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<document id="8EA0CEE56E7F7A8D8F55691291EBC8A8" ID-DOI="10.5281/zenodo.276822" ID-GBIF-Dataset="f5489381-8fe8-48a0-9fb6-5addeebaa3f8" ID-ISSN="1175-5326" ID-Zenodo-Dep="276822" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460237113383" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2011" docId="03C687E4FFDCFF91FF1BEA7D1565FE4D" docLanguage="en" docName="zt02763p033.pdf" docOrigin="Zootaxa 2763" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Hlavaciellus peninsularis Jałoszyński, 2011, sp. n." docType="treatment" docVersion="6" lastPageNumber="5" masterDocId="FFFFFF9CFFDFFF95FF8CEB58101EFFCA" masterDocTitle="Hlavaciellus Jałoszyński, 2006: eleven new species, detailed morphology and systematic position within Cephenniini (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="33" masterPageNumber="1" pageNumber="4" updateTime="1698239820214" updateUser="plazi">
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<mods:title id="3AE0AB8B67E369E247A77CB1399CAA4A">Hlavaciellus Jałoszyński, 2006: eleven new species, detailed morphology and systematic position within Cephenniini (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:namePart id="57DCB922556598636C2C967F77069BB4">Jałoszyński, Paweł</mods:namePart>
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<mods:date id="60065FA184505D49EADC6439F70CBDCD">2011</mods:date>
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<treatment id="03C687E4FFDCFF91FF1BEA7D1565FE4D" ID-DOI="http://doi.org/10.5281/zenodo.6192594" ID-GBIF-Taxon="119388728" ID-Zenodo-Dep="6192594" LSID="urn:lsid:plazi:treatment:03C687E4FFDCFF91FF1BEA7D1565FE4D" httpUri="http://treatment.plazi.org/id/03C687E4FFDCFF91FF1BEA7D1565FE4D" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<subSubSection id="C3756579FFDCFF96FF1BEA7D1178FE95" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="8BD036F2FFDCFF96FF1BEA7D120BFEF5" blockId="3.[151,533,293,351]" box="[151,533,293,319]" pageId="3" pageNumber="4">
<heading id="D098819EFFDCFF96FF1BEA7D120BFEF5" bold="true" box="[151,533,293,319]" fontSize="11" level="1" pageId="3" pageNumber="4" reason="1">
<emphasis id="B91BEAE0FFDCFF96FF1BEA7D120BFEF5" bold="true" box="[151,533,293,319]" pageId="3" pageNumber="4">
<taxonomicName id="4C6F4D71FFDCFF96FF1BEA7D11D5FEF5" ID-CoL="3M5PX" box="[151,459,293,319]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="peninsularis" status="sp. nov.">
<emphasis id="B91BEAE0FFDCFF96FF1BEA7D11D5FEF5" bold="true" box="[151,459,293,319]" italics="true" pageId="3" pageNumber="4">Hlavaciellus peninsularis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A228579BFFDCFF96FE5FEA7D120BFEF5" box="[467,533,293,319]" pageId="3" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD036F2FFDCFF96FF1BEA1F1178FE95" blockId="3.[151,533,293,351]" box="[151,358,327,351]" pageId="3" pageNumber="4">
(
<figureCitation id="13542A77FFDCFF96FF13EA1F10F3FE95" box="[159,237,327,351]" captionStart="FIGURES 1 3" captionStartId="4.[151,264,1341,1364]" captionTargetBox="[167,1385,459,1307]" captionTargetId="figure@4.[151,1436,447,1317]" captionTargetPageId="4" captionText="FIGURES 1 3. Dorsal view of holotypes of Hlavaciellus, males. 1, H. peninsularis sp. n. 2, H. cirrus sp. n. 3, H. angustifrons sp. n. Scale bars: 1 mm." httpUri="https://zenodo.org/record/276823/files/figure.png" pageId="3" pageNumber="4">Figs. 1</figureCitation>
,
<figureCitation id="13542A77FFDCFF96FF76EA1F1114FE95" box="[250,266,327,351]" captionStart="FIGURES 4 9" captionStartId="5.[151,264,1961,1984]" captionTargetBox="[158,1432,1102,1933]" captionTargetId="figure@5.[151,1436,1095,1937]" captionTargetPageId="5" captionText="FIGURES 4 9. Modifications of head in frontal view. 4, Hlavaciellus peninsularis sp. n. 5, H. cirrus sp. n. 6, H. angustifrons sp. n. 7, H. decoratus sp. n. 8, H. smetanai sp. n. 9, H. bifoveatus sp. n. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/276824/files/figure.png" pageId="3" pageNumber="4">4</figureCitation>
,
<figureCitation id="13542A77FFDCFF96FE9AEA1F1143FE95" box="[278,349,327,351]" captionStart="FIGURES 13 23" captionStartId="9.[151,264,1927,1950]" captionTargetBox="[171,1428,637,1895]" captionTargetId="figure@9.[151,1436,625,1907]" captionTargetPageId="9" captionText="FIGURES 13 23. Aedeagus in ventral (13, 15, 18, 20, 22) and lateral (14, 16, 19, 21, 23) views, and head of male in lateral view (17). 13, 14, Hlavaciellus peninsularis sp. n. 15 17, H. cirrus sp. n. 18, 19, H. angustifrons sp. n. 20, 21, H. decoratus sp. n. 22, 23, H. smetanai sp. n. Scale bars: 0.2 mm (13 16, 18 23), 0.1 mm (17)." httpUri="https://zenodo.org/record/276826/files/figure.png" pageId="3" pageNumber="4">13, 14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3756579FFDCFF96FF1BEAC91244FE3B" pageId="3" pageNumber="4" type="materials_examined">
<paragraph id="8BD036F2FFDCFF96FF1BEAC91244FE3B" blockId="3.[151,1437,400,2010]" pageId="3" pageNumber="4">
<emphasis id="B91BEAE0FFDCFF96FF1BEAC91279FE63" bold="true" box="[151,615,400,425]" pageId="3" pageNumber="4">
<typeStatus id="54D48850FFDCFF96FF1BEAC910CCFE63" box="[151,210,401,425]" pageId="3" pageNumber="4">Type</typeStatus>
material.
<typeStatus id="54D48850FFDCFF96FEC7EAC911A2FE63" box="[331,444,401,425]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F3787662FFDCFF96FE4BEAC8127CFE63" box="[455,610,400,425]" name="Malaysia" pageId="3" pageNumber="4">MALAYSIA</collectingCountry>
:
</emphasis>
ɗ, &quot;W.
<collectingCountry id="F3787662FFDCFF96FD33EAC81334FE62" box="[703,810,400,424]" name="Malaysia" pageId="3" pageNumber="4">Malaysia</collectingCountry>
: Pahang \
<collectingCountry id="F3787662FFDCFF96FC17EAC8141FFE62" box="[923,1025,400,424]" name="Cameroon" pageId="3" pageNumber="4">Cameron</collectingCountry>
Highl,
<quantity id="4C979B17FFDCFF96FBD9EAC814B7FE62" box="[1109,1193,400,424]" metricMagnitude="3" metricUnit="m" metricValue="1.72" pageId="3" pageNumber="4" unit="m" value="1720.0">1720 m</quantity>
\ Gunung Jasar # 19b \ Löbl &amp; Calame,
<date id="FFD11032FFDCFF96FEE1EAED11DAFE07" box="[365,452,437,461]" pageId="3" pageNumber="4" value="93-03-25">25.3.93</date>
&quot; [white, printed], &quot;
<taxonomicName id="4C6F4D71FFDCFF96FD28EAED1378FE07" box="[676,870,437,461]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEAE0FFDCFF96FD28EAED1378FE07" box="[676,870,437,461]" italics="true" pageId="3" pageNumber="4">HLAVACIELLUS</emphasis>
</taxonomicName>
\
<taxonomicName id="4C6F4D71FFDCFF96FC0DEAED1415FE07" box="[897,1035,437,461]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="peninsularis">
<emphasis id="B91BEAE0FFDCFF96FC0DEAED1415FE07" box="[897,1035,437,461]" italics="true" pageId="3" pageNumber="4">peninsularis</emphasis>
</taxonomicName>
m. \ det. P. JAŁOSZYŃSKI, '09 \
<typeStatus id="54D48850FFDCFF96FF1BEA81115FFE3B" box="[151,321,473,497]" pageId="3" pageNumber="4" type="holotype">HOLOTYPUS</typeStatus>
&quot; [red, printed] (
<collectionCode id="ED7EAE37FFDCFF96FE7EEA811250FE3B" box="[498,590,473,497]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="3" pageNumber="4">MHNG</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3756579FFDCFF96FF4BEAA51576FDF3" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="8BD036F2FFDCFF96FF4BEAA51576FDF3" blockId="3.[151,1437,400,2010]" pageId="3" pageNumber="4">
<emphasis id="B91BEAE0FFDCFF96FF4BEAA5115FFDDC" bold="true" box="[199,321,509,534]" pageId="3" pageNumber="4">Diagnosis.</emphasis>
Head in males with distinct, broadly U-shaped carina separating vertex from frontoclypeal region; antero-median part of vertex adjacent to carina concave and covered with extremely fine and dense punctures.
</paragraph>
</subSubSection>
<subSubSection id="C3756579FFDCFF96FF4BE91D11BCF917" pageId="3" pageNumber="4" type="description">
<paragraph id="8BD036F2FFDCFF96FF4BE91D10F7FD4B" blockId="3.[151,1437,400,2010]" pageId="3" pageNumber="4">
<emphasis id="B91BEAE0FFDCFF96FF4BE91D1147FD94" bold="true" box="[199,345,581,606]" pageId="3" pageNumber="4">Description.</emphasis>
BL 1.88. Body of male (
<figureCitation id="13542A77FFDCFF96FDFCE91D12AEFD97" box="[624,688,581,605]" captionStart="FIGURES 1 3" captionStartId="4.[151,264,1341,1364]" captionTargetBox="[167,1385,459,1307]" captionTargetId="figure@4.[151,1436,447,1317]" captionTargetPageId="4" captionText="FIGURES 1 3. Dorsal view of holotypes of Hlavaciellus, males. 1, H. peninsularis sp. n. 2, H. cirrus sp. n. 3, H. angustifrons sp. n. Scale bars: 1 mm." httpUri="https://zenodo.org/record/276823/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
) strongly convex, pigmentation dark reddish-brown, vestiture yellowish.
</paragraph>
<paragraph id="8BD036F2FFDCFF96FF4BE9D510E7FBC7" blockId="3.[151,1437,400,2010]" pageId="3" pageNumber="4">
Head (
<figureCitation id="13542A77FFDCFF96FE9DE9D5114BFD6F" box="[273,341,653,677]" captionStart="FIGURES 4 9" captionStartId="5.[151,264,1961,1984]" captionTargetBox="[158,1432,1102,1933]" captionTargetId="figure@5.[151,1436,1095,1937]" captionTargetPageId="5" captionText="FIGURES 4 9. Modifications of head in frontal view. 4, Hlavaciellus peninsularis sp. n. 5, H. cirrus sp. n. 6, H. angustifrons sp. n. 7, H. decoratus sp. n. 8, H. smetanai sp. n. 9, H. bifoveatus sp. n. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/276824/files/figure.png" pageId="3" pageNumber="4">Fig. 4</figureCitation>
) large, HL 0.28, HW 0.49; vertex long and regularly convex, anteriorly delimited by low but distinct broadly U-shaped carina; frons subtrapezoidal, nearly flat, clypeal region nearly rectangular, strongly convex; supraantennal tubercles prominent, distinctly separated from frons; eyes large, strongly convex and coarsely faceted. Punctures and setae as in
<figureCitation id="13542A77FFDCFF96FE7FE9A11226FCDB" box="[499,568,761,785]" captionStart="FIGURES 4 9" captionStartId="5.[151,264,1961,1984]" captionTargetBox="[158,1432,1102,1933]" captionTargetId="figure@5.[151,1436,1095,1937]" captionTargetPageId="5" captionText="FIGURES 4 9. Modifications of head in frontal view. 4, Hlavaciellus peninsularis sp. n. 5, H. cirrus sp. n. 6, H. angustifrons sp. n. 7, H. decoratus sp. n. 8, H. smetanai sp. n. 9, H. bifoveatus sp. n. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/276824/files/figure.png" pageId="3" pageNumber="4">Fig. 4</figureCitation>
. Antennae nearly as long as 0.7 BL, slender, gradually thickened up to antennomere X, uniformly covered with thin, long, dense and suberect setae, AnL 1.25, antennomere I 1.5x as long as broad; II distinctly narrower and shorter than I, 1.1x as long as broad; IIIV subequal in length and width, each as broad as II but slightly longer, 1.2x as long as broad; VI slightly larger than V, 1.2x as long as broad; VII slightly larger than VI, 1.1x as long as broad; VIII slightly broader but not longer than VII, nearly as broad as long; IX barely noticeably larger than VIII, as long as broad; X distinctly larger than IX, with slant apical margin (external lateral margin longer than internal one), as long as broad; XI slightly narrower than X, distinctly shorter than IXX together.
</paragraph>
<paragraph id="8BD036F2FFDCFF96FF4BEF4213C3FAC3" blockId="3.[151,1437,400,2010]" pageId="3" pageNumber="4">
Pronotum in dorsal view subrectangular with moderately convex anterior margin, equally broad from base up to anterior third, PL 0.58, PW 0.88; sides strongly rounded in anterior third, then nearly straight and parallel up to right and blunt hind angles; posterior margin shallowly bi-emarginate; ante-basal pits very shallow and diffused, each located slightly closer to posterior than to lateral margin of pronotum; lateral edges in posterior third slightly swollen to form indistinct lateral carinae. Pronotal disc glossy, in middle covered with very fine and shallow punctures separated by spaces 1
<date id="FFD11032FFDCFF96FE54EF9511F9FB2F" box="[472,487,1229,1253]" pageId="3" pageNumber="4">2</date>
x as wide as puncture diameters, punctures close to margins of pronotum are larger and denser, but not coarse. Setae long, moderately dense, suberect to erect.
</paragraph>
<paragraph id="8BD036F2FFDCFF96FF4BEE4D10F1F9CF" blockId="3.[151,1437,400,2010]" pageId="3" pageNumber="4">
Elytra oval, relatively short, very convex in anterior half and steeply lowering towards apices in posterior half, broadest near anterior fifth, EL 1.03, EW 0.93, EI 1.11; basal pit on each elytron small but distinct, located very close to scutellum; subhumeral lines sharply marked, as long as 0.34x EL, only slightly divergent, each developed as border between higher humeral region and lower adsutural area; apices of elytra rounded together. Punctures on basal third of each elytron small but more distinct than those on pronotum, gradually reducing in diameter and depth toward lateral margins and apices; setae similar to those on pronotum but slightly thicker.
<collectingCountry id="F3787662FFDCFF96FB67EE91153DFA2B" box="[1259,1315,1481,1505]" name="India" pageId="3" pageNumber="4">Hind</collectingCountry>
wings not studied.
</paragraph>
<paragraph id="8BD036F2FFDCFF96FF4BED4A1571F9E3" blockId="3.[151,1437,400,2010]" box="[199,1391,1553,1578]" pageId="3" pageNumber="4">Legs slender and long; protibiae barely noticeably bent outwards in middle; meso- and metatibiae straight.</paragraph>
<paragraph id="8BD036F2FFDCFF96FF4BED6D148FF987" blockId="3.[151,1437,400,2010]" box="[199,1169,1589,1613]" pageId="3" pageNumber="4">Metaventrite with deep postmesocoxal impressions, occupying nearly its entire length.</paragraph>
<paragraph id="8BD036F2FFDCFF96FF4BED011206F973" blockId="3.[151,1437,400,2010]" pageId="3" pageNumber="4">
Aedeagus (
<figureCitation id="13542A77FFDCFF96FEC4ED0111CBF9BB" box="[328,469,1625,1649]" captionStart="FIGURES 13 23" captionStartId="9.[151,264,1927,1950]" captionTargetBox="[171,1428,637,1895]" captionTargetId="figure@9.[151,1436,625,1907]" captionTargetPageId="9" captionText="FIGURES 13 23. Aedeagus in ventral (13, 15, 18, 20, 22) and lateral (14, 16, 19, 21, 23) views, and head of male in lateral view (17). 13, 14, Hlavaciellus peninsularis sp. n. 15 17, H. cirrus sp. n. 18, 19, H. angustifrons sp. n. 20, 21, H. decoratus sp. n. 22, 23, H. smetanai sp. n. Scale bars: 0.2 mm (13 16, 18 23), 0.1 mm (17)." httpUri="https://zenodo.org/record/276826/files/figure.png" pageId="3" pageNumber="4">Figs. 13, 14</figureCitation>
) very slender, AeL 0.49; median lobe with short, indistinctly separated apical part; internal armature slightly asymmetrical; parameres in lateral view very broad, with indistinctly expanded subapical region and strongly curved apices.
</paragraph>
<paragraph id="8BD036F2FFDCFF96FF4BED9D11BCF917" blockId="3.[151,1437,400,2010]" box="[199,418,1733,1757]" pageId="3" pageNumber="4">
<emphasis id="B91BEAE0FFDCFF96FF4BED9D113BF917" bold="true" box="[199,293,1733,1757]" pageId="3" pageNumber="4">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3756579FFDCFF96FF4BEDB11367F8CB" box="[199,889,1769,1794]" pageId="3" pageNumber="4" type="distribution">
<paragraph id="8BD036F2FFDCFF96FF4BEDB11367F8CB" blockId="3.[151,1437,400,2010]" box="[199,889,1769,1794]" pageId="3" pageNumber="4">
<emphasis id="B91BEAE0FFDCFF96FF4BEDB1117EF8C8" bold="true" box="[199,352,1769,1794]" pageId="3" pageNumber="4">Distribution.</emphasis>
West
<collectingCountry id="F3787662FFDCFF96FE29EDB1120EF8CB" box="[421,528,1769,1793]" name="Malaysia" pageId="3" pageNumber="4">Malaysia</collectingCountry>
:
<collectingCountry id="F3787662FFDCFF96FD96EDB1129FF8CB" box="[538,641,1769,1793]" name="Cameroon" pageId="3" pageNumber="4">Cameron</collectingCountry>
Highlands in Pahang.
</paragraph>
</subSubSection>
<subSubSection id="C3756579FFDCFF96FF4BEC55113DF883" pageId="3" pageNumber="4" type="etymology">
<paragraph id="8BD036F2FFDCFF96FF4BEC55113DF883" blockId="3.[151,1437,400,2010]" pageId="3" pageNumber="4">
<emphasis id="B91BEAE0FFDCFF96FF4BEC551150F8EF" bold="true" box="[199,334,1805,1829]" pageId="3" pageNumber="4">Etymology.</emphasis>
The name reflects the fact that this is the only
<taxonomicName id="4C6F4D71FFDCFF96FCD2EC5513F5F8EF" box="[862,1003,1805,1829]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEAE0FFDCFF96FCD2EC5513F5F8EF" box="[862,1003,1805,1829]" italics="true" pageId="3" pageNumber="4">Hlavaciellus</emphasis>
</taxonomicName>
known so far from the peninsular part of
<collectingCountry id="F3787662FFDCFF96FF39EC691101F883" box="[181,287,1841,1865]" name="Malaysia" pageId="3" pageNumber="4">Malaysia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3756579FFDCFF91FF4BEC0E1565FE4D" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="discussion">
<paragraph id="8BD036F2FFDCFF96FF4BEC0E11D8F813" blockId="3.[151,1437,400,2010]" pageId="3" pageNumber="4">
<emphasis id="B91BEAE0FFDCFF96FF4BEC0E1125F8A4" bold="true" box="[199,315,1878,1902]" pageId="3" pageNumber="4">Remarks.</emphasis>
Despite the unremarkable aedeagus highly similar to copulatory organs of several other species, the external morphology provides unique and unambiguous diagnostic characters. Within the
<taxonomicName id="4C6F4D71FFDCFF96FB1CEC211505F85B" box="[1168,1307,1913,1937]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="tribe" tribe="Cephenniini">Cephenniini</taxonomicName>
, the frontal U-shaped carina found in
<taxonomicName id="4C6F4D71FFDCFF96FE30ECC61274F87F" box="[444,618,1949,1973]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="peninsularis">
<emphasis id="B91BEAE0FFDCFF96FE30ECC61274F87F" box="[444,618,1949,1973]" italics="true" pageId="3" pageNumber="4">H. peninsularis</emphasis>
</taxonomicName>
can be seen only in several species of
<taxonomicName id="4C6F4D71FFDCFF96FBACECC514A9F87F" box="[1056,1207,1949,1973]" class="Insecta" family="Scydmaenidae" genus="Cephennodes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEAE0FFDCFF96FBACECC514A9F87F" box="[1056,1207,1949,1973]" italics="true" pageId="3" pageNumber="4">Cephennodes</emphasis>
</taxonomicName>
in the
<emphasis id="B91BEAE0FFDCFF96FA89ECC51582F87F" box="[1285,1436,1949,1973]" italics="true" pageId="3" pageNumber="4">impressifrons</emphasis>
group (
<bibRefCitation id="EFFE4B03FFDCFF96FF6BEC9811A4F813" author="Jaloszynski" box="[231,442,1984,2009]" pageId="3" pageNumber="29" refString="Jaloszynski, P. (2007 a) The Cephenniini (Coleoptera, Scydmaenidae) of China. II. Cephennodes Reitter of southern provinces, with taxonomic notes on the Cephennodes-Chelonoidum complex (Coleoptera, Scydmaenidae). Genus, 18 (1), 7 - 101." type="journal article" year="2007" yearSuffix="a">Jałoszyński 2007a</bibRefCitation>
).
</paragraph>
<paragraph id="8BD036F2FFDBFF91FF4BEBCF1565FE4D" blockId="4.[151,1436,151,391]" pageId="4" pageNumber="5">
The
<collectingCountry id="F3787662FFDBFF91FF75EBCF117EFF65" box="[249,352,151,175]" name="Cameroon" pageId="4" pageNumber="5">Cameron</collectingCountry>
Highlands is a popular place for collecting beetles in West
<collectingCountry id="F3787662FFDBFF91FC74EBCF147FFF65" box="[1016,1121,151,175]" name="Malaysia" pageId="4" pageNumber="5">Malaysia</collectingCountry>
, and specimens of Staphylinoidea taken on Mount Jasar (Gunung Jasar in Malay language) by sifting-based methods and flight intercept traps are relatively common and abundant in various museums (e.g., National Museum of Science, Tokyo and MHNG), and in private collection of the author, who visited this area in
<date id="FFD11032FFDBFF91FCD7EA5C140FFED6" box="[859,1041,260,284]" pageId="4" pageNumber="5" value="2003-12">December 2003</date>
. However, the male
<typeStatus id="54D48850FFDBFF91FB74EA5C1544FED6" box="[1272,1370,260,284]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C6F4D71FFDBFF91FA0DEA5D113CFE8A" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="peninsularis">
<emphasis id="B91BEAE0FFDBFF91FA0DEA5D113CFE8A" italics="true" pageId="4" pageNumber="5">H. peninsularis</emphasis>
</taxonomicName>
is the only specimen of
<taxonomicName id="4C6F4D71FFDBFF91FDB1EA7012D5FE8A" box="[573,715,296,320]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEAE0FFDBFF91FDB1EA7012D5FE8A" box="[573,715,296,320]" italics="true" pageId="4" pageNumber="5">Hlavaciellus</emphasis>
</taxonomicName>
found among large materials of
<taxonomicName id="4C6F4D71FFDBFF91FBCFEA7F14F8FEF5" box="[1091,1254,295,319]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="subFamily" subFamily="Scydmaeninae">Scydmaeninae</taxonomicName>
in these collections. This may suggest either a true rareness of this species, or, more likely, a biology or microhabitat preferences different from those of most
<taxonomicName id="4C6F4D71FFDBFF91FE55EA37127BFE4D" box="[473,613,367,391]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="tribe" tribe="Cephenniini">Cephenniini</taxonomicName>
, which are normally common in moist leaf litter in subtropical forests.
</paragraph>
</subSubSection>
</treatment>
</document>