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<document id="798D6D167E4E9EA615940C682C95429B" ID-DOI="10.11646/zootaxa.3920.1.1" ID-GBIF-Dataset="7289c86d-0ed1-4e93-b7b3-bc596042d854" ID-ISSN="1175-5326" ID-Zenodo-Dep="287886" ID-ZooBank="025F9918-F9F6-4049-B7CD-5AC632094F5E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460994190417" checkinUser="plazi" docAuthor="Kataev, Boris M. &amp; Liang, Hongbin" docDate="2015" docId="039787812E48FFC494FD4E1AEBDFE9F0" docLanguage="en" docName="zt03920p039.pdf" docOrigin="Zootaxa 3920 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Harpalus (Pseudoophonus) capito Morawitz 1862" docType="treatment" docVersion="7" lastPageNumber="6" masterDocId="FFAEFFF92E4DFFC1946A4A2BE927EF1A" masterDocTitle="Taxonomic review of Chinese species of ground beetles of the subgenus Pseudoophonus (genus Harpalus) (Coleoptera: Carabidae)" masterLastPageNumber="39" masterPageNumber="1" pageNumber="6" updateTime="1698562432667" updateUser="plazi">
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<mods:title id="CED3B99F2A35238D3C53329F69784E38">Taxonomic review of Chinese species of ground beetles of the subgenus Pseudoophonus (genus Harpalus) (Coleoptera: Carabidae)</mods:title>
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<mods:namePart id="91E55A724CA30A9B452D1273CF40A923">Kataev, Boris M.</mods:namePart>
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<mods:namePart id="2BE5A068E53B69E82950F95D8C44FE5B">Liang, Hongbin</mods:namePart>
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<mods:date id="77E01DF861B14A413B041F7007C24CF0">2015</mods:date>
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<mods:number id="A1875030A4C138C088023B459935FAEB">3920</mods:number>
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<treatment id="039787812E48FFC494FD4E1AEBDFE9F0" ID-DOI="http://doi.org/10.5281/zenodo.6114240" ID-GBIF-Taxon="127671656" ID-Zenodo-Dep="6114240" LSID="urn:lsid:plazi:treatment:039787812E48FFC494FD4E1AEBDFE9F0" httpUri="http://treatment.plazi.org/id/039787812E48FFC494FD4E1AEBDFE9F0" lastPageNumber="6" pageId="5" pageNumber="6">
<subSubSection id="C324651C2E48FFC494FD4E1AE802EB76" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="8B8136972E48FFC494FD4E1AEBD8EB51" blockId="5.[151,767,1073,1133]" box="[151,767,1073,1099]" pageId="5" pageNumber="6">
<heading id="D0C981FB2E48FFC494FD4E1AEBD8EB51" bold="true" box="[151,767,1073,1099]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
<taxonomicName id="4C3E4D142E48FFC494FD4E1AEBD8EB51" ID-CoL="3JSTD" authority="Morawitz, 1862" authorityName="Morawitz" authorityYear="1862" box="[151,767,1073,1099]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="capito" subGenus="Pseudoophonus">
<emphasis id="B94AEA852E48FFC494FD4E1AEBD8EB51" bold="true" box="[151,767,1073,1099]" pageId="5" pageNumber="6">
<emphasis id="B94AEA852E48FFC494FD4E1AE82EEB51" bold="true" box="[151,265,1073,1099]" italics="true" pageId="5" pageNumber="6">Harpalus</emphasis>
(
<emphasis id="B94AEA852E48FFC495704E1AE8FDEB51" bold="true" box="[282,474,1073,1099]" italics="true" pageId="5" pageNumber="6">Pseudoophonus</emphasis>
)
<emphasis id="B94AEA852E48FFC495814E1AEB12EB51" bold="true" box="[491,565,1073,1099]" italics="true" pageId="5" pageNumber="6">capito</emphasis>
Morawitz, 1862
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8136972E48FFC494FD4E7FE802EB76" blockId="5.[151,767,1073,1133]" box="[151,293,1108,1133]" pageId="5" pageNumber="6">
(
<figureCitation id="13052A122E48FFC494F54E7FE83AEB76" box="[159,285,1108,1133]" captionStart="FIGURES 11 – 14" captionStartId="6.[151,263,1553,1576]" captionTargetBox="[220,1362,388,1520]" captionTargetId="figure@6.[212,1374,379,1532]" captionTargetPageId="6" captionText="FIGURES 11 – 14. Harpalus, median lobe of aedeagus. 11 – 12, H. capito (Heilongjiang); 13 – 14, H. eous (Heilongjiang). 11, 13, dorsal view; 12, 14, view from the left side. Scale = 1.0 mm." httpUri="https://zenodo.org/record/287887/files/figure.png" pageId="5" pageNumber="6">Figs 11–12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E48FFC494FD4EB7EAA7EBCB" pageId="5" pageNumber="6" type="reference_group">
<paragraph id="8B8136972E48FFC494FD4EB7EAA7EBCB" blockId="5.[151,896,1180,1233]" pageId="5" pageNumber="6">
<treatmentCitationGroup id="AB2E11B92E48FFC494FD4EB7EAA7EBCB" pageId="5" pageNumber="6">
<taxonomicName id="4C3E4D142E48FFC494FD4EB7EB02EBA8" ID-CoL="3JSTD" authority="Morawitz, 1862a: 220" authorityName="Morawitz" authorityPageNumber="220" authorityYear="1862" box="[151,549,1180,1202]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="capito">
<emphasis id="B94AEA852E48FFC494FD4EB7E81BEBA8" box="[151,316,1180,1202]" italics="true" pageId="5" pageNumber="6">Harpalus capito</emphasis>
<treatmentCitation id="0A9F10862E48FFC495284EB7EB02EBA8" author="Morawitz" box="[322,549,1180,1202]" page="220" pageId="5" pageNumber="6" year="1862">
<bibRefCitation id="EFAF4B662E48FFC495284EB7E8D4EBA8" author="Morawitz" box="[322,499,1180,1202]" pageId="5" pageNumber="6" refString="Morawitz, A. (1862 a) Vorlaufige Diagnosen neuer Coleopteren aus Sudost-Sibirien. Bulletin de l'Academie Imperiale des Sciences de St. - Petersbourg, 5, 231 - 265." type="journal article" year="1862" yearSuffix="a">Morawitz, 1862a</bibRefCitation>
: 220
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="548588352E48FFC496454EB6EB46EBA8" box="[559,609,1181,1202]" pageId="5" pageNumber="6">Type</typeStatus>
locality: &quot;Ussuri&quot;,
<collectingCountry id="F32976072E48FFC4974E4EB7EA4CEBA8" box="[804,875,1180,1202]" name="Russia" pageId="5" pageNumber="6">Russia</collectingCountry>
.
<taxonomicName id="4C3E4D142E48FFC494FD4E90EBAAEBCB" ID-CoL="8KVHF" authority="Motschulsky, 1861: 3" authorityName="Motschulsky" authorityPageNumber="3" authorityYear="1861" box="[151,653,1211,1233]" class="Insecta" family="Carabidae" genus="Pseudoophonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="cephalotes">
<emphasis id="B94AEA852E48FFC494FD4E90E88EEBCA" box="[151,425,1211,1232]" italics="true" pageId="5" pageNumber="6">Pseudoophonus cephalotes</emphasis>
<treatmentCitation id="0A9F10862E48FFC495DA4E90EBAAEBCB" author="Motschulsky" box="[432,653,1211,1233]" page="3" pageId="5" pageNumber="6" year="1861">
<bibRefCitation id="EFAF4B662E48FFC495DA4E90EB53EBCB" author="Motschulsky" box="[432,628,1211,1233]" pageId="5" pageNumber="6" refString="Motschulsky, V. (1861) Insectes du Japon. Etudes Entomologiques, 10, 3 - 24." type="journal article" year="1861">Motschulsky, 1861</bibRefCitation>
: 3
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="548588352E48FFC496FD4E97EBEEEBCB" box="[663,713,1212,1233]" pageId="5" pageNumber="6">Type</typeStatus>
locality: &quot;
<collectingCountry id="F32976072E48FFC497584E90EA54EBCB" box="[818,883,1211,1233]" name="Japan" pageId="5" pageNumber="6">Japon</collectingCountry>
&quot;.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E48FFC494FD4ED6ED09EA44" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="8B8136972E48FFC494FD4ED6EB6BEA20" blockId="5.[151,1437,1277,1770]" pageId="5" pageNumber="6">
<emphasis id="B94AEA852E48FFC494FD4ED6E819EA0C" bold="true" box="[151,318,1277,1302]" pageId="5" pageNumber="6">
<typeStatus id="548588352E48FFC494FD4ED6E9F5EA0C" box="[151,210,1277,1302]" pageId="5" pageNumber="6">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="548588352E48FFC495274ED5E89AEA0C" box="[333,445,1278,1302]" pageId="5" pageNumber="6" type="lectotype">Lectotype</typeStatus>
(present designation) of
<taxonomicName id="4C3E4D142E48FFC496B34ED6EAB6EA0F" box="[729,913,1277,1301]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="capito">
<emphasis id="B94AEA852E48FFC496B34ED6EAB6EA0F" box="[729,913,1277,1301]" italics="true" pageId="5" pageNumber="6">Harpalus capito</emphasis>
</taxonomicName>
: ♂, labeled &quot;Ussuri, 223–1&quot;, and &quot;223.
<taxonomicName id="4C3E4D142E48FFC4913C4ED5E9F7EA20" authority="Mor." authorityName="Mor." class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="capito">
<emphasis id="B94AEA852E48FFC4913C4ED5ECBBEA0F" box="[1366,1436,1278,1301]" italics="true" pageId="5" pageNumber="6">capito</emphasis>
Mor.
</taxonomicName>
&quot; [Morawitz's handwriting] (
<collectionCode id="ED2FAE522E48FFC496664F09EB67EA20" box="[524,576,1314,1338]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="5" pageNumber="6">ZIN</collectionCode>
).
</paragraph>
<paragraph id="8B8136972E48FFC494AD4F6EED09EA44" blockId="5.[151,1437,1277,1770]" box="[199,1070,1349,1374]" pageId="5" pageNumber="6">
<emphasis id="B94AEA852E48FFC494AD4F6EEB0BEA44" bold="true" box="[199,556,1349,1374]" pageId="5" pageNumber="6">Additional material examined</emphasis>
. More than
<specimenCount id="9D38FD1E2E48FFC496DF4F6DEA7CEA44" box="[693,859,1349,1374]" pageId="5" pageNumber="6" type="generic">380 specimens</specimenCount>
(161 from
<collectingCountry id="F32976072E48FFC497B04F6EED05EA44" box="[986,1058,1349,1374]" name="China" pageId="5" pageNumber="6">China</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E48FFC494AD4F42EA21EAF7" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="8B8136972E48FFC494AD4F42EA21EAF7" blockId="5.[151,1437,1277,1770]" pageId="5" pageNumber="6">
<emphasis id="B94AEA852E48FFC494AD4F42E81DEA98" bold="true" box="[199,314,1385,1410]" pageId="5" pageNumber="6">Diagnosis</emphasis>
. This species is distinctly distinguished from other members of
<taxonomicName id="4C3E4D142E48FFC490754F42EDF7EA9B" box="[1055,1232,1385,1409]" class="Insecta" family="Carabidae" genus="Pseudoophonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA852E48FFC490754F42EDF7EA9B" box="[1055,1232,1385,1409]" italics="true" pageId="5" pageNumber="6">Pseudoophonus</emphasis>
</taxonomicName>
by having elytral intervals 1, 3, 5 and 7 with a row of discal pores, and pronotal basal margin not bordered. Ligular sclerite with apical plate and with several dorsal setae. Median lobe of aedeagus (
<figureCitation id="13052A122E48FFC497CD4F9AED0FEAD0" box="[935,1064,1457,1482]" captionStart="FIGURES 11 – 14" captionStartId="6.[151,263,1553,1576]" captionTargetBox="[220,1362,388,1520]" captionTargetId="figure@6.[212,1374,379,1532]" captionTargetPageId="6" captionText="FIGURES 11 – 14. Harpalus, median lobe of aedeagus. 11 – 12, H. capito (Heilongjiang); 13 – 14, H. eous (Heilongjiang). 11, 13, dorsal view; 12, 14, view from the left side. Scale = 1.0 mm." httpUri="https://zenodo.org/record/287887/files/figure.png" pageId="5" pageNumber="6">Figs 11–12</figureCitation>
) without any distinct sclerotized element in the internal sac. Body length 17.3–24.0 mm.
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E48FFC494AD4FD2EBDFE9F0" pageId="5" pageNumber="6" type="distribution">
<paragraph id="8B8136972E48FFC494AD4FD2EA42E940" blockId="5.[151,1437,1277,1770]" pageId="5" pageNumber="6">
<emphasis id="B94AEA852E48FFC494AD4FD2E87EE908" bold="true" box="[199,345,1529,1554]" pageId="5" pageNumber="6">Distribution</emphasis>
.
<collectingCountry id="F32976072E48FFC495034FD2E897E908" box="[361,432,1529,1554]" name="China" pageId="5" pageNumber="6">China</collectingCountry>
: the north-eastern and eastern parts; known from Heilongjiang, Jilin, Liaoning, Beijing, Hebei, Jiangsu,
<collectingCountry id="F32976072E48FFC495254C36E886E92C" box="[335,417,1565,1590]" name="Taiwan" pageId="5" pageNumber="6">Taiwan</collectingCountry>
(
<bibRefCitation id="EFAF4B662E48FFC495D84C35EBAAE92C" author="Kataev" box="[434,653,1565,1590]" pageId="5" pageNumber="6" refString="Kataev, B. M., Wrase, D. W. &amp; Ito, N. (2003) Harpalina. In: Lobl, L. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup, pp. 367 - 397." type="book" year="2003">
Kataev
<emphasis id="B94AEA852E48FFC496674C35EB66E92F" box="[525,577,1565,1589]" italics="true" pageId="5" pageNumber="6">et al</emphasis>
. 2003
</bibRefCitation>
), Nei Mongol, Shanxi, Ningxia, Shaanxi, Gansu, Henan, Shandong, Hubei, Anhui, Jiangxi, Zhejiang, Fujian and Hunan (
<bibRefCitation id="EFAF4B662E48FFC4968F4C69EA7EE940" author="Hua" box="[741,857,1602,1626]" pageId="5" pageNumber="6" refString="Hua, L. (2002) List of Chinese Insects. Vol. II. Zhongshan (Sun Yat-sen) University Press, Guangzhou, 612 pp." type="book" year="2002">Hua 2002</bibRefCitation>
).
</paragraph>
<paragraph id="8B8136972E48FFC494AD4C4EEBDFE9F0" blockId="5.[151,1437,1277,1770]" pageId="5" pageNumber="6">
The species is also distributed in the south of the Russian Far East (Amur Province, Khabarovsk Territory and Maritime Province, the south of Sakhalin Island, and Kunashir Island), North and
<collectingCountry id="F32976072E48FFC490284CA2EDF6E9BB" box="[1090,1233,1673,1698]" name="South Korea" pageId="5" pageNumber="6">South Korea</collectingCountry>
including Jejudo, and
<collectingCountry id="F32976072E48FFC494AD4C85E820E9DF" box="[199,263,1710,1733]" name="Japan" pageId="5" pageNumber="6">Japan</collectingCountry>
(Hokkaido, Honshu, Sikoku, and Kyushu Islands) (
<bibRefCitation id="EFAF4B662E48FFC497394C86EAF3E9DC" author="Habu" box="[851,980,1709,1734]" pageId="5" pageNumber="6" refString="Habu, A. (1973) Carabidae: Harpalini (Insecta, Coleoptera). Fauna Japonica. Keigaku Publishing Co., Tokio, xiii + 430 pp." type="book" year="1973">Habu 1973</bibRefCitation>
;
<bibRefCitation id="EFAF4B662E48FFC497884C86ED46E9DC" author="Lafer" box="[994,1121,1709,1734]" pageId="5" pageNumber="6" refString="Lafer, G. Sh. (1989) Fam. Carabidae-The ground-beetles. In: Lehr, P. A. (Ed.), Opredelitel' nasekomykh Dal'nego Vostoka SSSR v shesti tomakh. Tom III. Zhestkokrylye, ili zhuki. Chast' 1. Nauka, Leningrad, pp. 70 - 222. [in Russian]" type="journal article" year="1989">Lafer 1989</bibRefCitation>
;
<bibRefCitation id="EFAF4B662E48FFC490044C86ECB0E9DC" author="Kryzhanovskij" box="[1134,1431,1709,1734]" pageId="5" pageNumber="6" refString="Kryzhanovskij, O. L., Belousov, I. A., Kabak, I. I., Kataev, B. M., Makarov, K. V., Shilenkov, V. G. (1995) A checklist of the groundbeetles of Russia and adjacent lands (Insecta, Coleoptera, Carabidae). Pensoft Publishers, Sofia - Moscow, 272 pp." type="book" year="1995">
Kryzhanovskij
<emphasis id="B94AEA852E48FFC491714C85EC6AE9DF" box="[1307,1357,1709,1733]" italics="true" pageId="5" pageNumber="6">et al</emphasis>
. 1995
</bibRefCitation>
;
<bibRefCitation id="EFAF4B662E48FFC494FD4CF9E80FE9F0" author="Kataev" box="[151,296,1746,1770]" pageId="5" pageNumber="6" refString="Kataev, B. M. (1997) Ground beetles of the genus Harpalus (Insecta, Coleoptera, Carabidae) from East Asia. Steenstrupia, 23, 123 - 160." type="journal article" year="1997">Kataev 1997</bibRefCitation>
;
<bibRefCitation id="EFAF4B662E48FFC4955E4CF9EB22E9F0" author="Kataev" box="[308,517,1745,1770]" pageId="5" pageNumber="6" refString="Kataev, B. M., Wrase, D. W. &amp; Ito, N. (2003) Harpalina. In: Lobl, L. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup, pp. 367 - 397." type="book" year="2003">
Kataev
<emphasis id="B94AEA852E48FFC495E04CF9E89DE9F3" box="[394,442,1745,1769]" italics="true" pageId="5" pageNumber="6">et al</emphasis>
. 2003
</bibRefCitation>
; Moon &amp;
<bibRefCitation id="EFAF4B662E48FFC4961C4CFAEBCBE9F0" author="Paik" box="[630,748,1745,1770]" pageId="5" pageNumber="6" refString="Moon, Ch. - S. &amp; Paik, J. - Ch. (2006) Notes on Harpalini (Coleoptera, Carabidae) from Korea (1 - 3). Korean Journal of Soil Zoology, 11 (1 - 2), 30 - 84." type="journal article" year="2006">Paik 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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