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<document id="E76C1CAFB558DB7F8249F8B425BFC071" ID-CLB-Dataset="48132" ID-DOI="10.5281/zenodo.207073" ID-GBIF-Dataset="a3815507-6100-464e-aaf6-60d6820a1eec" ID-ISSN="1175-5326" ID-Zenodo-Dep="207073" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460228776049" checkinUser="plazi" docAuthor="Torres-Carvajal, Omar, Etheridge, Richard &amp; Queiroz, Kevin De" docDate="2011" docId="038287FDFFA3605D69838DC9D0F2F9EA" docLanguage="en" docName="zt02752p044.pdf" docOrigin="Zootaxa 2752" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Enyalioides palpebralis Boulenger 1883" docType="treatment" docVersion="11" lastPageNumber="27" masterDocId="FFBBFF85FFBB604769148B6ED174FF93" masterDocTitle="A systematic revision of Neotropical lizards in the clade Hoplocercinae (Squamata: Iguania)" masterLastPageNumber="44" masterPageNumber="1" pageNumber="25" updateTime="1720202091178" updateUser="ExternalLinkService">
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<mods:title id="D6E51906022EB9C5C543E802B3A94406">A systematic revision of Neotropical lizards in the clade Hoplocercinae (Squamata: Iguania)</mods:title>
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<mods:namePart id="40E7D4CD08F6BD2CAA4913526573BE16">Torres-Carvajal, Omar</mods:namePart>
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<mods:namePart id="8BA8DAE3DD117C45662C483EC50BBEF9">Etheridge, Richard</mods:namePart>
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<mods:namePart id="FB5A726FD6FA572AE9712CC69D58B17B">Queiroz, Kevin De</mods:namePart>
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<treatment id="038287FDFFA3605D69838DC9D0F2F9EA" ID-DOI="http://doi.org/10.5281/zenodo.6184967" ID-GBIF-Taxon="119387050" ID-Zenodo-Dep="6184967" LSID="urn:lsid:plazi:treatment:038287FDFFA3605D69838DC9D0F2F9EA" httpUri="http://treatment.plazi.org/id/038287FDFFA3605D69838DC9D0F2F9EA" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25">
<subSubSection id="C3316560FFA3605F69838DC9D3CCF96F" pageId="24" pageNumber="25" type="nomenclature">
<paragraph id="8B9436EBFFA3605F69838DC9D3FAF952" blockId="24.[151,654,1703,1729]" box="[151,654,1703,1729]" pageId="24" pageNumber="25">
<heading id="D0DC8187FFA3605F69838DC9D3FAF952" bold="true" box="[151,654,1703,1729]" fontSize="11" level="1" pageId="24" pageNumber="25" reason="1">
<taxonomicName id="4C2B4D68FFA3605F69838DC9D3FAF952" ID-CoL="6FH5Z" authority="Boulenger 1883" authorityName="Boulenger" authorityYear="1883" box="[151,654,1703,1729]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="palpebralis">
<emphasis id="B95FEAF9FFA3605F69838DC9D3FAF952" bold="true" box="[151,654,1703,1729]" pageId="24" pageNumber="25">
<emphasis id="B95FEAF9FFA3605F69838DC9D0C5F952" bold="true" box="[151,433,1703,1729]" italics="true" pageId="24" pageNumber="25">Enyalioides palpebralis</emphasis>
(
<bibRefCitation id="EFBA4B1AFFA3605F68D48DC9D3F2F952" author="Boulenger" box="[448,646,1703,1729]" pageId="24" pageNumber="38" refString="Boulenger, G. A. (1883) Description of a new species of lizard of the genus Enyalius. Proceedings of the Zoological Society of London, 1883, 46, 1 plate." type="book" year="1883">Boulenger 1883</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B9436EBFFA3605F69838D88D3CCF96F" blockId="24.[151,768,1766,1819]" box="[151,696,1766,1788]" pageId="24" pageNumber="25">Proposed standard English name: horned woodlizards</paragraph>
</subSubSection>
<subSubSection id="C3316560FFA3605F69838C6BD274F888" box="[151,768,1797,1819]" pageId="24" pageNumber="25" type="description">
<paragraph id="8B9436EBFFA3605F69838C6BD274F888" blockId="24.[151,768,1766,1819]" box="[151,768,1797,1819]" pageId="24" pageNumber="25">Proposed standard Spanish name: lagartijas de palo cornudas</paragraph>
</subSubSection>
<subSubSection id="C3316560FFA3605F69838C55D597F81C" pageId="24" pageNumber="25" type="reference_group">
<paragraph id="8B9436EBFFA3605F69838C55D385F8E3" blockId="24.[151,1436,1851,1935]" pageId="24" pageNumber="25">
<taxonomicName id="4C2B4D68FFA3605F69838C55D33BF8C2" authority="Boulenger (1883:46)" authorityName="Boulenger" authorityPageNumber="46" authorityYear="1883" box="[151,591,1851,1873]" class="Reptilia" family="Leiosauridae" genus="Enyalius" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="palpebralis">
<emphasis id="B95FEAF9FFA3605F69838C55D018F8C2" box="[151,364,1851,1873]" italics="true" pageId="24" pageNumber="25">Enyalius palpebralis</emphasis>
<bibRefCitation id="EFBA4B1AFFA3605F68618C55D33BF8C2" author="Boulenger" box="[373,591,1851,1873]" pageId="24" pageNumber="38" refString="Boulenger, G. A. (1883) Description of a new species of lizard of the genus Enyalius. Proceedings of the Zoological Society of London, 1883, 46, 1 plate." type="book" year="1883">Boulenger (1883:46)</bibRefCitation>
</taxonomicName>
.
<typeStatus id="54908849FFA3605F6B4F8C55D3C9F8C2" box="[603,701,1851,1873]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
: BMNH 81.5.13.25 (RR 1946.8.9.8), from “Cashiboya [Departamento Loreto,
<geoCoordinate id="EE1F502CFFA3605F68038C34D00AF8E3" box="[279,382,1882,1904]" direction="south" orientation="latitude" pageId="24" pageNumber="25" precision="15" value="-7.55">7°32'60''S</geoCoordinate>
,
<geoCoordinate id="EE1F502CFFA3605F689D8C34D370F8E3" box="[393,516,1882,1904]" direction="west" orientation="longitude" pageId="24" pageNumber="25" precision="15" value="-74.88334">74°52'60''W</geoCoordinate>
,
<quantity id="4CD39B0EFFA3605F6B1B8C34D327F8E3" box="[527,595,1882,1904]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="24" pageNumber="25" unit="m" value="160.0">160 m</quantity>
], eastern
<collectingCountry id="F33C767BFFA3605F6BA68C35D391F8E3" box="[690,741,1883,1904]" name="Peru" pageId="24" pageNumber="25">Peru</collectingCountry>
.”
</paragraph>
<paragraph id="8B9436EBFFA3605F69838C17D597F81C" blockId="24.[151,1436,1851,1935]" box="[151,1251,1913,1935]" pageId="24" pageNumber="25">
<taxonomicName id="4C2B4D68FFA3605F69838C17D305F81C" authority="Boulenger (1885:116)" authorityName="Boulenger" authorityPageNumber="116" authorityYear="1885" box="[151,625,1913,1935]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="palpebralis">
<emphasis id="B95FEAF9FFA3605F69838C17D0F3F81C" box="[151,391,1913,1935]" italics="true" pageId="24" pageNumber="25">Enyalioides palpebralis</emphasis>
<bibRefCitation id="EFBA4B1AFFA3605F689A8C17D305F81C" author="Boulenger" box="[398,625,1913,1935]" pageId="24" pageNumber="38" refString="Boulenger, G. A. (1885) Catalogue of the Lizards in the British Museum (Natural History) II. Taylor and Francis, London, 497 pp." type="book" year="1885">Boulenger (1885:116)</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="EFBA4B1AFFA3605F6B6F8C14D216F81C" author="Burt" box="[635,866,1913,1935]" pageId="24" pageNumber="38" refString="Burt, C. E. &amp; Burt, M. D. (1933) A preliminary check list of the lizards of South America. Transactions of the Academy of Sciences of St Louis, 28, 1 - 104." type="journal article" year="1933">Burt &amp; Burt (1933:24)</bibRefCitation>
;
<bibRefCitation id="EFBA4B1AFFA3605F6A798C14D5ABF81C" author="Peters" box="[877,1247,1913,1935]" pageId="24" pageNumber="40" refString="Peters, J. A. &amp; Donoso-Barros, R. (1970) Catalogue of the neotropical Squamata: Part II. Lizards and amphisbaenians. United States National Museum Bulletin, 297, 1 - 293." type="journal article" year="1970">Peters &amp; Donoso-Barros (1970:115)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3316560FFA3605E69838CDBD43BFF64" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="8B9436EBFFA3605E69838CDBD43BFF64" blockId="24.[151,1436,1973,2035]" lastBlockId="25.[151,1436,151,248]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
<emphasis id="B95FEAF9FFA3605F69838CDBD065F85D" bold="true" box="[151,273,1973,1998]" pageId="24" pageNumber="25">Diagnosis.</emphasis>
This species is different from other species of
<taxonomicName id="4C2B4D68FFA3605F6A328CD8D2DEF85D" box="[806,938,1974,1998]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="genus">
<emphasis id="B95FEAF9FFA3605F6A328CD8D2DEF85D" box="[806,938,1974,1998]" italics="true" pageId="24" pageNumber="25">Enyalioides</emphasis>
</taxonomicName>
in having a superciliary triangular flap projecting posterolaterally over each eye; a wide postorbital process of the squamosal (process narrow in other species); a pair of enlarged dorsal tubercles on the posterolateral aspect of the parietal roof; and in lacking a posteriorly projecting squamosal process of the postorbital (
<figureCitation id="13102A6EFFA2605E6BA78BD2D383FF47" box="[691,759,188,212]" captionStart="FIGURE 8" captionStartId="25.[151,250,716,739]" captionTargetBox="[246,1342,305,693]" captionTargetId="figure@25.[243,1344,301,695]" captionTargetPageId="25" captionText="FIGURE 8. Skulls of Enyalioides palpebralis (left) and E. laticeps (right) in lateral view. dt = dorsal tubercle of parietal bone; po = postorbital; sq = squamosal." httpUri="https://zenodo.org/record/207085/files/figure.png" pageId="25" pageNumber="26">Fig. 8</figureCitation>
;
<bibRefCitation id="EFBA4B1AFFA2605E6A178BD2D568FF47" author="Wiens" box="[771,1052,188,212]" pageId="25" pageNumber="41" refString="Wiens, J. J. &amp; Etheridge, R. (2003) Phylogenetic relationships of hoplocercid lizards: Coding and combining meristic, morphometric, and polymorphic data using step matrices. Herpetologica, 59, 375 - 398." type="journal article" year="2003">Wiens &amp; Etheridge 2003</bibRefCitation>
). In addition most specimens of
<taxonomicName id="4C2B4D68FFA2605E6C908BD3D063FF6B" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="palpebralis">
<emphasis id="B95FEAF9FFA2605E6C908BD3D063FF6B" italics="true" pageId="25" pageNumber="26">E. palpebralis</emphasis>
</taxonomicName>
have a discontinuous vertebral crest, with a small gap on the neck, and most lack femoral pores.
</paragraph>
</subSubSection>
<caption id="DF546663FFA2605E698389A2D09CFC92" httpUri="https://zenodo.org/record/207085/files/figure.png" pageId="25" pageNumber="26" targetBox="[246,1342,305,693]" targetPageId="25">
<paragraph id="8B9436EBFFA2605E698389A2D09CFC92" blockId="25.[151,1436,716,769]" pageId="25" pageNumber="26">
<emphasis id="B95FEAF9FFA2605E698389A2D066FD71" bold="true" box="[151,274,716,739]" pageId="25" pageNumber="26">FIGURE 8.</emphasis>
Skulls of
<taxonomicName id="4C2B4D68FFA2605E686E89A3D31FFD70" box="[378,619,717,739]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="palpebralis">
<emphasis id="B95FEAF9FFA2605E686E89A3D31FFD70" box="[378,619,717,739]" italics="true" pageId="25" pageNumber="26">Enyalioides palpebralis</emphasis>
</taxonomicName>
(left) and
<taxonomicName id="4C2B4D68FFA2605E6BC289A0D235FD70" box="[726,833,717,739]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="laticeps">
<emphasis id="B95FEAF9FFA2605E6BC289A0D235FD70" box="[726,833,717,739]" italics="true" pageId="25" pageNumber="26">E. laticeps</emphasis>
</taxonomicName>
(right) in lateral view. dt = dorsal tubercle of parietal bone; po = postorbital; sq = squamosal.
</paragraph>
</caption>
<subSubSection id="C3316560FFA2605E69D18843D3F6F8A5" pageId="25" pageNumber="26" type="description">
<paragraph id="8B9436EBFFA2605E69D18843D0F4FA89" blockId="25.[151,1437,813,1955]" pageId="25" pageNumber="26">
<emphasis id="B95FEAF9FFA2605E69D18843D023FCD5" bold="true" box="[197,343,813,838]" pageId="25" pageNumber="26">Description.</emphasis>
(1) dorsal head scales conical or multicarinate, strongly projecting dorsally; (2) posterior superciliaries enlarged, pointed, and projecting laterally; (3) scales on lateral edge of skull roof just posterior to superciliaries strongly projecting; the projection is more pronounced in adults; (4) one or two enlarged pretympanic scales present; (5) gular scales conical or multicarinate, strongly projecting ventrally; (6) dorsal and lateral neck scales heterogeneous in size, granular and conical scales of various sizes present; (7) vertebrals larger than adjacent dorsals, forming distinct raised middorsal crest that usually (85.9%) extends onto tail as a pair of crests; (8) nuchal region with discontinuous (85.7%) and single middorsal crest; (9) dorsals usually (71.4%) keeled and heterogeneous in size (85.7%); (10) discontinuous longitudinal row of large conical scales between dorsals and flank scales present; (11) scales on flanks keeled or smooth, heterogeneous in size; largest scales are conical in some specimens; (12) ventrals keeled; (13) fore limb scales mostly keeled dorsally and ventrally; (14) hind limb scales mostly keeled dorsally and ventrally; scattered enlarged scales present dorsally; dorsal scales of pes homogeneous in size; (15) caudals heterogeneous, slightly increasing in size posteriorly on each segment (4–6 scales in lateral view), not modified as conspicuous spines (
<figureCitation id="13102A6EFFA2605E6B1A8FB0D326FB65" box="[526,594,1246,1270]" captionStart="FIGURE 3" captionStartId="12.[151,250,1517,1540]" captionTargetBox="[280,1305,321,1491]" captionTargetId="figure@12.[274,1314,306,1505]" captionTargetPageId="12" captionText="FIGURE 3. Caudal segments from the anterior third of the tail of ten species of Hoplocercinae in lateral view. (A) Enyalioides cofanorum, (B) E. heterolepis, (C) E. laticeps, (D) E. microlepis, (E) E. oshaughnessyi, (F) E. palpebralis, (G) E. praestabilis, (H) E. rubrigularis, (I) E. touzeti, and (J) Morunasaurus annularis. Scale bars = 5 mm. Illustration by D. Paucar." httpUri="https://zenodo.org/record/207076/files/figure.png" pageId="25" pageNumber="26">Fig. 3</figureCitation>
); (16) tail compressed laterally. Meristic and morphometric characters are presented in
<tableCitation id="C6A90350FFA2605E68328E6CD00EFA89" box="[294,378,1282,1306]" captionStart="TABLE 1" captionStartId="6.[151,235,1304,1325]" captionTargetBox="[151,1436,1377,1892]" captionTargetPageId="6" captionText="TABLE 1. Summary of counts and measurements (mm) for species of Hoplocercinae. Range (first line) and mean ± standard" httpUri="http://table.plazi.org/id/DF546663FFBD604169838E76D4E8FABD" pageId="25" pageNumber="26" tableUuid="DF546663FFBD604169838E76D4E8FABD">Table 1</tableCitation>
.
</paragraph>
<paragraph id="8B9436EBFFA2605E69D18E48D38BF985" blockId="25.[151,1437,813,1955]" pageId="25" pageNumber="26">
<emphasis id="B95FEAF9FFA2605E69D18E48D0E5FAAC" bold="true" box="[197,401,1318,1343]" pageId="25" pageNumber="26">Coloration in life</emphasis>
(
<figureCitation id="13102A6EFFA2605E68B48E48D092FAAD" box="[416,486,1318,1342]" captionStart="FIGURE 4" captionStartId="14.[151,250,1897,1920]" captionTargetBox="[204,1383,190,1874]" captionTargetId="figure@14.[204,1384,184,1876]" captionTargetPageId="14" captionText="FIGURE 4. Photographs of live lizards representing twelve species of Hoplocercinae." httpUri="https://zenodo.org/record/207077/files/figure.png" pageId="25" pageNumber="26">Fig. 4</figureCitation>
). Dorsal background yellowish brown, dark brown, or greenish beige, with tan spots (males) or black marks forming reticulate bands; neck and shoulders with scattered dark brown spots; orange or red blotch on each side of neck in both sexes usually present; elongate tan blotch or irregular stripe extends from the dorsolateral surface of the head to a point dorsal to the insertion of the arm in females; flanks with reddish-brown reticulate pattern on greenish-gray background; venter white, whitish brown, or beige, with irregular black markings in juveniles; throat yellow in males and brownish in females; iris reddish tan (
<bibRefCitation id="EFBA4B1AFFA2605E6D4B8EB4D587FA61" author="Meede" box="[1119,1267,1498,1522]" pageId="25" pageNumber="40" refString="Meede, U. (1984) Herpetologische Studien uber Echsen (Sauria) in einem begrenzten Gebiet des Tropischen Regenwaldes in Peru: morphologische Kriterien, Autokologie und Zoogeographie. Artenliste der Reptilien im Untersuchungsgebiet., p. 1 - 189. Vol. Doctoral dissertation. Universitat Hamburg." type="book chapter" year="1984">Meede 1984</bibRefCitation>
;
<bibRefCitation id="EFBA4B1AFFA2605E6C158EB4D1A7F985" author="Reichle" pageId="25" pageNumber="40" refString="Reichle, S., Cortez, C., Padial, J. M. &amp; de la Riva, I. (2004) Notes on Enyalioides palpebralis (Boulenger, 1883) (Reptilia: Squamata: Hoplocercidae) in Bolivia. Boletin de la Asociacion Herpetologica Espanola, 15, 14 - 16." type="journal article" year="2004">
Reichle
<emphasis id="B95FEAF9FFA2605E6C758EB5D4E8FA61" box="[1377,1436,1498,1522]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
2004
</bibRefCitation>
;
<bibRefCitation id="EFBA4B1AFFA2605E69C98E90D0E4F985" author="Duellman" box="[221,400,1534,1558]" pageId="25" pageNumber="39" refString="Duellman, W. E. (2005) Cusco Amazonico, the lives of amphibian and reptiles in an Amazonian rainforest. Cornell University Press, Ithaca, xvi + 433 pp." type="book" year="2005">Duellman 2005</bibRefCitation>
); lining of mouth orange (
<figureCitation id="13102A6EFFA2605E6BA08E90D387F985" box="[692,755,1534,1558]" captionStart="FIGURE 4" captionStartId="14.[151,250,1897,1920]" captionTargetBox="[204,1383,190,1874]" captionTargetId="figure@14.[204,1384,184,1876]" captionTargetPageId="14" captionText="FIGURE 4. Photographs of live lizards representing twelve species of Hoplocercinae." httpUri="https://zenodo.org/record/207077/files/figure.png" pageId="25" pageNumber="26">Fig.4</figureCitation>
).
</paragraph>
<paragraph id="8B9436EBFFA2605E69D18D4CD3F6F8A5" blockId="25.[151,1437,813,1955]" pageId="25" pageNumber="26">
<emphasis id="B95FEAF9FFA2605E69D18D4CD0F2F9A9" bold="true" box="[197,390,1569,1594]" pageId="25" pageNumber="26">Natural history.</emphasis>
Based on observations of specimens in captivity, Schulte (1998) described fights between males in which one combatant bites the other’s superciliary flap or nuchal crest and shakes it strongly; a male will also bite and hold a female by her nuchal crests during copulation. Clutch consists of 2–
<specimenCount id="9D2DFD62FFA2605E6D7B8D04D509F911" box="[1135,1149,1642,1666]" pageId="25" pageNumber="26" type="egg">4</specimenCount>
eggs (ca.
<date id="FF95102BFFA2605E6DFA8D04D406F911" box="[1262,1394,1642,1666]" pageId="25" pageNumber="26" value="1912-10-24" valueMax="1912-10-28" valueMin="1912-10-24">
24–28 X
<quantity id="4CD39B0EFFA2605E6C428D04D406F911" box="[1366,1394,1642,1666]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.8" metricValueMin="1.2" pageId="25" pageNumber="26" unit="mm" value="15.0" valueMax="18.0" valueMin="12.0">12</quantity>
</date>
–18 mm), which are laid under leaf-litter (
<bibRefCitation id="EFBA4B1AFFA2605E6B2E8DE0D3BEF935" author="Meede" box="[570,714,1678,1702]" pageId="25" pageNumber="40" refString="Meede, U. (1984) Herpetologische Studien uber Echsen (Sauria) in einem begrenzten Gebiet des Tropischen Regenwaldes in Peru: morphologische Kriterien, Autokologie und Zoogeographie. Artenliste der Reptilien im Untersuchungsgebiet., p. 1 - 189. Vol. Doctoral dissertation. Universitat Hamburg." type="book chapter" year="1984">Meede 1984</bibRefCitation>
; Schulte 1998). This species is a sit-and-wait predator, and it has been found in leaf-litter or perching on the bases of tree trunks (no higher than
<quantity id="4CD39B0EFFA2605E6D048DDCD526F959" box="[1040,1106,1714,1738]" metricMagnitude="0" metricUnit="m" metricValue="1.7" pageId="25" pageNumber="26" unit="m" value="1.7">1.7 m</quantity>
) in primary terra firme forest and primary forest edges, sometimes close to water (
<bibRefCitation id="EFBA4B1AFFA2605E6BE18DB8D2CCF97D" author="Avila-Pires" box="[757,952,1750,1774]" pageId="25" pageNumber="38" refString="Avila-Pires, T. C. S. (1995) Lizards of Brazilian Amazonia (Reptilia: Squamata). Nationaal Natuurhistorisch Museum Zoologische Verhandelingen, 299, 1 - 706." type="journal article" year="1995">Avila-Pires 1995</bibRefCitation>
; Schulte 1998). Prey items of
<taxonomicName id="4C2B4D68FFA2605E6C0C8DB8D063F881" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="palpebralis">
<emphasis id="B95FEAF9FFA2605E6C0C8DB8D063F881" italics="true" pageId="25" pageNumber="26">Enyalioides palpebralis</emphasis>
</taxonomicName>
include caterpillars, spiders, aquatic dipteran larvae, diplopods, chilopods, beetle larvae, grasshoppers, and ants (
<bibRefCitation id="EFBA4B1AFFA2605E68168C70D0E6F8A5" author="Meede" box="[258,402,1822,1846]" pageId="25" pageNumber="40" refString="Meede, U. (1984) Herpetologische Studien uber Echsen (Sauria) in einem begrenzten Gebiet des Tropischen Regenwaldes in Peru: morphologische Kriterien, Autokologie und Zoogeographie. Artenliste der Reptilien im Untersuchungsgebiet., p. 1 - 189. Vol. Doctoral dissertation. Universitat Hamburg." type="book chapter" year="1984">Meede 1984</bibRefCitation>
;
<bibRefCitation id="EFBA4B1AFFA2605E68898C70D302F8A5" author="Reichle" box="[413,630,1822,1846]" pageId="25" pageNumber="40" refString="Reichle, S., Cortez, C., Padial, J. M. &amp; de la Riva, I. (2004) Notes on Enyalioides palpebralis (Boulenger, 1883) (Reptilia: Squamata: Hoplocercidae) in Bolivia. Boletin de la Asociacion Herpetologica Espanola, 15, 14 - 16." type="journal article" year="2004">
Reichle
<emphasis id="B95FEAF9FFA2605E68EE8C71D346F8A5" box="[506,562,1822,1846]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
2004
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3316560FFA2605E69D18C2FD585F831" pageId="25" pageNumber="26" type="distribution">
<paragraph id="8B9436EBFFA2605E69D18C2FD585F831" blockId="25.[151,1437,813,1955]" pageId="25" pageNumber="26">
<emphasis id="B95FEAF9FFA2605E69D18C2FD02AF8C9" bold="true" box="[197,350,1857,1882]" pageId="25" pageNumber="26">Distribution.</emphasis>
<taxonomicName id="4C2B4D68FFA2605E68708C2CD31AF8C9" box="[356,622,1858,1882]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="palpebralis">
<emphasis id="B95FEAF9FFA2605E68708C2CD31AF8C9" box="[356,622,1858,1882]" italics="true" pageId="25" pageNumber="26">Enyalioides palpebralis</emphasis>
</taxonomicName>
occurs on the eastern slopes of the Andes and adjacent lowlands in
<collectingCountry id="F33C767BFFA2605E6C768C2DD4ECF8C9" box="[1378,1432,1859,1882]" name="Peru" pageId="25" pageNumber="26">Peru</collectingCountry>
, northern
<collectingCountry id="F33C767BFFA2605E69E98C08D026F8ED" box="[253,338,1894,1918]" name="Bolivia" pageId="25" pageNumber="26">Bolivia</collectingCountry>
, and western
<collectingCountry id="F33C767BFFA2605E68FF8C08D35BF8ED" box="[491,559,1894,1918]" name="Brazil" pageId="25" pageNumber="26">Brazil</collectingCountry>
at elevations between
<quantity id="4CD39B0EFFA2605E6A3B8C08D2C8F8ED" box="[815,956,1894,1918]" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="13.0" metricValueMin="1.0" pageId="25" pageNumber="26" unit="m" value="700.0" valueMax="1300.0" valueMin="100.0">100–1300 m</quantity>
(
<figureCitation id="13102A6EFFA2605E6AD88C08D564F8ED" box="[972,1040,1894,1918]" captionStart="FIGURE 9" captionStartId="26.[151,250,1386,1409]" captionTargetBox="[157,1429,206,1356]" captionTargetId="figure@26.[151,1436,193,1365]" captionTargetPageId="26" captionText="FIGURE 9. Distribution of Enyalioides palpebralis (dots) and Hoplocercus spinosus (triangles)." httpUri="https://zenodo.org/record/207086/files/figure.png" pageId="25" pageNumber="26">Fig. 9</figureCitation>
). This species is known to occur in sympatry with
<taxonomicName id="4C2B4D68FFA2605E682B8CE5D0C1F831" box="[319,437,1930,1954]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="laticeps">
<emphasis id="B95FEAF9FFA2605E682B8CE5D0C1F831" box="[319,437,1930,1954]" italics="true" pageId="25" pageNumber="26">E. laticeps</emphasis>
</taxonomicName>
in southern
<collectingCountry id="F33C767BFFA2605E6B548CE5D307F831" box="[576,627,1931,1954]" name="Peru" pageId="25" pageNumber="26">Peru</collectingCountry>
(
<bibRefCitation id="EFBA4B1AFFA2605E6B968CE4D246F831" author="Duellman" box="[642,818,1930,1954]" pageId="25" pageNumber="39" refString="Duellman, W. E. (2005) Cusco Amazonico, the lives of amphibian and reptiles in an Amazonian rainforest. Cornell University Press, Ithaca, xvi + 433 pp." type="book" year="2005">Duellman 2005</bibRefCitation>
) and western
<collectingCountry id="F33C767BFFA2605E6ADB8CE4D567F831" box="[975,1043,1930,1954]" name="Brazil" pageId="25" pageNumber="26">Brazil</collectingCountry>
(
<bibRefCitation id="EFBA4B1AFFA2605E6D368CE4D590F831" author="Avila-Pires" box="[1058,1252,1930,1954]" pageId="25" pageNumber="38" refString="Avila-Pires, T. C. S. (1995) Lizards of Brazilian Amazonia (Reptilia: Squamata). Nationaal Natuurhistorisch Museum Zoologische Verhandelingen, 299, 1 - 706." type="journal article" year="1995">Avila-Pires 1995</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="DF546663FFA1605D69838E04D518FA12" box="[151,1132,1386,1409]" httpUri="https://zenodo.org/record/207086/files/figure.png" pageId="26" pageNumber="27" targetBox="[157,1429,206,1356]" targetPageId="26">
<paragraph id="8B9436EBFFA1605D69838E04D518FA12" blockId="26.[151,1132,1386,1409]" box="[151,1132,1386,1409]" pageId="26" pageNumber="27">
<emphasis id="B95FEAF9FFA1605D69838E04D066FA13" bold="true" box="[151,274,1386,1409]" pageId="26" pageNumber="27">FIGURE 9.</emphasis>
Distribution of
<taxonomicName id="4C2B4D68FFA1605D68A18E05D3D2FA12" box="[437,678,1387,1409]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="palpebralis">
<emphasis id="B95FEAF9FFA1605D68A18E05D3D2FA12" box="[437,678,1387,1409]" italics="true" pageId="26" pageNumber="27">Enyalioides palpebralis</emphasis>
</taxonomicName>
(dots) and
<taxonomicName id="4C2B4D68FFA1605D6A0D8E05D282FA12" box="[793,1014,1387,1409]" class="Reptilia" family="Hoplocercidae" genus="Hoplocercus" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="spinosus">
<emphasis id="B95FEAF9FFA1605D6A0D8E05D282FA12" box="[793,1014,1387,1409]" italics="true" pageId="26" pageNumber="27">Hoplocercus spinosus</emphasis>
</taxonomicName>
(triangles).
</paragraph>
</caption>
<subSubSection id="C3316560FFA1605D69D18EC0D0F2F9EA" pageId="26" pageNumber="27" type="discussion">
<paragraph id="8B9436EBFFA1605D69D18EC0D0F2F9EA" blockId="26.[151,1436,1454,1657]" pageId="26" pageNumber="27">
<emphasis id="B95FEAF9FFA1605D69D18EC0D04DFA55" bold="true" box="[197,313,1454,1478]" pageId="26" pageNumber="27">Remarks.</emphasis>
<taxonomicName id="4C2B4D68FFA1605D68578EC0D324FA55" box="[323,592,1454,1478]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="palpebralis">
<emphasis id="B95FEAF9FFA1605D68578EC0D324FA55" box="[323,592,1454,1478]" italics="true" pageId="26" pageNumber="27">Enyalioides palpebralis</emphasis>
</taxonomicName>
has been regarded as the only hoplocercine lacking femoral pores (
<bibRefCitation id="EFBA4B1AFFA1605D6C768EC0D054FA7A" author="Boulenger" pageId="26" pageNumber="38" refString="Boulenger, G. A. (1885) Catalogue of the Lizards in the British Museum (Natural History) II. Taylor and Francis, London, 497 pp." type="book" year="1885">Boulenger 1885</bibRefCitation>
;
<bibRefCitation id="EFBA4B1AFFA1605D68388EBFD099FA7A" author="Avila-Pires" box="[300,493,1489,1513]" pageId="26" pageNumber="38" refString="Avila-Pires, T. C. S. (1995) Lizards of Brazilian Amazonia (Reptilia: Squamata). Nationaal Natuurhistorisch Museum Zoologische Verhandelingen, 299, 1 - 706." type="journal article" year="1995">Avila-Pires 1995</bibRefCitation>
). Nonetheless, out of the
<specimenCount id="9D2DFD62FFA1605D6A1B8EBFD2D3FA7A" box="[783,935,1489,1513]" pageId="26" pageNumber="27" type="generic">20 specimens</specimenCount>
examined, we found an adult male (AMNH 56401, SVL =
<quantity id="4CD39B0EFFA1605D68298E98D0F9F99D" box="[317,397,1526,1550]" metricMagnitude="-2" metricUnit="m" metricValue="7.9" pageId="26" pageNumber="27" unit="mm" value="79.0">79 mm</quantity>
) from departamento Loreto in
<collectingCountry id="F33C767BFFA1605D6BF38E99D26EF99D" box="[743,794,1527,1550]" name="Peru" pageId="26" pageNumber="27">Peru</collectingCountry>
that had one femoral pore on each side. Examined specimens of all other species of
<taxonomicName id="4C2B4D68FFA1605D68C68D74D322F9A1" box="[466,598,1562,1586]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">
<emphasis id="B95FEAF9FFA1605D68C68D74D322F9A1" box="[466,598,1562,1586]" italics="true" pageId="26" pageNumber="27">Enyalioides</emphasis>
</taxonomicName>
have at least one femoral pore on each leg except for
<taxonomicName id="4C2B4D68FFA1605D6DA28D75D458F9A1" box="[1206,1324,1562,1586]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="laticeps">
<emphasis id="B95FEAF9FFA1605D6DA28D75D458F9A1" box="[1206,1324,1562,1586]" italics="true" pageId="26" pageNumber="27">E. laticeps</emphasis>
</taxonomicName>
, in which a few specimens also lack femoral pores (
<tableCitation id="C6A90350FFA1605D6B798D50D3B5F9C5" box="[621,705,1598,1622]" captionStart="TABLE 1" captionStartId="6.[151,235,1304,1325]" captionTargetBox="[151,1436,1377,1892]" captionTargetPageId="6" captionText="TABLE 1. Summary of counts and measurements (mm) for species of Hoplocercinae. Range (first line) and mean ± standard" httpUri="http://table.plazi.org/id/DF546663FFBD604169838E76D4E8FABD" pageId="26" pageNumber="27" tableUuid="DF546663FFBD604169838E76D4E8FABD">Table 1</tableCitation>
). Specimens of
<taxonomicName id="4C2B4D68FFA1605D6A6E8D50D57CF9C5" box="[890,1032,1598,1622]" class="Reptilia" family="Hoplocercidae" genus="Hoplocercus" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">
<emphasis id="B95FEAF9FFA1605D6A6E8D50D57CF9C5" box="[890,1032,1598,1622]" italics="true" pageId="26" pageNumber="27">Hoplocercus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2B4D68FFA1605D6D2B8D51D590F9C5" box="[1087,1252,1599,1622]" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">
<emphasis id="B95FEAF9FFA1605D6D2B8D51D590F9C5" box="[1087,1252,1599,1622]" italics="true" pageId="26" pageNumber="27">Morunasaurus</emphasis>
</taxonomicName>
have 2–5 femoral pores on each leg.
</paragraph>
</subSubSection>
</treatment>
</document>
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