1327 lines
300 KiB
XML
1327 lines
300 KiB
XML
<document id="D42422F07F5CC4E52C6165D9CB70C3CA" ID-CLB-Dataset="121763" ID-DOI="10.1093/zoolinnean/zlab097" ID-GBIF-Dataset="e4804bc4-8183-435b-8397-3be4c5537bb9" ID-ISSN="0024-4082" ID-Zenodo-Dep="6994456" ID-ZooBank="0BE2F196-846F-4AD2-8C4D-79639B7E409C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1660324114056" checkinUser="plazi" docAuthor="Boudinot, Brendon E., Richter, Adrian, Katzke, Julian, Chaul, Júlio C. M., Keller, Roberto A., Economo, Evan P., Beutel, Rolf Georg & Yamamoto, Shûhei" docDate="2022" docId="038287F4FFE8AA26D2FEFE34FC09947B" docLanguage="en" docName="zlab097.pdf" docOrigin="Zoological Journal of the Linnean Society 195" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Gerontoformica sternorhabda Boudinot & Richter & Katzke & Chaul & Keller & Economo & Beutel & Yamamoto 2022, SP. NOV." docType="treatment" docVersion="5" lastPageNumber="1369" masterDocId="FFBBFF8CFFE1AA28D237FF96FF979022" masterDocTitle="Evidence for the evolution of eusociality in stem ants and a systematic revision of † Gerontoformica (Hymenoptera: Formicidae)" masterLastPageNumber="1389" masterPageNumber="1355" pageNumber="1364" updateTime="1699366467079" updateUser="ExternalLinkService">
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<mods:title id="EA99942111298341C7CD41588B173EC4">Evidence for the evolution of eusociality in stem ants and a systematic revision of † Gerontoformica (Hymenoptera: Formicidae)</mods:title>
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<mods:namePart id="B468DE14CF78761B87323D5A9499755A">Boudinot, Brendon E.</mods:namePart>
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<mods:namePart id="32D49F65619208645484F361A7FCF38C">Richter, Adrian</mods:namePart>
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<mods:namePart id="796367A82E66A9C9B580F6907AA685B4">Katzke, Julian</mods:namePart>
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<mods:namePart id="E5C686C53A985B3466E4555EB86F7313">Chaul, Júlio C. M.</mods:namePart>
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<mods:namePart id="2D5A994DC6D3C3B6800C5193F50DB758">Keller, Roberto A.</mods:namePart>
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<mods:namePart id="53AC49DA7F1802710F969D453E8FC92C">Economo, Evan P.</mods:namePart>
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<mods:namePart id="09FBC762BB15BD85B1049CCBD0C66DBA">Beutel, Rolf Georg</mods:namePart>
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<mods:namePart id="F7B970974F72EDF59B5D2DBC69BF193A">Yamamoto, Shûhei</mods:namePart>
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<mods:title id="6C0E343D7025E896A8D072EABF2F5C7C">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="95F7B918AA83BE5A3F9B6F09C2FEF34B">2022</mods:date>
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<mods:number id="F857268623C423F6D756255794E8DF70">2022-02-05</mods:number>
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<treatment id="038287F4FFE8AA26D2FEFE34FC09947B" ID-DOI="http://doi.org/10.5281/zenodo.6994462" ID-GBIF-Taxon="199312769" ID-Zenodo-Dep="6994462" LSID="urn:lsid:plazi:treatment:038287F4FFE8AA26D2FEFE34FC09947B" httpUri="http://treatment.plazi.org/id/038287F4FFE8AA26D2FEFE34FC09947B" lastPageId="14" lastPageNumber="1369" pageId="9" pageNumber="1364">
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<subSubSection id="C3316569FFE8AA21D2FEFE34FD739198" box="[201,740,418,442]" pageId="9" pageNumber="1364" type="nomenclature">
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<paragraph id="8B9436E2FFE8AA21D2FEFE34FD739198" blockId="9.[201,740,418,442]" box="[201,740,418,442]" pageId="9" pageNumber="1364">
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<heading id="D0DC818EFFE8AA21D2FEFE34FD739198" box="[201,740,418,442]" centered="true" fontSize="9" level="2" pageId="9" pageNumber="1364" reason="2">
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<emphasis id="B95FEAF0FFE8AA21D2FEFE34FDED919B" bold="true" box="[201,634,418,442]" italics="true" pageId="9" pageNumber="1364">
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†
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<taxonomicName id="4C2B4D61FFE8AA21D2E2FE34FDED919B" authority="Boudinot & Richter & Katzke & Chaul & Keller & Economo & Beutel & Yamamoto, 2022" authorityName="Boudinot & Richter & Katzke & Chaul & Keller & Economo & Beutel & Yamamoto" authorityYear="2022" box="[213,634,418,442]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1381" phylum="Arthropoda" rank="species" species="sternorhabda" status="sp. nov.">
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<emphasis id="B95FEAF0FFE8AA21D2E2FE34FE22919B" bold="true" box="[213,437,418,442]" pageId="9" pageNumber="1364">GERONTOFORMICA</emphasis>
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STERNORHABDA
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</taxonomicName>
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</emphasis>
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<taxonomicNameLabel id="A26C578BFFE8AA21D0B6FE33FD739198" box="[641,740,418,442]" pageId="9" pageNumber="1364" rank="species">
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SP
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<emphasis id="B95FEAF0FFE8AA21D0A9FE34FD739198" bold="true" box="[670,740,418,442]" pageId="9" pageNumber="1364">. NOV.</emphasis>
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<subSubSection id="C3316569FFE8AA21D28FFE5FFD6B92A1" pageId="9" pageNumber="1364" type="description">
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<paragraph id="8B9436E2FFE8AA21D28FFE5FFE68923D" blockId="9.[179,763,457,543]" pageId="9" pageNumber="1364">
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(
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<figureCitation id="13102A67FFE8AA21D2F6FE5FFEAA91C0" box="[193,317,457,482]" captionStart="Figure 1" captionStartId="2.[145,223,1030,1052]" captionTargetBox="[145,1425,196,990]" captionTargetId="figure-370@2.[145,1425,195,990]" captionTargetPageId="2" captionText="Figure 1. Overview of the Kachin amber piece and its syninclusions.A, the pupa of †G. gracilis (CASENT01741231). B, the adult of †G. gracilis (CASENT0741232). C, the holotype of †G. sternorhabda sp. nov. (CASENT0741233). D, the paratype of †G. sternorhabda sp. nov. (CASENT0741234). E, the entire amber piece (AMNH SY-23)." figureDoi="http://doi.org/10.5281/zenodo.6994458" httpUri="https://zenodo.org/record/6994458/files/figure.png" pageId="9" pageNumber="1364">FIGS 1C, D</figureCitation>
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,
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<figureCitation id="13102A67FFE8AA21D37DFE5FFE2D91C0" box="[330,442,457,482]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="9" pageNumber="1364">2C, F, I, L</figureCitation>
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,
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<figureCitation id="13102A67FFE8AA21D3F0FE5FFE6491C0" box="[455,499,457,482]" captionStart-0="Figure 4" captionStart-1="Figure 5" captionStart-2="Figure 6" captionStartId-0="15.[163,242,1675,1697]" captionStartId-1="16.[145,227,1675,1697]" captionStartId-2="18.[145,223,1652,1674]" captionTargetBox-0="[229,1376,196,1635]" captionTargetBox-1="[223,1346,196,1628]" captionTargetBox-2="[146,1425,197,1609]" captionTargetId-0="figure-26@15.[229,1376,195,1635]" captionTargetId-1="figure-26@16.[223,1346,195,1635]" captionTargetId-2="figure-26@18.[145,1425,195,1612]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="18" captionText-0="Figure 4. Photomicrographs of †G. sternorhabda sp. nov. showing setation and certain key features. A–D, G–I, paratype, CASENT0741234. E, F, J, holotype, CASENT0741233.A, body, ventrolateral oblique view. B, head, dorsolateral oblique view. C–F, body and left legs: D, dorsolateral oblique; (F) is slightly offset from (E), revealing the prora and subpetiolar process. G, protarsus, posterodorsal oblique view. H, mesotarsus, dorsal view. I, metatarsus, dorsal view. J, metatarsus, posterior view. Abbreviations: ad, antennomere distorted, i.e. visibly distorted antennomere associated with decay bubble; cn, cinctus; ms, metanotal spiracle; pr, prora; psf, anterior flange of propodeal spiracle; ptb, probasitarsal brush; spp, subpetiolar process; ss, standing setae; tnc, transverse mesonotal carina." captionText-1="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" captionText-2="Figure 6. Detailsof thepronota andmetapleural glandsof †Gerontoformica.A,B,fluorescent lightmicroscopy;C–G,standard photomicroscopy. A, B, †G. sternorhabda sp. nov. paratype (UFV-LABECOL-009656). C, †G. pilosa (ANTWEB1038931). D, †G. sternorhabda sp. nov. non-type (JWJ-BU21). E, F, †G. gracilis holotype (JZC-Bu324A). G, †G. near or conspecific with gracilis (ANTWEB1032649). Abbreviations: msnttc, transverse carina of mesonotum; mtplglb, metapleural gland bulla; mtplgldf, metapleural gland dorsal flange; mtplglvf, metapleural gland ventral flange; mtvlc, ventrolateral carina of metathorax; mtntsp, metanotal spiracle; mtnt, metanotum; pntl, pronotal lobe; pntfl, lateral pronotal flange; pntfm, medial pronotal flange; ppdsf, anterior flange of propodeal spiracle; ppl, propleuron; pt, petiole." figureDoi-0="http://doi.org/10.5281/zenodo.6994466" figureDoi-1="http://doi.org/10.5281/zenodo.6994470" figureDoi-2="http://doi.org/10.5281/zenodo.6994472" httpUri-0="https://zenodo.org/record/6994466/files/figure.png" httpUri-1="https://zenodo.org/record/6994470/files/figure.png" httpUri-2="https://zenodo.org/record/6994472/files/figure.png" pageId="9" pageNumber="1364">
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4–
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<geoCoordinate id="EE1F5025FFE8AA21D3D2FE5FFE6491C0" box="[485,499,457,482]" degrees="6, 7" direction="east" orientation="longitude" pageId="9" pageNumber="1364" precision="5555" value="6.7">6</geoCoordinate>
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</figureCitation>
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,
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<figureCitation id="13102A67FFE8AA21D3C9FE5FFDAD91C0" box="[510,570,457,482]" captionStart="Figure 7" captionStartId="19.[164,241,1513,1535]" captionTargetBox="[163,1443,195,1470]" captionTargetId="figure-119@19.[163,1443,195,1473]" captionTargetPageId="19" captionText="Figure 7. Maxillary and labial palps of †G.gracilis and †G.sternorhabda sp. nov.. Small black stars indicate each palpomere on the right-hand side of the body.A, B, †G.gracilis adult, CASENT0741232, in anterior dorsolateral oblique view (A) and anterior view (B).C, D, †G.gracilis pupa, CASENT0741231, in anterior view (C) and anterior dorsolateral view (D).E, F, †G.sternorhabda sp. nov. holotype, CASENT0741233, and paratype, CASENT0741234, in anteroventral oblique view. Abbreviations: pl, labial palp; plp, process of the proximal labial palpomere; pm, maxillary palp; pm?, possible maxillary palpomere." figureDoi="http://doi.org/10.5281/zenodo.6994474" httpUri="https://zenodo.org/record/6994474/files/figure.png" pageId="9" pageNumber="1364">7E, F</figureCitation>
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,
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<figureCitation id="13102A67FFE8AA21D070FE5FFDF391C0" box="[583,612,457,482]" captionStart="Figure 11" captionStartId="27.[164,242,1506,1528]" captionTargetBox="[163,1443,196,1467]" captionTargetId="figure-137@27.[163,1443,195,1467]" captionTargetPageId="27" captionText="Figure 11. Couplet 1: mesonotum with (1-1) or without (1’-1) transverse ridge; abdominal segment IV with (1-2) or without (1’-2) cinctus; prora large (1-3) or small (1’-3). (1-1) †G. ufv-05, ANTWEB103848 (J. Chaul, AntWeb), mesosoma in profile view. (1’-1, 1’-3) †G. indet., JWJ-Bu19 (V. Perrichot, AntWeb), mesosoma in profile view. (1-2) †G. sternorhabda sp. nov. paratype, UFV-LABECOL-009656, metasoma in ventrolateral view. (1’-2) †G. gracilis, CASENT0741232, metasoma in lateral view. (1-3) †G. pilosa holotype, JZC-Bu225 (P. Barden, AntWeb), anterior metasoma in profile view." figureDoi="http://doi.org/10.5281/zenodo.6994482" httpUri="https://zenodo.org/record/6994482/files/figure.png" pageId="9" pageNumber="1364">11</figureCitation>
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(1-2), 12(2-1, 2-3, 2-4), 13(3’-1); SUPPORTING INFORMATION,
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<figureCitation id="13102A67FFE8AA21D0F2FE71FE5B923D" captionStart="Figure 1" captionStartId="2.[145,223,1030,1052]" captionTargetBox="[145,1425,196,990]" captionTargetId="figure-370@2.[145,1425,195,990]" captionTargetPageId="2" captionText="Figure 1. Overview of the Kachin amber piece and its syninclusions.A, the pupa of †G. gracilis (CASENT01741231). B, the adult of †G. gracilis (CASENT0741232). C, the holotype of †G. sternorhabda sp. nov. (CASENT0741233). D, the paratype of †G. sternorhabda sp. nov. (CASENT0741234). E, the entire amber piece (AMNH SY-23)." figureDoi="http://doi.org/10.5281/zenodo.6994458" httpUri="https://zenodo.org/record/6994458/files/figure.png" pageId="9" pageNumber="1364">FIGS S1</figureCitation>
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, S
|
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<figureCitation id="13102A67FFE8AA21D3DEFD90FE60923D" box="[489,503,518,543]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="9" pageNumber="1364">2</figureCitation>
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)
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</paragraph>
|
||
<paragraph id="8B9436E2FFE8AA21D294FDD9FD6B92A1" blockId="9.[163,779,590,643]" pageId="9" pageNumber="1364">
|
||
<emphasis id="B95FEAF0FFE8AA21D294FDD9FE679246" box="[163,496,590,612]" italics="true" pageId="9" pageNumber="1364">Z o o b a n k r e g i s t r a t i o n:</emphasis>
|
||
u r n: l s i d: z o o b a n k. org:act:
|
||
<uuid id="FF8D0C37FFE8AA21D2C5FDFBFD6B92A1" box="[242,764,621,643]" pageId="9" pageNumber="1364">E77C1A20-1398-465F-9607-AD11E97ADF1C</uuid>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3316569FFE8AA21D294FD34FF6F93AF" pageId="9" pageNumber="1364" type="materials_examined">
|
||
<paragraph id="8B9436E2FFE8AA21D294FD34FF6F93AF" blockId="9.[163,780,673,909]" pageId="9" pageNumber="1364">
|
||
<emphasis id="B95FEAF0FFE8AA21D294FD34FEE79295" box="[163,368,673,695]" italics="true" pageId="9" pageNumber="1364">Type material:</emphasis>
|
||
<materialsCitation id="3B433CBFFFE8AA21D3B5FD37FDE39330" ID-GBIF-Occurrence="3888700301" collectionCode="AMNH" country="Myanmar" location="Hukawng Valley" municipality="Wingless" pageId="9" pageNumber="1364" specimenCode="SY-23" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
|
||
<emphasis id="B95FEAF0FFE8AA21D3B5FD37FD9C9294" bold="true" box="[386,523,673,694]" pageId="9" pageNumber="1364">
|
||
<typeStatus id="54908840FFE8AA21D3B5FD37FD9C9294" box="[386,523,673,694]" pageId="9" pageNumber="1364" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
.
|
||
<collectingMunicipality id="6BF0AC98FFE8AA21D013FD34FD089295" box="[548,671,673,695]" pageId="9" pageNumber="1364">Wingless</collectingMunicipality>
|
||
female (w).
|
||
<collectingCountry id="F33C7672FFE8AA21D2D7FD56FECB92F7" box="[224,348,704,725]" name="Myanmar" pageId="9" pageNumber="1364">Myanmar</collectingCountry>
|
||
,
|
||
<collectingRegion id="49EFF800FFE8AA21D358FD56FDB792F7" box="[367,544,704,726]" country="Myanmar" name="Kachin" pageId="9" pageNumber="1364">Kachin State</collectingRegion>
|
||
:
|
||
<location id="8EF46039FFE8AA21D004FD56FC9E92F4" LSID="urn:lsid:plazi:treatment:038287F4FFE8AA26D2FEFE34FC09947B:8EF46039FFE8AA21D004FD56FC9E92F4" box="[563,777,704,726]" country="Myanmar" municipality="Wingless" name="Hukawng Valley" pageId="9" pageNumber="1364" stateProvince="Kachin">Hukawng Valley</location>
|
||
[CASENT 0741233 in an amber piece labelled
|
||
<collectionCode id="ED3AAE27FFE8AA21D294FD6BFF619330" box="[163,246,765,786]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="9" pageNumber="1364">AMNH</collectionCode>
|
||
<specimenCode id="DB8D9E99FFE8AA21D2C9FD6BFED69330" box="[254,321,765,786]" collectionCode="SY-" pageId="9" pageNumber="1364">SY-23</specimenCode>
|
||
and deposited at
|
||
<collectionCode id="ED3AAE27FFE8AA21D022FD6BFDF99330" box="[533,622,765,786]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="9" pageNumber="1364">AMNH</collectionCode>
|
||
]
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B433CBFFFE8AA21D0B7FD6BFF6393AF" ID-GBIF-Occurrence="3888700302" collectionCode="CELC" country="Myanmar" location="Hukawng Valley" municipality="Wingless" pageId="9" pageNumber="1364" specimenCount="3" stateProvince="Kachin" typeStatus="paratype">
|
||
<emphasis id="B95FEAF0FFE8AA21D0B7FD6BFC939330" bold="true" box="[640,772,765,786]" pageId="9" pageNumber="1364">
|
||
<typeStatus id="54908840FFE8AA21D0B7FD6BFC939330" box="[640,772,765,786]" pageId="9" pageNumber="1364" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
. Wingless females (w). Synincluded with
|
||
<typeStatus id="54908840FFE8AA21D095FC8AFC9D9310" box="[674,778,796,818]" pageId="9" pageNumber="1364" type="holotype">holotype</typeStatus>
|
||
[CASENT0741234]; same locality as synincluded holo- and
|
||
<typeStatus id="54908840FFE8AA21D2E1FCCCFEDE934D" box="[214,329,858,879]" pageId="9" pageNumber="1364" type="paratype">paratypes</typeStatus>
|
||
[UFV-LABECOL-009656, deposited in
|
||
<collectionCode id="ED3AAE27FFE8AA21D294FCEEFF7993AF" box="[163,238,888,909]" pageId="9" pageNumber="1364">CELC</collectionCode>
|
||
]
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3316569FFE8AA22D294FC51FC5E956C" lastPageId="10" lastPageNumber="1365" pageId="9" pageNumber="1364" type="diagnosis">
|
||
<paragraph id="8B9436E2FFE8AA21D294FC51FE6193FC" blockId="9.[163,502,966,990]" box="[163,502,966,990]" pageId="9" pageNumber="1364">
|
||
<emphasis id="B95FEAF0FFE8AA21D294FC51FE6193FC" box="[163,502,966,990]" italics="true" pageId="9" pageNumber="1364">Diagnosis (wingless female)</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFE8AA21D294FC78FE9D9593" blockId="9.[163,779,1006,1458]" pageId="9" pageNumber="1364">
|
||
Conforming to the diagnosis of the
|
||
<taxonomicName id="4C2B4D61FFE8AA21D07DFC78FD599426" authorityName="LATREILLE" authorityYear="1809" box="[586,718,1006,1028]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="family">Formicidae</taxonomicName>
|
||
(see:
|
||
<bibRefCitation id="EFBA4B13FFE8AA21D294FB9BFE0A9400" author="Boudinot BE & Khouri Z & Richter A & Van de Kamp T & Barden P & Perrichot V & Chaul JCM" box="[163,413,1037,1059]" pageId="9" pageNumber="1364" pagination="174 - 393" refId="ref23358" refString="Boudinot BE, Khouri Z, Richter A, Van de Kamp T, Barden P, Perrichot V, Chaul JCM. 2020 a. Chapter II. The evolution of the ants: extinct ant sister-group illuminates eusociality origin and post-K / Pg persistence. In: Boudinot BE, Systematic and evolutionary morphology: case studies on Formicidae, Mesozoic Aculeata, and hexapodan genitalia. Ph. D. Thesis, University of California, Davis, 174 - 393." type="book chapter" year="2020">
|
||
Boudinot
|
||
<emphasis id="B95FEAF0FFE8AA21D322FB98FEDF9400" box="[277,328,1037,1059]" italics="true" pageId="9" pageNumber="1364">et al</emphasis>
|
||
. 2020a
|
||
</bibRefCitation>
|
||
, b), including the following key conditions: (1) postgenal bridge elongated, thus head ‘prognathous’ (
|
||
<figureCitation id="13102A67FFE8AA21D36EFBDCFE269442" box="[345,433,1098,1120]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="9" pageNumber="1364">Fig. 2C</figureCitation>
|
||
); (2) cranial condyles of the anterior/ dorsal mandibular articulation enlarged (
|
||
<figureCitation id="13102A67FFE8AA21D29BFB1EFE9394BC" box="[172,260,1160,1182]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="9" pageNumber="1364">Fig. 5B</figureCitation>
|
||
); (3) toruli oriented dorsolaterally rather than simply dorsally (
|
||
<figureCitation id="13102A67FFE8AA21D393FB30FE60949E" box="[420,503,1190,1212]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="9" pageNumber="1364">Fig. 5A</figureCitation>
|
||
); (4) procoxae elongate, about twice as long as wide (
|
||
<figureCitation id="13102A67FFE8AA21D3DBFB53FDAA94F8" box="[492,573,1221,1243]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="9" pageNumber="1364">Fig. 2C</figureCitation>
|
||
); (5) prodisticoxal foramen ‘closed’ and protrochanter narrowly necked (
|
||
<figureCitation id="13102A67FFE8AA21D29CFA94FF69953A" box="[171,254,1282,1304]" captionStart="Figure 4" captionStartId="15.[163,242,1675,1697]" captionTargetBox="[229,1376,196,1635]" captionTargetId="figure-26@15.[229,1376,195,1635]" captionTargetPageId="15" captionText="Figure 4. Photomicrographs of †G. sternorhabda sp. nov. showing setation and certain key features. A–D, G–I, paratype, CASENT0741234. E, F, J, holotype, CASENT0741233.A, body, ventrolateral oblique view. B, head, dorsolateral oblique view. C–F, body and left legs: D, dorsolateral oblique; (F) is slightly offset from (E), revealing the prora and subpetiolar process. G, protarsus, posterodorsal oblique view. H, mesotarsus, dorsal view. I, metatarsus, dorsal view. J, metatarsus, posterior view. Abbreviations: ad, antennomere distorted, i.e. visibly distorted antennomere associated with decay bubble; cn, cinctus; ms, metanotal spiracle; pr, prora; psf, anterior flange of propodeal spiracle; ptb, probasitarsal brush; spp, subpetiolar process; ss, standing setae; tnc, transverse mesonotal carina." figureDoi="http://doi.org/10.5281/zenodo.6994466" httpUri="https://zenodo.org/record/6994466/files/figure.png" pageId="9" pageNumber="1364">Fig. 4F</figureCitation>
|
||
) [Note 1]; (6) meso- and metathoracicocoxal articulations ‘closed’, i.e. directed ventrally rather than laterally or ventrolaterally (
|
||
<figureCitation id="13102A67FFE8AA21D3CFFAD6FDDA9577" box="[504,589,1344,1366]" captionStart="Figure 4" captionStartId="15.[163,242,1675,1697]" captionTargetBox="[229,1376,196,1635]" captionTargetId="figure-26@15.[229,1376,195,1635]" captionTargetPageId="15" captionText="Figure 4. Photomicrographs of †G. sternorhabda sp. nov. showing setation and certain key features. A–D, G–I, paratype, CASENT0741234. E, F, J, holotype, CASENT0741233.A, body, ventrolateral oblique view. B, head, dorsolateral oblique view. C–F, body and left legs: D, dorsolateral oblique; (F) is slightly offset from (E), revealing the prora and subpetiolar process. G, protarsus, posterodorsal oblique view. H, mesotarsus, dorsal view. I, metatarsus, dorsal view. J, metatarsus, posterior view. Abbreviations: ad, antennomere distorted, i.e. visibly distorted antennomere associated with decay bubble; cn, cinctus; ms, metanotal spiracle; pr, prora; psf, anterior flange of propodeal spiracle; ptb, probasitarsal brush; spp, subpetiolar process; ss, standing setae; tnc, transverse mesonotal carina." figureDoi="http://doi.org/10.5281/zenodo.6994466" httpUri="https://zenodo.org/record/6994466/files/figure.png" pageId="9" pageNumber="1364">Fig. 4F</figureCitation>
|
||
); (7) abdominal segment II completely petiolated (
|
||
<figureCitation id="13102A67FFE8AA21D05AFAC8FD5D9556" box="[621,714,1374,1396]" captionStart="Figure 4" captionStartId="15.[163,242,1675,1697]" captionTargetBox="[229,1376,196,1635]" captionTargetId="figure-26@15.[229,1376,195,1635]" captionTargetPageId="15" captionText="Figure 4. Photomicrographs of †G. sternorhabda sp. nov. showing setation and certain key features. A–D, G–I, paratype, CASENT0741234. E, F, J, holotype, CASENT0741233.A, body, ventrolateral oblique view. B, head, dorsolateral oblique view. C–F, body and left legs: D, dorsolateral oblique; (F) is slightly offset from (E), revealing the prora and subpetiolar process. G, protarsus, posterodorsal oblique view. H, mesotarsus, dorsal view. I, metatarsus, dorsal view. J, metatarsus, posterior view. Abbreviations: ad, antennomere distorted, i.e. visibly distorted antennomere associated with decay bubble; cn, cinctus; ms, metanotal spiracle; pr, prora; psf, anterior flange of propodeal spiracle; ptb, probasitarsal brush; spp, subpetiolar process; ss, standing setae; tnc, transverse mesonotal carina." figureDoi="http://doi.org/10.5281/zenodo.6994466" httpUri="https://zenodo.org/record/6994466/files/figure.png" pageId="9" pageNumber="1364">Fig. 4F</figureCitation>
|
||
); (8) subpetiolar process present (
|
||
<figureCitation id="13102A67FFE8AA21D3DFFAEBFDA095B0" box="[488,567,1405,1427]" captionStart="Figure 4" captionStartId="15.[163,242,1675,1697]" captionTargetBox="[229,1376,196,1635]" captionTargetId="figure-26@15.[229,1376,195,1635]" captionTargetPageId="15" captionText="Figure 4. Photomicrographs of †G. sternorhabda sp. nov. showing setation and certain key features. A–D, G–I, paratype, CASENT0741234. E, F, J, holotype, CASENT0741233.A, body, ventrolateral oblique view. B, head, dorsolateral oblique view. C–F, body and left legs: D, dorsolateral oblique; (F) is slightly offset from (E), revealing the prora and subpetiolar process. G, protarsus, posterodorsal oblique view. H, mesotarsus, dorsal view. I, metatarsus, dorsal view. J, metatarsus, posterior view. Abbreviations: ad, antennomere distorted, i.e. visibly distorted antennomere associated with decay bubble; cn, cinctus; ms, metanotal spiracle; pr, prora; psf, anterior flange of propodeal spiracle; ptb, probasitarsal brush; spp, subpetiolar process; ss, standing setae; tnc, transverse mesonotal carina." figureDoi="http://doi.org/10.5281/zenodo.6994466" httpUri="https://zenodo.org/record/6994466/files/figure.png" pageId="9" pageNumber="1364">Fig. 4F</figureCitation>
|
||
); (9) prora present (
|
||
<figureCitation id="13102A67FFE8AA21D29CFA0AFF6D9593" box="[171,250,1436,1458]" captionStart="Figure 4" captionStartId="15.[163,242,1675,1697]" captionTargetBox="[229,1376,196,1635]" captionTargetId="figure-26@15.[229,1376,195,1635]" captionTargetPageId="15" captionText="Figure 4. Photomicrographs of †G. sternorhabda sp. nov. showing setation and certain key features. A–D, G–I, paratype, CASENT0741234. E, F, J, holotype, CASENT0741233.A, body, ventrolateral oblique view. B, head, dorsolateral oblique view. C–F, body and left legs: D, dorsolateral oblique; (F) is slightly offset from (E), revealing the prora and subpetiolar process. G, protarsus, posterodorsal oblique view. H, mesotarsus, dorsal view. I, metatarsus, dorsal view. J, metatarsus, posterior view. Abbreviations: ad, antennomere distorted, i.e. visibly distorted antennomere associated with decay bubble; cn, cinctus; ms, metanotal spiracle; pr, prora; psf, anterior flange of propodeal spiracle; ptb, probasitarsal brush; spp, subpetiolar process; ss, standing setae; tnc, transverse mesonotal carina." figureDoi="http://doi.org/10.5281/zenodo.6994466" httpUri="https://zenodo.org/record/6994466/files/figure.png" pageId="9" pageNumber="1364">Fig. 4F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFE8AA21D28FFA5DFAA89156" blockId="9.[184,779,1482,1903]" lastBlockId="9.[827,1444,197,1905]" pageId="9" pageNumber="1364">
|
||
<emphasis id="B95FEAF0FFE8AA21D28FFA5DFE5F95DD" bold="true" pageId="9" pageNumber="1364">
|
||
I. Among
|
||
<taxonomicName id="4C2B4D61FFE8AA21D357FA5CFE6495C2" authorityName="LATREILLE" authorityYear="1809" box="[352,499,1482,1504]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="family">Formicidae</taxonomicName>
|
||
, identifiable as †Sphecomyrminae:
|
||
</emphasis>
|
||
(1) Mandibles simple and bidentate (
|
||
<figureCitation id="13102A67FFE8AA21D397F99EFD97963C" box="[416,512,1544,1566]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="9" pageNumber="1364">Fig. 5A</figureCitation>
|
||
), without: (a) being strongly bowed and multidentate as in the †
|
||
<taxonomicName id="4C2B4D61FFE8AA21D2D5F9D3FEED9678" authorityName="Barden & Grimaldi" authorityYear="2016" box="[226,378,1605,1626]" class="Insecta" family="Formicidae" genus="Camelomecia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FEAF0FFE8AA21D2D5F9D3FEED9678" box="[226,378,1605,1626]" italics="true" pageId="9" pageNumber="1364">Camelomecia</emphasis>
|
||
</taxonomicName>
|
||
group, (b) the scythe-like blade as in the haidomyrmecine †
|
||
<taxonomicName id="4C2B4D61FFE8AA21D3CDF9F2FD0C965B" box="[506,667,1636,1657]" class="Insecta" family="Formicidae" genus="Haidomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FEAF0FFE8AA21D3CDF9F2FD0C965B" box="[506,667,1636,1657]" italics="true" pageId="9" pageNumber="1364">Haidomyrmex</emphasis>
|
||
</taxonomicName>
|
||
clade, (c) projecting anteriorly with numerous teeth as in the haidomyrmecine †
|
||
<taxonomicName id="4C2B4D61FFE8AA21D3E2F937FDED9694" authorityName="Perrichot, Wang & Barden" authorityYear="2020" box="[469,634,1697,1718]" class="Insecta" family="Formicidae" genus="Aquilomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FEAF0FFE8AA21D3E2F937FDED9694" box="[469,634,1697,1718]" italics="true" pageId="9" pageNumber="1364">Aquilomyrmex</emphasis>
|
||
</taxonomicName>
|
||
clade, or (d) the strong torsion of †
|
||
<taxonomicName id="4C2B4D61FFE8AA21D3D1F956FD1C96F4" box="[486,651,1728,1750]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="tribe" tribe="Zigrasimeciini">Zigrasimeciini</taxonomicName>
|
||
(the state in †
|
||
<taxonomicName id="4C2B4D61FFE8AA21D333F948FE0E96D1" authorityName="Borysenko" authorityYear="2017" box="[260,409,1758,1779]" class="Insecta" family="Formicidae" genus="Boltonimecia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FEAF0FFE8AA21D333F948FE0E96D1" box="[260,409,1758,1779]" italics="true" pageId="9" pageNumber="1364">Boltonimecia</emphasis>
|
||
</taxonomicName>
|
||
is uncertain); (2) the antennal scrobes on the face do not extend all the way to the compound eye (
|
||
<figureCitation id="13102A67FFE8AA21D39AF88AFE699713" box="[429,510,1820,1842]" captionStart="Figure 4" captionStartId="15.[163,242,1675,1697]" captionTargetBox="[229,1376,196,1635]" captionTargetId="figure-26@15.[229,1376,195,1635]" captionTargetPageId="15" captionText="Figure 4. Photomicrographs of †G. sternorhabda sp. nov. showing setation and certain key features. A–D, G–I, paratype, CASENT0741234. E, F, J, holotype, CASENT0741233.A, body, ventrolateral oblique view. B, head, dorsolateral oblique view. C–F, body and left legs: D, dorsolateral oblique; (F) is slightly offset from (E), revealing the prora and subpetiolar process. G, protarsus, posterodorsal oblique view. H, mesotarsus, dorsal view. I, metatarsus, dorsal view. J, metatarsus, posterior view. Abbreviations: ad, antennomere distorted, i.e. visibly distorted antennomere associated with decay bubble; cn, cinctus; ms, metanotal spiracle; pr, prora; psf, anterior flange of propodeal spiracle; ptb, probasitarsal brush; spp, subpetiolar process; ss, standing setae; tnc, transverse mesonotal carina." figureDoi="http://doi.org/10.5281/zenodo.6994466" httpUri="https://zenodo.org/record/6994466/files/figure.png" pageId="9" pageNumber="1364">Fig. 4B</figureCitation>
|
||
) (such scrobes observed in †Zigrasimeciinae, including †
|
||
<taxonomicName id="4C2B4D61FFE8AA21D059F8ACFC94976D" authorityName="Borysenko" authorityYear="2017" box="[622,771,1850,1871]" class="Insecta" family="Formicidae" genus="Boltonimecia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FEAF0FFE8AA21D059F8ACFC94976D" box="[622,771,1850,1871]" italics="true" pageId="9" pageNumber="1364">Boltonimecia</emphasis>
|
||
</taxonomicName>
|
||
) [Note 2]; (3) anterolateral corners of head not produced as robust triangles (
|
||
<figureCitation id="13102A67FFE8AA21D682FF53FA9490F9" box="[1205,1283,197,219]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="9" pageNumber="1364">Fig. 5A</figureCitation>
|
||
) (such corners observed in †
|
||
<taxonomicName id="4C2B4D61FFE8AA21D633FF72FBF090DB" authorityName="Lattke & Melo" authorityYear="2020" box="[1028,1127,228,249]" class="Insecta" family="Formicidae" genus="Dilobops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FEAF0FFE8AA21D633FF72FBF090DB" box="[1028,1127,228,249]" italics="true" pageId="9" pageNumber="1364">Dilobops</emphasis>
|
||
</taxonomicName>
|
||
of the †Haidomyrmecinae); and (4) (a) scapes shorter than the width of the head (
|
||
<figureCitation id="13102A67FFE8AA21D18AFEB7FB809115" box="[957,1047,289,311]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="9" pageNumber="1364">Fig. 5A</figureCitation>
|
||
) and (b) clypeus not extending posteriorly between the toruli (
|
||
<figureCitation id="13102A67FFE8AA21D6C2FED6FADA9177" box="[1269,1357,320,342]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="9" pageNumber="1364">Fig. 2L</figureCitation>
|
||
) (such extension observed in †
|
||
<taxonomicName id="4C2B4D61FFE8AA21D641FEC8FAA39156" box="[1142,1332,350,372]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="subFamily" subFamily="Brownimeciinae">Brownimeciinae</taxonomicName>
|
||
);
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFE8AA21D172FEEBFBB492A5" blockId="9.[827,1444,197,1905]" pageId="9" pageNumber="1364">
|
||
<emphasis id="B95FEAF0FFE8AA21D172FEEBFBD59193" bold="true" pageId="9" pageNumber="1364">
|
||
II. Within †Sphecomyrminae, identifiable as †
|
||
<taxonomicName id="4C2B4D61FFE8AA21D14DFE0DFBD59193" authorityName="Nel & Perrault" authorityYear="2004" box="[890,1090,411,433]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FEAF0FFE8AA21D14DFE0DFBD59193" bold="true" box="[890,1090,411,433]" italics="true" pageId="9" pageNumber="1364">Gerontoformica</emphasis>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
: (5) anteromedian clypeal margin not produced as distinct medial lobe (
|
||
<figureCitation id="13102A67FFE8AA21D141FE4FFC5191CC" box="[886,966,473,495]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="9" pageNumber="1364">Fig. 2L</figureCitation>
|
||
); vs. such a lobe present (†
|
||
<taxonomicName id="4C2B4D61FFE8AA21D6CCFE4FFA0B91CC" authorityName="Wilson & Brown" authorityYear="1967" box="[1275,1436,473,494]" class="Insecta" family="Formicidae" genus="Sphecomyrma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FEAF0FFE8AA21D6CCFE4FFA0B91CC" box="[1275,1436,473,494]" italics="true" pageId="9" pageNumber="1364">Sphecomyrma</emphasis>
|
||
</taxonomicName>
|
||
, †
|
||
<taxonomicName id="4C2B4D61FFE8AA21D14DFE6EFC76922F" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[890,993,504,525]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="species" species="tendir">
|
||
<emphasis id="B95FEAF0FFE8AA21D14DFE6EFC76922F" box="[890,993,504,525]" italics="true" pageId="9" pageNumber="1364">G. tendir</emphasis>
|
||
</taxonomicName>
|
||
) [Note 3]; and (6) (a) mandibles short and fitting against clypeus when closed (
|
||
<figureCitation id="13102A67FFE8AA21D74FFD80FC1A9269" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="9" pageNumber="1364">Fig. 5A</figureCitation>
|
||
) and (b) metanotum developed (
|
||
<figureCitation id="13102A67FFE8AA21D722FDA3FAFC9268" box="[1301,1387,565,587]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="9" pageNumber="1364">Fig. 2C</figureCitation>
|
||
); vs. mandibles elongate and metanotum not developed (†
|
||
<taxonomicName id="4C2B4D61FFE8AA21D1B5FDE5FB8392AA" authorityName="Engel & Grimaldi" authorityYear="2005" box="[898,1044,627,648]" class="Insecta" family="Formicidae" genus="Myanmyrma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FEAF0FFE8AA21D1B5FDE5FB8392AA" box="[898,1044,627,648]" italics="true" pageId="9" pageNumber="1364">Myanmyrma</emphasis>
|
||
</taxonomicName>
|
||
);
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFE8AA21D10CFD07FA1593B9" blockId="9.[827,1444,197,1905]" pageId="9" pageNumber="1364">
|
||
<emphasis id="B95FEAF0FFE8AA21D10CFD07FBF592E7" bold="true" pageId="9" pageNumber="1364">
|
||
III. Within †
|
||
<taxonomicName id="4C2B4D61FFE8AA21D1EDFD07FB359285" authorityName="Nel & Perrault" authorityYear="2004" box="[986,1186,657,679]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FEAF0FFE8AA21D1EDFD07FB359285" bold="true" box="[986,1186,657,679]" italics="true" pageId="9" pageNumber="1364">Gerontoformica</emphasis>
|
||
</taxonomicName>
|
||
, with the following unique condition
|
||
</emphasis>
|
||
: (7) the anteroventral (‘subpetiolar’) process of the petiolar sternum long, orthogonal to the longitudinal axis of the petiole, and more-or-less rod-like, i.e. with anterior and posterior margins parallel to subparallel (
|
||
<figureCitation id="13102A67FFE8AA21D141FCDFFC5C937C" box="[886,971,841,863]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="9" pageNumber="1364">Fig. 2C</figureCitation>
|
||
); vs. short and triangular (†
|
||
<taxonomicName id="4C2B4D61FFE8AA21D713FCDFFA0A937C" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[1316,1437,841,862]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B95FEAF0FFE8AA21D713FCDFFA0A937C" box="[1316,1437,841,862]" italics="true" pageId="9" pageNumber="1364">G. gracilis</emphasis>
|
||
</taxonomicName>
|
||
, †
|
||
<taxonomicName id="4C2B4D61FFE8AA21D14DFCFEFB8B935F" box="[890,1052,872,893]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="species" species="occidentalis">
|
||
<emphasis id="B95FEAF0FFE8AA21D14DFCFEFB8B935F" box="[890,1052,872,893]" italics="true" pageId="9" pageNumber="1364">G. occidentalis</emphasis>
|
||
</taxonomicName>
|
||
, †
|
||
<taxonomicName id="4C2B4D61FFE8AA21D604FCFEFB04935F" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[1075,1171,872,893]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="species" species="pilosa">
|
||
<emphasis id="B95FEAF0FFE8AA21D604FCFEFB04935F" box="[1075,1171,872,893]" italics="true" pageId="9" pageNumber="1364">G. pilosa</emphasis>
|
||
</taxonomicName>
|
||
, †
|
||
<taxonomicName id="4C2B4D61FFE8AA21D69DFCFEFA8A935F" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[1194,1309,872,893]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="species" species="robusta">
|
||
<emphasis id="B95FEAF0FFE8AA21D69DFCFEFA8A935F" box="[1194,1309,872,893]" italics="true" pageId="9" pageNumber="1364">G. robusta</emphasis>
|
||
</taxonomicName>
|
||
, †
|
||
<taxonomicName id="4C2B4D61FFE8AA21D704FCFEFA0A935F" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[1331,1437,872,893]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="species" species="rugosa">
|
||
<emphasis id="B95FEAF0FFE8AA21D704FCFEFA0A935F" box="[1331,1437,872,893]" italics="true" pageId="9" pageNumber="1364">G. rugosa</emphasis>
|
||
</taxonomicName>
|
||
, †
|
||
<taxonomicName id="4C2B4D61FFE8AA21D14DFC11FC7993B9" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[890,1006,902,924]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="species" species="spiralis">
|
||
<emphasis id="B95FEAF0FFE8AA21D14DFC11FC7993B9" box="[890,1006,902,924]" italics="true" pageId="9" pageNumber="1364">G. spiralis</emphasis>
|
||
</taxonomicName>
|
||
, †
|
||
<taxonomicName id="4C2B4D61FFE8AA21D630FC11FB0393B9" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[1031,1172,902,924]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="species" species="subcuspis">
|
||
<emphasis id="B95FEAF0FFE8AA21D630FC11FB0393B9" box="[1031,1172,902,924]" italics="true" pageId="9" pageNumber="1364">G. subcuspis</emphasis>
|
||
</taxonomicName>
|
||
, †
|
||
<taxonomicName id="4C2B4D61FFE8AA21D69AFC11FA8693B9" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[1197,1297,902,924]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="species" species="tendir">
|
||
<emphasis id="B95FEAF0FFE8AA21D69AFC11FA8693B9" box="[1197,1297,902,924]" italics="true" pageId="9" pageNumber="1364">G. tendir</emphasis>
|
||
</taxonomicName>
|
||
) [Note 4];
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFE8AA21D177FC33FC0B95A4" blockId="9.[827,1444,197,1905]" pageId="9" pageNumber="1364">
|
||
<emphasis id="B95FEAF0FFE8AA21D177FC33FAE693FB" bold="true" pageId="9" pageNumber="1364">
|
||
IV. Within †
|
||
<taxonomicName id="4C2B4D61FFE8AA21D1DBFC32FB239398" authorityName="Nel & Perrault" authorityYear="2004" box="[1004,1204,932,954]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FEAF0FFE8AA21D1DBFC32FB239398" bold="true" box="[1004,1204,932,954]" italics="true" pageId="9" pageNumber="1364">Gerontoformica</emphasis>
|
||
</taxonomicName>
|
||
, identifiable as a member of the
|
||
<taxonomicName id="4C2B4D61FFE8AA21D668FC55FB3A93FB" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[1119,1197,963,985]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="species" species="pilosa">
|
||
<emphasis id="B95FEAF0FFE8AA21D668FC55FB3A93FB" bold="true" box="[1119,1197,963,985]" italics="true" pageId="9" pageNumber="1364">pilosa</emphasis>
|
||
</taxonomicName>
|
||
species group
|
||
</emphasis>
|
||
: (8) mesoscutum with distinct, raised transverse carina separating mesoscutal and mesoscutellar regions (
|
||
<figureCitation id="13102A67FFE8AA21D1F8FB89FBB79417" box="[975,1056,1055,1077]" captionStart="Figure 6" captionStartId="18.[145,223,1652,1674]" captionTargetBox="[146,1425,197,1609]" captionTargetId="figure-26@18.[145,1425,195,1612]" captionTargetPageId="18" captionText="Figure 6. Detailsof thepronota andmetapleural glandsof †Gerontoformica.A,B,fluorescent lightmicroscopy;C–G,standard photomicroscopy. A, B, †G. sternorhabda sp. nov. paratype (UFV-LABECOL-009656). C, †G. pilosa (ANTWEB1038931). D, †G. sternorhabda sp. nov. non-type (JWJ-BU21). E, F, †G. gracilis holotype (JZC-Bu324A). G, †G. near or conspecific with gracilis (ANTWEB1032649). Abbreviations: msnttc, transverse carina of mesonotum; mtplglb, metapleural gland bulla; mtplgldf, metapleural gland dorsal flange; mtplglvf, metapleural gland ventral flange; mtvlc, ventrolateral carina of metathorax; mtntsp, metanotal spiracle; mtnt, metanotum; pntl, pronotal lobe; pntfl, lateral pronotal flange; pntfm, medial pronotal flange; ppdsf, anterior flange of propodeal spiracle; ppl, propleuron; pt, petiole." figureDoi="http://doi.org/10.5281/zenodo.6994472" httpUri="https://zenodo.org/record/6994472/files/figure.png" pageId="9" pageNumber="1364">Fig. 6B</figureCitation>
|
||
); vs. such a carina entirely absent or only poorly developed laterally, thus incomplete medially and not forming a distinct angle between the mesoscutal and mesoscutellar regions in profile view (
|
||
<taxonomicName id="4C2B4D61FFE8AA21D1C8FB0DFBF49492" authorityName="Nel & Perrault" authorityYear="2004" box="[1023,1123,1179,1200]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="species" species="cretacica">
|
||
<emphasis id="B95FEAF0FFE8AA21D1C8FB0DFBF49492" box="[1023,1123,1179,1200]" italics="true" pageId="9" pageNumber="1364">cretacica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2B4D61FFE8AA21D658FB0CFB53948D" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[1135,1220,1178,1199]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B95FEAF0FFE8AA21D658FB0CFB53948D" box="[1135,1220,1178,1199]" italics="true" pageId="9" pageNumber="1364">gracilis</emphasis>
|
||
</taxonomicName>
|
||
groups); (9) tergum of petiolar node anteroposteriorly longer than dorsoventrally tall (
|
||
<figureCitation id="13102A67FFE8AA21D657FB41FB2D94CF" box="[1120,1210,1239,1261]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="9" pageNumber="1364">Fig. 2C</figureCitation>
|
||
); vs. petiolar node tergum taller than long (
|
||
<taxonomicName id="4C2B4D61FFE8AA21D6B2FB61FB7E952E" authorityName="Nel & Perrault" authorityYear="2004" box="[1157,1257,1271,1292]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="species" species="cretacica">
|
||
<emphasis id="B95FEAF0FFE8AA21D6B2FB61FB7E952E" box="[1157,1257,1271,1292]" italics="true" pageId="9" pageNumber="1364">cretacica</emphasis>
|
||
</taxonomicName>
|
||
group); and (10) abdominal segment IV (metasomal III) with the cinctus distinct and impressed, i.e. divided into pre- and post-sclerites by a transverse sulcus (
|
||
<figureCitation id="13102A67FFE8AA21D74FFAC4FC1A95A5" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="9" pageNumber="1364">Fig. 5E</figureCitation>
|
||
);
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFE8AA22D17CFA06FDDC913A" blockId="9.[827,1444,197,1905]" lastBlockId="10.[147,762,197,1139]" lastPageId="10" lastPageNumber="1365" pageId="9" pageNumber="1364">
|
||
<emphasis id="B95FEAF0FFE8AA21D17CFA06FC7095C1" bold="true" pageId="9" pageNumber="1364">
|
||
V. Within the
|
||
<taxonomicName id="4C2B4D61FFE8AA21D67FFA19FB009587" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[1096,1175,1423,1445]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="species" species="pilosa">
|
||
<emphasis id="B95FEAF0FFE8AA21D67FFA19FB009587" bold="true" box="[1096,1175,1423,1445]" italics="true" pageId="9" pageNumber="1364">pilosa</emphasis>
|
||
</taxonomicName>
|
||
species group, distinguished from all species by the following
|
||
</emphasis>
|
||
: (11) cinctus developed, but transverse sulci weakly impressed, thus pre- and postsclerites of abdominal segment IV not meeting at strongly oblique angle (
|
||
<figureCitation id="13102A67FFE8AA21D722F9BFFAFC961D" box="[1301,1387,1577,1599]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="9" pageNumber="1364">Fig. 5E</figureCitation>
|
||
); vs. transverse sulci deeply impressed, thus pre- and postsclerites meeting at distinct oblique angle (†
|
||
<taxonomicName id="4C2B4D61FFE8AA21D1FBF913FBD196B8" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[972,1094,1669,1690]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="species" species="contega">
|
||
<emphasis id="B95FEAF0FFE8AA21D1FBF913FBD196B8" box="[972,1094,1669,1690]" italics="true" pageId="9" pageNumber="1364">G. contega</emphasis>
|
||
</taxonomicName>
|
||
, †
|
||
<taxonomicName id="4C2B4D61FFE8AA21D653F913FB4096B8" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[1124,1239,1669,1690]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="species" species="magna">
|
||
<emphasis id="B95FEAF0FFE8AA21D653F913FB4096B8" box="[1124,1239,1669,1690]" italics="true" pageId="9" pageNumber="1364">G. magna</emphasis>
|
||
</taxonomicName>
|
||
, †
|
||
<taxonomicName id="4C2B4D61FFE8AA21D6C3F913FACB96B8" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[1268,1372,1669,1690]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="species" species="pilosa">
|
||
<emphasis id="B95FEAF0FFE8AA21D6C3F913FACB96B8" box="[1268,1372,1669,1690]" italics="true" pageId="9" pageNumber="1364">G. pilosa</emphasis>
|
||
</taxonomicName>
|
||
); (12) head in full-face view broader lateromedially than long anteroposteriorly, excluding eyes (
|
||
<figureCitation id="13102A67FFE8AA21D740F954FC1C96D4" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="9" pageNumber="1364">Fig. 2L</figureCitation>
|
||
); vs. head longer than broad (†
|
||
<taxonomicName id="4C2B4D61FFE8AA21D728F977FA0A96D4" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[1311,1437,1761,1782]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="species" species="contega">
|
||
<emphasis id="B95FEAF0FFE8AA21D728F977FA0A96D4" box="[1311,1437,1761,1782]" italics="true" pageId="9" pageNumber="1364">G. contega</emphasis>
|
||
</taxonomicName>
|
||
, †
|
||
<taxonomicName id="4C2B4D61FFE8AA21D14DF896FC799737" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[890,1006,1792,1813]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="species" species="magna">
|
||
<emphasis id="B95FEAF0FFE8AA21D14DF896FC799737" box="[890,1006,1792,1813]" italics="true" pageId="9" pageNumber="1364">G. magna</emphasis>
|
||
</taxonomicName>
|
||
, †
|
||
<taxonomicName id="4C2B4D61FFE8AA21D63AF896FBE19736" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[1037,1142,1791,1813]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="species" species="pilosa">
|
||
<emphasis id="B95FEAF0FFE8AA21D63AF896FBE19736" box="[1037,1142,1791,1813]" italics="true" pageId="9" pageNumber="1364">G. pilosa</emphasis>
|
||
</taxonomicName>
|
||
) [Note 5]; (13) pretarsal claws edentate (
|
||
<figureCitation id="13102A67FFE8AA21D61BF888FB219711" box="[1068,1206,1822,1844]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="9" pageNumber="1364">Fig. 5G, I, J</figureCitation>
|
||
); vs. each claw with a single tooth of variable location (†
|
||
<taxonomicName id="4C2B4D61FFE8AA21D715F8ABFA0A9770" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[1314,1437,1853,1874]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="species" species="contega">
|
||
<emphasis id="B95FEAF0FFE8AA21D715F8ABFA0A9770" box="[1314,1437,1853,1874]" italics="true" pageId="9" pageNumber="1364">G. contega</emphasis>
|
||
</taxonomicName>
|
||
, †
|
||
<taxonomicName id="4C2B4D61FFE8AA21D14DF8CAFC799753" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[890,1006,1884,1905]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="species" species="magna">
|
||
<emphasis id="B95FEAF0FFE8AA21D14DF8CAFC799753" box="[890,1006,1884,1905]" italics="true" pageId="9" pageNumber="1364">G. magna</emphasis>
|
||
</taxonomicName>
|
||
, †
|
||
<taxonomicName id="4C2B4D61FFE8AA21D63BF8CAFBE29752" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[1036,1141,1883,1905]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="1364" phylum="Arthropoda" rank="species" species="pilosa">
|
||
<emphasis id="B95FEAF0FFE8AA21D63BF8CAFBE29752" box="[1036,1141,1883,1905]" italics="true" pageId="9" pageNumber="1364">G. pilosa</emphasis>
|
||
</taxonomicName>
|
||
) [Note 6]; and (14) body small, mesosoma length <
|
||
<quantity id="4CD39B07FFEBAA22D036FF53FDCD90F9" box="[513,602,197,219]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="1365" unit="mm" value="1.5">1.5 mm</quantity>
|
||
(
|
||
<figureCitation id="13102A67FFEBAA22D05AFF53FD5690F8" box="[621,705,197,219]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="10" pageNumber="1365">Fig. 2C</figureCitation>
|
||
); vs. body large, mesosoma length> 1.5 (†
|
||
<taxonomicName id="4C2B4D61FFEBAA22D04DFF72FD6490D8" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[634,755,228,250]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="contega">
|
||
<emphasis id="B95FEAF0FFEBAA22D04DFF72FD6490D8" box="[634,755,228,250]" italics="true" pageId="10" pageNumber="1365">G. contega</emphasis>
|
||
</taxonomicName>
|
||
, †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D2E7FE95FEA39135" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[208,308,258,280]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="pilosa">
|
||
<emphasis id="B95FEAF0FFEBAA22D2E7FE95FEA39135" box="[208,308,258,280]" italics="true" pageId="10" pageNumber="1365">G. pilosa</emphasis>
|
||
</taxonomicName>
|
||
) and> 2.5 (†
|
||
<taxonomicName id="4C2B4D61FFEBAA22D3FAFE95FDAB913A" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[461,572,259,280]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="magna">
|
||
<emphasis id="B95FEAF0FFEBAA22D3FAFE95FDAB913A" box="[461,572,259,280]" italics="true" pageId="10" pageNumber="1365">G. magna</emphasis>
|
||
</taxonomicName>
|
||
);
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFEBAA22D2A4FEB7FDF49450" blockId="10.[147,762,197,1139]" pageId="10" pageNumber="1365">
|
||
<emphasis id="B95FEAF0FFEBAA22D2A4FEB7FD259174" bold="true" pageId="10" pageNumber="1365">
|
||
VI. Further distinguished by species in the
|
||
<taxonomicName id="4C2B4D61FFEBAA22D335FEA9FEC79177" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[258,336,319,341]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="pilosa">
|
||
<emphasis id="B95FEAF0FFEBAA22D335FEA9FEC79177" bold="true" box="[258,336,319,341]" italics="true" pageId="10" pageNumber="1365">pilosa</emphasis>
|
||
</taxonomicName>
|
||
group by the following
|
||
</emphasis>
|
||
: (15) anteromedian clypeal margin distinctly evenly curved to an ‘incision’ at the point of contact with the rounded lateroclypeal lobes (
|
||
<figureCitation id="13102A67FFEBAA22D0F9FE0AFF7591F2" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="10" pageNumber="1365">Fig. 2L</figureCitation>
|
||
); vs. anteromedian clypeal margin weakly convex, without distinct incision between the medioclypeus and lateroclypeal lobes, which are themselves anterolaterally angled (†
|
||
<taxonomicName id="4C2B4D61FFEBAA22D2EFFDA3FEC29269" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[216,341,565,587]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="contega">
|
||
<emphasis id="B95FEAF0FFEBAA22D2EFFDA3FEC29269" box="[216,341,565,587]" italics="true" pageId="10" pageNumber="1365">G. contega</emphasis>
|
||
</taxonomicName>
|
||
); (16) in profile, pronotum evenly curved, mesoscutum more-or-less aligned with mesoscutellum and metanotum, and propodeal dorsal and posterior surfaces curving into one another obliquely (
|
||
<figureCitation id="13102A67FFEBAA22D3D6FD26FDAE92E7" box="[481,569,688,710]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="10" pageNumber="1365">Fig. 2C</figureCitation>
|
||
); vs. pronotum, mesonotum, and propodeum each shouldered in appearance, i.e. pronotum with strong anterior dorsolateral bulge, mesonotum with mesoscutal and mesoscutellar regions meeting at nearly a right angle, and propodeal dorsal and posterior surfaces also meeting at nearly a right angle (†
|
||
<taxonomicName id="4C2B4D61FFEBAA22D2EFFC11FEC093BE" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[216,343,903,924]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="contega">
|
||
<emphasis id="B95FEAF0FFEBAA22D2EFFC11FEC093BE" box="[216,343,903,924]" italics="true" pageId="10" pageNumber="1365">G. contega</emphasis>
|
||
</taxonomicName>
|
||
); (17) standing setation on body short, relatively sparse (with the exception of the propodeum and petiolar tergum) (
|
||
<figureCitation id="13102A67FFEBAA22D009FC52FDEC93F8" box="[574,635,964,986]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="10" pageNumber="1365">Fig.5</figureCitation>
|
||
); vs. longer setation present (†
|
||
<taxonomicName id="4C2B4D61FFEBAA22D3ADFC75FD9F93DA" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[410,520,995,1016]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="magna">
|
||
<emphasis id="B95FEAF0FFEBAA22D3ADFC75FD9F93DA" box="[410,520,995,1016]" italics="true" pageId="10" pageNumber="1365">G. magna</emphasis>
|
||
</taxonomicName>
|
||
, †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D016FC75FD1293D5" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[545,645,994,1016]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="pilosa">
|
||
<emphasis id="B95FEAF0FFEBAA22D016FC75FD1293D5" box="[545,645,994,1016]" italics="true" pageId="10" pageNumber="1365">G. pilosa</emphasis>
|
||
</taxonomicName>
|
||
); and (18) anteroventral process of abdominal sternum III (metasomal II) robust but short anteroposteriorly (
|
||
<figureCitation id="13102A67FFEBAA22D2FBFBA8FE899471" box="[204,286,1086,1108]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="10" pageNumber="1365">Fig. 2C</figureCitation>
|
||
); vs. prora anteroposteriorly long, sharkfin-like in form (†
|
||
<taxonomicName id="4C2B4D61FFEBAA22D3BAFBCBFE669450" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[397,497,1117,1138]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="pilosa">
|
||
<emphasis id="B95FEAF0FFEBAA22D3BAFBCBFE669450" box="[397,497,1117,1138]" italics="true" pageId="10" pageNumber="1365">G. pilosa</emphasis>
|
||
</taxonomicName>
|
||
) [Note 7].
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFEBAA22D2A6FB2AFE3494F1" blockId="10.[145,419,1211,1235]" box="[145,419,1211,1235]" pageId="10" pageNumber="1365">
|
||
<heading id="D0DC818EFFEBAA22D2A6FB2AFE3494F1" box="[145,419,1211,1235]" centered="true" fontSize="9" level="2" pageId="10" pageNumber="1365" reason="2">
|
||
<emphasis id="B95FEAF0FFEBAA22D2A6FB2AFE3494F1" box="[145,419,1211,1235]" italics="true" pageId="10" pageNumber="1365">Notes on the diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFEBAA22D2A6FB73FE2F95B1" blockId="10.[145,761,1252,1427]" pageId="10" pageNumber="1365">
|
||
<emphasis id="B95FEAF0FFEBAA22D2A6FB73FF7294DB" box="[145,229,1252,1274]" italics="true" pageId="10" pageNumber="1365">Note 1:</emphasis>
|
||
The right procoxa and protrochanter of the
|
||
<typeStatus id="54908840FFEBAA22D2A6FA94FF65953A" box="[145,242,1282,1304]" pageId="10" pageNumber="1365" type="holotype">holotype</typeStatus>
|
||
are slightly disarticulated. However, the focal details to evaluate are the circular shape of the prodisticoxal foramen and the crank-like (curved) and thin proximal neck of the protrochanter, which can be evaluated from
|
||
<figureCitation id="13102A67FFEBAA22D373FAEBFE2595B0" box="[324,434,1405,1427]" captionStart="Figure 4" captionStartId="15.[163,242,1675,1697]" captionTargetBox="[229,1376,196,1635]" captionTargetId="figure-26@15.[229,1376,195,1635]" captionTargetPageId="15" captionText="Figure 4. Photomicrographs of †G. sternorhabda sp. nov. showing setation and certain key features. A–D, G–I, paratype, CASENT0741234. E, F, J, holotype, CASENT0741233.A, body, ventrolateral oblique view. B, head, dorsolateral oblique view. C–F, body and left legs: D, dorsolateral oblique; (F) is slightly offset from (E), revealing the prora and subpetiolar process. G, protarsus, posterodorsal oblique view. H, mesotarsus, dorsal view. I, metatarsus, dorsal view. J, metatarsus, posterior view. Abbreviations: ad, antennomere distorted, i.e. visibly distorted antennomere associated with decay bubble; cn, cinctus; ms, metanotal spiracle; pr, prora; psf, anterior flange of propodeal spiracle; ptb, probasitarsal brush; spp, subpetiolar process; ss, standing setae; tnc, transverse mesonotal carina." figureDoi="http://doi.org/10.5281/zenodo.6994466" httpUri="https://zenodo.org/record/6994466/files/figure.png" pageId="10" pageNumber="1365">
|
||
Figure
|
||
<specimenCount id="9D2DFD6BFFEBAA22D3A1FAEBFE2595B0" box="[406,434,1405,1426]" pageId="10" pageNumber="1365" type="generic">4F</specimenCount>
|
||
</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFEBAA22D2A6FA2AFDBF9726" blockId="10.[145,761,1468,1796]" pageId="10" pageNumber="1365">
|
||
<emphasis id="B95FEAF0FFEBAA22D2A6FA2AFF7695F3" box="[145,225,1468,1489]" italics="true" pageId="10" pageNumber="1365">Note 2:</emphasis>
|
||
The only species of †Sphecomyrminae to have an elongate antennal scrobe is †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D03EFA4DFD1795D2" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[521,640,1499,1520]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="contega">
|
||
<emphasis id="B95FEAF0FFEBAA22D03EFA4DFD1795D2" box="[521,640,1499,1520]" italics="true" pageId="10" pageNumber="1365">G. contega</emphasis>
|
||
</taxonomicName>
|
||
. Although the scrobe of this species was observed via direct examination of the
|
||
<typeStatus id="54908840FFEBAA22D340F98EFE4B960C" box="[375,476,1560,1582]" pageId="10" pageNumber="1365" type="holotype">holotype</typeStatus>
|
||
, it remains a possibility that this is an artefact because other specimens of †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D2A9F9C3FECB9648" authorityName="Nel & Perrault" authorityYear="2004" box="[158,348,1621,1642]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FEAF0FFEBAA22D2A9F9C3FECB9648" box="[158,348,1621,1642]" italics="true" pageId="10" pageNumber="1365">Gerontoformica</emphasis>
|
||
</taxonomicName>
|
||
may have asymmetrical, sulcuslike distortions of the head corresponding to the position of the scrobe (see the ‘preservation’ section of the †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D2DFF927FEC896E4" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[232,351,1713,1734]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B95FEAF0FFEBAA22D2DFF927FEC896E4" box="[232,351,1713,1734]" italics="true" pageId="10" pageNumber="1365">G. gracilis</emphasis>
|
||
</taxonomicName>
|
||
description;
|
||
<figureCitation id="13102A67FFEBAA22D3C0F927FDD096E4" box="[503,583,1713,1735]" captionStart="Figure 10" captionStartId="24.[145,226,1060,1082]" captionTargetBox="[152,1397,206,1011]" captionTargetId="figure-386@24.[145,1425,195,1020]" captionTargetPageId="24" captionText="Figure 10. Features hidden by appendages, including artefacts caused by close proximity of limbs. †Gerontoformica gracilis: A, B, D, adult, CASENT0741232; C, pupa (CASENT0741231). A, C, mesosoma in lateral view with the legs distal to the coxae digitally removed, with the exception of the fore leg of (C), which is not cropped from the render. B, D, head in facial view with the scapes digitally rendered (B) and removed (D). Abbreviations: mspa, mesopectal artefact; sca, scapal artefact." figureDoi="http://doi.org/10.5281/zenodo.6994480" httpUri="https://zenodo.org/record/6994480/files/figure.png" pageId="10" pageNumber="1365">Fig. 10</figureCitation>
|
||
). Conducting a µ-CT scan of the type specimen and/ or the accrual of additional specimens are necessary.
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFEBAA22D2A6F8B8FCE49156" blockId="10.[145,760,1837,1890]" lastBlockId="10.[809,1424,197,372]" pageId="10" pageNumber="1365">
|
||
<emphasis id="B95FEAF0FFEBAA22D2A6F8B8FF669761" box="[145,241,1837,1859]" italics="true" pageId="10" pageNumber="1365">N o t e 3:</emphasis>
|
||
T h e a n t e r o m e d i a n c l y p e a l p r o c e s s o f †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D2A9F8DAFEDF9743" authorityName="Wilson & Brown" authorityYear="1967" box="[158,328,1868,1889]" class="Insecta" family="Formicidae" genus="Sphecomyrma" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FEAF0FFEBAA22D2A9F8DAFEDF9743" box="[158,328,1868,1889]" italics="true" pageId="10" pageNumber="1365">Sphecomyrma</emphasis>
|
||
</taxonomicName>
|
||
is lateromedially thin and at least as long as broad; it is distinctly bordered laterally by the medioclypeus, i.e. the clypeal ‘disc’ between the lateral clypeal lobes. In contrast, the entire medioclypeus of †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D1C8FEB7FBFD9114" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[1023,1130,289,310]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="tendir">
|
||
<emphasis id="B95FEAF0FFEBAA22D1C8FEB7FBFD9114" box="[1023,1130,289,310]" italics="true" pageId="10" pageNumber="1365">G. tendir</emphasis>
|
||
</taxonomicName>
|
||
is apparently produced. See Note 5 of the ‘Notes on classification’ section above.
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFEBAA22D11EFE08FA8291D2" blockId="10.[809,1426,413,496]" pageId="10" pageNumber="1365">
|
||
<emphasis id="B95FEAF0FFEBAA22D11EFE08FCE19190" box="[809,886,413,435]" italics="true" pageId="10" pageNumber="1365">Note 4:</emphasis>
|
||
The form of the subpetiolar process is unknown for four species within the genus: †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D734FE2AFA1D91F3" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[1283,1418,444,465]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="contega">
|
||
<emphasis id="B95FEAF0FFEBAA22D734FE2AFA1D91F3" box="[1283,1418,444,465]" italics="true" pageId="10" pageNumber="1365">G. contega</emphasis>
|
||
</taxonomicName>
|
||
, †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D102FE4DFC2F91D2" authorityName="Nel & Perrault" authorityYear="2004" box="[821,952,475,496]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="cretacica">
|
||
<emphasis id="B95FEAF0FFEBAA22D102FE4DFC2F91D2" box="[821,952,475,496]" italics="true" pageId="10" pageNumber="1365">G. cretacica</emphasis>
|
||
</taxonomicName>
|
||
, †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D1E6FE4DFBA891D2" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[977,1087,475,496]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="magna">
|
||
<emphasis id="B95FEAF0FFEBAA22D1E6FE4DFBA891D2" box="[977,1087,475,496]" italics="true" pageId="10" pageNumber="1365">G. magna</emphasis>
|
||
</taxonomicName>
|
||
and †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D6B4FE4DFA9991CD" baseAuthorityName="Engel & Grimaldi" baseAuthorityYear="2005" box="[1155,1294,474,496]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="orientalis">
|
||
<emphasis id="B95FEAF0FFEBAA22D6B4FE4DFA9991CD" box="[1155,1294,474,496]" italics="true" pageId="10" pageNumber="1365">G. orientalis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFEBAA22D11EFD8CFC589288" blockId="10.[809,1425,537,682]" pageId="10" pageNumber="1365">
|
||
<emphasis id="B95FEAF0FFEBAA22D11EFD8CFCEB920C" box="[809,892,537,559]" italics="true" pageId="10" pageNumber="1365">Note 5:</emphasis>
|
||
Although it is difficult to evaluate the head shape of †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D1AFFDAEFB9D926F" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[920,1034,568,589]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="contega">
|
||
<emphasis id="B95FEAF0FFEBAA22D1AFFDAEFB9D926F" box="[920,1034,568,589]" italics="true" pageId="10" pageNumber="1365">G. contega</emphasis>
|
||
</taxonomicName>
|
||
from the
|
||
<typeStatus id="54908840FFEBAA22D640FDAEFB4F926C" box="[1143,1240,568,590]" pageId="10" pageNumber="1365" type="holotype">holotype</typeStatus>
|
||
due to cut of the amber matrix, the head does appear to be longer than broad. Reevaluation of this condition through µ-CT is recommended.
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFEBAA22D11EFD45FC2893DE" blockId="10.[809,1426,722,1020]" pageId="10" pageNumber="1365">
|
||
<emphasis id="B95FEAF0FFEBAA22D11EFD45FC1492C5" box="[809,899,722,744]" italics="true" pageId="10" pageNumber="1365">Note 6:</emphasis>
|
||
Teeth on the pretarsal claws have been recorded for most species of †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D6F1FD67FA069325" authorityName="Nel & Perrault" authorityYear="2004" box="[1222,1425,753,775]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FEAF0FFEBAA22D6F1FD67FA069325" box="[1222,1425,753,775]" italics="true" pageId="10" pageNumber="1365">Gerontoformica</emphasis>
|
||
</taxonomicName>
|
||
(†
|
||
<taxonomicName id="4C2B4D61FFEBAA22D109FC86FC249307" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[830,947,784,805]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="contega">
|
||
<emphasis id="B95FEAF0FFEBAA22D109FC86FC249307" box="[830,947,784,805]" italics="true" pageId="10" pageNumber="1365">G. contega</emphasis>
|
||
</taxonomicName>
|
||
, †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D1FAFC86FBD59307" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[973,1090,784,805]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B95FEAF0FFEBAA22D1FAFC86FBD59307" box="[973,1090,784,805]" italics="true" pageId="10" pageNumber="1365">G. gracilis</emphasis>
|
||
</taxonomicName>
|
||
, †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D66BFC86FB5D9307" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[1116,1226,784,805]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="magna">
|
||
<emphasis id="B95FEAF0FFEBAA22D66BFC86FB5D9307" box="[1116,1226,784,805]" italics="true" pageId="10" pageNumber="1365">G. magna</emphasis>
|
||
</taxonomicName>
|
||
, †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D6D3FC86FA1C9307" box="[1252,1419,784,805]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="occidentalis">
|
||
<emphasis id="B95FEAF0FFEBAA22D6D3FC86FA1C9307" box="[1252,1419,784,805]" italics="true" pageId="10" pageNumber="1365">G. occidentalis</emphasis>
|
||
</taxonomicName>
|
||
, †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D102FCB9FC0F9361" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[821,920,814,836]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="pilosa">
|
||
<emphasis id="B95FEAF0FFEBAA22D102FCB9FC0F9361" box="[821,920,814,836]" italics="true" pageId="10" pageNumber="1365">G. pilosa</emphasis>
|
||
</taxonomicName>
|
||
, †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D187FCB9FBB29361" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[944,1061,814,836]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="robusta">
|
||
<emphasis id="B95FEAF0FFEBAA22D187FCB9FBB29361" box="[944,1061,814,836]" italics="true" pageId="10" pageNumber="1365">G. robusta</emphasis>
|
||
</taxonomicName>
|
||
, †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D60AFCB9FB259361" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[1085,1202,814,836]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="spiralis">
|
||
<emphasis id="B95FEAF0FFEBAA22D60AFCB9FB259361" box="[1085,1202,814,836]" italics="true" pageId="10" pageNumber="1365">G. spiralis</emphasis>
|
||
</taxonomicName>
|
||
and †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D6C1FCB9FA149361" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[1270,1411,814,836]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="subcuspis">
|
||
<emphasis id="B95FEAF0FFEBAA22D6C1FCB9FA149361" box="[1270,1411,814,836]" italics="true" pageId="10" pageNumber="1365">G. subcuspis</emphasis>
|
||
</taxonomicName>
|
||
). The condition of having reduced teeth on the pretarsal claws, however, is shared with †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D6F2FCFAFACF93A3" authorityName="Nel & Perrault" authorityYear="2004" box="[1221,1368,876,897]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="cretacica">
|
||
<emphasis id="B95FEAF0FFEBAA22D6F2FCFAFACF93A3" box="[1221,1368,876,897]" italics="true" pageId="10" pageNumber="1365">G. cretacica</emphasis>
|
||
</taxonomicName>
|
||
and †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D101FC1DFC0893BD" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[822,927,906,928]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="tendir">
|
||
<emphasis id="B95FEAF0FFEBAA22D101FC1DFC0893BD" box="[822,927,906,928]" italics="true" pageId="10" pageNumber="1365">G. tendir</emphasis>
|
||
</taxonomicName>
|
||
. Notably,
|
||
<bibRefCitation id="EFBA4B13FFEBAA22D62DFC1CFAB693BD" author="Perrichot V & Nel A & Neraudeau D & Lacau S & Guyot T" box="[1050,1313,906,928]" pageId="10" pageNumber="1365" pagination="91 - 97" refId="ref25043" refString="Perrichot V, Nel A, Neraudeau D, Lacau S, Guyot T. 2008. New fossil ants in French Cretaceous amber (Hymenoptera: Formicidae). Naturwissenschaften 95: 91 - 97." type="journal article" year="2008">
|
||
Perrichot
|
||
<emphasis id="B95FEAF0FFEBAA22D6A5FC1DFB5993BD" box="[1170,1230,906,928]" italics="true" pageId="10" pageNumber="1365">et al.</emphasis>
|
||
(2008)
|
||
</bibRefCitation>
|
||
recorded minute teeth on the pretarsal claws of †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D6D0FC3FFA1C939C" box="[1255,1419,937,958]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="occidentalis">
|
||
<emphasis id="B95FEAF0FFEBAA22D6D0FC3FFA1C939C" box="[1255,1419,937,958]" italics="true" pageId="10" pageNumber="1365">G. occidentalis</emphasis>
|
||
</taxonomicName>
|
||
, which suggests the need to re-evaluate the
|
||
<typeStatus id="54908840FFEBAA22D722FC5EFAE193FC" box="[1301,1398,968,990]" pageId="10" pageNumber="1365" type="holotype">holotype</typeStatus>
|
||
of †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D102FC71FC2F93DE" authorityName="Nel & Perrault" authorityYear="2004" box="[821,952,999,1020]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="cretacica">
|
||
<emphasis id="B95FEAF0FFEBAA22D102FC71FC2F93DE" box="[821,952,999,1020]" italics="true" pageId="10" pageNumber="1365">G. cretacica</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFEBAA22D11EFBB0FC5E956C" blockId="10.[809,1426,1061,1358]" pageId="10" pageNumber="1365">
|
||
<emphasis id="B95FEAF0FFEBAA22D11EFBB0FCEC9418" box="[809,891,1061,1083]" italics="true" pageId="10" pageNumber="1365">Note 7:</emphasis>
|
||
The prora is clearly an important structural feature for distinguishing stem ants but is not clearly visible in most specimens, including †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D738FBF5FA06945A" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[1295,1425,1123,1144]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="contega">
|
||
<emphasis id="B95FEAF0FFEBAA22D738FBF5FA06945A" box="[1295,1425,1123,1144]" italics="true" pageId="10" pageNumber="1365">G. contega</emphasis>
|
||
</taxonomicName>
|
||
and †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D159FB17FC7294B5" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[878,997,1153,1175]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="magna">
|
||
<emphasis id="B95FEAF0FFEBAA22D159FB17FC7294B5" box="[878,997,1153,1175]" italics="true" pageId="10" pageNumber="1365">G. magna</emphasis>
|
||
</taxonomicName>
|
||
in the
|
||
<taxonomicName id="4C2B4D61FFEBAA22D671FB17FB0794B4" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[1094,1168,1153,1174]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="pilosa">
|
||
<emphasis id="B95FEAF0FFEBAA22D671FB17FB0794B4" box="[1094,1168,1153,1174]" italics="true" pageId="10" pageNumber="1365">pilosa</emphasis>
|
||
</taxonomicName>
|
||
group, †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D737FB17FA0694B5" authorityName="Nel & Perrault" authorityYear="2004" box="[1280,1425,1153,1175]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="cretacica">
|
||
<emphasis id="B95FEAF0FFEBAA22D737FB17FA0694B5" box="[1280,1425,1153,1175]" italics="true" pageId="10" pageNumber="1365">G. cretacica</emphasis>
|
||
</taxonomicName>
|
||
and †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D15CFB36FC689497" baseAuthorityName="Engel & Grimaldi" baseAuthorityYear="2005" box="[875,1023,1184,1205]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="orientalis">
|
||
<emphasis id="B95FEAF0FFEBAA22D15CFB36FC689497" box="[875,1023,1184,1205]" italics="true" pageId="10" pageNumber="1365">G. orientalis</emphasis>
|
||
</taxonomicName>
|
||
in the
|
||
<taxonomicName id="4C2B4D61FFEBAA22D66EFB36FB559497" authorityName="Nel & Perrault" authorityYear="2004" box="[1113,1218,1184,1205]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="cretacica">
|
||
<emphasis id="B95FEAF0FFEBAA22D66EFB36FB559497" box="[1113,1218,1184,1205]" italics="true" pageId="10" pageNumber="1365">cretacica</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C2B4D61FFEBAA22D6FCFB36FAAA9497" baseAuthorityName="Engel & Grimaldi" baseAuthorityYear="2005" box="[1227,1341,1184,1205]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="orientalis">
|
||
<emphasis id="B95FEAF0FFEBAA22D6FCFB36FAAA9497" box="[1227,1341,1184,1205]" italics="true" pageId="10" pageNumber="1365">orientalis</emphasis>
|
||
</taxonomicName>
|
||
group, plus †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D158FB29FC4A94F6" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[879,989,1215,1236]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="rugosa">
|
||
<emphasis id="B95FEAF0FFEBAA22D158FB29FC4A94F6" box="[879,989,1215,1236]" italics="true" pageId="10" pageNumber="1365">G. rugosa</emphasis>
|
||
</taxonomicName>
|
||
and †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D611FB29FB1994F1" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[1062,1166,1214,1236]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="tendir">
|
||
<emphasis id="B95FEAF0FFEBAA22D611FB29FB1994F1" box="[1062,1166,1214,1236]" italics="true" pageId="10" pageNumber="1365">G. tendir</emphasis>
|
||
</taxonomicName>
|
||
. It is extremely small and nearly absent in †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D61DFB4BFB0994D0" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[1066,1182,1245,1266]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B95FEAF0FFEBAA22D61DFB4BFB0994D0" box="[1066,1182,1245,1266]" italics="true" pageId="10" pageNumber="1365">G. gracilis</emphasis>
|
||
</taxonomicName>
|
||
and †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D6D2FB4BFA1C94D0" box="[1253,1419,1245,1266]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="occidentalis">
|
||
<emphasis id="B95FEAF0FFEBAA22D6D2FB4BFA1C94D0" box="[1253,1419,1245,1266]" italics="true" pageId="10" pageNumber="1365">G. occidentalis</emphasis>
|
||
</taxonomicName>
|
||
, and it is developed, but short and comparatively inconspicuous in †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D62AFA8DFB34950D" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[1053,1187,1306,1328]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="robusta">
|
||
<emphasis id="B95FEAF0FFEBAA22D62AFA8DFB34950D" box="[1053,1187,1306,1328]" italics="true" pageId="10" pageNumber="1365">G. robusta</emphasis>
|
||
</taxonomicName>
|
||
, †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D6F1FA8DFAD9950D" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[1222,1358,1306,1328]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="spiralis">
|
||
<emphasis id="B95FEAF0FFEBAA22D6F1FA8DFAD9950D" box="[1222,1358,1306,1328]" italics="true" pageId="10" pageNumber="1365">G. spiralis</emphasis>
|
||
</taxonomicName>
|
||
, and †
|
||
<taxonomicName id="4C2B4D61FFEBAA22D102FAAFFC55956C" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[821,962,1337,1358]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="1365" phylum="Arthropoda" rank="species" species="subcuspis">
|
||
<emphasis id="B95FEAF0FFEBAA22D102FAAFFC55956C" box="[821,962,1337,1358]" italics="true" pageId="10" pageNumber="1365">G. subcuspis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3316569FFEBAA26D184FA07FD569748" lastPageId="14" lastPageNumber="1369" pageId="10" pageNumber="1365" type="description">
|
||
<paragraph id="8B9436E2FFEBAA22D184FA07FA90958A" blockId="10.[947,1287,1425,1450]" box="[947,1287,1425,1450]" pageId="10" pageNumber="1365">MEASUREMENTS AND INDICES</paragraph>
|
||
<paragraph id="8B9436E2FFEBAA22D11EFA2FFB419608" blockId="10.[809,1425,1465,1579]" pageId="10" pageNumber="1365">
|
||
<emphasis id="B95FEAF0FFEBAA22D11EFA2FFBD095EC" box="[809,1095,1465,1487]" italics="true" pageId="10" pageNumber="1365">
|
||
<typeStatus id="54908840FFEBAA22D11EFA2FFC1895EC" box="[809,911,1465,1486]" pageId="10" pageNumber="1365" type="holotype">Holotype</typeStatus>
|
||
, specimen C-33:
|
||
</emphasis>
|
||
HWed = 0.87; HWev = 0.95; EWl = 0.14; HD = 0.55; ML = 1.25; PnLi = 0.50; PnWa = 0.30; MnL = 0.27; AIIILm = 0.48; AIIILl = 0.53; HPI = 0.64; HSI = 0.69; AIIILI = 0.91.
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFEBAA22D11EF9C3FB4096E7" blockId="10.[809,1425,1620,1734]" pageId="10" pageNumber="1365">
|
||
<emphasis id="B95FEAF0FFEBAA22D11EF9C3FBDF964B" box="[809,1096,1620,1642]" italics="true" pageId="10" pageNumber="1365">
|
||
<typeStatus id="54908840FFEBAA22D11EF9C3FC079648" box="[809,912,1621,1642]" pageId="10" pageNumber="1365" type="paratype">Paratype</typeStatus>
|
||
, specimen C-34:
|
||
</emphasis>
|
||
HWed = 0.94; HWev = 1.06; EWl = 0.20; HD = 0.66; ML = 1.39; PnLi = 0.45; PnWa = 0.27; MnL = 0.28; AIIILm = 0.50; AIIILl = 0.64; HPI = 0.70; HSI = 0.67; AIIILI = 0.78.
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFEBAA22D11EF978FB419742" blockId="10.[809,1425,1774,1888]" pageId="10" pageNumber="1365">
|
||
<emphasis id="B95FEAF0FFEBAA22D11EF978FBD29721" box="[809,1093,1774,1795]" italics="true" pageId="10" pageNumber="1365">UFV-LABECOL-009656:</emphasis>
|
||
HWed = 0.79; HWev = 0.85; EWl = 0.11; HD = 0.46; ML = 1.05; PnLi = (~0.35–0.38); PnWa = –; MnL = 0.29; AIIILm = 0.4; AIIILl = 0.48; HPI = 0.58; HSI = 0.75; AIIILI = 0.83.
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFEAAA23D294FF50FEB890FF" blockId="11.[163,303,198,221]" box="[163,303,198,221]" pageId="11" pageNumber="1366">
|
||
<emphasis id="B95FEAF0FFEAAA23D294FF50FEB890FF" box="[163,303,198,221]" italics="true" pageId="11" pageNumber="1366">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFEAAA23D294FF78FAA39398" blockId="11.[163,780,237,1884]" lastBlockId="11.[827,1443,197,955]" pageId="11" pageNumber="1366">
|
||
<emphasis id="B95FEAF0FFEAAA23D294FF78FF679121" box="[163,240,237,259]" italics="true" pageId="11" pageNumber="1366">Head:</emphasis>
|
||
The
|
||
<emphasis id="B95FEAF0FFEAAA23D308FF7BFE139120" bold="true" box="[319,388,237,258]" pageId="11" pageNumber="1366">head</emphasis>
|
||
is broad in facial view, i. e. lateromedially wider than anteroposteriorly long from the anterior clypeal margin to the apparent posterior head margin (
|
||
<figureCitation id="13102A67FFEAAA23D378FEDFFE32917C" box="[335,421,329,351]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="11" pageNumber="1366">Fig. 2L</figureCitation>
|
||
); in posterior or lateral view, the head is dorsoventrally narrow, with the vertexal region not being particularly domed; standing setae are present on the vertexal and facial regions [Note 1]; there are short, decumbent setae distributed sparsely on the head capsule, a few longer, and suberect setae medially on the vertex near the ocelli. The
|
||
<emphasis id="B95FEAF0FFEAAA23D0B3FD97FF4B9217" bold="true" pageId="11" pageNumber="1366">compound eyes</emphasis>
|
||
are situated in the posterior third of the head (
|
||
<figureCitation id="13102A67FFEAAA23D29CFDA9FF6C9276" box="[171,251,575,597]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="11" pageNumber="1366">Fig. 2L</figureCitation>
|
||
); they bulge laterally, breaking the silhouette of the lateral head margins in facial view; their height above the surrounding surfaces of the cranium is comparatively low; they comprise over 100 ommatidia, but not more than 200 (
|
||
<figureCitation id="13102A67FFEAAA23D3FEFD2FFDB792EC" box="[457,544,697,719]" captionStart="Figure 4" captionStartId="15.[163,242,1675,1697]" captionTargetBox="[229,1376,196,1635]" captionTargetId="figure-26@15.[229,1376,195,1635]" captionTargetPageId="15" captionText="Figure 4. Photomicrographs of †G. sternorhabda sp. nov. showing setation and certain key features. A–D, G–I, paratype, CASENT0741234. E, F, J, holotype, CASENT0741233.A, body, ventrolateral oblique view. B, head, dorsolateral oblique view. C–F, body and left legs: D, dorsolateral oblique; (F) is slightly offset from (E), revealing the prora and subpetiolar process. G, protarsus, posterodorsal oblique view. H, mesotarsus, dorsal view. I, metatarsus, dorsal view. J, metatarsus, posterior view. Abbreviations: ad, antennomere distorted, i.e. visibly distorted antennomere associated with decay bubble; cn, cinctus; ms, metanotal spiracle; pr, prora; psf, anterior flange of propodeal spiracle; ptb, probasitarsal brush; spp, subpetiolar process; ss, standing setae; tnc, transverse mesonotal carina." figureDoi="http://doi.org/10.5281/zenodo.6994466" httpUri="https://zenodo.org/record/6994466/files/figure.png" pageId="11" pageNumber="1366">Fig. 4C</figureCitation>
|
||
) [Note 2]; they are apparently glabrous, i.e. lacking interstitial setation. The three
|
||
<emphasis id="B95FEAF0FFEAAA23D329FD60FEF1932E" bold="true" box="[286,358,758,780]" pageId="11" pageNumber="1366">ocelli</emphasis>
|
||
are completely developed (
|
||
<figureCitation id="13102A67FFEAAA23D09EFD61FD6C932E" box="[681,763,759,781]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="11" pageNumber="1366">Fig. 2L</figureCitation>
|
||
). The
|
||
<emphasis id="B95FEAF0FFEAAA23D2E1FC83FE0F9309" bold="true" box="[214,408,789,811]" pageId="11" pageNumber="1366">frontal carinae</emphasis>
|
||
diverge posterolaterally, toward but not reaching the compound eyes (
|
||
<figureCitation id="13102A67FFEAAA23D072FCA2FD0E936B" box="[581,665,820,842]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="11" pageNumber="1366">Figs 2L</figureCitation>
|
||
,
|
||
<figureCitation id="13102A67FFEAAA23D092FCA2FD529368" box="[677,709,820,842]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="11" pageNumber="1366">5A</figureCitation>
|
||
); they are sinuate in form, i.e. from their anterior margins they are directed relatively medially then bulge laterally before curving laterad; their anterior termini are close to the epistomal line, but not extending on to the clypeus; the minimum distance between the frontal carinae is about 0.29× maximum head width as measured in full face view. The
|
||
<emphasis id="B95FEAF0FFEAAA23D01CFB9DFC939402" bold="true" box="[555,772,1035,1056]" pageId="11" pageNumber="1366">antennal scrobes</emphasis>
|
||
, or depressed contact surfaces of the face laterad the frontal carinae, are parallel in orientation relative to the frontal carinae and are apparently longer than wide (
|
||
<figureCitation id="13102A67FFEAAA23D2DAFB13FEDE94B8" box="[237,329,1157,1179]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="11" pageNumber="1366">Figs 2L</figureCitation>
|
||
,
|
||
<figureCitation id="13102A67FFEAAA23D36EFB13FEED94B8" box="[345,378,1157,1178]" captionStart="Figure 4" captionStartId="15.[163,242,1675,1697]" captionTargetBox="[229,1376,196,1635]" captionTargetId="figure-26@15.[229,1376,195,1635]" captionTargetPageId="15" captionText="Figure 4. Photomicrographs of †G. sternorhabda sp. nov. showing setation and certain key features. A–D, G–I, paratype, CASENT0741234. E, F, J, holotype, CASENT0741233.A, body, ventrolateral oblique view. B, head, dorsolateral oblique view. C–F, body and left legs: D, dorsolateral oblique; (F) is slightly offset from (E), revealing the prora and subpetiolar process. G, protarsus, posterodorsal oblique view. H, mesotarsus, dorsal view. I, metatarsus, dorsal view. J, metatarsus, posterior view. Abbreviations: ad, antennomere distorted, i.e. visibly distorted antennomere associated with decay bubble; cn, cinctus; ms, metanotal spiracle; pr, prora; psf, anterior flange of propodeal spiracle; ptb, probasitarsal brush; spp, subpetiolar process; ss, standing setae; tnc, transverse mesonotal carina." figureDoi="http://doi.org/10.5281/zenodo.6994466" httpUri="https://zenodo.org/record/6994466/files/figure.png" pageId="11" pageNumber="1366">4B</figureCitation>
|
||
). The
|
||
<emphasis id="B95FEAF0FFEAAA23D3FDFB13FD0E94B9" bold="true" box="[458,665,1157,1179]" pageId="11" pageNumber="1366">antennal toruli</emphasis>
|
||
abut but do not indent the posterior clypeal margin [Note 3]; they are in the form of a low, more-or-less even ring. The
|
||
<emphasis id="B95FEAF0FFEAAA23D2EFFB77FEC494D4" bold="true" box="[216,339,1249,1270]" pageId="11" pageNumber="1366">antennae</emphasis>
|
||
are 12-merous (
|
||
<figureCitation id="13102A67FFEAAA23D022FB77FDFE94D4" box="[533,617,1249,1271]" captionStart="Figure 4" captionStartId="15.[163,242,1675,1697]" captionTargetBox="[229,1376,196,1635]" captionTargetId="figure-26@15.[229,1376,195,1635]" captionTargetPageId="15" captionText="Figure 4. Photomicrographs of †G. sternorhabda sp. nov. showing setation and certain key features. A–D, G–I, paratype, CASENT0741234. E, F, J, holotype, CASENT0741233.A, body, ventrolateral oblique view. B, head, dorsolateral oblique view. C–F, body and left legs: D, dorsolateral oblique; (F) is slightly offset from (E), revealing the prora and subpetiolar process. G, protarsus, posterodorsal oblique view. H, mesotarsus, dorsal view. I, metatarsus, dorsal view. J, metatarsus, posterior view. Abbreviations: ad, antennomere distorted, i.e. visibly distorted antennomere associated with decay bubble; cn, cinctus; ms, metanotal spiracle; pr, prora; psf, anterior flange of propodeal spiracle; ptb, probasitarsal brush; spp, subpetiolar process; ss, standing setae; tnc, transverse mesonotal carina." figureDoi="http://doi.org/10.5281/zenodo.6994466" httpUri="https://zenodo.org/record/6994466/files/figure.png" pageId="11" pageNumber="1366">Fig. 4B</figureCitation>
|
||
). The
|
||
<emphasis id="B95FEAF0FFEAAA23D083FB77FC9D94D4" bold="true" box="[692,778,1249,1270]" pageId="11" pageNumber="1366">scapes</emphasis>
|
||
are somewhat flattened and curved (
|
||
<figureCitation id="13102A67FFEAAA23D062FA96FD6C9537" box="[597,763,1280,1302]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="11" pageNumber="1366">Fig. 2C, F, I, L</figureCitation>
|
||
); they are about three to four times as long as wide (
|
||
<figureCitation id="13102A67FFEAAA23D0D7FA88FF2C9570" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="11" pageNumber="1366">Fig. 2I</figureCitation>
|
||
); their length is about half the width of the head, and less than half the length of the head; they bear a vestiture of short subdecumbent to suberect setae. The
|
||
<emphasis id="B95FEAF0FFEAAA23D2EEFA0FFED0958D" bold="true" box="[217,327,1433,1455]" pageId="11" pageNumber="1366">pedicels</emphasis>
|
||
are about twice as long as wide (
|
||
<figureCitation id="13102A67FFEAAA23D0E9FA0FFF5695EF" captionStart="Figure 4" captionStartId="15.[163,242,1675,1697]" captionTargetBox="[229,1376,196,1635]" captionTargetId="figure-26@15.[229,1376,195,1635]" captionTargetPageId="15" captionText="Figure 4. Photomicrographs of †G. sternorhabda sp. nov. showing setation and certain key features. A–D, G–I, paratype, CASENT0741234. E, F, J, holotype, CASENT0741233.A, body, ventrolateral oblique view. B, head, dorsolateral oblique view. C–F, body and left legs: D, dorsolateral oblique; (F) is slightly offset from (E), revealing the prora and subpetiolar process. G, protarsus, posterodorsal oblique view. H, mesotarsus, dorsal view. I, metatarsus, dorsal view. J, metatarsus, posterior view. Abbreviations: ad, antennomere distorted, i.e. visibly distorted antennomere associated with decay bubble; cn, cinctus; ms, metanotal spiracle; pr, prora; psf, anterior flange of propodeal spiracle; ptb, probasitarsal brush; spp, subpetiolar process; ss, standing setae; tnc, transverse mesonotal carina." figureDoi="http://doi.org/10.5281/zenodo.6994466" httpUri="https://zenodo.org/record/6994466/files/figure.png" pageId="11" pageNumber="1366">Fig. 4F</figureCitation>
|
||
); they are about one-third the length of the scapes, and somewhat more than half the length of the third antennomere; their setation is similar to that of the scapes. The
|
||
<emphasis id="B95FEAF0FFEAAA23D307F985FE37960B" bold="true" box="[304,416,1555,1577]" pageId="11" pageNumber="1366">flagellae</emphasis>
|
||
are longer than the mesosoma and are simple, i.e. not thickening distally (
|
||
<figureCitation id="13102A67FFEAAA23D09EF9A4FD6C966A" box="[681,763,1586,1608]" captionStart="Figure 4" captionStartId="15.[163,242,1675,1697]" captionTargetBox="[229,1376,196,1635]" captionTargetId="figure-26@15.[229,1376,195,1635]" captionTargetPageId="15" captionText="Figure 4. Photomicrographs of †G. sternorhabda sp. nov. showing setation and certain key features. A–D, G–I, paratype, CASENT0741234. E, F, J, holotype, CASENT0741233.A, body, ventrolateral oblique view. B, head, dorsolateral oblique view. C–F, body and left legs: D, dorsolateral oblique; (F) is slightly offset from (E), revealing the prora and subpetiolar process. G, protarsus, posterodorsal oblique view. H, mesotarsus, dorsal view. I, metatarsus, dorsal view. J, metatarsus, posterior view. Abbreviations: ad, antennomere distorted, i.e. visibly distorted antennomere associated with decay bubble; cn, cinctus; ms, metanotal spiracle; pr, prora; psf, anterior flange of propodeal spiracle; ptb, probasitarsal brush; spp, subpetiolar process; ss, standing setae; tnc, transverse mesonotal carina." figureDoi="http://doi.org/10.5281/zenodo.6994466" httpUri="https://zenodo.org/record/6994466/files/figure.png" pageId="11" pageNumber="1366">Fig. 4F</figureCitation>
|
||
); they bear a range of standing and appressed setae;
|
||
<emphasis id="B95FEAF0FFEAAA23D294F9F9FEF996A7" bold="true" box="[163,366,1647,1669]" pageId="11" pageNumber="1366">flagellomeres I</emphasis>
|
||
are the longest, being about four times as long as wide and more than half the length of the scapes;
|
||
<emphasis id="B95FEAF0FFEAAA23D372F93BFDAB96E0" bold="true" box="[325,572,1709,1731]" pageId="11" pageNumber="1366">flagellomeres II–IX</emphasis>
|
||
are about as long as the pedicel;
|
||
<emphasis id="B95FEAF0FFEAAA23D354F95DFDAC96C3" bold="true" box="[355,571,1739,1761]" pageId="11" pageNumber="1366">flagellomeres X</emphasis>
|
||
are longer than flagellomeres II–IX. The
|
||
<emphasis id="B95FEAF0FFEAAA23D3F2F97CFDBF96DD" bold="true" box="[453,552,1770,1791]" pageId="11" pageNumber="1366">clypeus</emphasis>
|
||
is about four times as wide (lateromedially) as long (anteroposteriorly), with the length measured from the midpoints of the anterior and posterior clypeal margins and the width measured between the lateralmost points of the clypeus (
|
||
<figureCitation id="13102A67FFEAAA23D1AAFF75FC7C90DB" box="[925,1003,227,249]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="11" pageNumber="1366">Fig. 2L</figureCitation>
|
||
). The
|
||
<emphasis id="B95FEAF0FFEAAA23D607FF72FAB590DB" bold="true" box="[1072,1314,228,249]" pageId="11" pageNumber="1366">lateroclypeal areas</emphasis>
|
||
are formed as lateral lobes (
|
||
<figureCitation id="13102A67FFEAAA23D63BFE94FBFD9135" box="[1036,1130,258,280]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="11" pageNumber="1366">Figs 2L</figureCitation>
|
||
,
|
||
<figureCitation id="13102A67FFEAAA23D64CFE94FB0C913A" box="[1147,1179,258,280]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="11" pageNumber="1366">5A</figureCitation>
|
||
). The
|
||
<emphasis id="B95FEAF0FFEAAA23D6DAFE94FCE29114" bold="true" pageId="11" pageNumber="1366">medioclypeal area</emphasis>
|
||
is anteriorly convex (
|
||
<figureCitation id="13102A67FFEAAA23D64FFEB7FB5C9114" box="[1144,1227,289,311]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="11" pageNumber="1366">Fig. 2L</figureCitation>
|
||
); its length at the midline of the head is about 0.21× head length also at midline, as measured in full-face view; it bears five or six long and flexuous setae that are situated near the anterior clypeal margin and anteriorly directed, consisting of one medial seta surrounded by a pair of setae and potentially a second, even more lateral pair [Note 4]; the anterior medioclypeal margin bears a row of chaetae. The
|
||
<emphasis id="B95FEAF0FFEAAA23D1CFFD80FB16920E" bold="true" box="[1016,1153,534,556]" pageId="11" pageNumber="1366">mandibles</emphasis>
|
||
are simple and apically bidentate (
|
||
<figureCitation id="13102A67FFEAAA23D183FDA3FB949269" box="[948,1027,565,587]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="11" pageNumber="1366">Fig. 5A</figureCitation>
|
||
). The
|
||
<emphasis id="B95FEAF0FFEAAA23D672FDA3FA9D9268" bold="true" box="[1093,1290,565,587]" pageId="11" pageNumber="1366">maxillary palps</emphasis>
|
||
are 5-merous (
|
||
<figureCitation id="13102A67FFEAAA23D174FDC5FC06924B" box="[835,913,595,617]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="11" pageNumber="1366">Figs 2I</figureCitation>
|
||
,
|
||
<figureCitation id="13102A67FFEAAA23D1A9FDC2FC29924B" box="[926,958,596,617]" captionStart="Figure 4" captionStartId="15.[163,242,1675,1697]" captionTargetBox="[229,1376,196,1635]" captionTargetId="figure-26@15.[229,1376,195,1635]" captionTargetPageId="15" captionText="Figure 4. Photomicrographs of †G. sternorhabda sp. nov. showing setation and certain key features. A–D, G–I, paratype, CASENT0741234. E, F, J, holotype, CASENT0741233.A, body, ventrolateral oblique view. B, head, dorsolateral oblique view. C–F, body and left legs: D, dorsolateral oblique; (F) is slightly offset from (E), revealing the prora and subpetiolar process. G, protarsus, posterodorsal oblique view. H, mesotarsus, dorsal view. I, metatarsus, dorsal view. J, metatarsus, posterior view. Abbreviations: ad, antennomere distorted, i.e. visibly distorted antennomere associated with decay bubble; cn, cinctus; ms, metanotal spiracle; pr, prora; psf, anterior flange of propodeal spiracle; ptb, probasitarsal brush; spp, subpetiolar process; ss, standing setae; tnc, transverse mesonotal carina." figureDoi="http://doi.org/10.5281/zenodo.6994466" httpUri="https://zenodo.org/record/6994466/files/figure.png" pageId="11" pageNumber="1366">4A</figureCitation>
|
||
,
|
||
<figureCitation id="13102A67FFEAAA23D1FDFDC2FB91924B" box="[970,1030,596,617]" captionStart="Figure 7" captionStartId="19.[164,241,1513,1535]" captionTargetBox="[163,1443,195,1470]" captionTargetId="figure-119@19.[163,1443,195,1473]" captionTargetPageId="19" captionText="Figure 7. Maxillary and labial palps of †G.gracilis and †G.sternorhabda sp. nov.. Small black stars indicate each palpomere on the right-hand side of the body.A, B, †G.gracilis adult, CASENT0741232, in anterior dorsolateral oblique view (A) and anterior view (B).C, D, †G.gracilis pupa, CASENT0741231, in anterior view (C) and anterior dorsolateral view (D).E, F, †G.sternorhabda sp. nov. holotype, CASENT0741233, and paratype, CASENT0741234, in anteroventral oblique view. Abbreviations: pl, labial palp; plp, process of the proximal labial palpomere; pm, maxillary palp; pm?, possible maxillary palpomere." figureDoi="http://doi.org/10.5281/zenodo.6994474" httpUri="https://zenodo.org/record/6994474/files/figure.png" pageId="11" pageNumber="1366">7E, F</figureCitation>
|
||
) [Notes 5, 6]; they are conspicuously short, with their total lengths shorter than the lengths of either the mandibles or scapes; with the exception of the apical palpomere, they are thick and bulging at about their midlengths; they are adorned with erect and appressed pilosity. The
|
||
<emphasis id="B95FEAF0FFEAAA23D644FD7BFA909320" bold="true" box="[1139,1287,749,771]" pageId="11" pageNumber="1366">labial palps</emphasis>
|
||
are 4-merous (
|
||
<figureCitation id="13102A67FFEAAA23D174FC9DFC049303" box="[835,915,779,801]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="11" pageNumber="1366">Figs 2I</figureCitation>
|
||
,
|
||
<figureCitation id="13102A67FFEAAA23D197FC9AFC579303" box="[928,960,780,801]" captionStart="Figure 4" captionStartId="15.[163,242,1675,1697]" captionTargetBox="[229,1376,196,1635]" captionTargetId="figure-26@15.[229,1376,195,1635]" captionTargetPageId="15" captionText="Figure 4. Photomicrographs of †G. sternorhabda sp. nov. showing setation and certain key features. A–D, G–I, paratype, CASENT0741234. E, F, J, holotype, CASENT0741233.A, body, ventrolateral oblique view. B, head, dorsolateral oblique view. C–F, body and left legs: D, dorsolateral oblique; (F) is slightly offset from (E), revealing the prora and subpetiolar process. G, protarsus, posterodorsal oblique view. H, mesotarsus, dorsal view. I, metatarsus, dorsal view. J, metatarsus, posterior view. Abbreviations: ad, antennomere distorted, i.e. visibly distorted antennomere associated with decay bubble; cn, cinctus; ms, metanotal spiracle; pr, prora; psf, anterior flange of propodeal spiracle; ptb, probasitarsal brush; spp, subpetiolar process; ss, standing setae; tnc, transverse mesonotal carina." figureDoi="http://doi.org/10.5281/zenodo.6994466" httpUri="https://zenodo.org/record/6994466/files/figure.png" pageId="11" pageNumber="1366">4A</figureCitation>
|
||
,
|
||
<figureCitation id="13102A67FFEAAA23D1FAFC9AFB9D9303" box="[973,1034,780,801]" captionStart="Figure 7" captionStartId="19.[164,241,1513,1535]" captionTargetBox="[163,1443,195,1470]" captionTargetId="figure-119@19.[163,1443,195,1473]" captionTargetPageId="19" captionText="Figure 7. Maxillary and labial palps of †G.gracilis and †G.sternorhabda sp. nov.. Small black stars indicate each palpomere on the right-hand side of the body.A, B, †G.gracilis adult, CASENT0741232, in anterior dorsolateral oblique view (A) and anterior view (B).C, D, †G.gracilis pupa, CASENT0741231, in anterior view (C) and anterior dorsolateral view (D).E, F, †G.sternorhabda sp. nov. holotype, CASENT0741233, and paratype, CASENT0741234, in anteroventral oblique view. Abbreviations: pl, labial palp; plp, process of the proximal labial palpomere; pm, maxillary palp; pm?, possible maxillary palpomere." figureDoi="http://doi.org/10.5281/zenodo.6994474" httpUri="https://zenodo.org/record/6994474/files/figure.png" pageId="11" pageNumber="1366">7E, F</figureCitation>
|
||
) [Note 6]; they are short, being just over half the length of the maxillary palps; with the exception of the apical palpomere, they are more-orless conical and thickening toward their apices; they are adorned with erect and appressed pilosity; the proximal palpomere lacks a distinct process.
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFEAAA24D10CFC7FFDB29476" blockId="11.[827,1444,1001,1881]" lastBlockId="12.[145,762,197,1108]" lastPageId="12" lastPageNumber="1367" pageId="11" pageNumber="1366">
|
||
<emphasis id="B95FEAF0FFEAAA23D10CFC7FFC2D93DC" box="[827,954,1001,1022]" italics="true" pageId="11" pageNumber="1366">Mesosoma:</emphasis>
|
||
The
|
||
<emphasis id="B95FEAF0FFEAAA23D633FC7FFB1093DC" bold="true" box="[1028,1159,1001,1022]" pageId="11" pageNumber="1366">pronotum</emphasis>
|
||
bears an anteromedian neck process, and lateral and posteromedian flanges (
|
||
<figureCitation id="13102A67FFEAAA23D173FBB0FC2A941E" box="[836,957,1062,1084]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="11" pageNumber="1366">Figs 2C, F</figureCitation>
|
||
,
|
||
<figureCitation id="13102A67FFEAAA23D1FBFBB0FC7B941E" box="[972,1004,1062,1084]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="11" pageNumber="1366">5C</figureCitation>
|
||
,
|
||
<figureCitation id="13102A67FFEAAA23D1CBFBB0FB8B941E" box="[1020,1052,1062,1084]" captionStart="Figure 6" captionStartId="18.[145,223,1652,1674]" captionTargetBox="[146,1425,197,1609]" captionTargetId="figure-26@18.[145,1425,195,1612]" captionTargetPageId="18" captionText="Figure 6. Detailsof thepronota andmetapleural glandsof †Gerontoformica.A,B,fluorescent lightmicroscopy;C–G,standard photomicroscopy. A, B, †G. sternorhabda sp. nov. paratype (UFV-LABECOL-009656). C, †G. pilosa (ANTWEB1038931). D, †G. sternorhabda sp. nov. non-type (JWJ-BU21). E, F, †G. gracilis holotype (JZC-Bu324A). G, †G. near or conspecific with gracilis (ANTWEB1032649). Abbreviations: msnttc, transverse carina of mesonotum; mtplglb, metapleural gland bulla; mtplgldf, metapleural gland dorsal flange; mtplglvf, metapleural gland ventral flange; mtvlc, ventrolateral carina of metathorax; mtntsp, metanotal spiracle; mtnt, metanotum; pntl, pronotal lobe; pntfl, lateral pronotal flange; pntfm, medial pronotal flange; ppdsf, anterior flange of propodeal spiracle; ppl, propleuron; pt, petiole." figureDoi="http://doi.org/10.5281/zenodo.6994472" httpUri="https://zenodo.org/record/6994472/files/figure.png" pageId="11" pageNumber="1366">6A</figureCitation>
|
||
); these flanges are flared in the
|
||
<typeStatus id="54908840FFEAAA23D10CFBD3FC3E9478" box="[827,937,1093,1114]" pageId="11" pageNumber="1366" type="paratype">paratype</typeStatus>
|
||
specimen C-34 (
|
||
<figureCitation id="13102A67FFEAAA23D6B6FBD3FB4C9478" box="[1153,1243,1093,1115]" captionStart="Figure 4" captionStartId="15.[163,242,1675,1697]" captionTargetBox="[229,1376,196,1635]" captionTargetId="figure-26@15.[229,1376,195,1635]" captionTargetPageId="15" captionText="Figure 4. Photomicrographs of †G. sternorhabda sp. nov. showing setation and certain key features. A–D, G–I, paratype, CASENT0741234. E, F, J, holotype, CASENT0741233.A, body, ventrolateral oblique view. B, head, dorsolateral oblique view. C–F, body and left legs: D, dorsolateral oblique; (F) is slightly offset from (E), revealing the prora and subpetiolar process. G, protarsus, posterodorsal oblique view. H, mesotarsus, dorsal view. I, metatarsus, dorsal view. J, metatarsus, posterior view. Abbreviations: ad, antennomere distorted, i.e. visibly distorted antennomere associated with decay bubble; cn, cinctus; ms, metanotal spiracle; pr, prora; psf, anterior flange of propodeal spiracle; ptb, probasitarsal brush; spp, subpetiolar process; ss, standing setae; tnc, transverse mesonotal carina." figureDoi="http://doi.org/10.5281/zenodo.6994466" httpUri="https://zenodo.org/record/6994466/files/figure.png" pageId="11" pageNumber="1366">Fig. 4C</figureCitation>
|
||
) but not in the
|
||
<typeStatus id="54908840FFEAAA23D10CFBF5FC0C945B" box="[827,923,1123,1145]" pageId="11" pageNumber="1366" type="holotype">holotype</typeStatus>
|
||
or other
|
||
<typeStatus id="54908840FFEAAA23D634FBF2FBFD945B" box="[1027,1130,1124,1145]" pageId="11" pageNumber="1366" type="paratype">paratype</typeStatus>
|
||
; the muscular node or ‘disc’ of the pronotum is almost spheroidal in shape, with the lateral margins strongly convex in dorsal view and the dorsal margin strongly convex in lateral view, and with an anteroposterior length approximating its dorsoventral height; pronotal setation is sparse, being represented by a few subdecumbent setae (
|
||
<figureCitation id="13102A67FFEAAA23D74FFA8DFCCD9572" captionStart="Figure 6" captionStartId="18.[145,223,1652,1674]" captionTargetBox="[146,1425,197,1609]" captionTargetId="figure-26@18.[145,1425,195,1612]" captionTargetPageId="18" captionText="Figure 6. Detailsof thepronota andmetapleural glandsof †Gerontoformica.A,B,fluorescent lightmicroscopy;C–G,standard photomicroscopy. A, B, †G. sternorhabda sp. nov. paratype (UFV-LABECOL-009656). C, †G. pilosa (ANTWEB1038931). D, †G. sternorhabda sp. nov. non-type (JWJ-BU21). E, F, †G. gracilis holotype (JZC-Bu324A). G, †G. near or conspecific with gracilis (ANTWEB1032649). Abbreviations: msnttc, transverse carina of mesonotum; mtplglb, metapleural gland bulla; mtplgldf, metapleural gland dorsal flange; mtplglvf, metapleural gland ventral flange; mtvlc, ventrolateral carina of metathorax; mtntsp, metanotal spiracle; mtnt, metanotum; pntl, pronotal lobe; pntfl, lateral pronotal flange; pntfm, medial pronotal flange; ppdsf, anterior flange of propodeal spiracle; ppl, propleuron; pt, petiole." figureDoi="http://doi.org/10.5281/zenodo.6994472" httpUri="https://zenodo.org/record/6994472/files/figure.png" pageId="11" pageNumber="1366">Fig. 6A</figureCitation>
|
||
). The
|
||
<emphasis id="B95FEAF0FFEAAA23D194FAACFBCA956D" bold="true" box="[931,1117,1338,1359]" pageId="11" pageNumber="1366">pronotal lobes</emphasis>
|
||
are well developed (
|
||
<figureCitation id="13102A67FFEAAA23D77EFAACFA0D9572" box="[1353,1434,1338,1360]" captionStart="Figure 6" captionStartId="18.[145,223,1652,1674]" captionTargetBox="[146,1425,197,1609]" captionTargetId="figure-26@18.[145,1425,195,1612]" captionTargetPageId="18" captionText="Figure 6. Detailsof thepronota andmetapleural glandsof †Gerontoformica.A,B,fluorescent lightmicroscopy;C–G,standard photomicroscopy. A, B, †G. sternorhabda sp. nov. paratype (UFV-LABECOL-009656). C, †G. pilosa (ANTWEB1038931). D, †G. sternorhabda sp. nov. non-type (JWJ-BU21). E, F, †G. gracilis holotype (JZC-Bu324A). G, †G. near or conspecific with gracilis (ANTWEB1032649). Abbreviations: msnttc, transverse carina of mesonotum; mtplglb, metapleural gland bulla; mtplgldf, metapleural gland dorsal flange; mtplglvf, metapleural gland ventral flange; mtvlc, ventrolateral carina of metathorax; mtntsp, metanotal spiracle; mtnt, metanotum; pntl, pronotal lobe; pntfl, lateral pronotal flange; pntfm, medial pronotal flange; ppdsf, anterior flange of propodeal spiracle; ppl, propleuron; pt, petiole." figureDoi="http://doi.org/10.5281/zenodo.6994472" httpUri="https://zenodo.org/record/6994472/files/figure.png" pageId="11" pageNumber="1366">Fig. 6A</figureCitation>
|
||
) [Note 7]. The
|
||
<emphasis id="B95FEAF0FFEAAA23D1EFFACFFBFA954C" bold="true" box="[984,1133,1369,1390]" pageId="11" pageNumber="1366">mesonotum</emphasis>
|
||
is divided into an anterior mesoscutal area and a posterior mesoscutellar area by the
|
||
<emphasis id="B95FEAF0FFEAAA23D150FA00FB42958E" bold="true" box="[871,1237,1430,1452]" pageId="11" pageNumber="1366">transverse mesonotal carina</emphasis>
|
||
(
|
||
<figureCitation id="13102A67FFEAAA23D6D2FA00FAC19589" box="[1253,1366,1430,1452]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="11" pageNumber="1366">Figs 2C, F</figureCitation>
|
||
,
|
||
<figureCitation id="13102A67FFEAAA23D754FA00FA0B9589" box="[1379,1436,1430,1451]" captionStart="Figure 4" captionStartId="15.[163,242,1675,1697]" captionTargetBox="[229,1376,196,1635]" captionTargetId="figure-26@15.[229,1376,195,1635]" captionTargetPageId="15" captionText="Figure 4. Photomicrographs of †G. sternorhabda sp. nov. showing setation and certain key features. A–D, G–I, paratype, CASENT0741234. E, F, J, holotype, CASENT0741233.A, body, ventrolateral oblique view. B, head, dorsolateral oblique view. C–F, body and left legs: D, dorsolateral oblique; (F) is slightly offset from (E), revealing the prora and subpetiolar process. G, protarsus, posterodorsal oblique view. H, mesotarsus, dorsal view. I, metatarsus, dorsal view. J, metatarsus, posterior view. Abbreviations: ad, antennomere distorted, i.e. visibly distorted antennomere associated with decay bubble; cn, cinctus; ms, metanotal spiracle; pr, prora; psf, anterior flange of propodeal spiracle; ptb, probasitarsal brush; spp, subpetiolar process; ss, standing setae; tnc, transverse mesonotal carina." figureDoi="http://doi.org/10.5281/zenodo.6994466" httpUri="https://zenodo.org/record/6994466/files/figure.png" pageId="11" pageNumber="1366">4C, F</figureCitation>
|
||
,
|
||
<figureCitation id="13102A67FFEAAA23D10CFA22FCCE95E8" box="[827,857,1460,1482]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="11" pageNumber="1366">5C</figureCitation>
|
||
,
|
||
<figureCitation id="13102A67FFEAAA23D151FA23FC1295E9" box="[870,901,1461,1483]" captionStart="Figure 6" captionStartId="18.[145,223,1652,1674]" captionTargetBox="[146,1425,197,1609]" captionTargetId="figure-26@18.[145,1425,195,1612]" captionTargetPageId="18" captionText="Figure 6. Detailsof thepronota andmetapleural glandsof †Gerontoformica.A,B,fluorescent lightmicroscopy;C–G,standard photomicroscopy. A, B, †G. sternorhabda sp. nov. paratype (UFV-LABECOL-009656). C, †G. pilosa (ANTWEB1038931). D, †G. sternorhabda sp. nov. non-type (JWJ-BU21). E, F, †G. gracilis holotype (JZC-Bu324A). G, †G. near or conspecific with gracilis (ANTWEB1032649). Abbreviations: msnttc, transverse carina of mesonotum; mtplglb, metapleural gland bulla; mtplgldf, metapleural gland dorsal flange; mtplglvf, metapleural gland ventral flange; mtvlc, ventrolateral carina of metathorax; mtntsp, metanotal spiracle; mtnt, metanotum; pntl, pronotal lobe; pntfl, lateral pronotal flange; pntfm, medial pronotal flange; ppdsf, anterior flange of propodeal spiracle; ppl, propleuron; pt, petiole." figureDoi="http://doi.org/10.5281/zenodo.6994472" httpUri="https://zenodo.org/record/6994472/files/figure.png" pageId="11" pageNumber="1366">6B</figureCitation>
|
||
). The
|
||
<emphasis id="B95FEAF0FFEAAA23D1FCFA23FB0F95E8" bold="true" box="[971,1176,1461,1482]" pageId="11" pageNumber="1366">mesoscutal area</emphasis>
|
||
is approximately in the form of a low saddle (i.e. is a low hyperbolic paraboloid), with a concave dorsal margin in lateral view, and with the anterior rim more upcurved than the posterior rim (
|
||
<figureCitation id="13102A67FFEAAA23D144F9B9FC539666" box="[883,964,1583,1605]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="11" pageNumber="1366">Fig. 2C</figureCitation>
|
||
). The
|
||
<emphasis id="B95FEAF0FFEAAA23D63BF9B9FA979666" bold="true" box="[1036,1280,1583,1604]" pageId="11" pageNumber="1366">mesoscutellar area</emphasis>
|
||
is convex, but sunken relative to the mesoscutal area (
|
||
<figureCitation id="13102A67FFEAAA23D738F9D8FAF69641" box="[1295,1377,1614,1636]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="11" pageNumber="1366">Fig. 2C</figureCitation>
|
||
). The
|
||
<emphasis id="B95FEAF0FFEAAA23D10CF9FBFC4596A0" bold="true" box="[827,978,1645,1666]" pageId="11" pageNumber="1366">mesopectus</emphasis>
|
||
is divided into dorsal and ventral areas by a longitudinal sulcus (
|
||
<figureCitation id="13102A67FFEAAA23D644F91DFB5D9682" box="[1139,1226,1675,1697]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="11" pageNumber="1366">Fig. 2C</figureCitation>
|
||
); its dorsoventral height is about equal to its anteroposterior length. The
|
||
<emphasis id="B95FEAF0FFEAAA23D10CF95FFBCC96FC" bold="true" box="[827,1115,1737,1758]" pageId="11" pageNumber="1366">dorsal mesopectal area</emphasis>
|
||
is approximately rectangular in shape, being somewhat more than twice as long anteroposteriorly as tall dorsoventrally (
|
||
<figureCitation id="13102A67FFEAAA23D72DF890FAF99739" box="[1306,1390,1798,1820]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="11" pageNumber="1366">Fig. 2C</figureCitation>
|
||
); its dorsoventral height is one-third the dorsoventral height of the lower mesopectal area as measured from the transverse sulcus directly ventrad to an imaginary line drawn parallel to the ventrolateral margins of the meso- and metapecta. The
|
||
<emphasis id="B95FEAF0FFEDAA24D3F4FE95FD6E913A" bold="true" box="[451,761,259,280]" pageId="12" pageNumber="1367">ventral mesopectal area</emphasis>
|
||
is approximately triangular in shape, being broad along is dorsal margin and narrowing posteroventrally to its ventrolateral margin (
|
||
<figureCitation id="13102A67FFEDAA24D3B3FEC8FE479156" box="[388,464,350,372]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="12" pageNumber="1367">Fig. 2C</figureCitation>
|
||
); its anteroposterior length along its dorsal margin is approximately equal to its dorsoventral height. The
|
||
<emphasis id="B95FEAF0FFEDAA24D39CFE0AFF7491F2" bold="true" pageId="12" pageNumber="1367">transverse mesometanotal sulcus</emphasis>
|
||
is anteroposteriorly short/ thin (
|
||
<figureCitation id="13102A67FFEDAA24D054FE2CFD2191F2" box="[611,694,442,464]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="12" pageNumber="1367">Fig. 2C</figureCitation>
|
||
). The
|
||
<emphasis id="B95FEAF0FFEDAA24D2A6FE4FFEB391CC" bold="true" box="[145,292,473,494]" pageId="12" pageNumber="1367">metanotum</emphasis>
|
||
is developed as a distinct bulge between the metanotal spiracles (
|
||
<figureCitation id="13102A67FFEDAA24D3F5FE6EFD8D922F" box="[450,538,504,526]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="12" pageNumber="1367">Fig. 2C</figureCitation>
|
||
). The
|
||
<emphasis id="B95FEAF0FFEDAA24D05DFE6EFE49920E" bold="true" pageId="12" pageNumber="1367">transverse metanotopropodeal sulcus</emphasis>
|
||
is anteroposteriorly long/ broad and continues ventrally toward the base of the mesocoxa, completely separating the lateral metapectal area from the lateral mesopectal area (
|
||
<figureCitation id="13102A67FFEDAA24D056FDE4FD2292AA" box="[609,693,626,648]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="12" pageNumber="1367">Fig. 2C</figureCitation>
|
||
). The
|
||
<emphasis id="B95FEAF0FFEDAA24D2A6FD07FE419285" bold="true" box="[145,470,657,679]" pageId="12" pageNumber="1367">metapleural gland orifice</emphasis>
|
||
is large, hairy, and in the form of a broad subelliptical pit (
|
||
<figureCitation id="13102A67FFEDAA24D3CCFD26FDDD92E4" box="[507,586,688,710]" captionStart="Figure 6" captionStartId="18.[145,223,1652,1674]" captionTargetBox="[146,1425,197,1609]" captionTargetId="figure-26@18.[145,1425,195,1612]" captionTargetPageId="18" captionText="Figure 6. Detailsof thepronota andmetapleural glandsof †Gerontoformica.A,B,fluorescent lightmicroscopy;C–G,standard photomicroscopy. A, B, †G. sternorhabda sp. nov. paratype (UFV-LABECOL-009656). C, †G. pilosa (ANTWEB1038931). D, †G. sternorhabda sp. nov. non-type (JWJ-BU21). E, F, †G. gracilis holotype (JZC-Bu324A). G, †G. near or conspecific with gracilis (ANTWEB1032649). Abbreviations: msnttc, transverse carina of mesonotum; mtplglb, metapleural gland bulla; mtplgldf, metapleural gland dorsal flange; mtplglvf, metapleural gland ventral flange; mtvlc, ventrolateral carina of metathorax; mtntsp, metanotal spiracle; mtnt, metanotum; pntl, pronotal lobe; pntfl, lateral pronotal flange; pntfm, medial pronotal flange; ppdsf, anterior flange of propodeal spiracle; ppl, propleuron; pt, petiole." figureDoi="http://doi.org/10.5281/zenodo.6994472" httpUri="https://zenodo.org/record/6994472/files/figure.png" pageId="12" pageNumber="1367">Fig. 6B</figureCitation>
|
||
); it is margined dorsally by a bulge, the
|
||
<emphasis id="B95FEAF0FFEDAA24D386FD59FD6592C6" bold="true" box="[433,754,719,740]" pageId="12" pageNumber="1367">metapleural gland bulla</emphasis>
|
||
, and ventrally by the
|
||
<emphasis id="B95FEAF0FFEDAA24D3A3FD7BFF749303" bold="true" pageId="12" pageNumber="1367">ventral metapleural gland flange</emphasis>
|
||
, which itself is a continuation of the sinuate
|
||
<emphasis id="B95FEAF0FFEDAA24D2A6FCBDFD339362" bold="true" box="[145,676,810,832]" pageId="12" pageNumber="1367">ventrolateral carina of the metapectus</emphasis>
|
||
and is spiniform and dorsally inclined. The
|
||
<emphasis id="B95FEAF0FFEDAA24D071FCDFFD4B937C" bold="true" box="[582,732,841,862]" pageId="12" pageNumber="1367">propodeum</emphasis>
|
||
is boxy (
|
||
<figureCitation id="13102A67FFEDAA24D2EEFCFEFEA3935F" box="[217,308,872,894]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="12" pageNumber="1367">Figs 2C</figureCitation>
|
||
,
|
||
<figureCitation id="13102A67FFEDAA24D374FCFEFEF4935C" box="[323,355,872,894]" captionStart="Figure 6" captionStartId="18.[145,223,1652,1674]" captionTargetBox="[146,1425,197,1609]" captionTargetId="figure-26@18.[145,1425,195,1612]" captionTargetPageId="18" captionText="Figure 6. Detailsof thepronota andmetapleural glandsof †Gerontoformica.A,B,fluorescent lightmicroscopy;C–G,standard photomicroscopy. A, B, †G. sternorhabda sp. nov. paratype (UFV-LABECOL-009656). C, †G. pilosa (ANTWEB1038931). D, †G. sternorhabda sp. nov. non-type (JWJ-BU21). E, F, †G. gracilis holotype (JZC-Bu324A). G, †G. near or conspecific with gracilis (ANTWEB1032649). Abbreviations: msnttc, transverse carina of mesonotum; mtplglb, metapleural gland bulla; mtplgldf, metapleural gland dorsal flange; mtplglvf, metapleural gland ventral flange; mtvlc, ventrolateral carina of metathorax; mtntsp, metanotal spiracle; mtnt, metanotum; pntl, pronotal lobe; pntfl, lateral pronotal flange; pntfm, medial pronotal flange; ppdsf, anterior flange of propodeal spiracle; ppl, propleuron; pt, petiole." figureDoi="http://doi.org/10.5281/zenodo.6994472" httpUri="https://zenodo.org/record/6994472/files/figure.png" pageId="12" pageNumber="1367">6B</figureCitation>
|
||
) [Note 8]; it bears standing setae and is the hairiest mesosomal region. The
|
||
<emphasis id="B95FEAF0FFEDAA24D041FC10FE919399" bold="true" pageId="12" pageNumber="1367">propodeal spiracles</emphasis>
|
||
are situated distant from the metanotum, but near the dorsal propodeal margin in lateral view (
|
||
<figureCitation id="13102A67FFEDAA24D2AEFC74FF6593D5" box="[153,242,994,1016]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="12" pageNumber="1367">Figs 2C</figureCitation>
|
||
,
|
||
<figureCitation id="13102A67FFEDAA24D337FC74FE8893DA" box="[256,287,994,1016]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="12" pageNumber="1367">5C</figureCitation>
|
||
); they are posteriorly to posterolaterally directed and protected anteriorly by the
|
||
<emphasis id="B95FEAF0FFEDAA24D0BFFB97FE539417" bold="true" pageId="12" pageNumber="1367">anterior flange of the propodeal</emphasis>
|
||
spiracles. The
|
||
<emphasis id="B95FEAF0FFEDAA24D043FBB6FF459471" bold="true" pageId="12" pageNumber="1367">propodeal lobes</emphasis>
|
||
are apparently not developed.
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFEDAA24D2A6FBE8FA8E9156" blockId="12.[145,761,1149,1876]" lastBlockId="12.[809,1426,197,372]" pageId="12" pageNumber="1367">
|
||
<emphasis id="B95FEAF0FFEDAA24D2A6FBE8FF5894B1" box="[145,207,1150,1171]" italics="true" pageId="12" pageNumber="1367">Legs:</emphasis>
|
||
The legs are developed as expected for the
|
||
<taxonomicName id="4C2B4D61FFEDAA24D2A6FB0AFE849490" authorityName="LATREILLE" authorityYear="1809" box="[145,275,1180,1202]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="1367" phylum="Arthropoda" rank="family">Formicidae</taxonomicName>
|
||
, with some notable characters and states. They are densely hairy, appearing shaggy, with the setae suberect (
|
||
<figureCitation id="13102A67FFEDAA24D308FB4FFE7294CC" box="[319,485,1241,1263]" captionStart="Figure 4" captionStartId="15.[163,242,1675,1697]" captionTargetBox="[229,1376,196,1635]" captionTargetId="figure-26@15.[229,1376,195,1635]" captionTargetPageId="15" captionText="Figure 4. Photomicrographs of †G. sternorhabda sp. nov. showing setation and certain key features. A–D, G–I, paratype, CASENT0741234. E, F, J, holotype, CASENT0741233.A, body, ventrolateral oblique view. B, head, dorsolateral oblique view. C–F, body and left legs: D, dorsolateral oblique; (F) is slightly offset from (E), revealing the prora and subpetiolar process. G, protarsus, posterodorsal oblique view. H, mesotarsus, dorsal view. I, metatarsus, dorsal view. J, metatarsus, posterior view. Abbreviations: ad, antennomere distorted, i.e. visibly distorted antennomere associated with decay bubble; cn, cinctus; ms, metanotal spiracle; pr, prora; psf, anterior flange of propodeal spiracle; ptb, probasitarsal brush; spp, subpetiolar process; ss, standing setae; tnc, transverse mesonotal carina." figureDoi="http://doi.org/10.5281/zenodo.6994466" httpUri="https://zenodo.org/record/6994466/files/figure.png" pageId="12" pageNumber="1367">Fig. 4A, D, G–J</figureCitation>
|
||
). Apparently, the
|
||
<emphasis id="B95FEAF0FFEDAA24D09CFB4FFE1F952F" bold="true" pageId="12" pageNumber="1367">apical protibial foramina</emphasis>
|
||
are open, i.e. without a bridge of sclerite dividing the calcar from the probasitarsus (
|
||
<figureCitation id="13102A67FFEDAA24D2ACFAA3FF6A9568" box="[155,253,1333,1355]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="12" pageNumber="1367">Fig. 5G</figureCitation>
|
||
) [Note 9]. The
|
||
<emphasis id="B95FEAF0FFEDAA24D3EEFAA3FD2D9568" bold="true" box="[473,698,1333,1355]" pageId="12" pageNumber="1367">mesoprefemora</emphasis>
|
||
and
|
||
<emphasis id="B95FEAF0FFEDAA24D2A6FAC5FECE954A" bold="true" box="[145,345,1363,1384]" pageId="12" pageNumber="1367">metaprefemora</emphasis>
|
||
are well developed and are broader ventrally than dorsally (
|
||
<figureCitation id="13102A67FFEDAA24D39BFAE4FD9795AA" box="[428,512,1394,1416]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="12" pageNumber="1367">Fig. 5D</figureCitation>
|
||
). The
|
||
<emphasis id="B95FEAF0FFEDAA24D07EFAE4FD2495AA" bold="true" box="[585,691,1394,1416]" pageId="12" pageNumber="1367">protibia</emphasis>
|
||
bears and anterior brush of dense suberect setae in their apical third, near the calcar (
|
||
<figureCitation id="13102A67FFEDAA24D3CBFA26FDC495E7" box="[508,595,1456,1478]" captionStart="Figure 4" captionStartId="15.[163,242,1675,1697]" captionTargetBox="[229,1376,196,1635]" captionTargetId="figure-26@15.[229,1376,195,1635]" captionTargetPageId="15" captionText="Figure 4. Photomicrographs of †G. sternorhabda sp. nov. showing setation and certain key features. A–D, G–I, paratype, CASENT0741234. E, F, J, holotype, CASENT0741233.A, body, ventrolateral oblique view. B, head, dorsolateral oblique view. C–F, body and left legs: D, dorsolateral oblique; (F) is slightly offset from (E), revealing the prora and subpetiolar process. G, protarsus, posterodorsal oblique view. H, mesotarsus, dorsal view. I, metatarsus, dorsal view. J, metatarsus, posterior view. Abbreviations: ad, antennomere distorted, i.e. visibly distorted antennomere associated with decay bubble; cn, cinctus; ms, metanotal spiracle; pr, prora; psf, anterior flange of propodeal spiracle; ptb, probasitarsal brush; spp, subpetiolar process; ss, standing setae; tnc, transverse mesonotal carina." figureDoi="http://doi.org/10.5281/zenodo.6994466" httpUri="https://zenodo.org/record/6994466/files/figure.png" pageId="12" pageNumber="1367">Fig. 4A</figureCitation>
|
||
). Each of the
|
||
<emphasis id="B95FEAF0FFEDAA24D2A6FA58FE9995C6" bold="true" box="[145,270,1486,1508]" pageId="12" pageNumber="1367">mesotibia</emphasis>
|
||
and
|
||
<emphasis id="B95FEAF0FFEDAA24D37FFA58FE5395C6" bold="true" box="[328,452,1486,1508]" pageId="12" pageNumber="1367">metatibia</emphasis>
|
||
bear a pair of apicoventral spurs (
|
||
<figureCitation id="13102A67FFEDAA24D2D7FA7BFEC49620" box="[224,339,1517,1539]" captionStart="Figure 4" captionStartId="15.[163,242,1675,1697]" captionTargetBox="[229,1376,196,1635]" captionTargetId="figure-26@15.[229,1376,195,1635]" captionTargetPageId="15" captionText="Figure 4. Photomicrographs of †G. sternorhabda sp. nov. showing setation and certain key features. A–D, G–I, paratype, CASENT0741234. E, F, J, holotype, CASENT0741233.A, body, ventrolateral oblique view. B, head, dorsolateral oblique view. C–F, body and left legs: D, dorsolateral oblique; (F) is slightly offset from (E), revealing the prora and subpetiolar process. G, protarsus, posterodorsal oblique view. H, mesotarsus, dorsal view. I, metatarsus, dorsal view. J, metatarsus, posterior view. Abbreviations: ad, antennomere distorted, i.e. visibly distorted antennomere associated with decay bubble; cn, cinctus; ms, metanotal spiracle; pr, prora; psf, anterior flange of propodeal spiracle; ptb, probasitarsal brush; spp, subpetiolar process; ss, standing setae; tnc, transverse mesonotal carina." figureDoi="http://doi.org/10.5281/zenodo.6994466" httpUri="https://zenodo.org/record/6994466/files/figure.png" pageId="12" pageNumber="1367">Figs 4I, H</figureCitation>
|
||
,
|
||
<figureCitation id="13102A67FFEDAA24D357FA7BFE149621" box="[352,387,1517,1539]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="12" pageNumber="1367">5H</figureCitation>
|
||
); the
|
||
<emphasis id="B95FEAF0FFEDAA24D3F2FA7BFD5B9620" bold="true" box="[453,716,1517,1539]" pageId="12" pageNumber="1367">anterior tibial spurs</emphasis>
|
||
are pectinate; the
|
||
<emphasis id="B95FEAF0FFEDAA24D303F99DFDD39603" bold="true" box="[308,580,1547,1569]" pageId="12" pageNumber="1367">posterior tibial spurs</emphasis>
|
||
are barbirulate to simple. The
|
||
<emphasis id="B95FEAF0FFEDAA24D37EF9BCFE0A961D" bold="true" box="[329,413,1578,1599]" pageId="12" pageNumber="1367">calcar</emphasis>
|
||
is apparently bifid apically, with one point being the apex of the elongate
|
||
<emphasis id="B95FEAF0FFEDAA24D09EF9DFFD6E967C" bold="true" box="[681,761,1609,1630]" pageId="12" pageNumber="1367">velum</emphasis>
|
||
and the other point a small array of hairs (
|
||
<figureCitation id="13102A67FFEDAA24D0A3F9FEFD7D965F" box="[660,746,1639,1662]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="12" pageNumber="1367">Fig. 5G</figureCitation>
|
||
); two stout setae are developed (inserted) posterior to the calcar. The
|
||
<emphasis id="B95FEAF0FFEDAA24D353F933FD859698" bold="true" box="[356,530,1701,1722]" pageId="12" pageNumber="1367">plantar lobes</emphasis>
|
||
of the tarsi are not developed (absent), but the tarsomeres have a brush of dense ventral setae, and are apically margined by thick, coarse chaetae (
|
||
<figureCitation id="13102A67FFEDAA24D3BAF897FD979734" box="[397,512,1793,1815]" captionStart="Figure 4" captionStartId="15.[163,242,1675,1697]" captionTargetBox="[229,1376,196,1635]" captionTargetId="figure-26@15.[229,1376,195,1635]" captionTargetPageId="15" captionText="Figure 4. Photomicrographs of †G. sternorhabda sp. nov. showing setation and certain key features. A–D, G–I, paratype, CASENT0741234. E, F, J, holotype, CASENT0741233.A, body, ventrolateral oblique view. B, head, dorsolateral oblique view. C–F, body and left legs: D, dorsolateral oblique; (F) is slightly offset from (E), revealing the prora and subpetiolar process. G, protarsus, posterodorsal oblique view. H, mesotarsus, dorsal view. I, metatarsus, dorsal view. J, metatarsus, posterior view. Abbreviations: ad, antennomere distorted, i.e. visibly distorted antennomere associated with decay bubble; cn, cinctus; ms, metanotal spiracle; pr, prora; psf, anterior flange of propodeal spiracle; ptb, probasitarsal brush; spp, subpetiolar process; ss, standing setae; tnc, transverse mesonotal carina." figureDoi="http://doi.org/10.5281/zenodo.6994466" httpUri="https://zenodo.org/record/6994466/files/figure.png" pageId="12" pageNumber="1367">Figs 4G–J</figureCitation>
|
||
,
|
||
<figureCitation id="13102A67FFEDAA24D03BF897FDDF9735" box="[524,584,1793,1815]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="12" pageNumber="1367">5G–J</figureCitation>
|
||
) [Note 10]. The
|
||
<emphasis id="B95FEAF0FFEDAA24D2A6F889FE139717" bold="true" box="[145,388,1823,1845]" pageId="12" pageNumber="1367">fourth tarsomeres</emphasis>
|
||
of each leg are deeply notched distally, thus appearing V-shaped (probasitarsi) or arrowhead-shaped (meso-, metabasitarsi) (
|
||
<figureCitation id="13102A67FFEDAA24D724FF53FA1590F8" box="[1299,1410,197,219]" captionStart="Figure 4" captionStartId="15.[163,242,1675,1697]" captionTargetBox="[229,1376,196,1635]" captionTargetId="figure-26@15.[229,1376,195,1635]" captionTargetPageId="15" captionText="Figure 4. Photomicrographs of †G. sternorhabda sp. nov. showing setation and certain key features. A–D, G–I, paratype, CASENT0741234. E, F, J, holotype, CASENT0741233.A, body, ventrolateral oblique view. B, head, dorsolateral oblique view. C–F, body and left legs: D, dorsolateral oblique; (F) is slightly offset from (E), revealing the prora and subpetiolar process. G, protarsus, posterodorsal oblique view. H, mesotarsus, dorsal view. I, metatarsus, dorsal view. J, metatarsus, posterior view. Abbreviations: ad, antennomere distorted, i.e. visibly distorted antennomere associated with decay bubble; cn, cinctus; ms, metanotal spiracle; pr, prora; psf, anterior flange of propodeal spiracle; ptb, probasitarsal brush; spp, subpetiolar process; ss, standing setae; tnc, transverse mesonotal carina." figureDoi="http://doi.org/10.5281/zenodo.6994466" httpUri="https://zenodo.org/record/6994466/files/figure.png" pageId="12" pageNumber="1367">Fig. 4G–J</figureCitation>
|
||
). The
|
||
<emphasis id="B95FEAF0FFEDAA24D16AFF72FBF490DB" bold="true" box="[861,1123,228,249]" pageId="12" pageNumber="1367">pretarsal claw teeth</emphasis>
|
||
are extremely small, only visible as weak bulges on the ventral margins of the claws (
|
||
<figureCitation id="13102A67FFEDAA24D14EFEB7FB959114" box="[889,1026,289,311]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="12" pageNumber="1367">Fig. 5G, I, J</figureCitation>
|
||
). The
|
||
<emphasis id="B95FEAF0FFEDAA24D67AFEB7FB3E9115" bold="true" box="[1101,1193,289,311]" pageId="12" pageNumber="1367">aroliae</emphasis>
|
||
of all legs are welldeveloped but comparatively small (
|
||
<figureCitation id="13102A67FFEDAA24D6F2FEA9FADE9177" box="[1221,1353,319,341]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="12" pageNumber="1367">Fig. 5G, I, J</figureCitation>
|
||
); they are ≤ 0.5× the length of the pretarsal claws.
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFEDAA24D11EFE08FA079776" blockId="12.[809,1426,413,1876]" pageId="12" pageNumber="1367">
|
||
<emphasis id="B95FEAF0FFEDAA24D11EFE08FC249191" box="[809,947,414,435]" italics="true" pageId="12" pageNumber="1367">Metasoma:</emphasis>
|
||
The
|
||
<emphasis id="B95FEAF0FFEDAA24D1C8FE0BFBF49191" bold="true" box="[1023,1123,413,435]" pageId="12" pageNumber="1367">petiole</emphasis>
|
||
is nodiform and lacks tergosternal fusion between its postsclerites (
|
||
<figureCitation id="13102A67FFEDAA24D703FE2DFA1C91F3" box="[1332,1419,443,465]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="12" pageNumber="1367">Figs 2C</figureCitation>
|
||
,
|
||
<figureCitation id="13102A67FFEDAA24D11EFE4CFCD291CD" box="[809,837,474,495]" captionStart="Figure 4" captionStartId="15.[163,242,1675,1697]" captionTargetBox="[229,1376,196,1635]" captionTargetId="figure-26@15.[229,1376,195,1635]" captionTargetPageId="15" captionText="Figure 4. Photomicrographs of †G. sternorhabda sp. nov. showing setation and certain key features. A–D, G–I, paratype, CASENT0741234. E, F, J, holotype, CASENT0741233.A, body, ventrolateral oblique view. B, head, dorsolateral oblique view. C–F, body and left legs: D, dorsolateral oblique; (F) is slightly offset from (E), revealing the prora and subpetiolar process. G, protarsus, posterodorsal oblique view. H, mesotarsus, dorsal view. I, metatarsus, dorsal view. J, metatarsus, posterior view. Abbreviations: ad, antennomere distorted, i.e. visibly distorted antennomere associated with decay bubble; cn, cinctus; ms, metanotal spiracle; pr, prora; psf, anterior flange of propodeal spiracle; ptb, probasitarsal brush; spp, subpetiolar process; ss, standing setae; tnc, transverse mesonotal carina." figureDoi="http://doi.org/10.5281/zenodo.6994466" httpUri="https://zenodo.org/record/6994466/files/figure.png" pageId="12" pageNumber="1367">4F</figureCitation>
|
||
,
|
||
<figureCitation id="13102A67FFEDAA24D165FE4CFCE791D2" box="[850,880,474,496]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="12" pageNumber="1367">5F</figureCitation>
|
||
) [Note 11]. The
|
||
<emphasis id="B95FEAF0FFEDAA24D61BFE4CFB6D91CD" bold="true" box="[1068,1274,474,496]" pageId="12" pageNumber="1367">petiolar tergum</emphasis>
|
||
is anteriorly narrowed, has the spiracles situated on anterolateral bulges, and has a collar posterior to its node (
|
||
<figureCitation id="13102A67FFEDAA24D769FD81FCDF9269" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="12" pageNumber="1367">Figs 2C</figureCitation>
|
||
,
|
||
<figureCitation id="13102A67FFEDAA24D160FDA0FCE1926E" box="[855,886,566,588]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="12" pageNumber="1367">5F</figureCitation>
|
||
); it bears several relatively long setae; setae on the remainder of the metasoma are gradually denser and longer from AIII–VII, with the greatest concentration around the sting. The
|
||
<emphasis id="B95FEAF0FFEDAA24D6E9FD04FA079285" bold="true" box="[1246,1424,658,680]" pageId="12" pageNumber="1367">petiolar node</emphasis>
|
||
is anteroposteriorly longer than dorsoventrally tall (
|
||
<figureCitation id="13102A67FFEDAA24D105FD59FC1092C7" box="[818,903,719,741]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="12" pageNumber="1367">Fig. 2C</figureCitation>
|
||
,
|
||
<figureCitation id="13102A67FFEDAA24D1A0FD59FC2192C7" box="[919,950,719,741]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="12" pageNumber="1367">5F</figureCitation>
|
||
); its anterior margin is longer than its posterior margin in lateral view, and these margins curve evenly into one another. The
|
||
<emphasis id="B95FEAF0FFEDAA24D682FC9BFAF19301" bold="true" box="[1205,1382,781,803]" pageId="12" pageNumber="1367">laterotergites</emphasis>
|
||
are developed (present) (
|
||
<figureCitation id="13102A67FFEDAA24D616FCBDFBE39363" box="[1057,1140,811,833]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="12" pageNumber="1367">Fig. 5F</figureCitation>
|
||
), approximately wedgeshaped and broadening posteriorly. The
|
||
<emphasis id="B95FEAF0FFEDAA24D714FCDCFC00935C" bold="true" pageId="12" pageNumber="1367">petiolar sternum</emphasis>
|
||
is anteriorly flat, with this region bearing stiff proprioceptor setae (
|
||
<figureCitation id="13102A67FFEDAA24D669FC11FB2393BF" box="[1118,1204,903,925]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="12" pageNumber="1367">Fig. 5F</figureCitation>
|
||
); its main portion is about twice as long anteroposteriorly as tall dorsoventrally (
|
||
<figureCitation id="13102A67FFEDAA24D1D0FC53FBD693F8" box="[999,1089,965,987]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="12" pageNumber="1367">Figs 2C</figureCitation>
|
||
,
|
||
<figureCitation id="13102A67FFEDAA24D667FC53FBF893F9" box="[1104,1135,965,987]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="12" pageNumber="1367">5F</figureCitation>
|
||
); it is broadly convex in cross-section at its midpoint (
|
||
<figureCitation id="13102A67FFEDAA24D64AFC75FB5F93DB" box="[1149,1224,995,1017]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="12" pageNumber="1367">Fig. 2I</figureCitation>
|
||
); anteroventrally it is produced as the subdigitate to almost triangular
|
||
<emphasis id="B95FEAF0FFEDAA24D11EFBB6FBBF9414" bold="true" box="[809,1064,1056,1078]" pageId="12" pageNumber="1367">subpetiolar process</emphasis>
|
||
, which is at least twice as long dorsoventrally as wide anteroposteriorly (
|
||
<figureCitation id="13102A67FFEDAA24D73CFBA9FAF59477" box="[1291,1378,1087,1109]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="12" pageNumber="1367">Figs 2C</figureCitation>
|
||
,
|
||
<figureCitation id="13102A67FFEDAA24D758FBD6FA1C9477" box="[1391,1419,1088,1109]" captionStart="Figure 4" captionStartId="15.[163,242,1675,1697]" captionTargetBox="[229,1376,196,1635]" captionTargetId="figure-26@15.[229,1376,195,1635]" captionTargetPageId="15" captionText="Figure 4. Photomicrographs of †G. sternorhabda sp. nov. showing setation and certain key features. A–D, G–I, paratype, CASENT0741234. E, F, J, holotype, CASENT0741233.A, body, ventrolateral oblique view. B, head, dorsolateral oblique view. C–F, body and left legs: D, dorsolateral oblique; (F) is slightly offset from (E), revealing the prora and subpetiolar process. G, protarsus, posterodorsal oblique view. H, mesotarsus, dorsal view. I, metatarsus, dorsal view. J, metatarsus, posterior view. Abbreviations: ad, antennomere distorted, i.e. visibly distorted antennomere associated with decay bubble; cn, cinctus; ms, metanotal spiracle; pr, prora; psf, anterior flange of propodeal spiracle; ptb, probasitarsal brush; spp, subpetiolar process; ss, standing setae; tnc, transverse mesonotal carina." figureDoi="http://doi.org/10.5281/zenodo.6994466" httpUri="https://zenodo.org/record/6994466/files/figure.png" pageId="12" pageNumber="1367">4F</figureCitation>
|
||
,
|
||
<figureCitation id="13102A67FFEDAA24D11EFBC8FCDF9456" box="[809,840,1118,1140]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="12" pageNumber="1367">5F</figureCitation>
|
||
); posteriorly, the sternum is concave, appearing notched, with the concavity receiving the prora. The
|
||
<emphasis id="B95FEAF0FFEDAA24D156FB0DFC599493" bold="true" box="[865,974,1179,1201]" pageId="12" pageNumber="1367">helcium</emphasis>
|
||
(presclerites of the third abdominal segment) is narrow relative to the third abdominal postsclerites (
|
||
<figureCitation id="13102A67FFEDAA24D1D4FB4FFBFD94CC" box="[995,1130,1241,1263]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="12" pageNumber="1367">Figs 2C, F</figureCitation>
|
||
,
|
||
<figureCitation id="13102A67FFEDAA24D64AFB4FFB5594CC" box="[1149,1218,1241,1262]" captionStart="Figure 4" captionStartId="15.[163,242,1675,1697]" captionTargetBox="[229,1376,196,1635]" captionTargetId="figure-26@15.[229,1376,195,1635]" captionTargetPageId="15" captionText="Figure 4. Photomicrographs of †G. sternorhabda sp. nov. showing setation and certain key features. A–D, G–I, paratype, CASENT0741234. E, F, J, holotype, CASENT0741233.A, body, ventrolateral oblique view. B, head, dorsolateral oblique view. C–F, body and left legs: D, dorsolateral oblique; (F) is slightly offset from (E), revealing the prora and subpetiolar process. G, protarsus, posterodorsal oblique view. H, mesotarsus, dorsal view. I, metatarsus, dorsal view. J, metatarsus, posterior view. Abbreviations: ad, antennomere distorted, i.e. visibly distorted antennomere associated with decay bubble; cn, cinctus; ms, metanotal spiracle; pr, prora; psf, anterior flange of propodeal spiracle; ptb, probasitarsal brush; spp, subpetiolar process; ss, standing setae; tnc, transverse mesonotal carina." figureDoi="http://doi.org/10.5281/zenodo.6994466" httpUri="https://zenodo.org/record/6994466/files/figure.png" pageId="12" pageNumber="1367">4C, F</figureCitation>
|
||
) [Note 12]; the helcial tergite conceals the helcial sternite in lateral view (
|
||
<figureCitation id="13102A67FFEDAA24D144FA80FC5E950E" box="[883,969,1302,1324]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="12" pageNumber="1367">Fig. 5F</figureCitation>
|
||
). The
|
||
<emphasis id="B95FEAF0FFEDAA24D62FFA80FAE39509" bold="true" box="[1048,1396,1302,1324]" pageId="12" pageNumber="1367">abdominal posttergite III</emphasis>
|
||
is somewhat constricted posteriorly and is not fused with the third abdominal poststernite (
|
||
<figureCitation id="13102A67FFEDAA24D6CFFAC5FADC954A" box="[1272,1355,1363,1385]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="12" pageNumber="1367">Fig. 2C</figureCitation>
|
||
); it is distinctly necked anteriorly, with a dorsoventrally short but distinct anterior surface which curves to the distinct ‘node’ of the sclerite; its ‘node’ is ‘shouldered’ in appearance, bulging anterolaterally around its ‘neck’, with the ‘shoulders’ visible over the ventrolateral tergal margins in ventral view (
|
||
<figureCitation id="13102A67FFEDAA24D70AF99DFA1E9602" box="[1341,1417,1547,1569]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="12" pageNumber="1367">Fig. 2I</figureCitation>
|
||
) [Note 13]. The
|
||
<emphasis id="B95FEAF0FFEDAA24D1E4F9BCFA06961D" bold="true" box="[979,1425,1578,1600]" pageId="12" pageNumber="1367">abdominal tergosternal margin III</emphasis>
|
||
is weakly curved, without distinct ‘shouldering’ as observed in various Formicinae and Dolichoderinae (
|
||
<figureCitation id="13102A67FFEDAA24D105F910FC1596B9" box="[818,898,1670,1692]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="12" pageNumber="1367">Fig. 2I</figureCitation>
|
||
) [Note 14]. The
|
||
<emphasis id="B95FEAF0FFEDAA24D667F910FCDC9698" bold="true" pageId="12" pageNumber="1367">abdominal poststernite III</emphasis>
|
||
is weakly constricted posteriorly, weakly angled lateromedially, and bears the prora anteroventrally (
|
||
<figureCitation id="13102A67FFEDAA24D105F974FC1596D5" box="[818,898,1762,1784]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="12" pageNumber="1367">Fig. 2I</figureCitation>
|
||
). The
|
||
<emphasis id="B95FEAF0FFEDAA24D1E5F974FB8996D5" bold="true" box="[978,1054,1762,1783]" pageId="12" pageNumber="1367">prora</emphasis>
|
||
is subdigitate in lateral view, being dorsoventrally long and anteroposteriorly thick (
|
||
<figureCitation id="13102A67FFEDAA24D143F889FC519716" box="[884,966,1823,1845]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="12" pageNumber="1367">Fig. 2C</figureCitation>
|
||
); lateromedially, it is relatively narrow, and approximately wedge-shaped in ventral view
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFECAA25D294FF53FC969321" blockId="13.[163,780,197,771]" pageId="13" pageNumber="1368">
|
||
(
|
||
<figureCitation id="13102A67FFECAA25D29BFF53FF6890F8" box="[172,255,197,219]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="13" pageNumber="1368">Fig. 2I</figureCitation>
|
||
). The
|
||
<emphasis id="B95FEAF0FFECAA25D363FF53FD0690F8" bold="true" box="[340,657,197,219]" pageId="13" pageNumber="1368">abdominal segment IV</emphasis>
|
||
is neatly divided into pre- and post-sclerites by the
|
||
<emphasis id="B95FEAF0FFECAA25D094FF72FC9390D8" bold="true" box="[675,772,228,250]" pageId="13" pageNumber="1368">cinctus</emphasis>
|
||
, or transverse sulci on the tergum and sternum (
|
||
<figureCitation id="13102A67FFECAA25D0E8FE94FF549115" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="13" pageNumber="1368">Fig. 5E</figureCitation>
|
||
); its constriction is comparatively shallow. The
|
||
<emphasis id="B95FEAF0FFECAA25D294FED6FD979177" bold="true" box="[163,512,320,342]" pageId="13" pageNumber="1368">abdominal presclerites IV</emphasis>
|
||
are slightly narrower dorsoventrally and lateromedially relative to the postsclerites (
|
||
<figureCitation id="13102A67FFECAA25D375FEEBFE0291B1" box="[322,405,381,403]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="13" pageNumber="1368">Fig. 5E</figureCitation>
|
||
). The
|
||
<emphasis id="B95FEAF0FFECAA25D3EAFEEBFF569193" bold="true" pageId="13" pageNumber="1368">abdominal poststernite IV</emphasis>
|
||
is shorter than the
|
||
<emphasis id="B95FEAF0FFECAA25D3AFFE0AFD429193" bold="true" box="[408,725,412,434]" pageId="13" pageNumber="1368">abdominal posttergite IV</emphasis>
|
||
. The
|
||
<emphasis id="B95FEAF0FFECAA25D294FE2CFE5991F2" bold="true" box="[163,462,442,464]" pageId="13" pageNumber="1368">abdominal segments V</emphasis>
|
||
and
|
||
<emphasis id="B95FEAF0FFECAA25D03CFE2DFDBE91F2" bold="true" box="[523,553,443,464]" pageId="13" pageNumber="1368">VI</emphasis>
|
||
are undivided and are homonomous in form, i.e. highly similar in shape, size, and other qualities of appearance (
|
||
<figureCitation id="13102A67FFECAA25D04BFE6EFD41922F" box="[636,726,504,526]" captionStart="Figure 2" captionStartId="5.[164,243,1414,1436]" captionTargetBox="[178,1430,214,1365]" captionTargetId="figure-147@5.[163,1443,195,1374]" captionTargetPageId="5" captionText="Figure 2. Volume renders of the pupa and two of the adult specimens. The black arrows indicate the subpetiolar process, while the white arrows indicate the prora. A–C, lateral view. D–F, dorsal view. G–I, ventral view. J–L, facial view. A, D, G, J. the pupa of †G. gracilis. B, E, H, K, the adult of †G. gracilis. C, F, I, L, the holotype of †G. sternorhabda sp. nov.. Abbreviations: AIII–VII, third through seventh abdominal segments; AIIIas, anterior surface of third abdominal segment; AIIIps, posterior surface of third abdominal segment; ce, compound eye; cl, clypeus; fc, frontal carina; lbr, labrum; md, mandible; msn, mesonotum; mspl, lower mesopectal region; mspu, upper mesopectal region; mtn, metanotum; oc, ocellus; pd, pedicel; pnt, pronotum; ppd, propodeum; pst, prosternum; pt, petiole; sc, scape." figureDoi="http://doi.org/10.5281/zenodo.6994460" httpUri="https://zenodo.org/record/6994460/files/figure.png" pageId="13" pageNumber="1368">Figs 2C</figureCitation>
|
||
,
|
||
<figureCitation id="13102A67FFECAA25D0D3FE6EFF23920E" captionStart="Figure 4" captionStartId="15.[163,242,1675,1697]" captionTargetBox="[229,1376,196,1635]" captionTargetId="figure-26@15.[229,1376,195,1635]" captionTargetPageId="15" captionText="Figure 4. Photomicrographs of †G. sternorhabda sp. nov. showing setation and certain key features. A–D, G–I, paratype, CASENT0741234. E, F, J, holotype, CASENT0741233.A, body, ventrolateral oblique view. B, head, dorsolateral oblique view. C–F, body and left legs: D, dorsolateral oblique; (F) is slightly offset from (E), revealing the prora and subpetiolar process. G, protarsus, posterodorsal oblique view. H, mesotarsus, dorsal view. I, metatarsus, dorsal view. J, metatarsus, posterior view. Abbreviations: ad, antennomere distorted, i.e. visibly distorted antennomere associated with decay bubble; cn, cinctus; ms, metanotal spiracle; pr, prora; psf, anterior flange of propodeal spiracle; ptb, probasitarsal brush; spp, subpetiolar process; ss, standing setae; tnc, transverse mesonotal carina." figureDoi="http://doi.org/10.5281/zenodo.6994466" httpUri="https://zenodo.org/record/6994466/files/figure.png" pageId="13" pageNumber="1368">4A, C</figureCitation>
|
||
,
|
||
<figureCitation id="13102A67FFECAA25D2F7FD80FF77920E" box="[192,224,534,556]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="13" pageNumber="1368">5E</figureCitation>
|
||
). The
|
||
<emphasis id="B95FEAF0FFECAA25D310FD80FDD3920E" bold="true" box="[295,580,534,556]" pageId="13" pageNumber="1368">abdominal tergum VII</emphasis>
|
||
is approximately dome-shaped;
|
||
<emphasis id="B95FEAF0FFECAA25D37DFDA0FE029269" box="[330,405,566,587]" italics="true" pageId="13" pageNumber="1368">in situ</emphasis>
|
||
, its external surface is oriented dorsoventrally relative to the preceding segments (
|
||
<figureCitation id="13102A67FFECAA25D0D7FDC2FF5492AA" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="13" pageNumber="1368">Fig. 5E</figureCitation>
|
||
). The
|
||
<emphasis id="B95FEAF0FFECAA25D33AFDE4FDA992AA" bold="true" box="[269,574,626,648]" pageId="13" pageNumber="1368">abdominal sternum VII</emphasis>
|
||
is lateromedially cupped and narrowed distally (
|
||
<figureCitation id="13102A67FFECAA25D027FD07FDF39285" box="[528,612,657,679]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="13" pageNumber="1368">Fig. 5E</figureCitation>
|
||
). The
|
||
<emphasis id="B95FEAF0FFECAA25D099FD07FD789285" bold="true" box="[686,751,657,679]" pageId="13" pageNumber="1368">sting</emphasis>
|
||
is long and narrow (
|
||
<figureCitation id="13102A67FFECAA25D34FFD26FE4592E7" box="[376,466,688,710]" captionStart="Figure 4" captionStartId="15.[163,242,1675,1697]" captionTargetBox="[229,1376,196,1635]" captionTargetId="figure-26@15.[229,1376,195,1635]" captionTargetPageId="15" captionText="Figure 4. Photomicrographs of †G. sternorhabda sp. nov. showing setation and certain key features. A–D, G–I, paratype, CASENT0741234. E, F, J, holotype, CASENT0741233.A, body, ventrolateral oblique view. B, head, dorsolateral oblique view. C–F, body and left legs: D, dorsolateral oblique; (F) is slightly offset from (E), revealing the prora and subpetiolar process. G, protarsus, posterodorsal oblique view. H, mesotarsus, dorsal view. I, metatarsus, dorsal view. J, metatarsus, posterior view. Abbreviations: ad, antennomere distorted, i.e. visibly distorted antennomere associated with decay bubble; cn, cinctus; ms, metanotal spiracle; pr, prora; psf, anterior flange of propodeal spiracle; ptb, probasitarsal brush; spp, subpetiolar process; ss, standing setae; tnc, transverse mesonotal carina." figureDoi="http://doi.org/10.5281/zenodo.6994466" httpUri="https://zenodo.org/record/6994466/files/figure.png" pageId="13" pageNumber="1368">Figs 4A</figureCitation>
|
||
,
|
||
<figureCitation id="13102A67FFECAA25D3D6FD26FD9592E4" box="[481,514,688,710]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="13" pageNumber="1368">5E</figureCitation>
|
||
). The
|
||
<emphasis id="B95FEAF0FFECAA25D079FD39FC9D92E7" bold="true" box="[590,778,687,709]" pageId="13" pageNumber="1368">third valvulae</emphasis>
|
||
(sting sheaths) are digitate in form and exserted to highly exserted as preserved (
|
||
<figureCitation id="13102A67FFECAA25D3CBFD7BFDC49320" box="[508,595,749,771]" captionStart="Figure 4" captionStartId="15.[163,242,1675,1697]" captionTargetBox="[229,1376,196,1635]" captionTargetId="figure-26@15.[229,1376,195,1635]" captionTargetPageId="15" captionText="Figure 4. Photomicrographs of †G. sternorhabda sp. nov. showing setation and certain key features. A–D, G–I, paratype, CASENT0741234. E, F, J, holotype, CASENT0741233.A, body, ventrolateral oblique view. B, head, dorsolateral oblique view. C–F, body and left legs: D, dorsolateral oblique; (F) is slightly offset from (E), revealing the prora and subpetiolar process. G, protarsus, posterodorsal oblique view. H, mesotarsus, dorsal view. I, metatarsus, dorsal view. J, metatarsus, posterior view. Abbreviations: ad, antennomere distorted, i.e. visibly distorted antennomere associated with decay bubble; cn, cinctus; ms, metanotal spiracle; pr, prora; psf, anterior flange of propodeal spiracle; ptb, probasitarsal brush; spp, subpetiolar process; ss, standing setae; tnc, transverse mesonotal carina." figureDoi="http://doi.org/10.5281/zenodo.6994466" httpUri="https://zenodo.org/record/6994466/files/figure.png" pageId="13" pageNumber="1368">Figs 4A</figureCitation>
|
||
,
|
||
<figureCitation id="13102A67FFECAA25D056FD7BFD179321" box="[609,640,749,771]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="13" pageNumber="1368">5E</figureCitation>
|
||
) [Note 15].
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFECAA25D294FCBAFDF495C6" blockId="13.[163,779,812,1508]" pageId="13" pageNumber="1368">
|
||
<emphasis id="B95FEAF0FFECAA25D294FCBAFEAB9363" box="[163,316,812,833]" italics="true" pageId="13" pageNumber="1368">Preservation:</emphasis>
|
||
Both the
|
||
<typeStatus id="54908840FFECAA25D38FFCBAFD8C9360" box="[440,539,812,834]" pageId="13" pageNumber="1368" type="holotype">holotype</typeStatus>
|
||
and the synincluded p a r a t y p e (Fi g. 4) h av e d e c o m p o s i t i o n b u b b l e s captured in the process of emanating from the head. The non-synincluded
|
||
<typeStatus id="54908840FFECAA25D39EFC1EFD8393BF" box="[425,532,904,925]" pageId="13" pageNumber="1368" type="paratype">paratype</typeStatus>
|
||
(
|
||
<figureCitation id="13102A67FFECAA25D011FC1EFDFD93BC" box="[550,618,904,926]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="13" pageNumber="1368">Fig. 5</figureCitation>
|
||
) has no such bubbles on the head, but some are present on the surface of the petiolar tergum, on some of the gastral tergites, and apparently a large one protrudes from the abdominal apex. The decomposition bubbles are associated with deformation of the cuticle, particularly for the
|
||
<typeStatus id="54908840FFECAA25D393FBD6FD879474" box="[420,528,1088,1110]" pageId="13" pageNumber="1368" type="holotype">holotype</typeStatus>
|
||
, where the affected cheek and fore tibia have bulged into the lumen of the bubble, as if displaced from compression. Three fracture lines are present around the head of the synincluded
|
||
<typeStatus id="54908840FFECAA25D372FB2DFE2094F2" box="[325,439,1211,1232]" pageId="13" pageNumber="1368" type="paratype">paratype</typeStatus>
|
||
, one approximately in the frontal plane, another at the base of the left antenna, and one between the head and pronotum. Finally, the metasoma of the synincluded
|
||
<typeStatus id="54908840FFECAA25D00EFA81FD33950E" box="[569,676,1303,1324]" pageId="13" pageNumber="1368" type="paratype">paratype</typeStatus>
|
||
appears weakly desiccated (wrinkled); it is surrounded by a bubble that makes rendering difficult, but it is otherwise preserved well enough to determine fine details and structural proportions. Neither the
|
||
<typeStatus id="54908840FFECAA25D294FA26FE9A95E4" box="[163,269,1456,1478]" pageId="13" pageNumber="1368" type="holotype">holotype</typeStatus>
|
||
nor the non-synincluded
|
||
<typeStatus id="54908840FFECAA25D060FA26FD5195E7" box="[599,710,1456,1477]" pageId="13" pageNumber="1368" type="paratype">paratype</typeStatus>
|
||
have fracture lines or apparent desiccation.
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFECAA25D294F983FE5F960F" blockId="13.[163,456,1557,1581]" box="[163,456,1557,1581]" pageId="13" pageNumber="1368">
|
||
<heading id="D0DC818EFFECAA25D294F983FE5F960F" box="[163,456,1557,1581]" centered="true" fontSize="9" level="2" pageId="13" pageNumber="1368" reason="2">
|
||
<emphasis id="B95FEAF0FFECAA25D294F983FE5F960F" box="[163,456,1557,1581]" italics="true" pageId="13" pageNumber="1368">Notes on the description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFECAA25D294F9A8FE589650" blockId="13.[163,779,1597,1650]" pageId="13" pageNumber="1368">
|
||
<emphasis id="B95FEAF0FFECAA25D294F9A8FF649670" box="[163,243,1597,1619]" italics="true" pageId="13" pageNumber="1368">Note 1:</emphasis>
|
||
The setation of the body, in general, could not be described in fine detail.
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFECAA25D294F90DFDF5974A" blockId="13.[163,779,1690,1896]" pageId="13" pageNumber="1368">
|
||
<emphasis id="B95FEAF0FFECAA25D294F90DFF659692" box="[163,242,1691,1712]" italics="true" pageId="13" pageNumber="1368">Note 2:</emphasis>
|
||
The ommatidia were counted for one eye of the
|
||
<typeStatus id="54908840FFECAA25D294F92CFE9A96ED" box="[163,269,1722,1743]" pageId="13" pageNumber="1368" type="paratype">paratype</typeStatus>
|
||
from a high-resolution photograph. It was not possible to count all of the ommatidia for this eye. Likewise, we counted 85 ommatidia in the left eye of the non-synincluded
|
||
<typeStatus id="54908840FFECAA25D3AEF880FD949709" box="[409,515,1814,1835]" pageId="13" pageNumber="1368" type="paratype">paratype</typeStatus>
|
||
; about 15% of this eye is concealed, so we expect that there are indeed> 100 ommatidia, but certainly less than 150.
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFECAA25D10CFF50FC5D9248" blockId="13.[827,1443,197,618]" pageId="13" pageNumber="1368">
|
||
<emphasis id="B95FEAF0FFECAA25D10CFF50FC1890F8" box="[827,911,197,219]" italics="true" pageId="13" pageNumber="1368">Note 3:</emphasis>
|
||
The position of the antennal toruli relative to the posterior clypeal margin is known to be of classificatory value above the species level (
|
||
<bibRefCitation id="EFBA4B13FFECAA25D174FEB7FC749114" author="Bolton B" box="[835,995,289,311]" pageId="13" pageNumber="1368" pagination="1 - 370" refId="ref23216" refString="Bolton B. 2003. Synopsis and classification of Formicidae. Memoirs of the American Entomological Institute 71: 1 - 370." type="journal article" year="2003">Bolton, 2003</bibRefCitation>
|
||
). We observe that the toruli contact the epistomal line in †
|
||
<emphasis id="B95FEAF0FFECAA25D667FED6FA8C9177" box="[1104,1307,320,341]" italics="true" pageId="13" pageNumber="1368">
|
||
<taxonomicName id="4C2B4D61FFECAA25D667FED6FA809177" authorityName="Boudinot & Richter & Katzke & Chaul & Keller & Economo & Beutel & Yamamoto" authorityYear="2022" box="[1104,1303,320,341]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="1368" phylum="Arthropoda" rank="species" species="sternorhabda">G. sternorhabda</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
but do not provide this in the diagnosis because the posterior limit of the clypeus is difficult to evaluate in some of the fossils, particularly in the
|
||
<typeStatus id="54908840FFECAA25D694FE0AFB419193" box="[1187,1238,412,433]" pageId="13" pageNumber="1368">type</typeStatus>
|
||
specimens of the
|
||
<taxonomicName id="4C2B4D61FFECAA25D10CFE2CFC0491ED" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[827,915,442,463]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="1368" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B95FEAF0FFECAA25D10CFE2CFC0491ED" box="[827,915,442,463]" italics="true" pageId="13" pageNumber="1368">gracilis</emphasis>
|
||
</taxonomicName>
|
||
group. However, among the
|
||
<taxonomicName id="4C2B4D61FFECAA25D6D4FE2CFABE91ED" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[1251,1321,442,463]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="1368" phylum="Arthropoda" rank="species" species="pilosa">
|
||
<emphasis id="B95FEAF0FFECAA25D6D4FE2CFABE91ED" box="[1251,1321,442,463]" italics="true" pageId="13" pageNumber="1368">pilosa</emphasis>
|
||
</taxonomicName>
|
||
group, we observed that the toruli are close to or abutting the clypeus in †
|
||
<taxonomicName id="4C2B4D61FFECAA25D1FDFE6EFBD1922F" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[970,1094,504,525]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="1368" phylum="Arthropoda" rank="species" species="contega">
|
||
<emphasis id="B95FEAF0FFECAA25D1FDFE6EFBD1922F" box="[970,1094,504,525]" italics="true" pageId="13" pageNumber="1368">G. contega</emphasis>
|
||
</taxonomicName>
|
||
and †
|
||
<taxonomicName id="4C2B4D61FFECAA25D6A6FE6EFB6D922F" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[1169,1274,504,525]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="1368" phylum="Arthropoda" rank="species" species="pilosa">
|
||
<emphasis id="B95FEAF0FFECAA25D6A6FE6EFB6D922F" box="[1169,1274,504,525]" italics="true" pageId="13" pageNumber="1368">G. pilosa</emphasis>
|
||
</taxonomicName>
|
||
. It is difficult to determine the posterior extent of the clypeus in †
|
||
<taxonomicName id="4C2B4D61FFECAA25D170FDA3FC209269" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[839,951,565,587]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="1368" phylum="Arthropoda" rank="species" species="magna">
|
||
<emphasis id="B95FEAF0FFECAA25D170FDA3FC209269" box="[839,951,565,587]" italics="true" pageId="13" pageNumber="1368">G. magna</emphasis>
|
||
</taxonomicName>
|
||
, thus no confident statement can be made at this time.
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFECAA25D10CFD05FBFA9326" blockId="13.[827,1442,658,772]" pageId="13" pageNumber="1368">
|
||
<emphasis id="B95FEAF0FFECAA25D10CFD05FC1D928A" box="[827,906,659,680]" italics="true" pageId="13" pageNumber="1368">Note 4:</emphasis>
|
||
The setae on the medioclypeal area could only be evaluated on the non-synincluded
|
||
<typeStatus id="54908840FFECAA25D6CAFD24FAFD92E5" box="[1277,1386,690,711]" pageId="13" pageNumber="1368" type="paratype">paratype</typeStatus>
|
||
. The lateralmost seta is probably paired, but the second hair could not be observed.
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFECAA25D10CFCB8FC42941A" blockId="13.[827,1444,813,1080]" pageId="13" pageNumber="1368">
|
||
<emphasis id="B95FEAF0FFECAA25D10CFCB8FC049360" box="[827,915,813,835]" italics="true" pageId="13" pageNumber="1368">Note 5:</emphasis>
|
||
We have recorded the maxillary palps as 5-merous based on scrutiny of the non-synincluded
|
||
<typeStatus id="54908840FFECAA25D10CFCFDFC3693A2" box="[827,929,875,896]" pageId="13" pageNumber="1368" type="paratype">paratype</typeStatus>
|
||
(
|
||
<figureCitation id="13102A67FFECAA25D19AFCFCFB8093A2" box="[941,1047,874,896]" captionStart="Figure 5" captionStartId="16.[145,227,1675,1697]" captionTargetBox="[223,1346,196,1628]" captionTargetId="figure-26@16.[223,1346,195,1635]" captionTargetPageId="16" captionText="Figure 5. Photomicrographic details of †G. sternorhabda sp. nov. Paratype specimen (UFV-LABECOL-009656). A, head in approximate facial view. B, mouthparts in anterolateral oblique view. C, head and mesosoma in lateral view. D, proximal segments of metathoracic leg in approximate anterior view. E, metasoma in approximate lateral view. F, petiole in lateral view. G, prothoracic tarsus, oblique posterior view. H, metatibial apex and metabasitarsal base in posterior view. I, propretarsus in oblique distal view. J, metapretarsus in posteroventral oblique view. Abbreviations: AI, propodeum; AII– AVII, second through seventh abdominal segments; ca, calcar; cav, velum (lamella) of calcar; ce, compound eye; cl, clypeus; cll, lateral lobe of clypeus; cns, cinctus of fourth abdominal sternum; cnt, cinctus of fourth abdominal tergum; fc, frontal carina; hetg, helcial tergite; msctl, mesoscutellar area (mesoscutellum); msctm, mesoscutal area (mesoscutum); msnttc, transverse" figureDoi="http://doi.org/10.5281/zenodo.6994470" httpUri="https://zenodo.org/record/6994470/files/figure.png" pageId="13" pageNumber="1368">Fig. 5B, C</figureCitation>
|
||
) and our renders of the synincluded types (
|
||
<figureCitation id="13102A67FFECAA25D1BEFC1FFC6D93BD" box="[905,1018,905,927]" captionStart="Figure 7" captionStartId="19.[164,241,1513,1535]" captionTargetBox="[163,1443,195,1470]" captionTargetId="figure-119@19.[163,1443,195,1473]" captionTargetPageId="19" captionText="Figure 7. Maxillary and labial palps of †G.gracilis and †G.sternorhabda sp. nov.. Small black stars indicate each palpomere on the right-hand side of the body.A, B, †G.gracilis adult, CASENT0741232, in anterior dorsolateral oblique view (A) and anterior view (B).C, D, †G.gracilis pupa, CASENT0741231, in anterior view (C) and anterior dorsolateral view (D).E, F, †G.sternorhabda sp. nov. holotype, CASENT0741233, and paratype, CASENT0741234, in anteroventral oblique view. Abbreviations: pl, labial palp; plp, process of the proximal labial palpomere; pm, maxillary palp; pm?, possible maxillary palpomere." figureDoi="http://doi.org/10.5281/zenodo.6994474" httpUri="https://zenodo.org/record/6994474/files/figure.png" pageId="13" pageNumber="1368">Fig. 7E, F</figureCitation>
|
||
). It is possible that there is a poorly developed proximal sixth palpomere (‘pm?’ in
|
||
<figureCitation id="13102A67FFECAA25D741FC3EFCEE93FE" captionStart="Figure 7" captionStartId="19.[164,241,1513,1535]" captionTargetBox="[163,1443,195,1470]" captionTargetId="figure-119@19.[163,1443,195,1473]" captionTargetPageId="19" captionText="Figure 7. Maxillary and labial palps of †G.gracilis and †G.sternorhabda sp. nov.. Small black stars indicate each palpomere on the right-hand side of the body.A, B, †G.gracilis adult, CASENT0741232, in anterior dorsolateral oblique view (A) and anterior view (B).C, D, †G.gracilis pupa, CASENT0741231, in anterior view (C) and anterior dorsolateral view (D).E, F, †G.sternorhabda sp. nov. holotype, CASENT0741233, and paratype, CASENT0741234, in anteroventral oblique view. Abbreviations: pl, labial palp; plp, process of the proximal labial palpomere; pm, maxillary palp; pm?, possible maxillary palpomere." figureDoi="http://doi.org/10.5281/zenodo.6994474" httpUri="https://zenodo.org/record/6994474/files/figure.png" pageId="13" pageNumber="1368">Fig. 7E, F</figureCitation>
|
||
). The palps are remarkably short compared to those of †
|
||
<taxonomicName id="4C2B4D61FFECAA25D19DFC73FBB693D8" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[938,1057,997,1018]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="1368" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B95FEAF0FFECAA25D19DFC73FBB693D8" box="[938,1057,997,1018]" italics="true" pageId="13" pageNumber="1368">G. gracilis</emphasis>
|
||
</taxonomicName>
|
||
, and they lack the process of the proximal labial palpomere which we observed in the other species.
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFECAA25D10CFBF4FB6A94F1" blockId="13.[827,1442,1121,1235]" pageId="13" pageNumber="1368">
|
||
<emphasis id="B95FEAF0FFECAA25D10CFBF4FC039454" box="[827,916,1121,1143]" italics="true" pageId="13" pageNumber="1368">Note 6:</emphasis>
|
||
The shape of the palpomeres is notable, as the proximal ones appear flattened to some degree. Extended µ-CT sampling is recommended to understand palpomere shape variation.
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFECAA25D10CFB6AFC76954C" blockId="13.[827,1442,1276,1390]" pageId="13" pageNumber="1368">
|
||
<emphasis id="B95FEAF0FFECAA25D10CFB6AFC069533" box="[827,913,1276,1297]" italics="true" pageId="13" pageNumber="1368">Note 7:</emphasis>
|
||
The pronotal lobes of †
|
||
<taxonomicName id="4C2B4D61FFECAA25D680FB6AFAE59533" authorityName="Nel & Perrault" authorityYear="2004" box="[1207,1394,1276,1297]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="1368" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FEAF0FFECAA25D680FB6AFAE59533" box="[1207,1394,1276,1297]" italics="true" pageId="13" pageNumber="1368">Gerontoformica</emphasis>
|
||
</taxonomicName>
|
||
are large compared to crown ants. Systematic evaluation of the development of these lobes among stem ants is recommended.
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFECAA25D10CFA34FB4796AC" blockId="13.[827,1443,1441,1678]" pageId="13" pageNumber="1368">
|
||
<emphasis id="B95FEAF0FFECAA25D10CFA34FC1C9594" box="[827,907,1441,1463]" italics="true" pageId="13" pageNumber="1368">Note 8:</emphasis>
|
||
The shape of the propodeum appears to differ between the two synincluded
|
||
<typeStatus id="54908840FFECAA25D682FA56FB6195F7" box="[1205,1270,1472,1493]" pageId="13" pageNumber="1368">types</typeStatus>
|
||
and the nonsynincluded
|
||
<typeStatus id="54908840FFECAA25D1F2FA49FC6F95D6" box="[965,1016,1503,1524]" pageId="13" pageNumber="1368">type</typeStatus>
|
||
.While the propodeum is rounded in the non-synincluded specimen, it is somewhat rectangular in the synincluded specimens, with the dorsal and posterior margins oriented nearly perpendicularly and narrowly rounding into one another in lateral view, and its posterior surface nearly flat.
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFECAA25D10CF921FC249744" blockId="13.[827,1443,1718,1894]" pageId="13" pageNumber="1368">
|
||
<emphasis id="B95FEAF0FFECAA25D10CF921FC1896E9" box="[827,911,1718,1740]" italics="true" pageId="13" pageNumber="1368">Note 9:</emphasis>
|
||
We have stated ‘apparently’, as we are not totally certain about this state. We have included it, however, in order to encourage future examination of this structure, which we know to be variable across the
|
||
<taxonomicName id="4C2B4D61FFECAA25D150F8A7FC719765" authorityName="LATREILLE" authorityYear="1809" box="[871,998,1841,1863]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="1368" phylum="Arthropoda" rank="family">Formicidae</taxonomicName>
|
||
and other groups of Aculeata (B. E. B., pers. obs.).
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFEFAA26D2A6FF50FDCD922F" blockId="14.[145,762,197,526]" pageId="14" pageNumber="1369">
|
||
<emphasis id="B95FEAF0FFEFAA26D2A6FF50FF6690F8" box="[145,241,197,219]" italics="true" pageId="14" pageNumber="1369">Note 10:</emphasis>
|
||
Plantar lobes are developed in †
|
||
<taxonomicName id="4C2B4D61FFEFAA26D04DFF50FD6490F8" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[634,755,197,219]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="1369" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B95FEAF0FFEFAA26D04DFF50FD6490F8" box="[634,755,197,219]" italics="true" pageId="14" pageNumber="1369">G. gracilis</emphasis>
|
||
</taxonomicName>
|
||
, which has far sparser ventral tarsomeral setae and proportionally larger aroliae. Because tarsal setae, lobes and aroliae are functional traits related to traction (e.g.
|
||
<bibRefCitation id="EFBA4B13FFEFAA26D30FFED6FDA79177" author="Beutel RG & Gorb SN" box="[312,560,320,342]" pageId="14" pageNumber="1369" pagination="177 - 207" refId="ref23057" refString="Beutel RG, Gorb SN. 2001. Ultrastructure of attachment specializations of hexapods (Arthropoda): evolutionary patterns inferred from a revised ordinal phylogeny. Journal of Zoological Systematics & Evolution Research 39: 177 - 207." type="journal article" year="2001">Beutel & Gorb 2001</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFBA4B13FFEFAA26D076FED6FF4F9156" author="Boudinot BE & Beutel RG & Gorb SN & Polilov AA" pageId="14" pageNumber="1369" pagination="99 - 113" refId="ref23481" refString="Boudinot BE, Beutel RG, Gorb SN, Polilov AA. 2021 a. Functional diversity of attachment and grooming structures is retained in all but the smallest insects. Journal of Zoology 313: 99 - 113." type="journal article" year="2021">
|
||
Boudinot
|
||
<emphasis id="B95FEAF0FFEFAA26D08CFED6FD6E9177" box="[699,761,320,341]" italics="true" pageId="14" pageNumber="1369">et al.</emphasis>
|
||
2021a
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFBA4B13FFEFAA26D2D1FEC8FE229156" author="Wohrl T & Richter A & Guo S & Reinhardt L & Nowotny M & Blickhan R" box="[230,437,350,372]" pageId="14" pageNumber="1369" pagination="242677" refId="ref25717" refString="Wohrl T, Richter A, Guo S, Reinhardt L, Nowotny M, Blickhan R. 2021. Comparative analysis of a geometric and an adhesive righting strategy against toppling in inclined hexapodal locomotion. Journal of Experimental Biology 224: jeb 242677." type="journal article" year="2021">
|
||
Wöhrl
|
||
<emphasis id="B95FEAF0FFEFAA26D30FFEC9FEE49151" box="[312,371,350,372]" italics="true" pageId="14" pageNumber="1369">et al.</emphasis>
|
||
2021
|
||
</bibRefCitation>
|
||
), the distinctions observed here suggest, or otherwise indicate, that there is niche separation between †
|
||
<taxonomicName id="4C2B4D61FFEFAA26D3B4FE0AFDAD9193" authorityName="Boudinot & Richter & Katzke & Chaul & Keller & Economo & Beutel & Yamamoto" authorityYear="2022" box="[387,570,412,433]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="1369" phylum="Arthropoda" rank="species" species="sternorhabda">
|
||
<emphasis id="B95FEAF0FFEFAA26D3B4FE0AFDAD9193" box="[387,570,412,433]" italics="true" pageId="14" pageNumber="1369">G. sternorhabda</emphasis>
|
||
</taxonomicName>
|
||
and †
|
||
<taxonomicName id="4C2B4D61FFEFAA26D048FE0AFD649193" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[639,755,412,433]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="1369" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B95FEAF0FFEFAA26D048FE0AFD649193" box="[639,755,412,433]" italics="true" pageId="14" pageNumber="1369">G. gracilis</emphasis>
|
||
</taxonomicName>
|
||
. Among
|
||
<taxonomicName id="4C2B4D61FFEFAA26D2D0FE2CFEBC91ED" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[231,299,442,463]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="1369" phylum="Arthropoda" rank="species" species="pilosa">
|
||
<emphasis id="B95FEAF0FFEFAA26D2D0FE2CFEBC91ED" box="[231,299,442,463]" italics="true" pageId="14" pageNumber="1369">pilosa</emphasis>
|
||
</taxonomicName>
|
||
group species, the fourth tarsomeres are notched in †
|
||
<taxonomicName id="4C2B4D61FFEFAA26D329FE4FFE0291CD" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[286,405,473,495]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="1369" phylum="Arthropoda" rank="species" species="contega">
|
||
<emphasis id="B95FEAF0FFEFAA26D329FE4FFE0291CD" box="[286,405,473,495]" italics="true" pageId="14" pageNumber="1369">G. contega</emphasis>
|
||
</taxonomicName>
|
||
, are
|
||
<taxonomicName id="4C2B4D61FFEFAA26D3E7FE4FFDB191CC" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[464,550,473,494]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="1369" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B95FEAF0FFEFAA26D3E7FE4FFDB191CC" box="[464,550,473,494]" italics="true" pageId="14" pageNumber="1369">gracilis</emphasis>
|
||
</taxonomicName>
|
||
-like in †
|
||
<taxonomicName id="4C2B4D61FFEFAA26D0BAFE4FFD6591CC" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[653,754,473,494]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="1369" phylum="Arthropoda" rank="species" species="pilosa">
|
||
<emphasis id="B95FEAF0FFEFAA26D0BAFE4FFD6591CC" box="[653,754,473,494]" italics="true" pageId="14" pageNumber="1369">G. pilosa</emphasis>
|
||
</taxonomicName>
|
||
, and are of uncertain form in †
|
||
<taxonomicName id="4C2B4D61FFEFAA26D3D1FE6EFDC4922F" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[486,595,504,525]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="1369" phylum="Arthropoda" rank="species" species="magna">
|
||
<emphasis id="B95FEAF0FFEFAA26D3D1FE6EFDC4922F" box="[486,595,504,525]" italics="true" pageId="14" pageNumber="1369">G. magna</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFEFAA26D2A6FDAAFF6F92EE" blockId="14.[145,761,572,716]" pageId="14" pageNumber="1369">
|
||
<emphasis id="B95FEAF0FFEFAA26D2A6FDAAFF6B9273" box="[145,252,572,593]" italics="true" pageId="14" pageNumber="1369">Note 11:</emphasis>
|
||
Whereas it was possible to determine whether or not the postsclerites of the metasoma were fused, it was not possible to evaluate such fusion for the presclerites of the petiole and third abdominal segment.
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFEFAA26D2A6FD6DFD0C934E" blockId="14.[145,760,762,876]" pageId="14" pageNumber="1369">
|
||
<emphasis id="B95FEAF0FFEFAA26D2A6FD6DFF799332" box="[145,238,763,784]" italics="true" pageId="14" pageNumber="1369">Note 12:</emphasis>
|
||
The helcium of †
|
||
<taxonomicName id="4C2B4D61FFEFAA26D381FD6DFDFB932D" authorityName="Boudinot & Richter & Katzke & Chaul & Keller & Economo & Beutel & Yamamoto" authorityYear="2022" box="[438,620,762,784]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="1369" phylum="Arthropoda" rank="species" species="sternorhabda">
|
||
<emphasis id="B95FEAF0FFEFAA26D381FD6DFDFB932D" box="[438,620,762,784]" italics="true" pageId="14" pageNumber="1369">G. sternorhabda</emphasis>
|
||
</taxonomicName>
|
||
is distinctly broader than that of †
|
||
<taxonomicName id="4C2B4D61FFEFAA26D392FC8FFDB3930C" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[421,548,793,814]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="1369" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B95FEAF0FFEFAA26D392FC8FFDB3930C" box="[421,548,793,814]" italics="true" pageId="14" pageNumber="1369">G. gracilis</emphasis>
|
||
</taxonomicName>
|
||
. However, it was not possible to evaluate this condition for the other previously described †
|
||
<taxonomicName id="4C2B4D61FFEFAA26D3BAFCC0FD0C934E" box="[397,667,854,876]" class="Insecta" family="Formicidae" genus="Gerontoformica" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="1369" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B95FEAF0FFEFAA26D3BAFCC0FDAA9349" box="[397,573,854,875]" italics="true" pageId="14" pageNumber="1369">Gerontoformica</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFEFAA26D2A6FC0DFF7F95FA" blockId="14.[145,761,922,1496]" pageId="14" pageNumber="1369">
|
||
<emphasis id="B95FEAF0FFEFAA26D2A6FC0DFF7A9392" box="[145,237,923,944]" italics="true" pageId="14" pageNumber="1369">Note 13:</emphasis>
|
||
Abdominal tergum III of the non-synincluded
|
||
<typeStatus id="54908840FFEFAA26D2A6FC2CFF6D93ED" box="[145,250,954,975]" pageId="14" pageNumber="1369" type="paratype">paratype</typeStatus>
|
||
is apparently not similarly shouldered. We are, therefore, less certain about the development and distribution of this state, i.e. whether it is artefactual or not, or whether the additional
|
||
<typeStatus id="54908840FFEFAA26D029FB80FD109409" box="[542,647,1046,1067]" pageId="14" pageNumber="1369" type="paratype">paratype</typeStatus>
|
||
is indeed conspecific. We choose to recognize the additional
|
||
<typeStatus id="54908840FFEFAA26D2A6FBC5FF69944A" box="[145,254,1107,1128]" pageId="14" pageNumber="1369" type="paratype">paratype</typeStatus>
|
||
as conspecific due to the high degree of shared conditions, as indicated by the diagnosis. We also consider the tergal shouldering to likely be a true developmental state of †
|
||
<taxonomicName id="4C2B4D61FFEFAA26D3DCFB39FD3E94E1" authorityName="Boudinot & Richter & Katzke & Chaul & Keller & Economo & Beutel & Yamamoto" authorityYear="2022" box="[491,681,1198,1220]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="1369" phylum="Arthropoda" rank="species" species="sternorhabda">
|
||
<emphasis id="B95FEAF0FFEFAA26D3DCFB39FD3E94E1" box="[491,681,1198,1220]" italics="true" pageId="14" pageNumber="1369">G. sternorhabda</emphasis>
|
||
</taxonomicName>
|
||
, as we observe it in both of the synincluded specimens, and as the synincluded †
|
||
<taxonomicName id="4C2B4D61FFEFAA26D340FB7AFE7D9523" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[375,490,1260,1281]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="1369" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B95FEAF0FFEFAA26D340FB7AFE7D9523" box="[375,490,1260,1281]" italics="true" pageId="14" pageNumber="1369">G. gracilis</emphasis>
|
||
</taxonomicName>
|
||
clearly does not display this condition. The third abdominal postsclerite is more-or-less evenly curved from anterior to posterior (†
|
||
<taxonomicName id="4C2B4D61FFEFAA26D291FADEFE9D957F" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[166,266,1352,1373]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="1369" phylum="Arthropoda" rank="species" species="pilosa">
|
||
<emphasis id="B95FEAF0FFEFAA26D291FADEFE9D957F" box="[166,266,1352,1373]" italics="true" pageId="14" pageNumber="1369">G. pilosa</emphasis>
|
||
</taxonomicName>
|
||
) or is entirely evenly curved from anterior to posterior (†
|
||
<taxonomicName id="4C2B4D61FFEFAA26D306FAF1FE309559" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[305,423,1382,1404]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="1369" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B95FEAF0FFEFAA26D306FAF1FE309559" box="[305,423,1382,1404]" italics="true" pageId="14" pageNumber="1369">G. gracilis</emphasis>
|
||
</taxonomicName>
|
||
, †
|
||
<taxonomicName id="4C2B4D61FFEFAA26D3F7FAF1FDA09559" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[448,567,1382,1404]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="1369" phylum="Arthropoda" rank="species" species="robusta">
|
||
<emphasis id="B95FEAF0FFEFAA26D3F7FAF1FDA09559" box="[448,567,1382,1404]" italics="true" pageId="14" pageNumber="1369">G. robusta</emphasis>
|
||
</taxonomicName>
|
||
, †
|
||
<taxonomicName id="4C2B4D61FFEFAA26D067FAF1FD509559" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[592,711,1382,1404]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="1369" phylum="Arthropoda" rank="species" species="spiralis">
|
||
<emphasis id="B95FEAF0FFEFAA26D067FAF1FD509559" box="[592,711,1382,1404]" italics="true" pageId="14" pageNumber="1369">G. spiralis</emphasis>
|
||
</taxonomicName>
|
||
and †
|
||
<taxonomicName id="4C2B4D61FFEFAA26D2AAFA13FEBB95B8" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[157,300,1413,1434]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="1369" phylum="Arthropoda" rank="species" species="subcuspis">
|
||
<emphasis id="B95FEAF0FFEFAA26D2AAFA13FEBB95B8" box="[157,300,1413,1434]" italics="true" pageId="14" pageNumber="1369">G. subcuspis</emphasis>
|
||
</taxonomicName>
|
||
); it could not be confidently determined for †
|
||
<taxonomicName id="4C2B4D61FFEFAA26D2FCFA32FE16959B" box="[203,385,1444,1465]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="1369" phylum="Arthropoda" rank="species" species="occidentalis">
|
||
<emphasis id="B95FEAF0FFEFAA26D2FCFA32FE16959B" box="[203,385,1444,1465]" italics="true" pageId="14" pageNumber="1369">G. occidentalis</emphasis>
|
||
</taxonomicName>
|
||
or for the four
|
||
<emphasis id="B95FEAF0FFEFAA26D07CFA32FD6E959B" box="[587,761,1444,1465]" italics="true" pageId="14" pageNumber="1369">nomina dubia</emphasis>
|
||
species.
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFEFAA26D2A6F994FE8096CC" blockId="14.[145,761,1537,1774]" pageId="14" pageNumber="1369">
|
||
<emphasis id="B95FEAF0FFEFAA26D2A6F994FE969635" box="[145,257,1537,1559]" italics="true" pageId="14" pageNumber="1369">Note 14:</emphasis>
|
||
Shouldering of the third abdominal tergosternal margin was used for generic identification and the tribe-level classifications of the Formicinae and Dolichoderinae by
|
||
<bibRefCitation id="EFBA4B13FFEFAA26D3A6F9CBFD889650" author="Bolton B" box="[401,543,1629,1651]" pageId="14" pageNumber="1369" refId="ref23193" refString="Bolton B. 1994. Identification guide to the ant genera of the world. Cambridge: Harvard University Press, 222." type="book" year="1994">Bolton (1994</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFBA4B13FFEFAA26D01DF9CBFDF39650" author="Bolton B" box="[554,612,1629,1650]" pageId="14" pageNumber="1369" pagination="1 - 370" refId="ref23216" refString="Bolton B. 2003. Synopsis and classification of Formicidae. Memoirs of the American Entomological Institute 71: 1 - 370." type="journal article" year="2003">2003</bibRefCitation>
|
||
). We perceive no difference in the form of these margins between †
|
||
<taxonomicName id="4C2B4D61FFEFAA26D2A9F90DFEFE968D" authorityName="Boudinot & Richter & Katzke & Chaul & Keller & Economo & Beutel & Yamamoto" authorityYear="2022" box="[158,361,1690,1712]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="1369" phylum="Arthropoda" rank="species" species="sternorhabda">
|
||
<emphasis id="B95FEAF0FFEFAA26D2A9F90DFEFE968D" box="[158,361,1690,1712]" italics="true" pageId="14" pageNumber="1369">G. sternorhabda</emphasis>
|
||
</taxonomicName>
|
||
and †
|
||
<taxonomicName id="4C2B4D61FFEFAA26D38DF90DFDAA968D" baseAuthorityName="Barden & Grimaldi" baseAuthorityYear="2014" box="[442,573,1690,1712]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="1369" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B95FEAF0FFEFAA26D38DF90DFDAA968D" box="[442,573,1690,1712]" italics="true" pageId="14" pageNumber="1369">G. gracilis</emphasis>
|
||
</taxonomicName>
|
||
; we encourage evaluation of the form of the margins for other stem
|
||
<taxonomicName id="4C2B4D61FFEFAA26D2A6F94EFE8496CC" authorityName="LATREILLE" authorityYear="1809" box="[145,275,1752,1774]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="1369" phylum="Arthropoda" rank="family">Formicidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFEFAA26D2A6F881FD569748" blockId="14.[145,761,1814,1898]" pageId="14" pageNumber="1369">
|
||
<emphasis id="B95FEAF0FFEFAA26D2A6F881FF7A9709" box="[145,237,1814,1836]" italics="true" pageId="14" pageNumber="1369">Note 15:</emphasis>
|
||
We anticipate that study of the sting apparati of stem ants using µ-CT to be fruitful. The degree to which the third valvulae are exserted is unusual.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3316569FFEFAA26D11EFF53FC3D9368" pageId="14" pageNumber="1369" type="discussion">
|
||
<paragraph id="8B9436E2FFEFAA26D11EFF53FBE090FF" blockId="14.[809,1143,197,221]" box="[809,1143,197,221]" pageId="14" pageNumber="1369">
|
||
<emphasis id="B95FEAF0FFEFAA26D11EFF53FBE090FF" box="[809,1143,197,221]" italics="true" pageId="14" pageNumber="1369">Remarks on the description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFEFAA26D11EFF78FC3D9368" blockId="14.[809,1425,237,842]" pageId="14" pageNumber="1369">
|
||
The non-synincluded
|
||
<typeStatus id="54908840FFEFAA26D6A2FF78FA899121" box="[1173,1310,238,259]" pageId="14" pageNumber="1369" type="paratype">paratype</typeStatus>
|
||
(UFV- LABECOL-009656) may not be conspecific with the synincluded type specimens. It differs from the synincluded specimens by the following states: (1) possibly in maxillary palpomere count (5 vs. 6; see Note 5 of the description); (2) the propodeal spiracle appears to be slightly higher on the propodeum; (3) the propodeum appears to be more rounded; (4) the petiolar node appears to be shorter; (5) the subpetiolar process appears to be straighter; and (6) the third abdominal posttergite does not appear to be shouldered. We note that these are ‘apparent’ differences because of the limited set of available specimens. Based on expectations from the neontological fauna, it is possible that the differences we observe may be infraspecific variation, but it will be necessary to evaluate more specimens. Because the specimens from the two pieces of amber are otherwise highly similar and equally diagnosable, we conservatively designate them as conspecific.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3316569FFEFAA26D11EFC14FC09947B" pageId="14" pageNumber="1369" type="etymology">
|
||
<paragraph id="8B9436E2FFEFAA26D11EFC14FC3D93B8" blockId="14.[809,938,898,922]" box="[809,938,898,922]" pageId="14" pageNumber="1369">
|
||
<emphasis id="B95FEAF0FFEFAA26D11EFC14FC3D93B8" box="[809,938,898,922]" italics="true" pageId="14" pageNumber="1369">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9436E2FFEFAA26D11EFC3DFC09947B" blockId="14.[809,1425,939,1114]" pageId="14" pageNumber="1369">
|
||
The specific epithet
|
||
<taxonomicName id="4C2B4D61FFEFAA26D603FC3DFB4B93E2" authorityName="Boudinot & Richter & Katzke & Chaul & Keller & Economo & Beutel & Yamamoto" authorityYear="2022" box="[1076,1244,939,960]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="1369" phylum="Arthropoda" rank="species" species="sternorhabda">
|
||
<emphasis id="B95FEAF0FFEFAA26D603FC3DFB4B93E2" box="[1076,1244,939,960]" italics="true" pageId="14" pageNumber="1369">sternorhabda</emphasis>
|
||
</taxonomicName>
|
||
combines the Ancient Greek words στερνών (sternum) and ράβδος (rod) in reference to the form of the subpetiolar process, which is unique among †
|
||
<taxonomicName id="4C2B4D61FFEFAA26D671FB91FB60943E" authorityName="Nel & Perrault" authorityYear="2004" box="[1094,1271,1031,1052]" class="Insecta" family="Formicidae" genus="Gerontoformica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="1369" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FEAF0FFEFAA26D671FB91FB60943E" box="[1094,1271,1031,1052]" italics="true" pageId="14" pageNumber="1369">Gerontoformica</emphasis>
|
||
</taxonomicName>
|
||
. The name is adjectival and feminine in form to match the gender of the genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |