431,530,1605]" captionTargetId="figure-126@26.[151,1436,525,1607]" captionTargetPageId="26" captionText="FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4403795" httpUri="https://zenodo.org/record/4403795/files/figure.png" pageId="7" pageNumber="158">6c</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFC1BB16FF1886E3FD38FD19" pageId="7" pageNumber="158" type="etymology">
<paragraph id="8B8736CEFFC1BB16FF1886E3FD38FD19" blockId="7.[151,1437,680,1535]" pageId="7" pageNumber="158">
<emphasis id="B94CEADCFFC1BB16FF1886E3FE83FD3C" bold="true" box="[151,286,681,707]" pageId="7" pageNumber="158">Etymology.</emphasis>
The species is named after my colleague Roland Gerstmeier who collected this new species, in appreciation of his support of my work on
<taxonomicName id="4C384D4DFFC1BB16FDB98686FD3DFD19" box="[566,672,716,742]" class="Insecta" family="Aderidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="family">Aderidae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFC1BB16FF4886BBFB4AFCF5" box="[199,1239,752,779]" pageId="7" pageNumber="158" type="materials_examined">
<paragraph id="8B8736CEFFC1BB16FF4886BBFB4AFCF5" blockId="7.[151,1437,680,1535]" box="[199,1239,752,779]" pageId="7" pageNumber="158">
<materialsCitation id="3B503C93FFC1BB16FF4886BBFB4EFCF5" ID-GBIF-Occurrence="3020160362" box="[199,1235,752,779]" country="Vietnam" location="Cuc Phyong National Park" municipality="Nho Quan District" pageId="7" pageNumber="158" specimenCount="1" stateProvince="Ninh Binh" typeStatus="holotype">
<emphasis id="B94CEADCFFC1BB16FF4886BBFEFBFCF5" bold="true" box="[199,358,752,779]" pageId="7" pageNumber="158">
<typeStatus id="5483886CFFC1BB16FF4886BBFE9FFCF4" box="[199,258,753,779]" pageId="7" pageNumber="158">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F32F765EFFC1BB16FEE386BAFE52FCF5" box="[364,463,752,778]" name="Vietnam" pageId="7" pageNumber="158">Vietnam</collectingCountry>
:
<collectingRegion id="49FCF82CFFC1BB16FE5686BAFD21FCF5" box="[473,700,752,778]" country="Vietnam" name="Ninh Binh" pageId="7" pageNumber="158">Ninh Bình Province</collectingRegion>
,
<collectingMunicipality id="6BE3ACB4FFC1BB16FD4886BAFC04FCF5" box="[711,921,752,779]" pageId="7" pageNumber="158">Nho Quan District</collectingMunicipality>
,
<location id="8EE76015FFC1BB16FC2B86BBFB4EFCF5" LSID="urn:lsid:plazi:treatment:039187D8FFC1BB16FF188676FBBFFA01:8EE76015FFC1BB16FC2B86BBFB4EFCF5" box="[932,1235,752,779]" country="Vietnam" municipality="Nho Quan District" name="Cuc Phyong National Park" pageId="7" pageNumber="158" stateProvince="Ninh Binh">Cúc Phýõng National Park</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFC1BB16FF48875EFCA9FCD0" box="[199,820,788,815]" pageId="7" pageNumber="158" type="distribution">
<paragraph id="8B8736CEFFC1BB16FF48875EFCA9FCD0" blockId="7.[151,1437,680,1535]" box="[199,820,788,815]" pageId="7" pageNumber="158">
<emphasis id="B94CEADCFFC1BB16FF48875EFEFCFCD1" bold="true" box="[199,353,788,814]" pageId="7" pageNumber="158">Distribution.</emphasis>
<collectingCountry id="F32F765EFFC1BB16FEE8875EFE57FCD1" box="[359,458,788,814]" name="Vietnam" pageId="7" pageNumber="158">Vietnam</collectingCountry>
:
<collectingRegion id="49FCF82CFFC1BB16FE5A875EFD28FCD1" box="[469,693,788,814]" country="Vietnam" name="Ninh Binh" pageId="7" pageNumber="158">Ninh Bình Province</collectingRegion>
(
<figureCitation id="13032A4BFFC1BB16FD4C875EFCBFFCD0" box="[707,802,788,815]" captionStart="FIGURE 1" captionStartId="3.[151,250,1474,1498]" captionTargetBox="[158,1432,565,1446]" captionTargetId="figure-172@3.[151,1437,561,1451]" captionTargetPageId="3" captionText="FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line." figureDoi="http://doi.org/10.5281/zenodo.4403787" httpUri="https://zenodo.org/record/4403787/files/figure.png" pageId="7" pageNumber="158">Figure 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFC1BB16FF488772FE68FA49" pageId="7" pageNumber="158" type="description">
<paragraph id="8B8736CEFFC1BB16FF488772FDC2FC89" blockId="7.[151,1437,680,1535]" pageId="7" pageNumber="158">
<emphasis id="B94CEADCFFC1BB16FF488772FEC4FCAD" bold="true" box="[199,345,824,850]" pageId="7" pageNumber="158">Description.</emphasis>
(
<figureCitation id="13032A4BFFC1BB16FEE78772FE44FCAC" box="[360,473,824,851]" captionStart="FIGURE 3" captionStartId="20.[151,250,188,212]" captionTargetId="figure-15@19.[244,1343,179,2016]" captionTargetPageId="19" captionText="FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4453388" httpUri="https://zenodo.org/record/4453388/files/figure.png" pageId="7" pageNumber="158">Figure 3d</figureCitation>
). Female unknown. Body length.
<specimenCount id="9D3EFD47FFC1BB16FCD98773FCF6FCAC" box="[854,875,825,851]" pageId="7" pageNumber="158" type="male">♂</specimenCount>
<typeStatus id="5483886CFFC1BB16FCFC8772FC44FCAD" box="[883,985,824,850]" pageId="7" pageNumber="158" type="holotype">holotype</typeStatus>
:
<quantity id="4CC09B2BFFC1BB16FC6B8772FBD6FCAC" box="[996,1099,824,851]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" pageId="7" pageNumber="158" unit="mm" value="3.25">3.25 mm</quantity>
. Robust, sturdy, entirely dark, covered with double golden pubescence.
</paragraph>
<paragraph id="8B8736CEFFC1BB16FF4887CAFC86FC1D" blockId="7.[151,1437,680,1535]" pageId="7" pageNumber="158">
<emphasis id="B94CEADCFFC1BB16FF4887CAFE99FC65" box="[199,260,896,922]" italics="true" pageId="7" pageNumber="158">Head</emphasis>
. Weakly transverse, barely wider than pronotum, covered with white pubescence. Pubescence particularly dense anteriorly, between the antennal space. Punctation coarse and very dense. Palpi orange, last article securiform. Posterior margin lined with a fringe of white pubescence.
</paragraph>
<paragraph id="8B8736CEFFC1BB16FF4887A7FDC4FBD5" blockId="7.[151,1437,680,1535]" pageId="7" pageNumber="158">
<emphasis id="B94CEADCFFC1BB16FF4887A7FEACFBF8" box="[199,305,1005,1031]" italics="true" pageId="7" pageNumber="158">Antennae</emphasis>
. Strong, serrated, as long as elytra, antennomeres 3–7 long, antennomeres 8–10 progressively shorter. Covered with hirsute blond pubescence.
</paragraph>
<paragraph id="8B8736CEFFC1BB16FF48807FFEDBFB21" blockId="7.[151,1437,680,1535]" pageId="7" pageNumber="158">
<emphasis id="B94CEADCFFC1BB16FF48807FFEABFBB1" box="[199,310,1077,1102]" italics="true" pageId="7" pageNumber="158">Pronotum</emphasis>
(
<figureCitation id="13032A4BFFC1BB16FEC9807EFE2BFBB0" box="[326,438,1076,1103]" captionStart="FIGURE 6" captionStartId="26.[151,250,1631,1655]" captionTargetBox="[158,1431,530,1605]" captionTargetId="figure-126@26.[151,1436,525,1607]" captionTargetPageId="26" captionText="FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4403795" httpUri="https://zenodo.org/record/4403795/files/figure.png" pageId="7" pageNumber="158">Figure 6c</figureCitation>
). Convex, subrectangular, slightly longer than wide. Sides parallel from base to anterior fifth, then converging markedly inward toward apical margin anteriorly. Densely punctuated, points nearly adjacent. Covered with double pubescence, moderately long blond hairs borne from the punctation and pulverulent interspersed hairs. Base bi-sinuated with a dense fringe of golden pubescence anterior to the scutellum and at each posterior angle.
</paragraph>
<paragraph id="8B8736CEFFC1BB16FF4880A2FBE3FAD8" blockId="7.[151,1437,680,1535]" pageId="7" pageNumber="158">
<emphasis id="B94CEADCFFC1BB16FF4880A2FE90FAFD" box="[199,269,1256,1282]" italics="true" pageId="7" pageNumber="158">Elytra</emphasis>
. Length:
<quantity id="4CC09B2BFFC1BB16FEF580A2FE7CFAFC" box="[378,481,1256,1283]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="7" pageNumber="158" unit="mm" value="1.15">1.15 mm</quantity>
. Punctation strong, points well delineated, deep, space between points larger than the points themselves. Densely covered with double pubescence (
<figureCitation id="13032A4BFFC1BB16FCC48146FC2BFAD8" box="[843,950,1292,1319]" captionStart="FIGURE 5" captionStartId="25.[151,250,1602,1626]" captionTargetBox="[170,1408,188,1563]" captionTargetId="figure-15@25.[169,1419,181,1578]" captionTargetPageId="25" captionText="FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4403791" httpUri="https://zenodo.org/record/4403791/files/figure.png" pageId="7" pageNumber="158">Figure 5j</figureCitation>
). Weakly convex.
</paragraph>
<paragraph id="8B8736CEFFC1BB16FF48817BFCB4FAB5" blockId="7.[151,1437,680,1535]" box="[199,809,1328,1355]" pageId="7" pageNumber="158">
<emphasis id="B94CEADCFFC1BB16FF48817BFF61FAB5" box="[199,252,1329,1354]" italics="true" pageId="7" pageNumber="158">Legs</emphasis>
. Short, sturdy, dark. No visible sexual characters.
</paragraph>
<paragraph id="8B8736CEFFC1BB16FF48811EFE68FA49" blockId="7.[151,1437,680,1535]" pageId="7" pageNumber="158">
<emphasis id="B94CEADCFFC1BB16FF48811EFEA9FA91" box="[199,308,1364,1390]" italics="true" pageId="7" pageNumber="158">Aedeagus</emphasis>
(
<figureCitation id="13032A4BFFC1BB16FECC811EFE2CFA90" box="[323,433,1364,1391]" captionStart="FIGURE 4" captionStartId="22.[151,250,186,210]" captionTargetId="figure-15@21.[275,1312,181,1998]" captionTargetPageId="21" captionText="FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4453392" httpUri="https://zenodo.org/record/4453392/files/figure.png" pageId="7" pageNumber="158">Figure 4f</figureCitation>
). Very sturdy, strongly sclerotized, sides sinuated in dorsal view. Apex complex, lozenge shaped, terminated with a stinger. Strongly bent in lateral view, with an upward kink in apical fifth and a brief downward deflection at the tip.
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFC1BB16FF48818BFBBFFA01" pageId="7" pageNumber="158" type="materials_examined">
<paragraph id="8B8736CEFFC1BB16FF48818BFBBFFA01" blockId="7.[151,1437,680,1535]" pageId="7" pageNumber="158">
<emphasis id="B94CEADCFFC1BB16FF48818BFEE8FA25" bold="true" box="[199,373,1472,1499]" pageId="7" pageNumber="158">Type material.</emphasis>
<materialsCitation id="3B503C93FFC1BB16FEF3818AFBBFFA01" ID-GBIF-Occurrence="3020160370" collectingDate="2017-05-06" collectionCode="NME" collectorName="R. Gerstmeier" country="Vietnam" elevation="375" latitude="20.350279" location="Cuc Phyong National Park" longLatPrecision="21" longitude="105.59334" municipality="Nho Quan District" pageId="7" pageNumber="158" specimenCount="1" specimenCount-male="1" stateProvince="Ninh Binh" typeStatus="holotype">
<typeStatus id="5483886CFFC1BB16FEF3818AFE79FA25" box="[380,484,1472,1498]" pageId="7" pageNumber="158" type="holotype">Holotype</typeStatus>
(
<specimenCount id="9D3EFD47FFC1BB16FE7B818BFD96FA24" box="[500,523,1473,1499]" pageId="7" pageNumber="158" type="male">♂</specimenCount>
,
<collectionCode id="ED29AE0BFFC1BB16FD98818BFDC7FA24" box="[535,602,1473,1499]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="7" pageNumber="158">NME</collectionCode>
).
<emphasis id="B94CEADCFFC1BB16FDE2818AFD4EFA25" bold="true" box="[621,723,1472,1498]" pageId="7" pageNumber="158">
<collectingCountry id="F32F765EFFC1BB16FDE2818AFD4EFA25" box="[621,723,1472,1498]" name="Vietnam" pageId="7" pageNumber="158">Vietnam</collectingCountry>
</emphasis>
:
<collectingRegion id="49FCF82CFFC1BB16FD6E818AFC5EFA25" box="[737,963,1472,1498]" country="Vietnam" name="Ninh Binh" pageId="7" pageNumber="158">Ninh Bình Province</collectingRegion>
,
<collectingMunicipality id="6BE3ACB4FFC1BB16FC40818AFB3DFA25" box="[975,1184,1472,1499]" pageId="7" pageNumber="158">Nho Quan District</collectingMunicipality>
,
<location id="8EE76015FFC1BB16FB24818BFF50FA01" LSID="urn:lsid:plazi:treatment:039187D8FFC1BB16FF188676FBBFFA01:8EE76015FFC1BB16FB24818BFF50FA01" country="Vietnam" latitude="20.350279" longLatPrecision="21" longitude="105.59334" municipality="Nho Quan District" name="Cuc Phyong National Park" pageId="7" pageNumber="158" stateProvince="Ninh Binh">Cúc Phýõng National Park</location>
, end of road,
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<geoCoordinate id="EE0C5009FFC1BB16FE7281AEFD05FA00" box="[509,664,1508,1535]" degrees="105" direction="east" minutes="35" orientation="longitude" pageId="7" pageNumber="158" precision="15" seconds="36" value="105.59334">105°35’36” E</geoCoordinate>
(
<quantity id="4CC09B2BFFC1BB16FD2881AEFD6CFA00" box="[679,753,1508,1535]" metricMagnitude="2" metricUnit="m" metricValue="3.75" pageId="7" pageNumber="158" unit="m" value="375.0">
<elevation id="0015D1FDFFC1BB16FD2881AEFD6CFA00" box="[679,753,1508,1535]" metricMagnitude="2" metricUnit="m" metricValue="3.75" pageId="7" pageNumber="158" unit="m" value="375.0">375 m</elevation>
</quantity>
) (
<collectorName id="26CD5318FFC1BB16FC8681AFFC35FA00" box="[777,936,1508,1535]" pageId="7" pageNumber="158">R. Gerstmeier</collectorName>
)
<date id="FF86100EFFC1BB16FC3981AEFB83FA01" box="[950,1054,1508,1534]" pageId="7" pageNumber="158" value="2017-05-06">
<collectingDate id="EFC2E9E6FFC1BB16FC3981AEFB83FA01" box="[950,1054,1508,1534]" pageId="7" pageNumber="158" value="2017-05-06">6.V.2017</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>
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